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<document id="2FA0F03E33F4A4723636D64E72B76715" ID-DOI="10.1649/0010-065x-69.2.183" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104758" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630169988497" checkinUser="diego" docAuthor="Ratcliffe, Brett C." docDate="2015" docId="0389C677FFAC1673FD09EA8F1F66F99C" docLanguage="en" docName="0010-065X-69.2.183.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-69.2.183" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Marmarina parvula Ratcliffe 2015, new species" docType="treatment" docVersion="3" lastPageNumber="195" masterDocId="FFB0BE0FFFA6167FFFE8E9401B31FF5D" masterDocTitle="A Review of the Neotropical GenusMarmarinaKirby, 1827 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini), with Description of a New Species from Argentina" masterLastPageNumber="201" masterPageNumber="183" pageNumber="193" updateTime="1699630745546" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="99B8FEC8723B5A6C148E3919FD8B2D0A">A Review of the Neotropical GenusMarmarinaKirby, 1827 (Coleoptera: Scarabaeidae: Cetoniinae: Gymnetini), with Description of a New Species from Argentina</mods:title>
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<mods:namePart id="03323CD0232849F7BD5873F82F1C79A3">Ratcliffe, Brett C.</mods:namePart>
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<mods:title id="4D8E03803C20C50FBDECA94AD4615E03">The Coleopterists Bulletin</mods:title>
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<mods:date id="530171ECF2D169F2A4B8A1C0D5C05054">2015</mods:date>
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<mods:number id="5D4A1E5026D3C8A2DD65FF79AB14F08C">2015-06-30</mods:number>
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<mods:number id="303831F4A9E2ADDEF714B633AC693D90">69</mods:number>
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<mods:classification id="E59C8847EA1FAEF3C823678A6D6E2EFE">journal article</mods:classification>
<mods:identifier id="A0556F4BCDA3D46CEBF406DBD0F81F7F" type="DOI">10.1649/0010-065x-69.2.183</mods:identifier>
<mods:identifier id="57FD98D91430C0FA0C3F26DD1C731EFF" type="ISSN">1938-4394</mods:identifier>
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<treatment id="0389C677FFAC1673FD09EA8F1F66F99C" LSID="urn:lsid:plazi:treatment:0389C677FFAC1673FD09EA8F1F66F99C" httpUri="http://treatment.plazi.org/id/0389C677FFAC1673FD09EA8F1F66F99C" lastPageId="12" lastPageNumber="195" pageId="10" pageNumber="193">
<subSubSection id="C33A24EAFFAC1675FD09EA8F18C6FCA2" pageId="10" pageNumber="193" type="nomenclature">
<paragraph id="8B9F7761FFAC1675FD09EA8F18C6FCA2" blockId="10.[737,1162,975,1023]" pageId="10" pageNumber="193">
<emphasis id="B954AB73FFAC1675FD09EA8F1FBBFCB9" bold="true" box="[737,1162,975,997]" pageId="10" pageNumber="193">
<taxonomicName id="4C200CE2FFAC1675FD09EA8F1F36FCB9" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2015" box="[737,1031,975,997]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="193" phylum="Arthropoda" rank="species" species="parvula" status="sp. nov.">
<emphasis id="B954AB73FFAC1675FD09EA8F1897FCB8" bold="true" box="[737,934,975,997]" italics="true" pageId="10" pageNumber="193">Marmarina parvula</emphasis>
Ratcliffe
</taxonomicName>
,
<taxonomicNameLabel id="A2671608FFAC1675FBFCEA8F1FBBFCB9" box="[1044,1162,975,996]" pageId="10" pageNumber="193" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B6BE4FFAC1675FC93EAAA18C0FCA2" box="[891,1009,1001,1023]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="11.[175,215,877,896]" captionStartId-1="12.[197,229,982,1001]" captionTargetBox-0="[173,1151,193,853]" captionTargetBox-1="[173,1193,194,960]" captionTargetId-0="figure-407@11.[151,1171,183,869]" captionTargetId-1="figure-359@12.[173,1193,194,960]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" captionText-1="Fig. 28. Topotype of Marmarina parvula at Cabra Corral, Salta, Argentina. Photograph by Gonzalo Rivero via Wikimedia Commons." figureDoi-0="http://doi.org/10.5281/zenodo.10104772" figureDoi-1="http://doi.org/10.5281/zenodo.10104776" httpUri-0="https://zenodo.org/record/10104772/files/figure.png" httpUri-1="https://zenodo.org/record/10104776/files/figure.png" pageId="10" pageNumber="193">Figs. 2328</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33A24EAFFAC1674FD33ED541878F92F" lastPageId="11" lastPageNumber="194" pageId="10" pageNumber="193" type="materials_examined">
<paragraph id="8B9F7761FFAC1675FD33ED541F4DFAE5" blockId="10.[707,1194,1044,1729]" pageId="10" pageNumber="193">
<emphasis id="B954AB73FFAC1675FD33ED5418A5FB74" bold="true" box="[731,916,1044,1065]" pageId="10" pageNumber="193">Type Material.</emphasis>
<materialsCitation id="3B487D3CFFAC1675FC4CED5418ABFBC9" collectorName="Coll. Martinez &amp; A. Martinez" location="Salta" pageId="10" pageNumber="193" specimenCount="2" stateProvince="Salta" typeStatus="holotype">
<typeStatus id="549BC9C3FFAC1675FC4CED541F37FB74" box="[932,1030,1044,1065]" pageId="10" pageNumber="193" type="holotype">Holotype</typeStatus>
male, labeled “Argentina/
<collectingRegion id="49E4B983FFAC1675FCABED6F1849FB19" box="[835,888,1071,1092]" country="Argentina" name="Salta" pageId="10" pageNumber="193">Salta</collectingRegion>
/D
<superScript id="7C55DA29FFAC1675FC7CED6918ACFB6A" attach="left" box="[916,925,1065,1079]" fontSize="6" pageId="10" pageNumber="193">o</superScript>
La Viña/Cabra Corral/
<collectorName id="26D512B7FFAC1675FD2BED091853FB03" box="[707,866,1097,1118]" pageId="10" pageNumber="193">Coll. Martinez</collectorName>
/Ene. 991// H &amp;A HOWDEN/ COLLECTION/ex
<collectorName id="26D512B7FFAC1675FC6DED241F30FB24" box="[901,1025,1124,1145]" pageId="10" pageNumber="193">A. Martinez</collectorName>
coll.” and with my red
<typeStatus id="549BC9C3FFAC1675FCE5ED3F1853FBC9" box="[781,866,1151,1172]" pageId="10" pageNumber="193" type="holotype">holotype</typeStatus>
label
</materialsCitation>
.
<typeStatus id="549BC9C3FFAC1675FC4CED3F18C8FBC9" box="[932,1017,1151,1172]" pageId="10" pageNumber="193" type="allotype">Allotype</typeStatus>
female with same data except date of “Marz. 91” and with my red
<typeStatus id="549BC9C3FFAC1675FD2BEDF41828FB94" box="[707,793,1204,1225]" pageId="10" pageNumber="193" type="allotype">allotype</typeStatus>
label.
<materialsCitation id="3B487D3CFFAC1675FC81EDF41F33FADE" collectingDate="1982-02-12" collectorName="Coll. Martinez" country="Argentina" county="El Carril" location="Argentina" municipality="Vinaco" pageId="10" pageNumber="193" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Salta" typeStatus="paratype">
<typeStatus id="549BC9C3FFAC1675FC81EDF418E1FB94" box="[873,976,1204,1225]" pageId="10" pageNumber="193" type="paratype">Paratypes</typeStatus>
labeled as follows:
<specimenCount id="9D26BCE8FFAC1675FD2BED8E183FFBBE" box="[707,782,1230,1251]" pageId="10" pageNumber="193" type="male">2 males</specimenCount>
with same data as holotype but with date of “Marz. 91”, and
<specimenCount id="9D26BCE8FFAC1675FC79EDA918E7FBA3" box="[913,982,1257,1278]" pageId="10" pageNumber="193" type="male">1 male</specimenCount>
labeled “
<location id="8EFF21BAFFAC1675FBD5EDA91F90FBA3" LSID="urn:lsid:plazi:treatment:0389C677FFAC1673FD09EA8F1F66F99C:8EFF21BAFFAC1675FBD5EDA91F90FBA3" box="[1085,1185,1257,1278]" country="Argentina" county="El Carril" municipality="Vinaco" name="Argentina" pageId="10" pageNumber="193" stateProvince="Salta">Argentina</location>
/
<collectingRegion id="49E4B983FFAC1675FD2BEC4319C7FA45" box="[707,758,1283,1304]" country="Argentina" name="Salta" pageId="10" pageNumber="193">Salta</collectingRegion>
/D
<superScript id="7C55DA29FFAC1675FCF9EDBE182BFA51" attach="left" box="[785,794,1278,1292]" fontSize="6" pageId="10" pageNumber="193">o</superScript>
<location id="8EFF21BAFFAC1675FCCCEC43184AFA45" LSID="urn:lsid:plazi:treatment:0389C677FFAC1673FD09EA8F1F66F99C:8EFF21BAFFAC1675FCCCEC43184AFA45" box="[804,891,1283,1304]" country="Argentina" county="El Carril" municipality="Vinaco" name="La Vina" pageId="10" pageNumber="193" stateProvince="Salta">La Viña</location>
/
<location id="8EFF21BAFFAC1675FC6AEC431F35FA45" LSID="urn:lsid:plazi:treatment:0389C677FFAC1673FD09EA8F1F66F99C:8EFF21BAFFAC1675FC6AEC431F35FA45" box="[898,1028,1283,1304]" country="Argentina" county="El Carril" municipality="Vinaco" name="Ampascachi" pageId="10" pageNumber="193" stateProvince="Salta">Ampascachi</location>
/
<collectorName id="26D512B7FFAC1675FBE4EC431F93FA45" box="[1036,1186,1283,1304]" pageId="10" pageNumber="193">Coll. Martinez</collectorName>
/ Ene. 993”;
<specimenCount id="9D26BCE8FFAC1675FCC7EC5E18B3FA6E" box="[815,898,1310,1331]" pageId="10" pageNumber="193" type="female">1 female</specimenCount>
labeled “
<collectingCountry id="F33737F1FFAC1675FC36EC5E1F5CFA6E" box="[990,1133,1310,1331]" name="Argentina" pageId="10" pageNumber="193">ARGENTINA</collectingCountry>
, Prov.
<collectingRegion id="49E4B983FFAC1675FD2BEC7819C4FA10" box="[707,757,1336,1357]" country="Argentina" name="Salta" pageId="10" pageNumber="193">Salta</collectingRegion>
/
<collectingMunicipality id="6BFBED1BFFAC1675FD15EC781876FA10" box="[765,839,1336,1357]" pageId="10" pageNumber="193">Viñaco</collectingMunicipality>
, 15kmS.
<collectingCounty id="62FE0FEDFFAC1675FC41EC781F36FA10" box="[937,1031,1336,1357]" pageId="10" pageNumber="193">El Carril</collectingCounty>
/
<date id="FF9E51A1FFAC1675FBE7EC781F43FA10" box="[1039,1138,1336,1357]" pageId="10" pageNumber="193" value="1982-02-12">
<collectingDate id="EFDAA849FFAC1675FBE7EC781F43FA10" box="[1039,1138,1336,1357]" pageId="10" pageNumber="193" value="1982-02-12">12.II.1982</collectingDate>
</date>
/H &amp; A Howden//Deposit/Ant Refuse”. All
<typeStatus id="549BC9C3FFAC1675FBA1EC131F99FA35" box="[1097,1192,1363,1384]" pageId="10" pageNumber="193" type="paratype">paratypes</typeStatus>
with my yellow
<typeStatus id="549BC9C3FFAC1675FC87EC2E18F5FADE" box="[879,964,1390,1411]" pageId="10" pageNumber="193" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
<materialsCitation id="3B487D3CFFAC1675FBE7EC2E1F47FAC0" collectionCode="CMNC" country="Argentina" pageId="10" pageNumber="193" specimenCount="1" stateProvince="Salta" typeStatus="holotype">
<typeStatus id="549BC9C3FFAC1675FBE7EC2E1F5FFADE" box="[1039,1134,1390,1411]" pageId="10" pageNumber="193" type="holotype">Holotype</typeStatus>
,
<typeStatus id="549BC9C3FFAC1675FB94EC2E19DFFAC0" pageId="10" pageNumber="193" type="allotype">allotype</typeStatus>
, and
<specimenCount id="9D26BCE8FFAC1675FCCCECC81898FAC0" box="[804,937,1416,1437]" pageId="10" pageNumber="193" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
deposited at
<collectionCode id="ED31EFA4FFAC1675FBC5ECC81F47FAC0" box="[1069,1142,1416,1437]" pageId="10" pageNumber="193">CMNC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B487D3CFFAC1675FB69ECC81F49FAE5" collectionCode="UNSM, BCRC" country="Argentina" pageId="10" pageNumber="193" specimenCount="1" stateProvince="Salta" typeStatus="paratype">
One
<typeStatus id="549BC9C3FFAC1675FD2BECE31823FAE5" box="[707,786,1443,1464]" pageId="10" pageNumber="193" type="paratype">paratype</typeStatus>
deposited each in
<collectionCode id="ED31EFA4FFAC1675FC2AECE31F37FAE5" box="[962,1030,1443,1464]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="10" pageNumber="193">UNSM</collectionCode>
and
<collectionCode id="ED31EFA4FFAC1675FBDEECE31F49FAE5" box="[1078,1144,1443,1464]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="10" pageNumber="193">BCRC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F7761FFAC1675FD33ECFD1F98F99C" blockId="10.[707,1194,1044,1729]" pageId="10" pageNumber="193">
<emphasis id="B954AB73FFAC1675FD33ECFD180FFA8F" bold="true" box="[731,830,1469,1490]" pageId="10" pageNumber="193">
<typeStatus id="549BC9C3FFAC1675FD33ECFD180BFA8F" box="[731,826,1469,1490]" pageId="10" pageNumber="193" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Length
<quantity id="4CD8DA84FFAC1675FC39ECFD1F19FA8F" box="[977,1064,1469,1490]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="193" unit="mm" value="11.4">11.4 mm</quantity>
; width across humeri
<quantity id="4CD8DA84FFAC1675FCE4EC981866FAB0" box="[780,855,1496,1517]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="193" unit="mm" value="6.1">6.1 mm</quantity>
. Ground color and punctures black, weakly shiny, with enamel-like yellowish orange on head and pronotum and enamel-like pale greenish yellow on pygidium and elytra, with opaque yellowish orange marks as follows: Head completely covered except for black spot on center of occiput and on apical rim of clypeus. Pronotum completely covered except for thick, broken, black bands radiating anteriorly from base
</paragraph>
<caption id="DF5F27E9FFAD1674FF47EA2D1FA4FCDD" ID-DOI="http://doi.org/10.5281/zenodo.10104772" ID-Zenodo-Dep="10104772" box="[175,1173,876,896]" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194" startId="11.[175,215,877,896]" targetBox="[173,1151,193,853]" targetPageId="11">
<paragraph id="8B9F7761FFAD1674FF47EA2D1FA4FCDD" blockId="11.[151,1173,876,920]" box="[175,1173,876,896]" pageId="11" pageNumber="194">
<emphasis id="B954AB73FFAD1674FF47EA2D1A10FCDD" bold="true" box="[175,289,876,896]" pageId="11" pageNumber="194">Figs. 2327.</emphasis>
<taxonomicName id="4C200CE2FFAD1674FEDEEA2D1AD6FC22" authorityName="Ratcliffe" authorityYear="2015" box="[310,487,877,895]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="194" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAD1674FEDEEA2D1AD6FC22" box="[310,487,877,895]" italics="true" pageId="11" pageNumber="194">Marmarina parvula</emphasis>
</taxonomicName>
.
<emphasis id="B954AB73FFAD1674FE1FEA2D1927FCDD" bold="true" box="[503,534,877,896]" pageId="11" pageNumber="194">23)</emphasis>
Habitus;
<emphasis id="B954AB73FFAD1674FD9CEA2D1984FCDD" bold="true" box="[628,693,876,896]" pageId="11" pageNumber="194">2425)</emphasis>
Mesometasternal process, lateral and ventral views;
</paragraph>
</caption>
<paragraph id="8B9F7761FFAD1674FF7FEAC51A08FCC5" blockId="11.[151,1173,876,920]" box="[151,313,900,920]" pageId="11" pageNumber="194">
<emphasis id="B954AB73FFAD1674FF7FEAC51BE7FCC5" bold="true" box="[151,214,900,920]" pageId="11" pageNumber="194">2627)</emphasis>
Parameres.
</paragraph>
<paragraph id="8B9F7761FFAD1674FF7FEA841F7EFA35" blockId="11.[151,639,964,1729]" lastBlockId="11.[685,1173,964,1729]" pageId="11" pageNumber="194">
of pronotal lobe (
<figureCitation id="131B6BE4FFAD1674FEA7EA841AACFC84" box="[335,413,964,985]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194">Fig. 23</figureCitation>
). Elytra each covered except on median half and short, incomplete, oblique black bands on lateral half (
<figureCitation id="131B6BE4FFAD1674FDC9EABA1943FB52" box="[545,626,1018,1039]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194">Fig. 23</figureCitation>
). Pygidium intermittently covered on apical half. Venter shiny black with intermittent, enamel-like pale yellowish green on mesepimera, metepisterna, metasternum, and posterior margins of abdominal sternites 15. Metacoxae on lateral edge, all metafemora, and meso- and metatibiae each with intermittent, enamel-like pale yellowish green.
<emphasis id="B954AB73FFAD1674FF7FED8E1BE9FBBE" bold="true" box="[151,216,1230,1251]" pageId="11" pageNumber="194">Head:</emphasis>
Frons with punctures small, moderately dense. Clypeus with small, sparse punctures; apex reflexed, subtruncate in dorsal view, emarginate in anterior view. Interocular width equals 7.0 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to entire stem.
<emphasis id="B954AB73FFAD1674FF7FEC2E1A34FADE" bold="true" box="[151,261,1390,1411]" pageId="11" pageNumber="194">Pronotum:</emphasis>
Surface with small, sparse punctures on yellow areas, punctures larger, slightly denser on black areas. Lateral margins arcuate with slender marginal line not reaching basal angle.
<emphasis id="B954AB73FFAD1674FF7FEC981BEFFAB0" bold="true" box="[151,222,1496,1517]" pageId="11" pageNumber="194">Elytra:</emphasis>
Surface similar to that of pronotum except punctures on median half larger, horseshoeshaped. Sutural costa and costae on disc each strongly elevated on apical halves. Apices at suture subquadrate.
<emphasis id="B954AB73FFAD1674FE93EF021AC1F90A" bold="true" box="[379,496,1602,1623]" pageId="11" pageNumber="194">Pygidium:</emphasis>
Black areas densely rugopunctate, with minute, black setae; surface on colored areas usually with sparse, moderately large punctures. In lateral view, surface evenly convex.
<emphasis id="B954AB73FFAD1674FE98EFEC1A8AF99C" bold="true" box="[368,443,1708,1729]" pageId="11" pageNumber="194">Venter:</emphasis>
Metasternum with large, dense, round, setigerous punctures, setae long, black; mesometasternal process with small, sparse punctures. Mesometasternal process, in lateral view, short with apex projecting slightly downwards from ventral axis of body, apex rounded (
<figureCitation id="131B6BE4FFAD1674FD5CED0919CDFB03" box="[692,764,1097,1118]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194">Fig. 24</figureCitation>
); in ventral view, sides tapering to rounded apex (
<figureCitation id="131B6BE4FFAD1674FD04ED241806FB24" box="[748,823,1124,1145]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194">Fig. 25</figureCitation>
). Abdominal sternites 15 longitudinally depressed at center, mostly impunctate, each with small, sparse punctures on extreme lateral margins and with irregular row of small punctures on apical margin of sternites; sternite 6 with more punctures along apical margin.
<emphasis id="B954AB73FFAD1674FC1EEDA91F01FBA3" bold="true" box="[1014,1072,1257,1278]" pageId="11" pageNumber="194">Legs:</emphasis>
Protibiae weakly tridentate, basal tooth nearly obsolete.
<emphasis id="B954AB73FFAD1674FD45EC5E1813FA6E" bold="true" box="[685,802,1310,1331]" pageId="11" pageNumber="194">Parameres:</emphasis>
In caudal view, form elongate, subrectangular, apices blunt and with small tooth on both mesal and lateral edges (
<figureCitation id="131B6BE4FFAD1674FC27EC131F72FA35" box="[975,1091,1362,1384]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="11" pageNumber="194">Figs. 2627</figureCitation>
).
</paragraph>
<paragraph id="8B9F7761FFAD1674FD2DEC2E1878F92F" blockId="11.[685,1173,964,1729]" pageId="11" pageNumber="194">
<emphasis id="B954AB73FFAD1674FD2DEC2E1812FADE" bold="true" box="[709,803,1390,1411]" pageId="11" pageNumber="194">
<typeStatus id="549BC9C3FFAD1674FD2DEC2E182EFADE" box="[709,799,1390,1411]" pageId="11" pageNumber="194" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Length
<quantity id="4CD8DA84FFAD1674FC22EC2E1F25FADE" box="[970,1044,1390,1411]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="11" pageNumber="194" unit="mm" value="9.9">9.9 mm</quantity>
; width across humeri
<quantity id="4CD8DA84FFAD1674FD15ECC8187FFAC0" box="[765,846,1416,1437]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="194" unit="mm" value="5.8">5.8 mm</quantity>
. As
<typeStatus id="549BC9C3FFAD1674FC6AECC818E6FAC0" box="[898,983,1416,1437]" pageId="11" pageNumber="194" type="holotype">holotype</typeStatus>
except in the following respects:
<emphasis id="B954AB73FFAD1674FCB6ECE318ACFAE5" bold="true" box="[862,925,1443,1464]" pageId="11" pageNumber="194">Head:</emphasis>
Clypeus on apical half black; frons and base of clypeus with narrow, longitudinal, black line; clypeal apex barely emarginate. Antenna with club slightly longer than antennomeres 27.
<emphasis id="B954AB73FFAD1674FC49EF4D18DDF97F" bold="true" box="[929,1004,1549,1570]" pageId="11" pageNumber="194">Venter:</emphasis>
Yellowish green color on abdominal sternites reduced.
<emphasis id="B954AB73FFAD1674FBB0EF671FA5F961" bold="true" box="[1112,1172,1575,1596]" pageId="11" pageNumber="194">Legs:</emphasis>
Protibiae broader, strongly tridentate, teeth subequally spaced.
</paragraph>
</subSubSection>
<subSubSection id="C33A24EAFFAD1673FD2DEF371932FA10" lastPageId="12" lastPageNumber="195" pageId="11" pageNumber="194" type="description">
<paragraph id="8B9F7761FFAD1673FD2DEF371A80FBBE" blockId="11.[685,1173,964,1729]" lastBlockId="12.[173,660,1097,1729]" lastPageId="12" lastPageNumber="195" pageId="11" pageNumber="194">
<emphasis id="B954AB73FFAD1674FD2DEF37181BF9D1" bold="true" box="[709,810,1655,1676]" pageId="11" pageNumber="194">Variation.</emphasis>
Males (
<specimenCount id="9D26BCE8FFAD1674FC6BEF3718CAF9D1" box="[899,1019,1655,1676]" pageId="11" pageNumber="194" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
). Length
<quantity id="4CD8DA84FFAD1674FBB4EF37183AF9FA" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.08" metricValueMin="1.06" pageId="11" pageNumber="194" unit="mm" value="10.7" valueMax="10.8" valueMin="10.6">10.6 10.8 mm</quantity>
; width across humeri
<quantity id="4CD8DA84FFAD1674FC07EFD21F5CF9FA" box="[1007,1133,1681,1703]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="11" pageNumber="194" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.56.5 mm</quantity>
. As
<typeStatus id="549BC9C3FFAD1674FD45EFEC1830F99C" box="[685,769,1708,1729]" pageId="11" pageNumber="194" type="holotype">holotype</typeStatus>
except in the following respects:
<emphasis id="B954AB73FFAD1674FBBDEFEC1FA2F99C" bold="true" box="[1109,1171,1708,1729]" pageId="11" pageNumber="194">Head:</emphasis>
Clypeus with extended black area near apex in
<specimenCount id="9D26BCE8FFAA1673FD87ED091A24FB24" pageId="12" pageNumber="195" type="generic">two specimens</specimenCount>
, frons with extended area of black at center in
<specimenCount id="9D26BCE8FFAA1673FEE4ED3E1AA7FBCE" box="[268,406,1150,1171]" pageId="12" pageNumber="195" type="generic">one specimen</specimenCount>
.
<emphasis id="B954AB73FFAA1673FE4AED3E1921FBCE" bold="true" box="[418,528,1150,1171]" pageId="12" pageNumber="195">Pronotum:</emphasis>
Lateral margins slightly flattened between middle and basal angle.
<emphasis id="B954AB73FFAA1673FF04EDF31A62FB95" bold="true" box="[236,339,1203,1224]" pageId="12" pageNumber="195">Pygidium:</emphasis>
One specimen with surface completely black, rugopunctate.
</paragraph>
<caption id="DF5F27E9FFAA1673FF2DEA961A5FFB5C" ID-DOI="http://doi.org/10.5281/zenodo.10104776" ID-Zenodo-Dep="10104776" httpUri="https://zenodo.org/record/10104776/files/figure.png" pageId="12" pageNumber="195" startId="12.[197,229,982,1001]" targetBox="[173,1193,194,960]" targetPageId="12">
<paragraph id="8B9F7761FFAA1673FF2DEA961F98FCB4" blockId="12.[173,1193,982,1025]" box="[197,1193,982,1001]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FF2DEA961A3DFCB4" bold="true" box="[197,268,982,1001]" pageId="12" pageNumber="195">Fig. 28.</emphasis>
Topotype of
<taxonomicName id="4C200CE2FFAA1673FE7CEA971971FCB4" authorityName="Ratcliffe" authorityYear="2015" box="[404,576,983,1001]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FE7CEA971971FCB4" box="[404,576,983,1001]" italics="true" pageId="12" pageNumber="195">Marmarina parvula</emphasis>
</taxonomicName>
at Cabra Corral, Salta, Argentina. Photograph by Gonzalo Rivero via
</paragraph>
<paragraph id="8B9F7761FFAA1673FF45EAAE1A5FFB5C" blockId="12.[173,1193,982,1025]" box="[173,366,1006,1025]" pageId="12" pageNumber="195">Wikimedia Commons.</paragraph>
</caption>
<paragraph id="8B9F7761FFAA1673FF2DEDA91932FA10" blockId="12.[173,660,1097,1729]" pageId="12" pageNumber="195">
Female (
<specimenCount id="9D26BCE8FFAA1673FEF1EDA91AB1FBA3" box="[281,384,1257,1278]" pageId="12" pageNumber="195" type="generic" typeStatus="paratype">1 paratype</specimenCount>
). The specimen (found dead) is in four pieces that are card-mounted. A length measurement is not possible, but all characters are similar to those of the
<typeStatus id="549BC9C3FFAA1673FE47EC781ACFFA10" box="[431,510,1336,1357]" pageId="12" pageNumber="195" type="allotype">allotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A24EAFFAA1673FF2DEC131AB7FAE5" pageId="12" pageNumber="195" type="etymology">
<paragraph id="8B9F7761FFAA1673FF2DEC131AB7FAE5" blockId="12.[173,660,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FF2DEC131A06FA35" bold="true" box="[197,311,1363,1384]" pageId="12" pageNumber="195">Etymology.</emphasis>
The epithet is adjectival and from the Latin
<emphasis id="B954AB73FFAA1673FEE4EC2E1A7CFADE" box="[268,333,1390,1411]" italics="true" pageId="12" pageNumber="195">parvus</emphasis>
, meaning little or small, in reference to its small size. Used here in the diminutive to indicate very small.
</paragraph>
</subSubSection>
<subSubSection id="C33A24EAFFAA1673FF2DECFD1862FADE" pageId="12" pageNumber="195" type="distribution">
<paragraph id="8B9F7761FFAA1673FF2DECFD18F6FBBE" blockId="12.[173,660,1097,1729]" lastBlockId="12.[707,1193,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FF2DECFD1A76FA8F" bold="true" box="[197,327,1469,1490]" pageId="12" pageNumber="195">Distribution.</emphasis>
<taxonomicName id="4C200CE2FFAA1673FEBDECFD1916FA8F" authorityName="Ratcliffe" authorityYear="2015" box="[341,551,1469,1490]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FEBDECFD1916FA8F" box="[341,551,1469,1490]" italics="true" pageId="12" pageNumber="195">Marmarina parvula</emphasis>
</taxonomicName>
is known only from central and northwestern
<collectingCountry id="F33737F1FFAA1673FDC5EC9819A3FAB0" box="[557,658,1496,1517]" name="Argentina" pageId="12" pageNumber="195">Argentina</collectingCountry>
in
<collectingRegion id="49E4B983FFAA1673FF21ECB21A6DF95A" box="[201,348,1522,1543]" country="Argentina" name="Salta" pageId="12" pageNumber="195">Salta Province</collectingRegion>
. At present, there are no other
<taxonomicName id="4C200CE2FFAA1673FF45EF4D1A2AF97F" authorityName="KIRBY" authorityYear="1827" box="[173,283,1549,1570]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B954AB73FFAA1673FF45EF4D1A2AF97F" box="[173,283,1549,1570]" italics="true" pageId="12" pageNumber="195">Marmarina</emphasis>
</taxonomicName>
collecting records from the type localities, but the collecting data are too fragmentary to say if
<taxonomicName id="4C200CE2FFAA1673FEE2EF021A4BF90A" authorityName="Ratcliffe" authorityYear="2015" box="[266,378,1602,1623]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FEE2EF021A4BF90A" box="[266,378,1602,1623]" italics="true" pageId="12" pageNumber="195">M. parvula</emphasis>
</taxonomicName>
is truly isolated from other species.
<bibRefCitation id="EFB10A90FFAA1673FEE8EF1D1AA0F92F" author="Di Iorio" box="[256,401,1629,1650]" pageId="12" pageNumber="195" pagination="1 - 87" refId="ref13271" refString="Di Iorio, O. 2013. A review of the Cetoniinae (Coleoptera: Scarabaeidae) from Argentina and adjacent countries: systematics and geographic distributions. Zootaxa 3668: 1 - 87." type="journal article" year="2013">Di Iorio (2013</bibRefCitation>
, figs. 129132) illustrated four additional specimens from
<collectingRegion id="49E4B983FFAA1673FDEEEF371952F9D1" box="[518,611,1655,1676]" country="Argentina" name="San Luis" pageId="12" pageNumber="195">San Luis</collectingRegion>
and
<collectingRegion id="49E4B983FFAA1673FF45EFD21A30F9FA" box="[173,257,1682,1703]" country="Argentina" name="La Rioja" pageId="12" pageNumber="195">La Rioja</collectingRegion>
in western
<collectingCountry id="F33737F1FFAA1673FE90EFD21AE7F9FA" box="[376,470,1682,1703]" name="Argentina" pageId="12" pageNumber="195">Argentina</collectingCountry>
that could be conspecific with
<taxonomicName id="4C200CE2FFAA1673FEC1EFEC1AA0F99C" authorityName="Ratcliffe" authorityYear="2015" box="[297,401,1708,1729]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FEC1EFEC1AA0F99C" box="[297,401,1708,1729]" italics="true" pageId="12" pageNumber="195">M. parvula</emphasis>
</taxonomicName>
. I attempted to obtain these on loan so that I might designate them
<typeStatus id="549BC9C3FFAA1673FBD3ED091FA2FB03" box="[1083,1171,1097,1118]" pageId="12" pageNumber="195" type="paratype">paratypes</typeStatus>
if they represented
<taxonomicName id="4C200CE2FFAA1673FC8EED2418FEFB24" authorityName="Ratcliffe" authorityYear="2015" box="[870,975,1124,1145]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FC8EED2418FEFB24" box="[870,975,1124,1145]" italics="true" pageId="12" pageNumber="195">M. parvula</emphasis>
</taxonomicName>
, but this was not possible due to the extreme difficulties of securing permits simply for sending specimens from and to
<collectingCountry id="F33737F1FFAA1673FD35EDF31808FB95" box="[733,825,1203,1224]" name="Argentina" pageId="12" pageNumber="195">Argentina</collectingCountry>
(O. Di Iorio, personal communication to BCR,
<date id="FF9E51A1FFAA1673FCF1ED8E188DFBBE" box="[793,956,1230,1251]" pageId="12" pageNumber="195" value="2014-10-27">27 October 2014</date>
).
</paragraph>
<paragraph id="8B9F7761FFAA1673FD33EDA91896FA10" blockId="12.[707,1193,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FD33EDA918A1FBA3" bold="true" box="[731,912,1257,1278]" pageId="12" pageNumber="195">Locality Records.</emphasis>
<materialsCitation id="3B487D3CFFAA1673FC75EDA91895FA10" collectionCode="AMIC, CMNC" collectorName="El Carril" country="Argentina" location="SALTA" pageId="12" pageNumber="195" specimenCount="6" stateProvince="Salta">
<specimenCount id="9D26BCE8FFAA1673FC75EDA91F22FBA3" box="[925,1043,1257,1278]" pageId="12" pageNumber="195" type="generic">6 specimens</specimenCount>
originally from
<collectionCode id="ED31EFA4FFAA1673FD2BEC431833FA45" box="[707,770,1283,1304]" pageId="12" pageNumber="195">AMIC</collectionCode>
and
<collectionCode id="ED31EFA4FFAA1673FCDFEC4318B0FA45" box="[823,897,1283,1304]" pageId="12" pageNumber="195">CMNC</collectionCode>
.
<emphasis id="B954AB73FFAA1673FC66EC431F67FA45" bold="true" box="[910,1110,1283,1304]" pageId="12" pageNumber="195">
<collectingCountry id="F33737F1FFAA1673FC66EC431F14FA45" box="[910,1061,1283,1304]" name="Argentina" pageId="12" pageNumber="195">ARGENTINA</collectingCountry>
(6):
</emphasis>
<collectingRegion id="49E4B983FFAA1673FBB7EC431F98FA45" box="[1119,1193,1283,1304]" country="Argentina" name="Salta" pageId="12" pageNumber="195">SALTA</collectingRegion>
(6): Ampascachi,
<collectorName id="26D512B7FFAA1673FC92EC5E18E3FA6E" box="[890,978,1310,1331]" pageId="12" pageNumber="195">El Carril</collectorName>
(
<quantity id="4CD8DA84FFAA1673FC0CEC5E1F14FA6E" box="[996,1061,1310,1331]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="195" unit="km" value="15.0">15 km</quantity>
S), Embalse Cabra Corral (
<figureCitation id="131B6BE4FFAA1673FCB9EC7818ADFA10" box="[849,924,1336,1357]" captionStart="Fig" captionStartId="12.[197,229,982,1001]" captionTargetBox="[173,1193,194,960]" captionTargetId="figure-359@12.[173,1193,194,960]" captionTargetPageId="12" captionText="Fig. 28. Topotype of Marmarina parvula at Cabra Corral, Salta, Argentina. Photograph by Gonzalo Rivero via Wikimedia Commons." figureDoi="http://doi.org/10.5281/zenodo.10104776" httpUri="https://zenodo.org/record/10104776/files/figure.png" pageId="12" pageNumber="195">Fig. 28</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F7761FFAA1673FD33EC131862FADE" blockId="12.[707,1193,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FD33EC1318F4FA35" bold="true" box="[731,965,1363,1384]" pageId="12" pageNumber="195">Temporal Distribution.</emphasis>
January (2), February (1), March (3).
</paragraph>
</subSubSection>
<subSubSection id="C33A24EAFFAA1673FD33ECC81F66F99C" pageId="12" pageNumber="195" type="diagnosis">
<paragraph id="8B9F7761FFAA1673FD33ECC818D3F92F" blockId="12.[707,1193,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FD33ECC81875FAC0" bold="true" box="[731,836,1416,1437]" pageId="12" pageNumber="195">Diagnosis.</emphasis>
<taxonomicName id="4C200CE2FFAA1673FCB9ECC81F3DFAC0" authorityName="Ratcliffe" authorityYear="2015" box="[849,1036,1416,1437]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FCB9ECC81F3DFAC0" box="[849,1036,1416,1437]" italics="true" pageId="12" pageNumber="195">Marmarina parvula</emphasis>
</taxonomicName>
is unique in the genus because of its small size (less than
<quantity id="4CD8DA84FFAA1673FBA0ECE31FAEFAE5" box="[1096,1183,1443,1464]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="195" unit="mm" value="11.5">11.5 mm</quantity>
), abdominal sternites longitudinally depressed at the center, and the distinctive form of the parameres with small teeth on both the mesal and lateral edges of the apex of each paramere (
<figureCitation id="131B6BE4FFAA1673FC0BEF4D1F1BF97F" box="[995,1066,1549,1570]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="12" pageNumber="195">Fig. 26</figureCitation>
). These characters and those of the mesometasternal process will distinguish
<taxonomicName id="4C200CE2FFAA1673FCB6EF0218F8F90A" authorityName="Ratcliffe" authorityYear="2015" box="[862,969,1602,1623]" class="Insecta" family="Cetoniidae" genus="Marmarina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B954AB73FFAA1673FCB6EF0218F8F90A" box="[862,969,1602,1623]" italics="true" pageId="12" pageNumber="195">M. parvula</emphasis>
</taxonomicName>
from
<taxonomicName id="4C200CE2FFAA1673FBEFEF021F58F90A" box="[1031,1129,1602,1623]" class="Insecta" family="Cetoniidae" genus="Gymnetis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="195" phylum="Arthropoda" rank="species" species="tigrina">
<emphasis id="B954AB73FFAA1673FBEFEF021F58F90A" box="[1031,1129,1602,1623]" italics="true" pageId="12" pageNumber="195">M. tigrina</emphasis>
</taxonomicName>
(compare
<figureCitation id="131B6BE4FFAA1673FD1CEF1D1857F92F" box="[756,870,1628,1650]" captionStart="Figs" captionStartId="11.[175,215,877,896]" captionTargetBox="[173,1151,193,853]" captionTargetId="figure-407@11.[151,1171,183,869]" captionTargetPageId="11" captionText="Figs. 2327. Marmarina parvula. 23) Habitus; 2425) Mesometasternal process, lateral and ventral views;" figureDoi="http://doi.org/10.5281/zenodo.10104772" httpUri="https://zenodo.org/record/10104772/files/figure.png" pageId="12" pageNumber="195">Figs. 2427</figureCitation>
and
<figureCitation id="131B6BE4FFAA1673FC71EF1D18E6F92F" box="[921,983,1628,1650]" captionStart="Figs" captionStartId="14.[197,237,798,817]" captionTargetBox="[173,659,194,776]" captionTargetId="figure-634@14.[173,659,194,776]" captionTargetPageId="14" captionText="Figs. 3235. Marmarina tigrina. 3233) Mesometasternal process, lateral and ventral views; 3435) Parameres." figureDoi="http://doi.org/10.5281/zenodo.10104780" httpUri="https://zenodo.org/record/10104780/files/figure.png" pageId="12" pageNumber="195">3235</figureCitation>
).
</paragraph>
<paragraph id="8B9F7761FFAA1673FD33EF371F66F99C" blockId="12.[707,1193,1097,1729]" pageId="12" pageNumber="195">
<emphasis id="B954AB73FFAA1673FD33EF37184FF9D1" bold="true" box="[731,894,1655,1676]" pageId="12" pageNumber="195">Natural History.</emphasis>
The specimens were collected from habitats that are all xeric. One specimen was taken from an ant refuse pile (label data).
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</subSubSection>
</treatment>
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