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<document id="4D815859A66A15C24DAA18CD5520082B" ID-DOI="10.11646/phytotaxa.439.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13873370" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727814559806" checkinUser="felipe" docAuthor="Dayanti, Eka P., Pitopang, Ramadanil, Ardi, Wisnu H. &amp; Thomas, Daniel C." docDate="2020" docId="E2688345FFEAFFEDD6D8FF22A7AD9CEF" docLanguage="en" docName="phytotaxa.439.2.4.pdf" docOrigin="Phytotaxa 439 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.439.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia sidolensis Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D 2020, spec. nov." docType="treatment" docVersion="2" lastPageNumber="140" masterDocId="1E51FB3DFFEBFFE9D650FFD6A65D9E3B" masterDocTitle="Two New Species of Begonia (Begoniaceae, Section Petermannia) from Mount Sidole, Central Sulawesi, Indonesia" masterLastPageNumber="142" masterPageNumber="136" pageNumber="137" updateTime="1727816323394" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="11493FBBF79CE3C94D7983C1CA1CCB9A">Two New Species of Begonia (Begoniaceae, Section Petermannia) from Mount Sidole, Central Sulawesi, Indonesia</mods:title>
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<mods:namePart id="E9AD4AD2C585875F5D0536A488FADCC5">Dayanti, Eka P.</mods:namePart>
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<mods:affiliation id="2057CB4F1939C0B68F33C0D79B4DE8E4">Department of Biology, Faculty of Mathematics and Natural Sciences, Tadulako University; Jl. Sukarno Hatta Km 10, Kampus Bumi Tadulako, Tondo Palu, Indonesia, 94117.</mods:affiliation>
<mods:nameIdentifier id="747A3F2E6D32F2B7978C1178B1FD01CB" type="email">salukieka@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="E61E1CAE82802691D7CCBBA3C9B67DB7">Pitopang, Ramadanil</mods:namePart>
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<mods:affiliation id="5584F8380B03B05610216DDE8CEEC922">Department of Biology, Faculty of Mathematics and Natural Sciences, Tadulako University; Jl. Sukarno Hatta Km 10, Kampus Bumi Tadulako, Tondo Palu, Indonesia, 94117.</mods:affiliation>
<mods:nameIdentifier id="BFB97948EAD6BFA83491843589FDE00D" type="email">pitopang_64@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="C66C870BE07893319F9B530249E2B1A5">Ardi, Wisnu H.</mods:namePart>
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<mods:affiliation id="4800C375C7B302C03D87FE9E292CC0F0">Research Center for Plant Conservation and Botanic Gardens-Indonesian Institute of Science (LIPI), Jl. Ir. H. Juanda no. 13 PO. BOX 309, Bogor 16122, West Java, Indonesia;</mods:affiliation>
<mods:nameIdentifier id="A0558B22DE660759A0BA42F5C7464CD0" type="email">wisnu.handoyo.ardi@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="352BFFD0BEE8072FD5AE322F2D0531EC">Thomas, Daniel C.</mods:namePart>
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<mods:affiliation id="C56415A22D605A8CA72726B2DFFE5E4D">Herbarium, Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569;</mods:affiliation>
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<mods:title id="C7214AD8EB4861A4A0715244FF0367C6">Phytotaxa</mods:title>
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<mods:date id="37F81CEE4F9F78FE76EB53D49E563DA1">2020</mods:date>
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<mods:number id="CE9A5DFB408CDAFBF667F1AF66DB5D02">2020-04-16</mods:number>
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<emphasis id="58B5EE41FFEAFFE8D6D8FF22A6C39F35" bold="true" box="[136,158,244,270]" italics="true" pageId="1" pageNumber="137">1.</emphasis>
<taxonomicName id="ADC149D0FFEAFFE8D6EDFF22A4899F35" ID-CoL="7SHJ7" authority="Dayanti, Ramadanil &amp; Ardi" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[189,724,244,271]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="137" phylum="Tracheophyta" rank="species" species="sidolensis" status="sp. nov.">
<emphasis id="58B5EE41FFEAFFE8D6EDFF22A7C89F34" bold="true" box="[189,405,244,271]" italics="true" pageId="1" pageNumber="137">Begonia sidolensis</emphasis>
Dayanti, Ramadanil &amp; Ardi
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, 
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(
<figureCitation id="F2FA2ED6FFEAFFE8D50DFF22A59B9F35" box="[861,966,244,270]" captionStart="FIGURE 1" captionStartId="1.[136,229,1568,1590]" captionTargetBox="[182,1402,670,1523]" captionTargetId="figure-177@1.[162,1425,655,1543]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the distribution of Begonia sidolensis (circle) and B. rolandfadlii (square)." figureDoi="http://doi.org/10.5281/zenodo.13873372" httpUri="https://zenodo.org/record/13873372/files/figure.png" pageId="1" pageNumber="137">Figures 1</figureCitation>
&amp; 
<figureCitation id="F2FA2ED6FFEAFFE8D5BAFF22A5A49F35" box="[1002,1017,244,270]" captionStart="FIGURE 2" captionStartId="2.[136,229,1985,2007]" captionTargetBox="[262,1321,605,1960]" captionTargetId="figure-231@2.[262,1326,605,1960]" captionTargetPageId="2" captionText="FIGURE 2. Begonia sidolensis Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D–E. Male inflorescence. F. Male flowers (front view). G. Female flower (side view). H. Female flower (front view). I. Dry fruit.J. Ovary cross section.All photos from E.P. Dayanti ED 12. Photos: A–I: Eka Dayanti; J: Wisnu Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873374" httpUri="https://zenodo.org/record/13873374/files/figure.png" pageId="1" pageNumber="137">2</figureCitation>
)
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<emphasis id="58B5EE41FFEAFFE8D6D8FECEA6829F09" bold="true" box="[136,223,280,306]" pageId="1" pageNumber="137">Section</emphasis>
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<emphasis id="58B5EE41FFEAFFE8D6B6FECFA72A9F09" box="[230,375,281,306]" italics="true" pageId="1" pageNumber="137">Petermannia</emphasis>
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<emphasis id="58B5EE41FFEAFFE8D6D8FEB5A6AE9F42" bold="true" box="[136,243,355,377]" pageId="1" pageNumber="137">Diagnosis:</emphasis>
— 
<taxonomicName id="ADC149D0FFEAFFE8D75BFEB2A7DB9F42" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[267,390,356,377]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="137" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFEAFFE8D75BFEB2A7DB9F42" box="[267,390,356,377]" italics="true" pageId="1" pageNumber="137">B. sidolensis</emphasis>
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is a species from montane forest of Gunung Sidole, 
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. It resembles 
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<emphasis id="58B5EE41FFEAFFE8D23AFEB2A3509F42" box="[1130,1293,356,377]" italics="true" pageId="1" pageNumber="137">Begonia gemella</emphasis>
Warb. ex 
<bibRefCitation id="0E504FA2FFEAFFE8D322FEB2A7949FA6" author="Smith, L. B. &amp; Wasshausen, D. C." pageId="1" pageNumber="137" pagination="441 - 451" refId="ref4553" refString="Smith, L. B. &amp; Wasshausen, D. C. (1983) Notes on Begonia I. Phytologia 52 (7): 441 - 451." type="journal article" year="1983">Smith &amp; Wasshausen (1983: 443)</bibRefCitation>
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, known from lowland limestone forests of 
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, in the creeping habit and small lamina size, but can be easily distinguished by its shorter petioles (0.5–2vs 
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), male inflorescences comprised of 2−3 monochasial partial inflorescences with caducous bracteoles (vs male inflorescence a monochasium, bracteoles persistent), shorter male flower pedicels (13–16 vs 
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) and larger male flower tepals (11–13 × 14–15 vs 5–9 × 
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), and solitary female flowers with 5 tepals (vs female flowers 2-tepalled and usually paired with a single male flower).
</paragraph>
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<emphasis id="58B5EE41FFEAFFE8D6D8FDEAA69F9C6A" bold="true" box="[136,194,572,593]" pageId="1" pageNumber="137">Type:</emphasis>
— 
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. 
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, 
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: 
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, 
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, 
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<collectingDate id="0E3BED7BFFEAFFE8D5C8FDEAA20D9C6A" box="[920,1104,572,594]" pageId="1" pageNumber="137" value="2018-12-27">27 December 2018</collectingDate>
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, 
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. 
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. 
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12
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(
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!; 
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!)
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.
</paragraph>
<caption id="3EBE62DBFFEAFFE8D6D8F9F6A208980D" ID-DOI="http://doi.org/10.5281/zenodo.13873372" ID-Zenodo-Dep="13873372" box="[136,1109,1568,1591]" httpUri="https://zenodo.org/record/13873372/files/figure.png" pageId="1" pageNumber="137" startId="1.[136,229,1568,1590]" targetBox="[182,1402,670,1523]" targetPageId="1" targetType="figure">
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<emphasis id="58B5EE41FFEAFFE8D6D8F9F6A6A3980E" bold="true" box="[136,254,1568,1590]" pageId="1" pageNumber="137">FIGURE 1.</emphasis>
Map showing the distribution of 
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<emphasis id="58B5EE41FFEAFFE8D46FF9F7A4A9980D" box="[575,756,1569,1590]" italics="true" pageId="1" pageNumber="137">Begonia sidolensis</emphasis>
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(circle) and 
<taxonomicName id="ADC149D0FFEAFFE8D53DF9F7A5A7980D" box="[877,1018,1569,1590]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="137" phylum="Tracheophyta" rank="species" species="rolandfadlii">
<emphasis id="58B5EE41FFEAFFE8D53DF9F7A5A7980D" box="[877,1018,1569,1590]" italics="true" pageId="1" pageNumber="137">B. rolandfadlii</emphasis>
</taxonomicName>
(square).
</paragraph>
</caption>
<paragraph id="6A7E3253FFEAFFEBD6D8F9B3A31A9C7F" blockId="1.[136,1452,1637,2060]" lastBlockId="2.[136,1452,158,581]" lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137">
Perennial, monoecious herb, up to 
<quantity id="AD399FB6FFEAFFE8D44CF9B3A43998BB" box="[540,612,1637,1664]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="137" unit="cm" value="10.0">10 cm</quantity>
tall; stem creeping, rooting at the nodes when in contact with substrate, with microscopic glandular hairs and sparsely hairy with macroscopic hairs to glabrescent on stems. 
<emphasis id="58B5EE41FFEAFFE8D2ACF95FA36B9898" bold="true" box="[1276,1334,1673,1699]" italics="true" pageId="1" pageNumber="137">Stem</emphasis>
branched, internodes 
<quantity id="AD399FB6FFEAFFE8D755F97BA70598F3" box="[261,344,1709,1736]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="137" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long, reddish–brownish. 
<emphasis id="58B5EE41FFEAFFE8D428F97BA49A98FC" bold="true" box="[632,711,1709,1735]" italics="true" pageId="1" pageNumber="137">Leaves</emphasis>
alternate; 
<emphasis id="58B5EE41FFEAFFE8D510F97BA5CB98FC" box="[832,918,1709,1735]" italics="true" pageId="1" pageNumber="137">stipules</emphasis>
persistent, ovate to elliptic, ca. 4–5 × 
<quantity id="AD399FB6FFEAFFE8D317F97BA3FA98F3" box="[1351,1447,1709,1736]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="137" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, reddish, glabrous, translucent, midrib abaxially slightly prominent, margin entire, apex acuminate and narrowed into bristle up to ca. 
<quantity id="AD399FB6FFEAFFE8D76CF923A720992B" box="[316,381,1781,1808]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="137" unit="mm" value="2.0">2 mm</quantity>
long; 
<emphasis id="58B5EE41FFEAFFE8D794F923A4469934" box="[452,539,1781,1807]" italics="true" pageId="1" pageNumber="137">petioles</emphasis>
<quantity id="AD399FB6FFEAFFE8D473F923A4D3992B" box="[547,654,1781,1808]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="1" pageNumber="137" unit="mm" value="12.5" valueMax="20.0" valueMin="5.0">5–20 mm</quantity>
long, reddish, terete, sparsely hairy; 
<emphasis id="58B5EE41FFEAFFE8D27CF923A2279934" box="[1068,1146,1781,1807]" italics="true" pageId="1" pageNumber="137">lamina</emphasis>
basifixed, ovate to elliptic, 2.5–4.7 × 
<quantity id="AD399FB6FFEAFFE8D6ADF8CFA723990F" box="[253,382,1817,1844]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="1" pageNumber="137" unit="cm" value="2.15" valueMax="2.8" valueMin="1.5">1.5–2.8 cm</quantity>
, very asymmetric, base cordate, lobes not or sometimes slightly overlapping, apex acuminate, margin serrulate to double serrate to shallowly lobed (up to 20% of lamina width), ciliate, adaxially green-reddish, sometimes irregularly variegated with silvery blotches or silvery on and along the main veins, glabrous to very sparsely hairy between the veins, abaxially pale green, hairy on the veins, venation palmate-pinnate, primary veins 4–6, actinodromous, secondary veins craspedodromous. 
<emphasis id="58B5EE41FFEAFFE8D496F87EA53399F8" bold="true" box="[710,878,1960,1987]" italics="true" pageId="1" pageNumber="137">Inflorescences</emphasis>
terminal, protogynous; female flowers solitary, one node basal to the male inflorescence or further separated, 
<emphasis id="58B5EE41FFEAFFE8D55BF81BA52D99DC" box="[779,880,1997,2023]" italics="true" pageId="1" pageNumber="137">peduncle</emphasis>
ca. 
<quantity id="AD399FB6FFEAFFE8D5CEF81BA5AE99D3" box="[926,1011,1997,2024]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="137" unit="mm" value="1.5">1.5 mm</quantity>
long, red, glabrous; male inflorescence composed of up to 3 partial inflorescences, each a simple monochasium with 2–3 flowers, peduncle ca. 
<quantity id="AD399FB6FFEAFFE8D346F827A3319637" box="[1302,1388,2033,2060]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="137" unit="mm" value="2.5">2.5 mm</quantity>
long, red, glabrous; bracts stipule-like, persistent, ovate, 4–5 × 
<quantity id="AD399FB6FFE9FFEBD570FF48A5DF9E82" box="[800,898,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="138" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, translucent, midrib slightly prominent, reddish, glabrous, apex acuminate, and narrowed into bristle ca. 
<quantity id="AD399FB6FFE9FFEBD4A4FF14A5149EE6" box="[756,841,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="138" unit="mm" value="1.5">1.5 mm</quantity>
long, bracteoles caducous. 
<emphasis id="58B5EE41FFE9FFEBD22BFF14A3269EE7" box="[1147,1403,194,221]" italics="true" pageId="2" pageNumber="138">
<emphasis id="58B5EE41FFE9FFEBD22BFF14A3479EE6" bold="true" box="[1147,1306,194,221]" italics="true" pageId="2" pageNumber="138">Male flowers:</emphasis>
pedicels
</emphasis>
<quantity id="AD399FB6FFE9FFEBD3D1FF14A68B9F3A" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="2" pageNumber="138" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13– 16 mm</quantity>
long, pink, glabrous; 
<emphasis id="58B5EE41FFE9FFEBD798FF30A4569F3B" box="[456,523,230,256]" italics="true" pageId="2" pageNumber="138">tepals</emphasis>
2, pale pink, broadly ovate to suborbicular, 11–13 × 
<quantity id="AD399FB6FFE9FFEBD202FF30A2909F3A" box="[1106,1229,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="138" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
, abaxially glabrous, margin entire, apex rounded; 
<emphasis id="58B5EE41FFE9FFEBD786FEDCA4069F1F" box="[470,603,266,292]" italics="true" pageId="2" pageNumber="138">androecium</emphasis>
yellow, stamens 36–40, free filaments 
<quantity id="AD399FB6FFE9FFEBD247FEDCA2C29F1E" box="[1047,1183,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="2" pageNumber="138" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.3–0.6 mm</quantity>
long, fused at the base, anthers 
<quantity id="AD399FB6FFE9FFEBD6B1FEF8A7099F72" box="[225,340,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="138" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, dehiscing through unilaterally positioned slits ca. ½ as long as the anthers. 
<emphasis id="58B5EE41FFE9FFEBD2BFFEF8A6B99F57" italics="true" pageId="2" pageNumber="138">
<emphasis id="58B5EE41FFE9FFEBD2BFFEF8A3F19F72" bold="true" box="[1263,1452,302,329]" italics="true" pageId="2" pageNumber="138">Female flowers:</emphasis>
pedicels
</emphasis>
<quantity id="AD399FB6FFE9FFEBD6BDFE84A7109F56" box="[237,333,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="138" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long, glabrous, reddish; 
<emphasis id="58B5EE41FFE9FFEBD43DFE84A4ED9F57" box="[621,688,338,364]" italics="true" pageId="2" pageNumber="138">tepals</emphasis>
5, white tinged pink, one smaller, elliptic, 7–9 × 
<quantity id="AD399FB6FFE9FFEBD2A0FE84A3269F56" box="[1264,1403,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.5" metricValueMin="4.5" pageId="2" pageNumber="138" unit="mm" value="6.0" valueMax="7.5" valueMin="4.5">4.5–7.5 mm</quantity>
, the other four larger, broadly ovate, 8–12.5 × 
<quantity id="AD399FB6FFE9FFEBD407FEA0A49E9FAA" box="[599,707,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="2" pageNumber="138" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
, abaxially glabrous, margin entire, apex rounded; 
<emphasis id="58B5EE41FFE9FFEBD2BEFEA1A3739FAB" box="[1262,1326,375,400]" italics="true" pageId="2" pageNumber="138">ovary</emphasis>
(excluding wings) ellipsoid, 6–8 × 
<quantity id="AD399FB6FFE9FFEBD7CCFE4CA7A39F8E" box="[412,510,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="138" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, reddish-greenish, glabrous, locules 3, placentae bilamellate, wings 3, equal, base rounded, apex truncate to subtruncate, widest point 
<quantity id="AD399FB6FFE9FFEBD495FE68A57F9FE2" box="[709,802,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="138" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
(at the middle to subapically); 
<emphasis id="58B5EE41FFE9FFEBD22AFE68A2F09FE3" box="[1146,1197,446,472]" italics="true" pageId="2" pageNumber="138">style</emphasis>
up to ca. 
<quantity id="AD399FB6FFE9FFEBD348FE68A3309FE2" box="[1304,1389,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="138" unit="mm" value="2.5">2.5 mm</quantity>
long, basally fused, 3-branched, each stylodium bifurcate in the stigma region, stigmatic surface a spirally twisted papillose band, orange. 
<emphasis id="58B5EE41FFE9FFEBD778FDD0A7BB9C1B" box="[296,486,518,544]" italics="true" pageId="2" pageNumber="138">
<emphasis id="58B5EE41FFE9FFEBD778FDD0A7339C1B" bold="true" box="[296,366,518,544]" italics="true" pageId="2" pageNumber="138">Fruit:</emphasis>
peduncles
</emphasis>
ca. 
<quantity id="AD399FB6FFE9FFEBD445FDD0A4319C1A" box="[533,620,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="138" unit="mm" value="1.5">1.5 mm</quantity>
long; 
<emphasis id="58B5EE41FFE9FFEBD4E4FDD0A54D9C1B" box="[692,784,518,544]" italics="true" pageId="2" pageNumber="138">pedicels</emphasis>
<quantity id="AD399FB6FFE9FFEBD548FDD0A5289C1A" box="[792,885,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="138" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
long; seed-bearing part ellipsoid, 8–10 × 
<quantity id="AD399FB6FFE9FFEBD31EFDD0A3F19C1A" box="[1358,1452,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="138" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
(excluding the wings), wing shape as for ovary, widest point 
<quantity id="AD399FB6FFE9FFEBD563FDFCA5CE9C7E" box="[819,915,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="2" pageNumber="138" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
. 
<emphasis id="58B5EE41FFE9FFEBD5CDFDFCA5809C7F" box="[925,989,554,580]" italics="true" pageId="2" pageNumber="138">Seeds</emphasis>
barrel-shaped, ca. 
<quantity id="AD399FB6FFE9FFEBD2E1FDFCA35A9C7E" box="[1201,1287,554,581]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="138" unit="mm" value="0.3">0.3 mm</quantity>
long.
</paragraph>
</subSubSection>
<caption id="3EBE62DBFFE9FFEBD6D8F817A43D9624" ID-DOI="http://doi.org/10.5281/zenodo.13873374" ID-Zenodo-Dep="13873374" httpUri="https://zenodo.org/record/13873374/files/figure.png" pageId="2" pageNumber="138" startId="2.[136,229,1985,2007]" targetBox="[262,1321,605,1960]" targetPageId="2" targetType="figure">
<paragraph id="6A7E3253FFE9FFEBD6D8F817A43D9624" blockId="2.[136,1451,1985,2079]" pageId="2" pageNumber="138">
<emphasis id="58B5EE41FFE9FFEBD6D8F817A6A399ED" bold="true" box="[136,254,1985,2007]" pageId="2" pageNumber="138">FIGURE 2.</emphasis>
<taxonomicName id="ADC149D0FFE9FFEBD754F814A27599ED" authority="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[260,1064,1985,2007]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="2" pageNumber="138" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFE9FFEBD754F814A7E799ED" box="[260,442,1985,2007]" italics="true" pageId="2" pageNumber="138">Begonia sidolensis</emphasis>
Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D
</taxonomicName>
–E. Male inflorescence. F. Male flowers (front view). G. Female flower (side view). H. Female flower (front view). I. Dry fruit. J. Ovary cross section.All photos from 
<emphasis id="58B5EE41FFE9FFEBD363F830A69B9625" italics="true" pageId="2" pageNumber="138">E.P. Dayanti ED 12</emphasis>
. Photos: A–I: Eka Dayanti; J: Wisnu Ardi.
</paragraph>
</caption>
<caption id="3EBE62DBFFE8FFEAD6D8F903A2C69934" ID-DOI="http://doi.org/10.5281/zenodo.13873376" ID-Zenodo-Dep="13873376" httpUri="https://zenodo.org/record/13873376/files/figure.png" pageId="3" pageNumber="139" startId="3.[136,229,1749,1771]" targetBox="[240,1360,190,1725]" targetPageId="3" targetType="figure">
<paragraph id="6A7E3253FFE8FFEAD6D8F903A2C69934" blockId="3.[136,1452,1749,1807]" pageId="3" pageNumber="139">
<emphasis id="58B5EE41FFE8FFEAD6D8F903A6A098D1" bold="true" box="[136,253,1749,1771]" pageId="3" pageNumber="139">FIGURE 3.</emphasis>
<taxonomicName id="ADC149D0FFE8FFEAD753F900A5FA98D0" authority="Dayanti, Ramadanil &amp; D. C. Thomas. A. Habit. B." box="[259,935,1749,1771]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="139" phylum="Tracheophyta" rank="species" species="rolandfadlii">
<emphasis id="58B5EE41FFE8FFEAD753F900A79698D1" box="[259,459,1749,1771]" italics="true" pageId="3" pageNumber="139">Begonia rolandfadlii</emphasis>
Dayanti, Ramadanil &amp; D.C.Thomas. A. Habit. B.
</taxonomicName>
Male Inflorescence. C. Female flower (side view). D. Male flower (front view). E. Female flower (front view). All photos from 
<emphasis id="58B5EE41FFE8FFEAD51DF92CA5A59935" box="[845,1016,1785,1807]" italics="true" pageId="3" pageNumber="139">E.P. Dayanti ED9</emphasis>
by Eka Dayanti.
</paragraph>
</caption>
<subSubSection id="22DB61D8FFE8FFEAD6EDF8EBA6AB99A4" pageId="3" pageNumber="139" type="distribution">
<paragraph id="6A7E3253FFE8FFEAD6EDF8EBA25B996C" blockId="3.[136,1452,1853,2060]" box="[189,1030,1853,1880]" pageId="3" pageNumber="139">
<emphasis id="58B5EE41FFE8FFEAD6EDF8EBA704996C" bold="true" box="[189,345,1853,1879]" pageId="3" pageNumber="139">Distribution:</emphasis>
—Endemic to Sidole, 
<collectingRegion id="A805FCB1FFE8FFEAD402F8EBA54E996C" box="[594,787,1853,1879]" country="Indonesia" name="Sulawesi Tengah" pageId="3" pageNumber="139">Central Sulawesi</collectingRegion>
, 
<collectingRegion id="A805FCB1FFE8FFEAD54EF8EBA5DB996C" box="[798,902,1853,1879]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="139">Sulawesi</collectingRegion>
, 
<collectingCountry id="12D672C3FFE8FFEAD5C2F8EBA25E996C" box="[914,1027,1853,1879]" name="Indonesia" pageId="3" pageNumber="139">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="6A7E3253FFE8FFEAD6EDF8B7A6AB99A4" blockId="3.[136,1452,1853,2060]" pageId="3" pageNumber="139">
<emphasis id="58B5EE41FFE8FFEAD6EDF8B7A77E9940" bold="true" box="[189,291,1889,1915]" pageId="3" pageNumber="139">Habitat:</emphasis>
—Montane rainforests, growing terrestrially on steep slopes, in dense shade, at ca. 
<quantity id="AD399FB6FFE8FFEAD35BF8B4A3F19947" box="[1291,1452,1890,1916]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="3" pageNumber="139" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700–1900 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="22DB61D8FFE8FFEDD6EDF87CA7AD9CEF" lastPageId="4" lastPageNumber="140" pageId="3" pageNumber="139" type="etymology">
<paragraph id="6A7E3253FFE8FFEAD6EDF87CA6A899DC" blockId="3.[136,1452,1853,2060]" pageId="3" pageNumber="139">
<emphasis id="58B5EE41FFE8FFEAD6EDF87CA71599FF" bold="true" box="[189,328,1962,1988]" pageId="3" pageNumber="139">Etymology:</emphasis>
—The species epithet refers to Mount Sidole in 
<collectingRegion id="A805FCB1FFE8FFEAD5D8F87FA21399F8" box="[904,1102,1961,1987]" country="Indonesia" name="Sulawesi Tengah" pageId="3" pageNumber="139">Central Sulawesi</collectingRegion>
, where the 
<typeStatus id="B57A8CF1FFE8FFEAD28CF87CA35199FF" box="[1244,1292,1962,1988]" pageId="3" pageNumber="139">type</typeStatus>
material was collected.
</paragraph>
<paragraph id="6A7E3253FFE8FFEDD6EDF824A28E9F3B" blockId="3.[136,1452,1853,2060]" lastBlockId="4.[136,1452,158,725]" lastPageId="4" lastPageNumber="140" pageId="3" pageNumber="139">
<emphasis id="58B5EE41FFE8FFEAD6EDF824A7559637" bold="true" box="[189,264,2034,2060]" pageId="3" pageNumber="139">Notes:</emphasis>
— 
<taxonomicName id="ADC149D0FFE8FFEAD774F824A7AC9630" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[292,497,2033,2059]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="139" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFE8FFEAD774F824A7AC9630" box="[292,497,2033,2059]" italics="true" pageId="3" pageNumber="139">Begonia sidolensis</emphasis>
</taxonomicName>
is an understorey herb that grows on steep slopes. The prostrate-creeping habit is a rare character in 
<taxonomicName id="ADC149D0FFEFFFEDD75FFF49A40E9E83" authorityName="de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" box="[271,595,158,184]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="section" section="Petermannia">
<emphasis id="58B5EE41FFEFFFEDD75FFF49A7369E83" box="[271,363,159,184]" italics="true" pageId="4" pageNumber="140">Begonia</emphasis>
section 
<emphasis id="58B5EE41FFEFFFEDD795FF49A40E9E83" box="[453,595,159,184]" italics="true" pageId="4" pageNumber="140">Petermannia</emphasis>
</taxonomicName>
, although some other species with this growth habit are known from 
<collectingRegion id="A805FCB1FFEFFFEDD318FF48A3F69E83" box="[1352,1451,158,184]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="140">Sulawesi</collectingRegion>
and the 
<collectingCountry id="12D672C3FFEFFFEDD68EFF14A7149EE7" box="[222,329,194,220]" name="Indonesia" pageId="4" pageNumber="140">Moluccas</collectingCountry>
(
<collectingRegion id="A805FCB1FFEFFFEDD707FF14A7E29EE7" box="[343,447,194,220]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="140">Sulawesi</collectingRegion>
: 
<taxonomicName id="ADC149D0FFEFFFEDD799FF15A5169EE7" authority="Warb. ex Sm. &amp; Wassh." authorityName="Warb. ex Smith &amp; Wasshausen" authorityYear="1983" box="[457,843,194,221]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="gemella">
<emphasis id="58B5EE41FFEFFFEDD799FF15A4639EE7" box="[457,574,194,220]" italics="true" pageId="4" pageNumber="140">B. gemella</emphasis>
Warb. ex Sm. &amp; Wassh.
</taxonomicName>
; 
<taxonomicName id="ADC149D0FFEFFFEDD505FF15A2EE9EE7" authority="Irmscher (1953 a: 486)" authorityName="Irmscher" authorityPageNumber="486" authorityYear="1953" box="[853,1203,194,221]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="flacca">
<emphasis id="58B5EE41FFEFFFEDD505FF15A5EB9EE7" box="[853,950,194,220]" italics="true" pageId="4" pageNumber="140">B. flacca</emphasis>
<bibRefCitation id="0E504FA2FFEFFFEDD5ECFF14A2EE9EE7" author="Irmscher, E." box="[956,1203,194,221]" pageId="4" pageNumber="140" pagination="477 - 503" refId="ref3921" refString="Irmscher, E. (1953 a) Neue Begoniaceen von O. Beccari in Malesien Gesammelt. Webbia 9: 477 - 503. https: // doi. org / 10.1080 / 00837792.1954.10669621" type="journal article" year="1953">Irmscher (1953a: 486)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="ADC149D0FFEFFFEDD2EEFF15A7F19F3B" authority="Ardi &amp; D. C. Thomas (2019: 23)" authorityName="Ardi &amp; D. C. Thomas" authorityPageNumber="23" authorityYear="2019" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="mattampensis">
<emphasis id="58B5EE41FFEFFFEDD2EEFF15A32E9EE7" box="[1214,1395,195,220]" italics="true" pageId="4" pageNumber="140">B. mattampensis</emphasis>
Ardi &amp; D.C. 
<bibRefCitation id="0E504FA2FFEFFFEDD68EFF30A7F19F3B" author="Thomas, D. C. &amp; Ardi, W. H." box="[222,428,230,256]" pageId="4" pageNumber="140" pagination="217 - 223" refId="ref4638" refString="Thomas, D. C. &amp; Ardi, W. H. (2019) Begonia mabberleyana (Begoniaceae), a new species fromCentral Sulawesi, Indonesia. Gardens' Bulletin Singapore 71 (Suppl. 2): 217 - 223. https: // doi. org / 10.26492 / gbs 71 (suppl. 2). 2019 - 16" type="journal article" year="2019">Thomas (2019:23)</bibRefCitation>
</taxonomicName>
; 
<collectingCountry id="12D672C3FFEFFFEDD7E8FF30A47A9F3B" box="[440,551,230,256]" name="Indonesia" pageId="4" pageNumber="140">Moluccas</collectingCountry>
: 
<taxonomicName id="ADC149D0FFEFFFEDD460FF31A2929F3B" authority="Ardhaka &amp; Ardi (Ardhaka et al. 2016: 61))" box="[560,1231,230,256]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="manuselaensis">
<emphasis id="58B5EE41FFEFFFEDD460FF31A4B29F3B" box="[560,751,230,256]" italics="true" pageId="4" pageNumber="140">B. manuselaensis</emphasis>
Ardhaka &amp; Ardi (
<bibRefCitation id="0E504FA2FFEFFFEDD5E5FF30A2E69F3B" author="Ardhaka, I. M. &amp; Ardi, W. H. &amp; Undaharta, N. K. E. &amp; Tirta, I. G." box="[949,1211,230,256]" pageId="4" pageNumber="140" pagination="61 - 64" refId="ref3485" refString="Ardhaka, I. M., Ardi, W. H., Undaharta, N. K. E. &amp; Tirta, I. G. (2016) A new species of Begonia (Begoniaceae) from Manusela National Park, Seram. Reinwardtia 15 (1): 61 - 64. https: // doi. org / 10.14203 / reinwardtia. v 15 i 1.2443" type="journal article" year="2016">
Ardhaka 
<emphasis id="58B5EE41FFEFFFEDD24CFF31A20E9F3B" box="[1052,1107,230,256]" italics="true" pageId="4" pageNumber="140">et al.</emphasis>
2016: 61
</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
<paragraph id="6A7E3253FFEFFFEDD6EDFEDCA7389C1B" blockId="4.[136,1452,158,725]" pageId="4" pageNumber="140">
Apart from similarities to 
<taxonomicName id="ADC149D0FFEFFFEDD7BBFEDDA4389F1F" authorityName="Warb. ex Smith &amp; Wasshausen" authorityYear="1983" box="[491,613,266,292]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="gemella">
<emphasis id="58B5EE41FFEFFFEDD7BBFEDDA4389F1F" box="[491,613,266,292]" italics="true" pageId="4" pageNumber="140">B. gemella</emphasis>
</taxonomicName>
(see diagnosis), 
<taxonomicName id="ADC149D0FFEFFFEDD578FEDDA5EA9F1F" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[808,951,266,292]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFEFFFEDD578FEDDA5EA9F1F" box="[808,951,266,292]" italics="true" pageId="4" pageNumber="140">B. sidolensis</emphasis>
</taxonomicName>
also has morphological similarities with 
<emphasis id="58B5EE41FFEFFFEDD3C3FEDDA76F9F73" italics="true" pageId="4" pageNumber="140">
<taxonomicName id="ADC149D0FFEFFFEDD3C3FEDDA7739F73" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="manuselaensis">B. manuselaensis</taxonomicName>
,
</emphasis>
a creeping species described from lowland limestone forest from Seram, but 
<taxonomicName id="ADC149D0FFEFFFEDD2DAFEF9A34B9F73" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[1162,1302,302,328]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFEFFFEDD2DAFEF9A34B9F73" box="[1162,1302,302,328]" italics="true" pageId="4" pageNumber="140">B. sidolensis</emphasis>
</taxonomicName>
can be easily distinguished by stems and leaves that have a sparse indumentum of simple hairs or are glabrescent (vs a very sparse indumentum of small flattened red scales), shorter petioles (0.5–2 vs 
<quantity id="AD399FB6FFEFFFEDD5D9FEA0A5AB9FAB" box="[905,1014,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="4" pageNumber="140" unit="cm" value="5.75" valueMax="7.5" valueMin="4.0">4−7.5 cm</quantity>
), shorter male flower pedicels (1.3−1.6 vs. 
<quantity id="AD399FB6FFEFFFEDD6FFFE4CA75B9F8F" box="[175,262,410,436]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="4" pageNumber="140" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">4−7 cm</quantity>
) and shorter female flower pedicels (0.5−0.7 vs 
<quantity id="AD399FB6FFEFFFEDD576FE4CA5C89F8F" box="[806,917,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="4" pageNumber="140" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.5−4 cm</quantity>
). 
<taxonomicName id="ADC149D0FFEFFFEDD5F8FE4DA20C9F8F" authorityName="Irmscher" authorityYear="1953" box="[936,1105,410,436]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="flacca">
<emphasis id="58B5EE41FFEFFFEDD5F8FE4DA20C9F8F" box="[936,1105,410,436]" italics="true" pageId="4" pageNumber="140">Begonia flacca</emphasis>
</taxonomicName>
and 
<taxonomicName id="ADC149D0FFEFFFEDD2D8FE4DA31C9F8F" authorityName="Ardi &amp; D. C. Thomas" authorityYear="2019" box="[1160,1345,411,436]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="mattampensis">
<emphasis id="58B5EE41FFEFFFEDD2D8FE4DA31C9F8F" box="[1160,1345,411,436]" italics="true" pageId="4" pageNumber="140">B. mattampensis</emphasis>
</taxonomicName>
are other allied species from 
<collectingRegion id="A805FCB1FFEFFFEDD732FE68A7979FE3" box="[354,458,446,472]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="140">Sulawesi</collectingRegion>
, similar in their creeping habit, but 
<taxonomicName id="ADC149D0FFEFFFEDD506FE69A5BE9FE3" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[854,995,446,472]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFEFFFEDD506FE69A5BE9FE3" box="[854,995,446,472]" italics="true" pageId="4" pageNumber="140">B. sidolensis</emphasis>
</taxonomicName>
differs in having a much smaller stature. Further, both 
<taxonomicName id="ADC149D0FFEFFFEDD74EFE35A7229FC7" authorityName="Irmscher" authorityYear="1953" box="[286,383,482,508]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="flacca">
<emphasis id="58B5EE41FFEFFFEDD74EFE35A7229FC7" box="[286,383,482,508]" italics="true" pageId="4" pageNumber="140">B. flacca</emphasis>
</taxonomicName>
and 
<taxonomicName id="ADC149D0FFEFFFEDD7E4FE35A4369FC7" authorityName="Ardi &amp; D. C. Thomas" authorityYear="2019" box="[436,619,483,508]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="mattampensis">
<emphasis id="58B5EE41FFEFFFEDD7E4FE35A4369FC7" box="[436,619,483,508]" italics="true" pageId="4" pageNumber="140">B. mattampensis</emphasis>
</taxonomicName>
are found in lowland limestone karst habitats and have never been reported from high altitudes.
</paragraph>
<paragraph id="6A7E3253FFEFFFEDD6EDFDFCA7AD9CEF" blockId="4.[136,1452,158,725]" pageId="4" pageNumber="140">
<emphasis id="58B5EE41FFEFFFEDD6EDFDFCA4229C7F" bold="true" box="[189,639,554,580]" pageId="4" pageNumber="140">Provisional conservation assessment:</emphasis>
—VuD2, Vulnerable. 
<taxonomicName id="ADC149D0FFEFFFEDD52CFDFDA20F9C7F" authorityName="Dayanti, Ramadanil &amp; Ardi. A. Habit. B. Leaves. C. Stipules. D" authorityYear="2020" box="[892,1106,554,580]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="140" phylum="Tracheophyta" rank="species" species="sidolensis">
<emphasis id="58B5EE41FFEFFFEDD52CFDFDA20F9C7F" box="[892,1106,554,580]" italics="true" pageId="4" pageNumber="140">Begonia sidolensis</emphasis>
</taxonomicName>
is known from only a single collection from Gunung Sidole. The forest condition at the 
<typeStatus id="B57A8CF1FFEFFFEDD57DFD99A5009C52" box="[813,861,591,617]" pageId="4" pageNumber="140">type</typeStatus>
locality is good, and currently there are no signs of anthropogenic threats in the montane forest of Gunung Sidole. However, due to the very restricted distribution and limited number of observed mature individuals, this species may be prone to stochastic effects and natural disasters such as tree falls and landslides.
</paragraph>
</subSubSection>
</treatment>
</document>