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<document id="50307286BD182A7FCBE14FD0B25CA863" ID-CLB-Dataset="21820" ID-DOI="10.5852/ejt.2012.32" ID-GBIF-Dataset="0dbfd769-022a-4f18-9a99-447e5e865f89" ID-ISSN="2118-9773" ID-Zenodo-Dep="3858570" ID-ZooBank="778FFF1D-32E1-466B-82DB-BF9B30C2ABE1" IM.bibliography_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1590542125421" checkinUser="valdenar" docAuthor="Camacho, Miguel Domínguez &amp; Vandenspiegel, Didier" docDate="2012" docId="D20C87ABAF524847DB60A638FC83FE75" docLanguage="en" docName="ejt-2012-32.pdf.imf" docOrigin="European Journal of Taxonomy 32" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Remysymphyla spinosa Camacho &amp; Vandenspiegel 2012, sp. nov." docType="treatment" docUuid="2F1D7455-D3F6-4DC5-A731-04C893267D6A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="23" masterDocId="2E35FFD3AF40485FD93CA047FFA0FFFA" masterDocTitle="Scolopendrellidae (Myriapoda, Symphyla) from the Afrotropics with descriptions of seven new species" masterLastPageNumber="28" masterPageNumber="1" pageNumber="19" updateTime="1733833526089" updateUser="julia" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="9D840260E1BAE04D093AF9CA0CA1371B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E383937652E101D9C91D5A71C8E054E2">
<mods:title id="092CFAE49D4CBBD3C21490BC90AFEC7F">Scolopendrellidae (Myriapoda, Symphyla) from the Afrotropics with descriptions of seven new species</mods:title>
</mods:titleInfo>
<mods:name id="F6371296FC806D72547BB04EA782B069" type="personal">
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</mods:role>
<mods:namePart id="BE1F7DB5B0E35DF20B679E43AA783E51">Camacho, Miguel Domínguez</mods:namePart>
<mods:nameIdentifier id="835161B0707E6E99631C17B778854438" type="LSID">866695CE-585A-4469-B59A-0FFFEFD79683</mods:nameIdentifier>
<mods:affiliation id="E8DB8A51600DB8EE92949D0608ABCCAD">Puentecesures 1 B, 6 C. 28029 Madrid (Spain). Corresponding author, Email: migueldomcam @ gmail. com &amp; urn: lsid: zoobank. org: author: 866695 CE- 585 A- 4469 - B 59 A- 0 FFFEFD 79683</mods:affiliation>
</mods:name>
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<mods:roleTerm id="FA3A3505AA1EE742291522DD7B467E57">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E7EBF7EB8B59C64BB32FB8A752D57B0">Vandenspiegel, Didier</mods:namePart>
<mods:nameIdentifier id="353A82B22495A7E00D3E01026A1146F1" type="LSID">CE8C3D01-28AD-43F7-9D4F-04802E68CB1A</mods:nameIdentifier>
<mods:affiliation id="2A300012951C2EDFF491DE50FF41A339">Invertebrate Section, Royal Museum for Central Africa. B- 3080 Tervuren (Belgium). Email: dvdspiegel @ africamuseum. be &amp; urn: lsid: zoobank. org: author: CE 8 C 3 D 01 - 28 AD- 43 F 7 - 9 D 4 F- 04802 E 68 CB 1 A</mods:affiliation>
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<mods:title id="2B2CAA6D5077015928ABD24155862073">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="39446B0DB3A34905018579C21A3CB44C">
<mods:date id="62F4AAFA9048EF6AF8BEF82A5166AB94">2012</mods:date>
<mods:detail id="5BDCBE7062B92CD3B871ECAD88574DA2" type="pubDate">
<mods:number id="81F7678D0319198C010699AB4604AB86">2012-12-13</mods:number>
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<mods:detail id="AF745DF517B5B2CC7574F189E0C0B288" type="volume">
<mods:number id="A78D6A39186571C6609BBE854778DA90">32</mods:number>
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<mods:start id="485E039A87EDBC82A45321E897944B54">1</mods:start>
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<mods:classification id="58724EF654C739737CBC850EFBDF4E9D">journal article</mods:classification>
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<treatment id="D20C87ABAF524847DB60A638FC83FE75" ID-DOI="http://doi.org/10.5281/zenodo.3859022" ID-GBIF-Taxon="164358757" ID-Zenodo-Dep="3859022" LSID="urn:lsid:zoobank.org:act:2F1D7455-D3F6-4DC5-A731-04C893267D6A" httpUri="http://treatment.plazi.org/id/D20C87ABAF524847DB60A638FC83FE75" lastPageId="24" lastPageNumber="23" pageId="18" pageNumber="19" scope_class="Symphyla" scope_family="Scolopendrellidae">
<subSubSection id="12BF6536AF52484DDB60A638FC77F960" box="[604,983,1663,1690]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="5A1A36BDAF52484DDB60A638FC77F960" blockId="18.[367,1221,1663,1760]" box="[604,983,1663,1690]" pageId="18" pageNumber="19">
<heading id="015281D1AF52484DDB60A638FC77F960" box="[604,983,1663,1690]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="9DA54D3EAF52484DDB60A638FCD5F963" ID-CoL="9B8QS" authority="Camacho &amp; Vandenspiegel, 2012" authorityName="Camacho &amp; Vandenspiegel" authorityYear="2012" box="[604,885,1663,1690]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="68D1EAAFAF52484DDB60A638FCD5F963" bold="true" box="[604,885,1663,1690]" italics="true" pageId="18" pageNumber="19">Remysymphyla spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E257D4AF52484DDA41A6C7FC77F960" box="[893,983,1664,1690]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12BF6536AF52484DD853A6E5FCF4F91A" pageId="18" pageNumber="19" type="description">
<paragraph id="5A1A36BDAF52484DD853A6E5FB65F947" blockId="18.[367,1221,1663,1760]" box="[367,1221,1698,1725]" pageId="18" pageNumber="19">
<uri id="2E343ABFAF52484DD853A6E5FB65F947" box="[367,1221,1698,1725]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="2E030C68AF52484DDBAEA6E5FB65F947" box="[658,1221,1698,1725]" pageId="18" pageNumber="19">2F1D7455-D3F6-4DC5-A731-04C893267D6A</uuid>
</uri>
</paragraph>
<paragraph id="5A1A36BDAF52484DDBE3A681FCF4F91A" blockId="18.[367,1221,1663,1760]" box="[735,852,1734,1760]" pageId="18" pageNumber="19">
<figureCitation id="C29E2A38AF52484DDBE3A681FC8AF91A" box="[735,810,1734,1760]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="18" pageNumber="19">Figs 9</figureCitation>
,
<figureCitation id="C29E2A38AF52484DDA0BA681FCF4F91A" box="[823,852,1734,1760]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="18" pageNumber="19">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="12BF6536AF52484BD981A740FD26FE82" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="5A1A36BDAF52484DD981A740FECCF8DB" blockId="18.[189,364,1799,1825]" box="[189,364,1799,1825]" pageId="18" pageNumber="19">
<heading id="015281D1AF52484DD981A740FECCF8DB" bold="true" box="[189,364,1799,1825]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="68D1EAAFAF52484DD981A740FECCF8DB" bold="true" box="[189,364,1799,1825]" pageId="18" pageNumber="19">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF52484DD9D6A771FAEEF86D" blockId="18.[189,1399,1846,1943]" pageId="18" pageNumber="19">
<materialsCitation id="EACD3CE0AF52484DD9D6A771FAEEF86D" ID-GBIF-Occurrence="2626001304" collectingDate="1959-08" collectionCode="MRAC" collectorName="Leleup N." country="Republic of the Congo" county="Sex" elevation="2500" latitude="-3.5" location="terr. D'Uvira" longLatPrecision="1307" longitude="29.0" municipality="Itombwe" pageId="18" pageNumber="19" specimenCode="MRAC 22161" specimenCount="1" stateProvince="Kivu" typeStatus="holotype">
<emphasis id="68D1EAAFAF52484DD9D6A771FEFCF8AA" bold="true" box="[234,348,1846,1872]" pageId="18" pageNumber="19">
<typeStatus id="851E881FAF52484DD9D6A771FEFCF8AA" box="[234,348,1846,1872]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
Sex unknown,
<collectingCountry id="22B2762DAF52484DD851A71EFE75F889" box="[365,469,1881,1907]" name="Democratic Republic of the Congo" pageId="18" pageNumber="19">CONGO</collectingCountry>
D.R.,
<collectingRegion id="9861F85FAF52484DDB1FA71EFDC1F889" box="[547,609,1881,1907]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="18" pageNumber="19">Kivu</collectingRegion>
,
<collectingMunicipality id="BA7EACC7AF52484DDB51A71EFD78F889" box="[621,728,1881,1907]" pageId="18" pageNumber="19">Itombwe</collectingMunicipality>
,
<location id="5F7A6066AF52484DDBD8A71EFCDFF889" LSID="urn:lsid:plazi:treatment:D20C87ABAF524847DB60A638FC83FE75:5F7A6066AF52484DDBD8A71EFCDFF889" box="[740,895,1881,1907]" country="Republic of the Congo" county="Sex" latitude="-3.5" longLatPrecision="1307" longitude="29.0" municipality="Itombwe" name="terr. D'Uvira" pageId="18" pageNumber="19" stateProvince="Kivu">terr. D’Uvira</location>
,
<location id="5F7A6066AF52484DDAB7A71EFB5BF889" LSID="urn:lsid:plazi:treatment:D20C87ABAF524847DB60A638FC83FE75:5F7A6066AF52484DDAB7A71EFB5BF889" box="[907,1275,1881,1907]" country="Republic of the Congo" county="Sex" latitude="-3.5" longLatPrecision="1307" longitude="29.0" municipality="Itombwe" name="tete de source de la Nyalengwe" pageId="18" pageNumber="19" stateProvince="Kivu">tête de source de la Nyalengwe</location>
(
<geoCoordinate id="3F91507AAF52484DDC37A71EFAD1F889" box="[1291,1393,1881,1907]" degrees="03" direction="south" minutes="30" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="-3.5">03º30’ S</geoCoordinate>
,
<geoCoordinate id="3F91507AAF52484DD981A73BFE93F86D" box="[189,307,1916,1943]" degrees="029" direction="east" minutes="00" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="29.0">029º00’ E</geoCoordinate>
), in a mountain forest with bamboo,
<quantity id="9D5D9B58AF52484DDBD8A73BFC9FF86D" box="[740,831,1916,1943]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="m" value="2500.0">
<elevation id="D188D18EAF52484DDBD8A73BFC9FF86D" box="[740,831,1916,1943]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="m" value="2500.0">2500 m</elevation>
</quantity>
, coll.
<collectorName id="F750536BAF52484DDABAA73BFC5DF86D" box="[902,1021,1916,1943]" pageId="18" pageNumber="19">Leleup N.</collectorName>
,
<date id="2E1B107DAF52484DDD35A73BFB25F86D" box="[1033,1157,1916,1943]" pageId="18" pageNumber="19" value="1959-08">
<collectingDate id="3E5FE995AF52484DDD35A73BFB25F86D" box="[1033,1157,1916,1943]" pageId="18" pageNumber="19" value="1959-08">Aug. 1959</collectingDate>
</date>
(
<specimenCode id="0A039EC6AF52484DDDA8A73BFAE1F86D" box="[1172,1345,1916,1943]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="18" pageNumber="19" type="Museum">MRAC 22161</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5A1A36BDAF52484DD9D6A7F9FD49F806" blockId="18.[189,745,1982,2044]" pageId="18" pageNumber="19">
<materialsCitation id="EACD3CE0AF52484DD9D6A7F9FD49F806" collectionCode="MRAC" pageId="18" pageNumber="19" specimenCode="MRAC 22162" specimenCount="2" typeStatus="paratype">
<emphasis id="68D1EAAFAF52484DD9D6A7F9FEC7F822" bold="true" box="[234,359,1982,2008]" pageId="18" pageNumber="19">
<typeStatus id="851E881FAF52484DD9D6A7F9FEC7F822" box="[234,359,1982,2008]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="4CA3FD34AF52484DD981A7A6FF6EF801" box="[189,206,2017,2043]" pageId="18" pageNumber="19">2</specimenCount>
, sex?, same data as holotype (
<specimenCode id="0A039EC6AF52484DDB12A7A6FD7DF806" box="[558,733,2017,2044]" pageId="18" pageNumber="19">MRAC 22162</specimenCode>
).
</materialsCitation>
</paragraph>
<caption id="0EDA6635AF53484CD981A680FCD6F823" ID-DOI="http://doi.org/10.5281/zenodo.3858600" ID-Zenodo-Dep="3858600" httpUri="https://zenodo.org/record/3858600/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1735,1761]" targetBox="[189,1398,265,1713]" targetPageId="19">
<paragraph id="5A1A36BDAF53484CD981A680FCD6F823" blockId="19.[189,1399,1735,2009]" pageId="19" pageNumber="20">
<emphasis id="68D1EAAFAF53484CD981A680FEACF918" bold="true" box="[189,268,1735,1762]" pageId="19" pageNumber="20">Fig. 8.</emphasis>
<taxonomicName id="9DA54D3EAF53484CD82AA68FFC8FF918" authority="Aubry &amp; Masson, 1952" authorityName="Aubry &amp; Masson" authorityYear="1952" box="[278,815,1735,1762]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="68D1EAAFAF53484CD82AA68FFDA8F918" box="[278,520,1736,1762]" italics="true" pageId="19" pageNumber="20">Remysymphyla hova</emphasis>
Aubry &amp; Masson, 1952
</taxonomicName>
. Paratypes (MNHN K002) from Ambanja (A-D) and Ambatolampy (E and F), Madagascar.
<emphasis id="68D1EAAFAF53484CDB8BA6ACFD6CF8FF" bold="true" box="[695,716,1771,1797]" pageId="19" pageNumber="20">A</emphasis>
. Head and anterior half of the trunk, dorsal view.
<emphasis id="68D1EAAFAF53484CDC27A6ACFAD7F8FF" box="[1307,1399,1771,1797]" italics="true" pageId="19" pageNumber="20">T1-T11,</emphasis>
tergites 1-11;
<emphasis id="68D1EAAFAF53484CD858A749FE6CF8D2" box="[356,460,1806,1832]" italics="true" pageId="19" pageNumber="20">TA1-TA9</emphasis>
, tergal areas 1-9 (
<emphasis id="68D1EAAFAF53484CDB97A749FD75F8D2" box="[683,725,1806,1832]" italics="true" pageId="19" pageNumber="20">fide</emphasis>
<bibRefCitation id="3E344B4CAF53484CDBE3A749FB89F8D2" author="Dominguez Camacho" box="[735,1065,1806,1832]" pageId="19" pageNumber="20" refId="ref11856" refString="Dominguez Camacho, M. 2009. Phylogeny of the Symphyla. PhD Thesis, Freie Universitat Berlin. http: // www. diss. fu-berlin. de / diss / servlets / MCRFileNodeServlet / FUDISS _ derivate _ 000000007088 /" type="url" year="2009">Domínguez Camacho 2009</bibRefCitation>
, fig. 4, page 108). Doubled labeling of tergites shows partial sclerotization.
<emphasis id="68D1EAAFAF53484CDBD1A775FCA1F8B6" bold="true" box="[749,769,1842,1868]" pageId="19" pageNumber="20">B</emphasis>
. Head and first pair of legs, ventral view.
<emphasis id="68D1EAAFAF53484CDDD4A776FA80F8B6" bold="true" box="[1256,1312,1841,1868]" pageId="19" pageNumber="20">C, D</emphasis>
. Detail of the first leg of two different specimens.
<emphasis id="68D1EAAFAF53484CDB88A712FD68F895" bold="true" box="[692,712,1877,1903]" pageId="19" pageNumber="20">E</emphasis>
. Cerci (
<emphasis id="68D1EAAFAF53484CDA14A712FC9CF895" box="[808,828,1877,1903]" italics="true" pageId="19" pageNumber="20">C</emphasis>
) last tergal areas (
<emphasis id="68D1EAAFAF53484CDD2AA712FB00F895" box="[1046,1184,1877,1903]" italics="true" pageId="19" pageNumber="20">TA14, TA15</emphasis>
) (
<emphasis id="68D1EAAFAF53484CDD87A713FB45F894" box="[1211,1253,1876,1902]" italics="true" pageId="19" pageNumber="20">fide</emphasis>
<bibRefCitation id="3E344B4CAF53484CDDD2A712FED1F868" author="Dominguez Camacho" pageId="19" pageNumber="20" refId="ref11856" refString="Dominguez Camacho, M. 2009. Phylogeny of the Symphyla. PhD Thesis, Freie Universitat Berlin. http: // www. diss. fu-berlin. de / diss / servlets / MCRFileNodeServlet / FUDISS _ derivate _ 000000007088 /" type="url" year="2009">Domínguez Camacho 2009</bibRefCitation>
, fig. 4, page 108) corresponding to tergites 18-21 (
<emphasis id="68D1EAAFAF53484CDAFAA73FFB8BF868" box="[966,1067,1912,1938]" italics="true" pageId="19" pageNumber="20">T18-T21</emphasis>
), dorsal view. Only exterior parts of tergites 17 and 18 are sclerotized (doubled labeled).
<emphasis id="68D1EAAFAF53484CDAB8A7DCFC36F84F" bold="true" box="[900,918,1947,1973]" pageId="19" pageNumber="20">F</emphasis>
. Last tergite (
<emphasis id="68D1EAAFAF53484CDD07A7DCFBC9F84F" box="[1083,1129,1947,1973]" italics="true" pageId="19" pageNumber="20">T21</emphasis>
) and cerci (
<emphasis id="68D1EAAFAF53484CDDCAA7DCFAAAF84F" box="[1270,1290,1947,1973]" italics="true" pageId="19" pageNumber="20">C</emphasis>
) of other specimen.
<emphasis id="68D1EAAFAF53484CD806A7F8FE2FF823" box="[314,399,1983,2009]" italics="true" pageId="19" pageNumber="20">Arrows</emphasis>
, diagonal basis of the terminal areas (
<emphasis id="68D1EAAFAF53484CDA72A7F8FCC5F822" box="[846,869,1983,2008]" italics="true" pageId="19" pageNumber="20">ta</emphasis>
).
</paragraph>
</caption>
<paragraph id="5A1A36BDAF54484BD981A14BFEFDFEDF" blockId="20.[189,349,267,294]" box="[189,349,267,294]" pageId="20" pageNumber="21">
<heading id="015281D1AF54484BD981A14BFEFDFEDF" bold="true" box="[189,349,267,294]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="68D1EAAFAF54484BD981A14BFEFDFEDF" bold="true" box="[189,349,267,294]" pageId="20" pageNumber="21">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A17DFD26FE82" blockId="20.[189,1398,314,376]" pageId="20" pageNumber="21">
<collectingCountry id="22B2762DAF54484BD981A17DFE85FEAE" box="[189,293,314,340]" name="Democratic Republic of the Congo" pageId="20" pageNumber="21">CONGO</collectingCountry>
D.R.,
<collectingRegion id="9861F85FAF54484BD84EA17DFE10FEAE" box="[370,432,314,340]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="20" pageNumber="21">Kivu</collectingRegion>
, Itombwe, terr. D’Uvira, tête de source de la Nyalengwe (
<geoCoordinate id="3F91507AAF54484BDD6CA17DFB15FEAE" box="[1104,1205,314,340]" degrees="03" direction="south" minutes="30" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-3.5">03º30’ S</geoCoordinate>
,
<geoCoordinate id="3F91507AAF54484BDDFDA17DFA97FEAE" box="[1217,1335,314,340]" degrees="029" direction="east" minutes="00" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="29.0">029º00’ E</geoCoordinate>
), in a mountain forest with bamboo,
<quantity id="9D5D9B58AF54484BDB1AA11AFD22FE82" box="[550,642,349,376]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="m" value="2500.0">2500 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="12BF6536AF54484BD981A1D8FE65FDF1" pageId="20" pageNumber="21" type="etymology">
<paragraph id="5A1A36BDAF54484BD981A1D8FEE4FE43" blockId="20.[189,324,415,441]" box="[189,324,415,441]" pageId="20" pageNumber="21">
<heading id="015281D1AF54484BD981A1D8FEE4FE43" bold="true" box="[189,324,415,441]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="68D1EAAFAF54484BD981A1D8FEE4FE43" bold="true" box="[189,324,415,441]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A18AFE65FDF1" blockId="20.[189,1398,461,523]" pageId="20" pageNumber="21">The species name is an adjective emphasizing the long and erect setae of the tergites, which give this species a spiny aspect.</paragraph>
</subSubSection>
<subSubSection id="12BF6536AF54484BD981A276FBC3F843" pageId="20" pageNumber="21" type="description">
<paragraph id="5A1A36BDAF54484BD981A276FE74FDB1" blockId="20.[189,468,561,587]" box="[189,468,561,587]" pageId="20" pageNumber="21">
<heading id="015281D1AF54484BD981A276FE74FDB1" bold="true" box="[189,468,561,587]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="68D1EAAFAF54484BD981A276FE74FDB1" bold="true" box="[189,468,561,587]" pageId="20" pageNumber="21">
Description (
<typeStatus id="851E881FAF54484BD861A276FE6CFDB1" box="[349,460,561,587]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A226FE25FD81" blockId="20.[189,389,608,635]" box="[189,389,608,635]" pageId="20" pageNumber="21">
LENGTH.
<quantity id="9D5D9B58AF54484BD81FA227FE20FD81" box="[291,384,608,635]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="21" unit="mm" value="2.1">2.1 mm</quantity>
.
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A2E5FE1EFCB3" blockId="20.[189,1399,674,841]" pageId="20" pageNumber="21">
HEAD. 1.3 times as long as broad with broadest part behind the middle on a level with lateral protuberances (
<figureCitation id="C29E2A38AF54484BD9FAA282FE84FD25" box="[198,292,709,735]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
). Lateral margins nearly straight. Central rod well-developed and not broken. Frontal branches also well-developed. Median branches not very distinct in the
<typeStatus id="851E881FAF54484BDAF8A2AFFB8EFCF8" box="[964,1070,744,770]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
, but clearly distinct in the
<typeStatus id="851E881FAF54484BD981A34BFE8DFCDC" box="[189,301,780,806]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
(
<figureCitation id="C29E2A38AF54484BD802A34CFE3CFCDF" box="[318,412,779,805]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9B</figureCitation>
). Dorsal surface with very long and spiny setae of equal length. Cuticle with dense granulation (
<figureCitation id="C29E2A38AF54484BD86FA369FE11FCB3" box="[339,433,814,841]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9G</figureCitation>
).
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A337FE44FB7A" blockId="20.[189,1399,880,1152]" pageId="20" pageNumber="21">
ANTENNAE. Broken in
<typeStatus id="851E881FAF54484BD8FCA337FD8BFC70" box="[448,555,880,906]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
. Rather short, in
<typeStatus id="851E881FAF54484BDBCBA337FCFCFC70" box="[759,860,880,906]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
left antenna with 27 segments, right with 29, (
<figureCitation id="C29E2A38AF54484BD9FAA3D4FE81FC57" box="[198,289,915,941]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9C</figureCitation>
). All segments of equal length and pretty broad, 1.8-1.9 times as wide as long. Antennae densely setose, with a spiny appearance; setae erect and long. Longest setae in the inner part of the median segments, with length of 0.5-0.7 times the diameter of segments. Secondary whorl of setae appears in the 6
<superScript id="ADD09BF5AF54484BD9C6A3BCFEB0FBED" attach="none" box="[250,272,1019,1047]" fontSize="6" pageId="20" pageNumber="21">th-</superScript>
8
<superScript id="ADD09BF5AF54484BD81CA3BCFE8DFBF3" attach="left" box="[288,301,1019,1033]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
segments. Size and number of setae decreases gradually in the seven distal segments, and are totally absent in the last three ones. Apical segment flattened, more than 3 times as wide as long. It presents a transversal depression, bestowing a coffee-bean shape. All segments with pretty long and conspicuous pubescence.
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A4EFFE54FA21" blockId="20.[189,1399,1192,1500]" pageId="20" pageNumber="21">
TERGITES. Not all well-visible in the studied specimens, at least 17 can be counted in the
<typeStatus id="851E881FAF54484BDDDBA4EFFAF1FB38" box="[1255,1361,1192,1218]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
, of which 13 present triangular processes (
<figureCitation id="C29E2A38AF54484BDBBAA48CFD49FB1F" box="[646,745,1227,1253]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Figs 9A</figureCitation>
,
<figureCitation id="C29E2A38AF54484BDBC8A48CFC8BFB1F" box="[756,811,1227,1253]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">10A</figureCitation>
). All tergites – except the first and the two latest – sclerotized only in the lateral parts. All tergal setae very long; shortest setae at least as long as triangular processes. Triangular processes longer than broad in the anterior tergites, with digitiform ends and no apical setae; processes of the posterior tergites thinner and reduced to small digitiform projections (
<figureCitation id="C29E2A38AF54484BDC7BA573FF52FA88" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10C</figureCitation>
). First tergite with six setae arranged in two groups of three. Second tergite comparatively very large (
<figureCitation id="C29E2A38AF54484BD836A53CFE2FFA6F" box="[266,399,1403,1429]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9A, E</figureCitation>
), with five posteromarginal, one central and six lateromarginal setae. Anterolateral setae not longer than the rest. All tergites with pubescence. Last tergite subelliptical, with posterior margin convex (
<figureCitation id="C29E2A38AF54484BD840A586FE47FA21" box="[380,487,1473,1499]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A644FD12F97C" blockId="20.[189,1398,1539,1671]" pageId="20" pageNumber="21">
LEGS. First pair of legs reduced to two oviform knobs densely covered by numerous spines (
<figureCitation id="C29E2A38AF54484BDDD3A644FAC9F9E7" box="[1263,1385,1539,1565]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9F, H</figureCitation>
). Two longer setae on the laterals, about twice as long as the whole knob. Last pair of legs with thick and long setae at the distal part of the joints (
<figureCitation id="C29E2A38AF54484BDBA5A60EFC86F999" box="[665,806,1609,1635]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10B, C</figureCitation>
). Femur and tibia with three setae; tarsus with two protruding dorsal setae. Cuticle pubescent.
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A6E9FC1AF932" blockId="20.[189,954,1710,1736]" box="[189,954,1710,1736]" pageId="20" pageNumber="21">
STYLI. Small, elongate and spike-like, covered by hairs (
<figureCitation id="C29E2A38AF54484BDA73A6E9FC0DF932" box="[847,941,1710,1736]" captionStart="Fig" captionStartId="21.[189,232,1734,1760]" captionTargetBox="[263,1318,265,1713]" captionTargetId="figure@21.[263,1318,265,1713]" captionTargetPageId="21" captionText="Fig. 9. Remysymphyla spinosa sp. nov. A. Head and first 3 tergites (T1, T2, T3) of holotype, dorsal view. Arrows, digitiform end of triangular processes; al, anterolateral setae; cr, central rod; fb, frontal branches; lp, lateral protuberance of the head; mb, median branches. B. Head and first tergite (T1) of paratype, dorsal view. Cr, central rod; lp, lateral protuberance of the head; mb, median branches. C. Antennae of paratype. Arrows, transversal depression of apical segments. D. Stylus of the leg 11 (arrow) of paratype. E. Detail of first (T1) and second (T2) tergites of holotype. Arrows, digitiform end of triangular processes; al, anterolateral seta; cs, central seta. F. Head and first legs of paratype, lateral view. G. Detail of the surface of the head of paratype. H. Detail of the first leg of paratype." figureDoi="http://doi.org/10.5281/zenodo.3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="20" pageNumber="21">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="5A1A36BDAF54484BD981A6A8FBC3F843" blockId="20.[189,1399,1775,1978]" pageId="20" pageNumber="21">
CERCI. 2.8 times as long as wide, with straight margins (
<figureCitation id="C29E2A38AF54484BDA66A6A8FC4AF8F3" box="[858,1002,1775,1801]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10B, C</figureCitation>
). About 25 visible setae from the dorsal view, slightly curved and depressed. Cuticle with faint pubescence. Limit between cerci
<emphasis id="68D1EAAFAF54484BDC09A754FEA8F8B5" italics="true" pageId="20" pageNumber="21">sensu stricto</emphasis>
and terminal areas not transversal, but diagonal, as a continuation of the outer margin towards the inner margin (
<figureCitation id="C29E2A38AF54484BD858A71EFE70F889" box="[356,464,1881,1907]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10B</figureCitation>
). Terminal area small, 0.1 times as long as the total length of the cerci, conical, longer in the outer margin than in the inner with six diagonal striae, parallel to the basis of the terminal area. Apical seta about as long as terminal areas, curved inwards (
<figureCitation id="C29E2A38AF54484BDAF4A7D8FBF6F843" box="[968,1110,1951,1977]" captionStart="Fig" captionStartId="22.[189,232,1756,1782]" captionTargetBox="[189,1398,620,1739]" captionTargetId="figure@22.[189,1398,620,1739]" captionTargetPageId="22" captionText="Fig. 10. Remysymphyla spinosa sp. nov. A. Posterior part of the trunk of the holotype, tegal areas 8-15 (TA8-TA15) (fide Domínguez Camacho 2009: 108, fig. 4), corresponding to tergites 9-21 (T9-T21). Doubled labeling of tergites shows partial sclerotization. B. Last tergite (T21), last pair of legs and cerci (C) of holotype, dorsal view. Arrows, diagonal limit between cerci sensu stricto and terminal areas (tar); as, apical seta; f, femur; t, tibia; ta, tarsus of the last right leg. C. Last tergites, last pair of legs and cerci (C) of holotype, lateral view. Arrow, digitiform projection of tergite 19; as, apical seta; f, femur; t, tibia; ta, tarsus of the last left leg; tar, terminal area of the cerci." figureDoi="http://doi.org/10.5281/zenodo.3858608" httpUri="https://zenodo.org/record/3858608/files/figure.png" pageId="20" pageNumber="21">Fig. 10B, C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="0EDA6635AF55484AD981A681FB48F82D" ID-DOI="http://doi.org/10.5281/zenodo.3858604" ID-Zenodo-Dep="3858604" httpUri="https://zenodo.org/record/3858604/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1734,1760]" targetBox="[263,1318,265,1713]" targetPageId="21">
<paragraph id="5A1A36BDAF55484AD981A681FB48F82D" blockId="21.[189,1399,1734,2007]" pageId="21" pageNumber="22">
<emphasis id="68D1EAAFAF55484AD981A681FEAEF91A" bold="true" box="[189,270,1734,1760]" pageId="21" pageNumber="22">Fig. 9.</emphasis>
<taxonomicName id="9DA54D3EAF55484AD824A681FD8DF91A" authority="Camacho &amp; Vandenspiegel, 2012" authorityName="Camacho &amp; Vandenspiegel" authorityYear="2012" box="[280,557,1734,1760]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="68D1EAAFAF55484AD824A681FD8DF91A" box="[280,557,1734,1760]" italics="true" pageId="21" pageNumber="22">Remysymphyla spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E257D4AF55484ADB0BA681FD35F91A" box="[567,661,1734,1760]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="68D1EAAFAF55484ADBA3A681FD14F91A" bold="true" box="[671,692,1734,1760]" pageId="21" pageNumber="22">A</emphasis>
. Head and first 3 tergites (
<emphasis id="68D1EAAFAF55484ADAC2A681FBDFF91A" box="[1022,1151,1734,1760]" italics="true" pageId="21" pageNumber="22">T1, T2, T3</emphasis>
) of holotype, dorsal view.
<emphasis id="68D1EAAFAF55484AD838A6AEFEF9F8F9" box="[260,345,1769,1795]" italics="true" pageId="21" pageNumber="22">Arrows</emphasis>
, digitiform end of triangular processes;
<emphasis id="68D1EAAFAF55484ADA06A6AEFCF1F8F9" box="[826,849,1769,1795]" italics="true" pageId="21" pageNumber="22">al</emphasis>
, anterolateral setae;
<emphasis id="68D1EAAFAF55484ADD74A6ADFBC0F8F9" box="[1096,1120,1770,1795]" italics="true" pageId="21" pageNumber="22">cr</emphasis>
, central rod;
<emphasis id="68D1EAAFAF55484ADC3DA6AEFAB8F8F9" box="[1281,1304,1769,1795]" italics="true" pageId="21" pageNumber="22">fb</emphasis>
, frontal branches;
<emphasis id="68D1EAAFAF55484AD805A74BFEF0F8DC" box="[313,336,1804,1830]" italics="true" pageId="21" pageNumber="22">lp</emphasis>
, lateral protuberance of the head;
<emphasis id="68D1EAAFAF55484ADBC7A74BFCBFF8DC" box="[763,799,1804,1830]" italics="true" pageId="21" pageNumber="22">mb</emphasis>
, median branches.
<emphasis id="68D1EAAFAF55484ADD33A74BFB83F8DC" bold="true" box="[1039,1059,1804,1830]" pageId="21" pageNumber="22">B</emphasis>
. Head and first tergite (
<emphasis id="68D1EAAFAF55484ADC72A74BFACDF8DC" box="[1358,1389,1804,1830]" italics="true" pageId="21" pageNumber="22">T1</emphasis>
) of paratype, dorsal view.
<emphasis id="68D1EAAFAF55484AD8CDA777FDB0F8B0" box="[497,528,1840,1866]" italics="true" pageId="21" pageNumber="22">Cr</emphasis>
, central rod;
<emphasis id="68D1EAAFAF55484ADB8AA768FD6DF8B3" box="[694,717,1839,1865]" italics="true" pageId="21" pageNumber="22">lp</emphasis>
, lateral protuberance of the head;
<emphasis id="68D1EAAFAF55484ADD4DA768FB35F8B3" box="[1137,1173,1839,1865]" italics="true" pageId="21" pageNumber="22">mb</emphasis>
, median branches.
<emphasis id="68D1EAAFAF55484AD981A715FF72F896" bold="true" box="[189,210,1874,1900]" pageId="21" pageNumber="22">C</emphasis>
. Antennae of paratype.
<emphasis id="68D1EAAFAF55484AD8CAA714FDEBF897" box="[502,587,1875,1901]" italics="true" pageId="21" pageNumber="22">Arrows</emphasis>
, transversal depression of apical segments.
<emphasis id="68D1EAAFAF55484ADD5DA714FBD6F897" bold="true" box="[1121,1142,1875,1901]" pageId="21" pageNumber="22">D</emphasis>
. Stylus of the leg 11 (
<emphasis id="68D1EAAFAF55484AD9FBA730FEAEF86A" box="[199,270,1911,1936]" italics="true" pageId="21" pageNumber="22">arrow</emphasis>
) of paratype.
<emphasis id="68D1EAAFAF55484AD88EA731FE66F86A" bold="true" box="[434,454,1910,1936]" pageId="21" pageNumber="22">E</emphasis>
. Detail of first (
<emphasis id="68D1EAAFAF55484ADBBEA731FD01F86A" box="[642,673,1910,1936]" italics="true" pageId="21" pageNumber="22">T1</emphasis>
) and second (
<emphasis id="68D1EAAFAF55484ADA7BA731FCC5F86A" box="[839,869,1910,1936]" italics="true" pageId="21" pageNumber="22">T2</emphasis>
) tergites of holotype.
<emphasis id="68D1EAAFAF55484ADD56A731FB1FF86A" box="[1130,1215,1910,1936]" italics="true" pageId="21" pageNumber="22">Arrows</emphasis>
, digitiform end of triangular processes;
<emphasis id="68D1EAAFAF55484AD8E4A7DEFE4FF849" box="[472,495,1945,1971]" italics="true" pageId="21" pageNumber="22">al</emphasis>
, anterolateral seta;
<emphasis id="68D1EAAFAF55484ADBEAA7DDFD4EF849" box="[726,750,1946,1971]" italics="true" pageId="21" pageNumber="22">cs</emphasis>
, central seta.
<emphasis id="68D1EAAFAF55484ADAAEA7DEFC04F849" bold="true" box="[914,932,1945,1971]" pageId="21" pageNumber="22">F</emphasis>
. Head and first legs of paratype, lateral view.
<emphasis id="68D1EAAFAF55484AD83FA7FBFEBAF82C" bold="true" box="[259,282,1980,2006]" pageId="21" pageNumber="22">G</emphasis>
. Detail of the surface of the head of paratype.
<emphasis id="68D1EAAFAF55484ADA07A7FBFCF2F82C" bold="true" box="[827,850,1980,2006]" pageId="21" pageNumber="22">H</emphasis>
. Detail of the first leg of paratype.
</paragraph>
</caption>
<subSubSection id="12BF6536AF564849D981A14CFBF1FE60" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="5A1A36BDAF564849D981A14CFDD4FEDF" blockId="22.[189,628,267,293]" box="[189,628,267,293]" pageId="22" pageNumber="23">
<heading id="015281D1AF564849D981A14CFDD4FEDF" bold="true" box="[189,628,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="68D1EAAFAF564849D981A14CFDD4FEDF" bold="true" box="[189,628,267,293]" pageId="22" pageNumber="23">Affinities and differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF564849D981A17EFBF1FE60" blockId="22.[189,1399,313,410]" pageId="22" pageNumber="23">
The morphology of the first pair of legs, which does not present evidence of several joints, makes
<taxonomicName id="9DA54D3EAF564849D981A11AFE6FFE8C" authority="Camacho &amp; Vandenspiegel, 2012" authorityName="Camacho &amp; Vandenspiegel" authorityYear="2012" box="[189,463,349,375]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="68D1EAAFAF564849D981A11AFE6FFE8C" box="[189,463,349,375]" italics="true" pageId="22" pageNumber="23">Remysymphyla spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E257D4AF564849D8EBA11AFD92FE8D" box="[471,562,349,375]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
more similar to
<taxonomicName id="9DA54D3EAF564849DBCAA11AFBD2FE8D" authority="Scheller, 1971" authorityName="Scheller" authorityYear="1971" box="[758,1138,348,375]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hebetocornuta">
<emphasis id="68D1EAAFAF564849DBCAA11AFC61FE8D" box="[758,961,349,375]" italics="true" pageId="22" pageNumber="23">R. hebetocornuta</emphasis>
<bibRefCitation id="3E344B4CAF564849DAF5A11BFBD2FE8D" author="Scheller U." box="[969,1138,348,375]" pageId="22" pageNumber="23" pagination="98 - 187" refId="ref12284" refString="Scheller U. 1971. Symphyla from Ceylon and Peninsular India. Entomologica Scandinavica, Supplement 1: 98 - 187." type="journal article" year="1971">Scheller, 1971</bibRefCitation>
</taxonomicName>
than to the remaining African species. The spiny aspect of the body is characteristic for this species.
</paragraph>
</subSubSection>
<subSubSection id="12BF6536AF564847D981A186FC83FE75" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="discussion">
<paragraph id="5A1A36BDAF564849D981A186FE8FFE21" blockId="22.[189,303,449,475]" box="[189,303,449,475]" pageId="22" pageNumber="23">
<heading id="015281D1AF564849D981A186FE8FFE21" bold="true" box="[189,303,449,475]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="68D1EAAFAF564849D981A186FE8FFE21" bold="true" box="[189,303,449,475]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="5A1A36BDAF564847D981A1B7FC83FE75" blockId="22.[189,1399,496,593]" lastBlockId="24.[189,1399,267,399]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
The genus attribution of this species has been problematic, mainly due to the above-explained confusion regarding the definition of
<taxonomicName id="9DA54D3EAF564849D8C6A254FD0AFDD7" authorityName="Aubry &amp; Masson" authorityYear="1952" box="[506,682,531,557]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="68D1EAAFAF564849D8C6A254FD0AFDD7" box="[506,682,531,557]" italics="true" pageId="22" pageNumber="23">Remysymphyla</emphasis>
</taxonomicName>
. The study of
<typeStatus id="851E881FAF564849DA6BA254FC2AFDD7" box="[855,906,531,557]" pageId="22" pageNumber="23">type</typeStatus>
specimens of
<taxonomicName id="9DA54D3EAF564849DD08A254FEDFFDAB" authority="Aubry &amp; Masson, 1952" authorityName="Aubry &amp; Masson" authorityYear="1952" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="68D1EAAFAF564849DD08A254FA85FDD7" box="[1076,1317,531,557]" italics="true" pageId="22" pageNumber="23">Remysymphyla hova</emphasis>
Aubry &amp; Masson, 1952
</taxonomicName>
allowed the recognition of the first pair of legs of
<taxonomicName id="9DA54D3EAF564849DAF7A270FBE5FDAA" authority="Camacho &amp; Vandenspiegel, 2012" authorityName="Camacho &amp; Vandenspiegel" authorityYear="2012" box="[971,1093,567,592]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="68D1EAAFAF564849DAF7A270FBE5FDAA" box="[971,1093,567,592]" italics="true" pageId="22" pageNumber="23">R. spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E257D4AF564849DD70A270FB06FDAB" box="[1100,1190,567,593]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
as typical for this genus (
<figureCitation id="C29E2A38AF584847D82EA14CFEFCFEDF" box="[274,348,267,293]" captionStart="Fig" captionStartId="19.[189,232,1735,1761]" captionTargetBox="[189,1398,265,1713]" captionTargetId="figure@19.[189,1398,265,1713]" captionTargetPageId="19" captionText="Fig. 8. Remysymphyla hova Aubry &amp; Masson, 1952. Paratypes (MNHN K002) from Ambanja (A-D) and Ambatolampy (E and F), Madagascar. A. Head and anterior half of the trunk, dorsal view. T1-T11, tergites 1-11; TA1-TA9, tergal areas 1-9 (fide Domínguez Camacho 2009, fig. 4, page 108). Doubled labeling of tergites shows partial sclerotization. B. Head and first pair of legs, ventral view. C, D. Detail of the first leg of two different specimens. E. Cerci (C) last tergal areas (TA14, TA15) (fide Domínguez Camacho 2009, fig. 4, page 108) corresponding to tergites 18-21 (T18-T21), dorsal view. Only exterior parts of tergites 17 and 18 are sclerotized (doubled labeled). F. Last tergite (T21) and cerci (C) of other specimen. Arrows, diagonal basis of the terminal areas (ta)." figureDoi="http://doi.org/10.5281/zenodo.3858600" httpUri="https://zenodo.org/record/3858600/files/figure.png" pageId="24" pageNumber="25">Figs 8</figureCitation>
B-D; 9F, H). However, observations using SEM seems to be suboptimal for detection of one of its major diagnostic characters: the number of joints on the first pair of legs. Although no joints were discernible in this species, the shape of the oviforms knobs of
<taxonomicName id="9DA54D3EAF584847DAD7A114FBC7FE96" authority="Camacho &amp; Vandenspiegel, 2012" authorityName="Camacho &amp; Vandenspiegel" authorityYear="2012" box="[1003,1127,339,364]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="68D1EAAFAF584847DAD7A114FBC7FE96" box="[1003,1127,339,364]" italics="true" pageId="24" pageNumber="25">R. spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E257D4AF584847DD4CA115FB6CFE96" box="[1136,1228,338,364]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
resembles the two-jointed legs of
<taxonomicName id="9DA54D3EAF584847D89EA131FCBFFE75" authority="Scheller, 1971" authorityName="Scheller" authorityYear="1971" box="[418,799,373,399]" class="Symphyla" family="Scolopendrellidae" genus="Remysymphyla" higherTaxonomySource="CoL" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hebetocornuta">
<emphasis id="68D1EAAFAF584847D89EA131FDCBFE75" box="[418,619,373,399]" italics="true" pageId="24" pageNumber="25">R. hebetocornuta</emphasis>
<bibRefCitation id="3E344B4CAF584847DB4FA132FCBFFE75" author="Scheller U." box="[627,799,373,399]" pageId="24" pageNumber="25" pagination="98 - 187" refId="ref12284" refString="Scheller U. 1971. Symphyla from Ceylon and Peninsular India. Entomologica Scandinavica, Supplement 1: 98 - 187." type="journal article" year="1971">Scheller, 1971</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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