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<document id="DA310F2D78736D022136B140261DD04E" ID-CLB-Dataset="35231" ID-DOI="http://dx.doi.org/10.3897/zookeys.393.6717" ID-GBIF-Dataset="ca1977b6-7464-4c67-999d-761e0faa67ab" ID-PMC="PMC3974427" ID-Pensoft-Pub="1313-2970-393-1" ID-PubMed="24715783" ID-ZooBank="5B035E76BBD14A44A5D8C2140E6168F1" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-393-1" ModsDocOrigin="ZooKeys 393" ModsDocTitle="A cybertaxonomic revision of the micro-landsnail genus Plectostoma Adam (Mollusca, Caenogastropoda, Diplommatinidae), from Peninsular Malaysia, Sumatra and Indochina" checkinTime="1451246065565" checkinUser="pensoft" docAuthor="Liew, Thor-Seng, Vermeulen, Jaap Jan, Marzuki, Mohammad Effendi bin &amp; Schilthuizen, Menno" docDate="2014" docId="E573215260DDCD0E978CA9341D22D75B" docLanguage="en" docName="ZooKeys 393: 1-107" docOrigin="ZooKeys 393" docSource="http://dx.doi.org/10.3897/zookeys.393.6717" docTitle="Plectostoma crassipupa van Benthem Jutting 1952" docType="treatment" docVersion="5" lastPageNumber="40" masterDocId="A935FFF6FFE56726FFC5FFA8FFCE7A2A" masterDocTitle="A cybertaxonomic revision of the micro-landsnail genus Plectostoma Adam (Mollusca, Caenogastropoda, Diplommatinidae), from Peninsular Malaysia, Sumatra and Indochina" masterLastPageNumber="107" masterPageNumber="1" pageNumber="39" updateTime="1701373668194" updateUser="ExternalLinkService">
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<mods:title id="A7F38C53D0C60D6762A8E275F8E448F6">A cybertaxonomic revision of the micro-landsnail genus Plectostoma Adam (Mollusca, Caenogastropoda, Diplommatinidae), from Peninsular Malaysia, Sumatra and Indochina</mods:title>
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<mods:namePart id="647E3447C70401CA22123CE3401A5517">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="B671666D6559F6FF7AF0B80EDE77E264">Vermeulen, Jaap Jan</mods:namePart>
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<mods:namePart id="5845F97F40C459DB3EA1C40F38EDC002">Marzuki, Mohammad Effendi bin</mods:namePart>
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<mods:namePart id="7539481B61E5FDD94E07F2FBA82DC48D">Schilthuizen, Menno</mods:namePart>
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<treatment id="E573215260DDCD0E978CA9341D22D75B" ID-GBIF-Taxon="152052221" LSID="urn:lsid:plazi:treatment:E573215260DDCD0E978CA9341D22D75B" httpUri="http://treatment.plazi.org/id/E573215260DDCD0E978CA9341D22D75B" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<subSubSection id="8D9D3760DC5BED92F0836323C23E924E" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="7A3E0C58ECDA473392A23C24756065D5" pageId="38" pageNumber="39">
<taxonomicName id="1FC4E4912D31866C23DA705F43E5F255" ID-CoL="7SR9N" LSID="http://species-id.net/wiki/Plectostoma_crassipupa" authority="van Benthem Jutting, 1952" authorityName="van Benthem Jutting" authorityYear="1952" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma crassipupa" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="crassipupa">
<pageBreakToken id="AAFF1DF070579004BC708DDCF6141523" pageId="38" pageNumber="39" start="start">Plectostoma</pageBreakToken>
crassipupa (van Benthem Jutting, 1952)
</taxonomicName>
Figures 17I, M, 26, Appendix 6
</paragraph>
</subSubSection>
<subSubSection id="F14B00E3617D504B0060A0450F244032" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="C992E455AF671CE1AB9A4C61B3BE99D0" pageId="38" pageNumber="39">
<taxonomicName id="8A09BDF98EFD59C0EABD152BE5A7CA72" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma crassipupa" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="crassipupa">Opisthostoma crassipupa</taxonomicName>
van Benthem Jutting, 1952: 51, figure 26 (original description).
</paragraph>
<paragraph id="9A95B67D86BA261412B085F470E29968" pageId="38" pageNumber="39">
<taxonomicName id="F6C6F7C1FB43B2D6CC562879343CCBE5" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma crassipupa" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="crassipupa">Opisthostoma crassipupa</taxonomicName>
van Benthem Jutting,
<bibRefCitation id="79C60BE2ABCC13D933B3CABB2A8E3F6D" author="van Benthem Jutting, WSS" journalOrPublisher="The Bulletin of the National Museum" pageId="72" pageNumber="73" pagination="34 - 48" title="Additional new species and new localities of the family Vertiginidae and the genera Oophana and Opisthostoma from Malaya." url="http://rmbr.nus.edu.sg/rbz/biblio/26/26brm034-048.pdf" volume="26" year="1961">van Benthem Jutting (1961</bibRefCitation>
: 39).
</paragraph>
<paragraph id="3DF85919496BA6E44D472828796BC8E4" pageId="38" pageNumber="39">
<taxonomicName id="654A9776CE62BDE6C69F731DCA412521" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma crassipupa" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="crassipupa">Opisthostoma crassipupa</taxonomicName>
van Benthem Jutting,
<bibRefCitation id="DF405D647A5D793C37127BB672F18829" author="Clements, GR" journalOrPublisher="National University of Singapore" pageId="73" pageNumber="74" title="Conservation biogeography of terrestrial molluscs on tropical limestone karsts." year="2007">Clements (2007</bibRefCitation>
: 74).
</paragraph>
<paragraph id="01B0200598A2FF0A55BB70F632DB97E9" pageId="38" pageNumber="39">
<taxonomicName id="0931446975387CAAD9A37943895FDF91" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma crassipupa" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="crassipupa">Opisthostoma crassipupa</taxonomicName>
van Benthem Jutting, Clements et al. (2008: 2760).
</paragraph>
</subSubSection>
<subSubSection id="D6421E8A25FF9F2BF91F9F31EACD0883" pageId="38" pageNumber="39" type="type material">
<paragraph id="7C1EFB2C0CBD6FEBBD62CFAA58CE65CE" pageId="38" pageNumber="39">Type material.</paragraph>
<paragraph id="764B91873721C23D9F42E65F233F9F44" pageId="38" pageNumber="39">Holotype: ZMA 135994(1) (Seen). Paratype: ZMA 135993(&gt;50) (Seen).</paragraph>
</subSubSection>
<subSubSection id="99DD79C801B69F292236059AD6808924" pageId="38" pageNumber="39" type="other examined materials">
<paragraph id="17658565A2D05C3D6280B6569C3A7552" pageId="38" pageNumber="39">Other examined materials.</paragraph>
<paragraph id="229B164782634A73CB948DB493918B26" pageId="38" pageNumber="39">BOR 5512(&gt;50), BOR 5515(2), BOR 5624(&gt;10), BOR 5629(3), V 8392(&gt;10), V 8407(10), V 8408(4), V 8437(&gt;10), V 8898(7). V 8912(1), V 8956(1), V 9097(9), V 9157(&gt;10), V 9326(&gt;10), V 9353(3), V 9366(10).</paragraph>
<paragraph id="14BDD1C783DC2A2B6BA854C5E5D5F2D8" pageId="38" pageNumber="39">Genetic distance between BOR 5512 and BOR 5515 is 14%.</paragraph>
</subSubSection>
<subSubSection id="1E9C4C7FA6E1D1389985EF9F9C9EE16C" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="DF628BEBA40CB003E18180D27E5DD248" pageId="38" pageNumber="39">Diagnosis.</paragraph>
<paragraph id="750B7AE96A9722C06EB48627766C7B4E" pageId="38" pageNumber="39">
Shares with
<taxonomicName id="DB8FADFFA401BBB8655AEBB64D7F4890" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma sciaphilum" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="sciaphilum">Plectostoma sciaphilum</taxonomicName>
,
<taxonomicName id="AFC3147FA29FF3A8E3745253B79A6C9D" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma senex" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="senex">Plectostoma senex</taxonomicName>
, and
<taxonomicName id="AAF1737C6027ED2F9C90064FFC71D256" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma turriforme" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="turriforme">Plectostoma turriforme</taxonomicName>
the tuba form, but differs by having a slightly or moderately convex apex and depressed apical spire.
</paragraph>
</subSubSection>
<subSubSection id="6C19C792712E385A45CE6384BF427D62" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="63F4EA821299C3374717A3C2AB1F2630" pageId="38" pageNumber="39">Description.</paragraph>
<paragraph id="F8A219A5B60264886687548908AB1E91" pageId="38" pageNumber="39">Apex. Shape: slightly or moderately convex.</paragraph>
<paragraph id="360E3A2FB210AD83630AF1138252CBE0" pageId="38" pageNumber="39">Spire. Height: 1.3-3.0 mm. Width: 1.2-1.4 mm. Number of whorls: 3 1/2-4 1/2. Apical spire shape: depressed conical. Basal spire shape: ovoid. Whorl periphery: moderately or distinctly convex. Umbilicus: closed by tuba (common) or partially open (rare).</paragraph>
<paragraph id="A005F849FBC8EFA7B32E5DBE9BEC43C7" pageId="38" pageNumber="39">Constriction. Parietal teeth: none. Basal teeth: none.</paragraph>
<paragraph id="FB061DFC0DCAA3A6DD2C41BDA2DCD88F" pageId="38" pageNumber="39">Tuba. Coiling direction: type 3 and aperture visible from left lateral view. Tuba whorl length in proportion to spire last whorl: ca. 1/2-3/4 Proportion of tuba that attaches to spire: whole.</paragraph>
<paragraph id="20BEBEF462889E651D217C241B9BA206" pageId="39" pageNumber="40">
<pageBreakToken id="353F16152672C10C4E7B314BF1C05001" pageId="39" pageNumber="40" start="start">Aperture</pageBreakToken>
and peristome. Peristome: double peristomes. Outer peristome shape: same as inner peristome and uniformly projected all around, except the posterior part.
</paragraph>
<paragraph id="D1EA31DF070F112A4D9D2D7BEF74D368" pageId="39" pageNumber="40">Spiral lines. Thick lines: present. Thin lines: present.</paragraph>
<paragraph id="FB05CCB2441265F346B499F8415FAAD9" pageId="39" pageNumber="40">Radial ribs. Rib density: 6-8 ribs per mm. Rib intensity: thin. Shape: straight. Inclination: moderately prosoclin.</paragraph>
<caption id="F513EDDB675A178AA6C3415B0C06DFE0" pageId="39" pageNumber="40">
<paragraph id="4ED1307748F25D65A3E0E042579F01C8" pageId="39" pageNumber="40">
Figures 26.
<taxonomicName id="B4F7CD2F43FB48C491A63F51D759AAF2" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma crassipupa" order="Architaenioglossa" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="crassipupa">Plectostoma crassipupa</taxonomicName>
(van Benthem Jutting, 1952) BOR 5512. A frontal view B left lateral view C back view D left lateral view E top view F bottom view G parietal part of constriction inner whorl H basal part of constriction inner whorl. Scale bar = 1 mm (for
<normalizedToken id="0C009D9FC726150686572D2552805EAB" originalValue="AF">A-F</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="649AC3C4C18C883676DC138951165DD0" pageId="39" pageNumber="40" type="distribution">
<paragraph id="5154C1707E8B613C9950E1F4AA9CC66A" pageId="39" pageNumber="40">Distribution.</paragraph>
<paragraph id="5417A11931708CF8F4E463A5E135AD79" pageId="39" pageNumber="40">
Type locality. Gua Musang (
<geoCoordinate id="13191C4171176273F33E6B0F1A5F078D" direction="north" orientation="latitude" precision="15" value="4.8830557">4°52'59&quot;N</geoCoordinate>
,
<geoCoordinate id="E2A9E5972F278D2B46BA1AE88004B5F4" direction="east" orientation="longitude" precision="15" value="101.97">101°58'12&quot;E</geoCoordinate>
).
</paragraph>
<paragraph id="B813B4681D300CF8F61D6B84EF71DF0C" pageId="39" pageNumber="40">Distribution range. This species mainly occurs in the limestome hills that are located between Gua Musang and as far as 30 km radius of Gua Musang. One populations exists at a limestone hill that is located about 90 km north of Gua Musang (Figure 18A).</paragraph>
</subSubSection>
<subSubSection id="C92E71B35A5A870112467487E25F5091" pageId="39" pageNumber="40" type="conservation status">
<paragraph id="BAACA9775149DE398120D0C1116C9FCF" pageId="39" pageNumber="40">Conservation status.</paragraph>
<paragraph id="8D3E94120B8FD6126CE652D3DDF138D8" pageId="39" pageNumber="40">Least Concern. Almost all the limestone hills are located along main roads and/or surrounded by oil palm plantation or cleared for urban development. However, several large limestone hills that hold the species are located in the well-protected National Park (Taman Negara), State of Pahang, Malaysia.</paragraph>
</subSubSection>
<subSubSection id="0EA70F31E60FCA6DE382B8406223403A" pageId="39" pageNumber="40" type="discussion">
<paragraph id="91827ECEE272ADDB88073BBC5DE7CD29" pageId="39" pageNumber="40">Discussion.</paragraph>
<paragraph id="55B33007809452EDD75F8FC736123D38" pageId="39" pageNumber="40">The species is well-characterised, although it displays considerable variability in the shell shape (Appendix 6).</paragraph>
</subSubSection>
</treatment>
</document>