treatments-xml/data/E2/54/D6/E254D6E8B09D5BAFBBB8C5CE2EBC9D21.xml

<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.57578" ID-GBIF-Dataset="6e5a6293-b6f4-4f2c-9e69-4745f5d88870" ID-GBIF-Taxon="170398987" ID-Pensoft-Pub="2535-0730-2-79" ID-Pensoft-UUID="25215E4779855297B5CEA4AB9DE9BC33" ID-ZooBank="361BA656C8DC4F1DA7B8A167E95B2BB9" LSID="urn:lsid:zoobank.org:act:ADE1DCF5-09B3-45A5-B107-FBF536F2FC72" ModsDocID="2535-0730-2-79" checkinTime="1605780343973" checkinUser="pensoft" docAuthor="Venegas, Pablo J., Garcia-Ayachi, Luis A., Chavez-Arribasplata, Juan C., Chavez, German, Wong, Ivan &amp; Garcia-Bravo, Antonio" docDate="2020" docId="E254D6E8B09D5BAFBBB8C5CE2EBC9D21" docLanguage="en" docName="EvolutSyst 4(2): 79-108" docOrigin="Evolutionary Systematics 4 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.4.57578" docTitle="Stenocercus catherineae Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo 2020, sp. nov." docType="treatment" docUuid="ADE1DCF5-09B3-45A5-B107-FBF536F2FC72" docUuidSource="ZooBank" docVersion="3" id="25215E4779855297B5CEA4AB9DE9BC33" lastPageNumber="79" masterDocId="25215E4779855297B5CEA4AB9DE9BC33" masterDocTitle="Four new species of Stenocercus Dumeril &amp; Bibron, 1837 (Squamata, Iguania) from the Department of Amazonas in northeastern Peru" masterLastPageNumber="108" masterPageNumber="79" pageId="0" pageNumber="79" updateTime="1643445256845" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Four new species of Stenocercus Dumeril &amp; Bibron, 1837 (Squamata, Iguania) from the Department of Amazonas in northeastern Peru</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Venegas, Pablo J.</mods:namePart>
<mods:affiliation>Instituto Peruano de Herpetologia (IPH), Augusto Salazar Bondy 136, Urb. Higuereta, Surco, Lima, Peru &amp; Division de Herpetologia, Centro de Ornitologia y Biodiversidad (CORBIDI), Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Peru</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6501-4492</mods:nameIdentifier>
<mods:nameIdentifier type="email">pvenegas@corbidi.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garcia-Ayachi, Luis A.</mods:namePart>
<mods:affiliation>Instituto Peruano de Herpetologia (IPH), Augusto Salazar Bondy 136, Urb. Higuereta, Surco, Lima, Peru &amp; Division de Herpetologia, Centro de Ornitologia y Biodiversidad (CORBIDI), Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Peru</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3529-1261</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chavez-Arribasplata, Juan C.</mods:namePart>
<mods:affiliation>Instituto Peruano de Herpetologia (IPH), Augusto Salazar Bondy 136, Urb. Higuereta, Surco, Lima, Peru &amp; Division de Herpetologia, Centro de Ornitologia y Biodiversidad (CORBIDI), Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Peru</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chavez, German</mods:namePart>
<mods:affiliation>Instituto Peruano de Herpetologia (IPH), Augusto Salazar Bondy 136, Urb. Higuereta, Surco, Lima, Peru &amp; Division de Herpetologia, Centro de Ornitologia y Biodiversidad (CORBIDI), Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Peru</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9291-156X</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wong, Ivan</mods:namePart>
<mods:affiliation>Macanche RAM, Urb. 06 de Setiembre Mz. H, lote 20, Piura, Peru</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garcia-Bravo, Antonio</mods:namePart>
<mods:affiliation>Division de Herpetologia, Centro de Ornitologia y Biodiversidad (CORBIDI), Santa Rita No. 105 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, Peru &amp; Escuela de Ingenieria Forestal, Facultad de Ingenieria y Ciencias Agrarias, Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas. Calle Higos Urco N 342 - 350 - 356. Amazonas, Peru</mods:affiliation>
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<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>79</mods:start>
<mods:end>108</mods:end>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.4.57578</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.4.57578</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-2-79</mods:identifier>
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<treatment ID-GBIF-Taxon="170398987" LSID="urn:lsid:zoobank.org:act:ADE1DCF5-09B3-45A5-B107-FBF536F2FC72" httpUri="http://treatment.plazi.org/id/E254D6E8B09D5BAFBBB8C5CE2EBC9D21" lastPageNumber="79" pageId="0" pageNumber="79">
<subSubSection pageId="0" pageNumber="79" type="nomenclature">
<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="E254D6E8-B09D-5BAF-BBB8-C5CE2EBC9D21" authority="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo, 2020" authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae" status="sp. nov.">Stenocercus catherineae</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stenocercus catherineae sp. nov. preserved holotype, adult male, SVL 82 mm (CORBIDI 21365): dorsal (A), lateral (B), and ventral (C) views of the head; dorsal (D) and ventral (E) views of the entire specimen. Photographs by Luis A. Garcia-Ayachi. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/476348" pageId="0" pageNumber="79">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsolateral and ventral views of Stenocercus catherineae sp. nov. in life: (A, B) holotype, adult male, SVL 82 mm (CORBIDI 21365); (C, D) adult male, SVL 79 mm (CORBIDI 21366); (E, F) adult female, SVL 75 mm (CORBIDI 21367); (G, H) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas." httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral view of the postfemoral mite pocket in three species of Stenocercus: (A) S. catherineae sp. nov. (CORBIDI 21365), (B) S. philmayi sp. nov. (CORBIDI 21092), and (C) S. huancabambae (CORBIDI 19747). Photographs by Luis A. Garcia-Ayachi. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/476350" pageId="0" pageNumber="79">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="type material">
<paragraph pageId="0" pageNumber="79">Type material.</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Holotype:</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="79">
PERU • ♂, adult; Amazonas Department, 
<normalizedToken originalValue="Bongará">Bongara</normalizedToken>
Province, Florida District, Huembo; 
<geoCoordinate degrees="5" direction="south" minutes="51.47" orientation="latitude" precision="9" value="-5.8578334">5°51.47'S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="58.75" orientation="longitude" precision="9" value="-77.979164">77°58.75'W</geoCoordinate>
; 2090 m a.s.l.; 09 Dec. 2019; P.J. Venegas and L.A. 
<normalizedToken originalValue="García-Ayachi">Garcia-Ayachi</normalizedToken>
leg.; CORBIDI 21365.
</paragraph>
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Paratypes</emphasis>
:
</paragraph>
<paragraph pageId="0" pageNumber="79">
PERU • 1 ♂, 1 ♀, adults, collected with the holotype; CORBIDI 21366-67 • 1 ♂, juvenile; Amazonas Department, 
<normalizedToken originalValue="Bongará">Bongara</normalizedToken>
Province, Cuispes District, Cuispes; 
<geoCoordinate degrees="5" direction="south" minutes="55.49" orientation="latitude" precision="9" value="-5.9248333">5°55.49'S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="56.94" orientation="longitude" precision="9" value="-77.949">77°56.94'W</geoCoordinate>
; 1850 m a.s.l.; 8 Mar. 2017; G. 
<normalizedToken originalValue="Chávez">Chavez</normalizedToken>
leg.; CORBIDI 18662 • 1 ♀, 1 ♂, adults; Amazonas Department, 
<normalizedToken originalValue="Bongará">Bongara</normalizedToken>
Province, Valera District, Cocachimba; 
<geoCoordinate degrees="6" direction="south" minutes="2.76" orientation="latitude" precision="9" value="-6.046">6°2.76'S</geoCoordinate>
, 
<geoCoordinate degrees="77" direction="west" minutes="53.55" orientation="longitude" precision="9" value="-77.8925">77°53.55'W</geoCoordinate>
; 1460 m a.s.l.; 22 May 2007; by P.J. Venegas leg.; CORBIDI 501-02.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/476348" pageId="0" pageNumber="79" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 1.</emphasis>
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. preserved holotype, adult male, SVL 82 mm (CORBIDI 21365): dorsal (
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
), and ventral (
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
) views of the head; dorsal (
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
) views of the entire specimen. Photographs by Luis A. 
<normalizedToken originalValue="García-Ayachi">Garcia-Ayachi</normalizedToken>
. Scale bars: 10 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 2.</emphasis>
Dorsolateral and ventral views of 
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. in life: (
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
) holotype, adult male, SVL 82 mm (CORBIDI 21365); (
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
) adult male, SVL 79 mm (CORBIDI 21366); (
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
) adult female, SVL 75 mm (CORBIDI 21367); (
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. differs from other species of 
<taxonomicName authorityName="A.M.C.Dumeril &amp; Bibron" authorityYear="1837" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus</emphasis>
</taxonomicName>
except for 
<taxonomicName genus="S." lsidName="S. aculeatus" pageId="0" pageNumber="79" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">S. aculeatus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
Werner, 1901, 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
Cadle, 1991, 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
Cadle, 2001, and 
<taxonomicName genus="S." lsidName="S. scapularis" pageId="0" pageNumber="79" rank="species" species="scapularis">
<emphasis italics="true" pageId="0" pageNumber="79">S. scapularis</emphasis>
</taxonomicName>
Boulenger, 1901 by having: (1) projecting-angulate temporals, (2) laterally oriented nostrils; (3) dorsal and lateral scales of body similar in size, and (3) scales on posterior surface of thighs keeled and imbricate.
</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1879" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus aculeatus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus aculeatus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
, 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="S." lsidName="S. scapularis" pageId="0" pageNumber="79" rank="species" species="scapularis">
<emphasis italics="true" pageId="0" pageNumber="79">S. scapularis</emphasis>
</taxonomicName>
can be easily distinguished from 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. by having strongly keeled ventrals, whereas in the new species ventrals are feebly to moderately keeled. 
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1879" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus aculeatus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus aculeatus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
and 
<taxonomicName genus="S." lsidName="S. scapularis" pageId="0" pageNumber="79" rank="species" species="scapularis">
<emphasis italics="true" pageId="0" pageNumber="79">S. scapularis</emphasis>
</taxonomicName>
also have a dorsolateral crest (absent in 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov.); and 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
lacks a postfemoral mite pocket (present in 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov.). 
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov. by having smaller dorsal scales with 43 to 53 vertebrals and 46 to 59 scales around midbody versus 32 to 38 vertebrals and 34 to 45 scales around midbody in 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov. The gular region in adult males of 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. is black, whereas in 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov. it is cream.
</paragraph>
<paragraph pageId="0" pageNumber="79">
The new species is most similar to 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsolateral and ventral views of adult males of three species of Stenocercus: (A, B) S. huancabambae, (C, D) S. aculeatus, (E, F) S. prionotus from northern-central Peru, and (G, H) S. prionotus from southern Peru. Photographs by: (A - D) Pablo J. Venegas, (E) Andy Barboza, (F) Diego Vasquez, and (G, H) German Chavez." httpUri="https://binary.pensoft.net/fig/476351" pageId="0" pageNumber="79">4A, B</figureCitation>
) with which it shares ventral scales strongly keeled, like the rest of the aforementioned species, and a strongly compressed tail. Furthermore, both species are geographically close at the northern extreme of the central Andes in the Department of Amazonas (
<bibRefCitation author="Cadle, JE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="79" pagination="1 - 96" refId="B10" refString="Cadle, JE, 1991. Systematics of lizards of the genus Stenocercus (Iguania: Tropiduridae) from northern Peru: New species and comments on relationships and distribution patterns. Proceedings of the Academy of Natural Sciences of Philadelphia 143: 1 - 96" title="Systematics of lizards of the genus Stenocercus (Iguania: Tropiduridae) from northern Peru: New species and comments on relationships and distribution patterns." volume="143" year="1991">Cadle 1991</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.2006.00284.x" author="Torres-Carvajal, O" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="79" pagination="311 - 326" refId="B86" refString="Torres-Carvajal, O, 2007b. Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus). Zoologica Scripta 36: 311 - 326, DOI: https://doi.org/10.1111/j.1463-6409.2006.00284.x" title="Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus)." url="https://doi.org/10.1111/j.1463-6409.2006.00284.x" volume="36" year="2007 b">Torres-Carvajal 2007b</bibRefCitation>
). Nevertheless, the new species can be distinguished from 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
(characters in parentheses) by having the parietal eye not visible through the interparietal cornea (visible); postfemoral mite pocket as a distinct, deep slit-like opening (shallow slit-like opening, Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral view of the postfemoral mite pocket in three species of Stenocercus: (A) S. catherineae sp. nov. (CORBIDI 21365), (B) S. philmayi sp. nov. (CORBIDI 21092), and (C) S. huancabambae (CORBIDI 19747). Photographs by Luis A. Garcia-Ayachi. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/476350" pageId="0" pageNumber="79">3</figureCitation>
); dark patch covering most of the gular region of adult males present (absent); black patch on ventral surface of neck absent (present as a circular or elongate blotch, Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsolateral and ventral views of adult males of three species of Stenocercus: (A, B) S. huancabambae, (C, D) S. aculeatus, (E, F) S. prionotus from northern-central Peru, and (G, H) S. prionotus from southern Peru. Photographs by: (A - D) Pablo J. Venegas, (E) Andy Barboza, (F) Diego Vasquez, and (G, H) German Chavez." httpUri="https://binary.pensoft.net/fig/476351" pageId="0" pageNumber="79">4B</figureCitation>
); and a strongly compressed tail in adult males (very strongly compressed tail as a tape-like along its proximal two thirds).
</paragraph>
<paragraph pageId="0" pageNumber="79">
At the locality of Cuispes, and probably also in Huembo, 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName genus="S." lsidName="S. flagracanthus" pageId="0" pageNumber="79" rank="species" species="flagracanthus">
<emphasis italics="true" pageId="0" pageNumber="79">S. flagracanthus</emphasis>
</taxonomicName>
sp. nov. exist in sympatry; however, both species are strikingly different. 
<taxonomicName genus="S." lsidName="S. flagracanthus" pageId="0" pageNumber="79" rank="species" species="flagracanthus">
<emphasis italics="true" pageId="0" pageNumber="79">S. flagracanthus</emphasis>
</taxonomicName>
sp. nov. has a relatively short tail armed with projecting spines, whereas 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. has a long tail, compressed laterally, and a projecting vertebral crest.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/476350" pageId="0" pageNumber="79" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 3.</emphasis>
Lateral view of the postfemoral mite pocket in three species of 
<taxonomicName authorityName="A.M.C.Dumeril &amp; Bibron" authorityYear="1837" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus</emphasis>
</taxonomicName>
: (
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
) 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. (CORBIDI 21365), (
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
) 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov. (CORBIDI 21092), and (
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
) 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
(CORBIDI 19747). Photographs by Luis A. 
<normalizedToken originalValue="García-Ayachi">Garcia-Ayachi</normalizedToken>
. Scale bars: 5 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/476351" pageId="0" pageNumber="79" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 4.</emphasis>
Dorsolateral and ventral views of adult males of three species of 
<taxonomicName authorityName="A.M.C.Dumeril &amp; Bibron" authorityYear="1837" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus</emphasis>
</taxonomicName>
: (
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
) 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
, (
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
) 
<taxonomicName genus="S." lsidName="S. aculeatus" pageId="0" pageNumber="79" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">S. aculeatus</emphasis>
</taxonomicName>
, (
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
) 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
from northern-central Peru, and (
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
) 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
from southern Peru. Photographs by: (
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
) Pablo J. Venegas, (
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
) Andy Barboza, (
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
) Diego Vasquez, and (
<emphasis bold="true" pageId="0" pageNumber="79">G, H</emphasis>
) German 
<normalizedToken originalValue="Chávez">Chavez</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="definition">
<paragraph pageId="0" pageNumber="79">Definition.</paragraph>
<paragraph pageId="0" pageNumber="79">
(1) Maximum SVL in males 83 mm (
<emphasis italics="true" pageId="0" pageNumber="79">n</emphasis>
= 3); (2) SVL in females 75 mm (
<emphasis italics="true" pageId="0" pageNumber="79">n</emphasis>
= 2); (3) vertebrals 43-53; (4) paravertebrals 62-73; (5) scales around midbody 46-59; (6) supraoculars 4; (7) internasals 4-6; (8) postrostrals 2-4; (9) loreals 5-6; (10) gulars 22-26; (11) subdigitals on Finger IV 18-21; (12) subdigitals on Toe IV 25-30; (13) posthumeral mite pocket present as a deep depression with a wide opening [Type 3 of 
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.2006.00284.x" author="Torres-Carvajal, O" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="79" pagination="311 - 326" refId="B86" refString="Torres-Carvajal, O, 2007b. Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus). Zoologica Scripta 36: 311 - 326, DOI: https://doi.org/10.1111/j.1463-6409.2006.00284.x" title="Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus)." url="https://doi.org/10.1111/j.1463-6409.2006.00284.x" volume="36" year="2007 b">Torres-Carvajal (2007b)</bibRefCitation>
]; (14) postfemoral mite pocket present as a distinct pocket with a posteroventrally oriented slit-like opening [Type 2 of 
<bibRefCitation DOI="https://doi.org/10.1111/j.1463-6409.2006.00284.x" author="Torres-Carvajal, O" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="79" pagination="311 - 326" refId="B86" refString="Torres-Carvajal, O, 2007b. Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus). Zoologica Scripta 36: 311 - 326, DOI: https://doi.org/10.1111/j.1463-6409.2006.00284.x" title="Phylogeny and biogeography of a large radiation of Andean Lizards (Iguania, Stenocercus)." url="https://doi.org/10.1111/j.1463-6409.2006.00284.x" volume="36" year="2007 b">Torres-Carvajal (2007b)</bibRefCitation>
]; (15) parietal eye not visible through interparietal cornea in any specimens (
<emphasis italics="true" pageId="0" pageNumber="79">n</emphasis>
= 6); (16) scales on occipitoparietal region large, multicarinate, not imbricate; (17) projecting angulate temporals present; (18) row of enlarged supraoculars present, occupying most of supraocular region; (19) scales on frontonasal region and supraoculars slightly imbricate, multicarinate; (20) preauricular fringe present, short; (21) neck folds absent; (22) lateral and dorsal nuchals similar in size; (23) posterior gulars rhomboidal, projected posteriorly, keeled and imbricate, not notched; (24) lateral and dorsal body scales similar in size; (25) vertebrals larger than adjacent paravertebrals, forming a distinct vertebral crest; (26) dorsolateral crest absent; (27) ventrals keeled, imbricate, mucronate; (28) scales on posterior surfaces of thighs keeled, imbricate, mucronate; (29) inguinal granular pocket absent; (30) inguinal groove absent; (31) preanals not projecting; (32) tail strongly compressed laterally in adult males; (33) tail length 65-70% of total length; (34) caudal whorls per autotomic segment three; (35) caudals not spinose; (36) dark brown stripe extending anterodorsally from subocular region to supraciliaries always present; (37) dark patch extensively covering gular region of females present; (38) dark patch covering gular region in adult males present; (39) black patch on ventral surface of neck in adult males absent; (40) dark midventral longitudinal mark such as faint line, conspicuous stripe, or extensive patch in adult males absent; (41) dark patches on ventral surface of thighs in adult males absent; (42) two xiphisternal and three postxiphisternal pairs of inscriptional ribs fused medially, forming three chevrons (Pattern 6A of 
<bibRefCitation DOI="https://doi.org/10.1655/03-15" author="Torres-Carvajal, O" journalOrPublisher="Herpetologica" pageId="0" pageNumber="79" pagination="75 - 83" refId="B84" refString="Torres-Carvajal, O, 2004. The abdominal skeleton of Tropidurid Lizards (Squamata: Tropiduridae). Herpetologica 60: 75 - 83, DOI: https://doi.org/10.1655/03-15" title="The abdominal skeleton of Tropidurid Lizards (Squamata: Tropiduridae)." url="https://doi.org/10.1655/03-15" volume="60" year="2004">Torres-Carvajal 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description of the holotype.</paragraph>
<paragraph pageId="0" pageNumber="79">
Adult male (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stenocercus catherineae sp. nov. preserved holotype, adult male, SVL 82 mm (CORBIDI 21365): dorsal (A), lateral (B), and ventral (C) views of the head; dorsal (D) and ventral (E) views of the entire specimen. Photographs by Luis A. Garcia-Ayachi. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/476348" pageId="0" pageNumber="79">1</figureCitation>
); SVL 82 mm; TL 156 mm; maximum head width 15.7 mm; head length 18.7 mm; head height 12.8 mm; occipitals, parietals, interparietals, and postparietals large, multicarinate, slightly imbricate; parietal eye not visible; supraoculars in four rows, multicarinate, slightly imbricate, subequal in size; one canthal; canthal not in contact with the nasal; scales on frontonasal region slightly imbricate and multicarinate; internasals four; postrostrals three, two most lateral wider than long on each side, medial postrostrals as long as wide; supralabials four; infralabials five; loreals four; lorilabials in one row; preocular one, in contact with canthal; lateral temporals keeled, imbricate; gulars in 22 rows between tympanic openings; all gulars keeled, imbricate, apical pit absent; second infralabial not in contact with second and third sublabials; mental in contact with first pair of infralabials; lateral and dorsal scales of body and neck keeled, imbricate, mucronate; lateral and dorsal body scales similar in size; scales around midbody 46; vertebrals larger than dorsals, 43 scales on vertebral row, serrate vertebral crest present; paravertebrals 65; ventrals broad, rhomboidal, keeled, imbricate; preauricular fringe short, composed of three enlarged scales, all similar in size; antegular, gular, postauricular, oblique, supraauricular, longitudinal and antehumeral neck folds absent; limb scales keeled, imbricate; ventral scales of hindlimbs keeled and ventral scales of upper arms keeled, mucronate; lamellae on Finger IV 18; lamellae on Toe IV 27; tail strongly compressed laterally; caudals keeled, imbricate, mucronate; basal subcaudals keeled, imbricate; tail length 1.9 times SVL; posthumeral mite pocket present as a deep depression with a wide opening; postfemoral mite pocket present as a distinct deep pocket with a posteroventrally oriented slit-like opening; postfemoral region composed of imbricate, smooth scales that become keeled towards the tail.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration in life">
<paragraph pageId="0" pageNumber="79">Coloration in life</paragraph>
<paragraph pageId="0" pageNumber="79">
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsolateral and ventral views of Stenocercus catherineae sp. nov. in life: (A, B) holotype, adult male, SVL 82 mm (CORBIDI 21365); (C, D) adult male, SVL 79 mm (CORBIDI 21366); (E, F) adult female, SVL 75 mm (CORBIDI 21367); (G, H) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas." httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79">2A, B</figureCitation>
). Dorsum pale brown with the first two chevrons over the vertebral line black and the rest slightly darker than the background; cream line extending vertically from the arm insertion to the scapular region surrounded by a black blotch; dorsal surface of limbs darker than the dorsum with faint dirty cream transverse bars; flanks reddish brown, including the tail, becoming red toward the venter and dotted with white; subocular and loreal regions creamy white; dorsal surface of head black with the superciliaries and rostral cream; labials, sublabials and mental black, extending as an irregular longitudinal stripe to the neck; gular region black with cream irregular blotches to the sides; a black irregular stripe extends from the gular region ventrolaterally to the arm insertion; ventral surface of neck, chest and forelimbs are dirty cream with a black spot ventrolaterally in the arm; belly and ventral surface of tail pink; pelvic region and ventral surface of hindlimbs dirty cream. The iris is reddish brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration in preservative">
<paragraph pageId="0" pageNumber="79">Coloration in preservative</paragraph>
<paragraph pageId="0" pageNumber="79">
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stenocercus catherineae sp. nov. preserved holotype, adult male, SVL 82 mm (CORBIDI 21365): dorsal (A), lateral (B), and ventral (C) views of the head; dorsal (D) and ventral (E) views of the entire specimen. Photographs by Luis A. Garcia-Ayachi. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/476348" pageId="0" pageNumber="79">1D, E</figureCitation>
). It is similar to the coloration in life with the dorsal surface of trunk, limbs and tail darker than in life. Moreover, the reddish coloration of the flanks almost disappears.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="intraspecific variation">
<paragraph pageId="0" pageNumber="79">Intraspecific variation.</paragraph>
<paragraph pageId="0" pageNumber="79">
Measurement and scutellation characters of 
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. are presented in Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Variation in scutellation and sexual dimorphism in snout-vent length (mm) of Stenocercus aculeatus, S. catherineae sp. nov., S. dracopennatus sp. nov., S. philmayi sp. nov., and their most similar species S. angulifer and S. huancabambae. Range followed by mean + / - standard deviation is given for quantitative characters if applicable." httpUri="http://table.plazi.org/id/D432C09A71DA6BF568AF606FA5DC9345" pageId="0" pageNumber="79" tableUuid="D432C09A71DA6BF568AF606FA5DC9345">1</tableCitation>
. The second infralabial is in contact with the third sublabial in all specimens, and the first pair of postmentals are not in contact medially in two specimens (CORBIDI 501 and 21367). The three male paratypes are identical to the holotype, including a juvenile (CORBIDI 18662), varying only by having few scattered white dots on the head and, the black patch on the gular region extends to the ventral surface of the neck as a bold band (CORBIDI 21366) (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsolateral and ventral views of Stenocercus catherineae sp. nov. in life: (A, B) holotype, adult male, SVL 82 mm (CORBIDI 21365); (C, D) adult male, SVL 79 mm (CORBIDI 21366); (E, F) adult female, SVL 75 mm (CORBIDI 21367); (G, H) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas." httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79">2C, D</figureCitation>
). Ventral scales in the juvenile male are strongly keeled and mucronate.
</paragraph>
<paragraph pageId="0" pageNumber="79">
Sexual dimorphism is evident in adult individuals. In two adult female paratypes (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsolateral and ventral views of Stenocercus catherineae sp. nov. in life: (A, B) holotype, adult male, SVL 82 mm (CORBIDI 21365); (C, D) adult male, SVL 79 mm (CORBIDI 21366); (E, F) adult female, SVL 75 mm (CORBIDI 21367); (G, H) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas." httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79">2E, H</figureCitation>
) dorsal coloration is dusty brown with cinnamon vertebral chevrons along the back and tail, and cinnamon blotches along the flanks; hindlimbs with scattered dark brown transverse stripes; head in both specimens are darker than the rest of body, being dark gray (CORBIDI 501) or dark brown (CORBIDI 21367); sides of head grayish white (CORBIDI 21367) or dark gray (CORBIDI 501) with the loreal and subocular region white, and labials brown. Ventral coloration is pale brown with the gular region dark brown (CORBIDI 21367) or dusty cream with the gular region dark gray (CORBIDI 501) but both specimens have a faint pink hue on belly and base of tail (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsolateral and ventral views of Stenocercus catherineae sp. nov. in life: (A, B) holotype, adult male, SVL 82 mm (CORBIDI 21365); (C, D) adult male, SVL 79 mm (CORBIDI 21366); (E, F) adult female, SVL 75 mm (CORBIDI 21367); (G, H) adult female, SVL 75 mm (CORBIDI 501). Photographs by Pablo J. Venegas." httpUri="https://binary.pensoft.net/fig/476349" pageId="0" pageNumber="79">2F, H</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="D432C09A71DA6BF568AF606FA5DC9345" httpUri="http://table.plazi.org/id/D432C09A71DA6BF568AF606FA5DC9345" pageId="0" pageNumber="79" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Table 1.</emphasis>
Variation in scutellation and sexual dimorphism in snout-vent length (mm) of 
<taxonomicName authorityName="O'Shaughnessy" authorityYear="1879" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus aculeatus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus aculeatus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName genus="S." lsidName="S. dracopennatus" pageId="0" pageNumber="79" rank="species" species="dracopennatus">
<emphasis italics="true" pageId="0" pageNumber="79">S. dracopennatus</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov., and their most similar species 
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
and 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
. Range followed by mean 
<normalizedToken originalValue="±">+/-</normalizedToken>
standard deviation is given for quantitative characters if applicable.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="79">
<table pageId="0" pageNumber="79">
<tr pageId="0" pageNumber="79">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="79" rowspan="1">-</td>
<td colspan="3" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. aculeatus" pageId="0" pageNumber="79" rank="species" species="aculeatus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. aculeatus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. dracopennatus" pageId="0" pageNumber="79" rank="species" species="dracopennatus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. dracopennatus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">Character</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">CORBIDI 1712</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">CORBIDI 11483</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 38</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 6</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 3</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 32</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">n = 8</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Scales around midbody</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">41</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">37</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">32-37</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">38-50</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">46-59</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">39-45</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">37-53</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">34-45</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
35.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.00
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
42.10 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.37
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
53.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
5.44
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
41.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.21
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
43.94 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.20
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
41.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.70
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Vertebrals</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">37</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">38</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">31-39</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">37-47</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">43-53</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">38-40</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">37-51</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">32-38</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
35.20 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.27
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
41.63 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.25
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
47.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
4.76
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
39.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.15
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
43.39 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.40
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
35.88 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.96
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Paravertebrals</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">50</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">53</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">41-48</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">37-57</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">62-73</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">53-57</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">48-64</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">49-59</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
45.20 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.95
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
50.42 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.81
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
68.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
4.37
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
54.67 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.08
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
56.31 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.74
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
55.14 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.48
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Gulars</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">20</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">18</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">15-18</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">16-20</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">22-26</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">19-20</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">18-28</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">20-24</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
16.60 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.14
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
18.47 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.76
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
24.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.63
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
19.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.58
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
20.93 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.64
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
21.75 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.75
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Supraoculars</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">3-6</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-5</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">5</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Internasals</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-8</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-6</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4-7</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">3-5</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">4</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Subdigitals Finger IV</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">21</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">21</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">18-19</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">15-19</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">18-21</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">19-21</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">14-20</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">18-21</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
18.80 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.45
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
17.28 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.11
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
19.33 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.21
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
20.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.00
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
16.81 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.41
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
19.25 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.16
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Subdigitals Toe IV</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">31</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">27</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">23-27</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">19-25</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">25-30</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">26-28</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">20-28</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">28-31</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
25.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.58
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
22.84 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.44
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
27.50 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.64
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
27.00 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.00
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
24.62 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.56
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
29.25 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.16
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">Tail length/total length</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">0.72</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="2">0.72</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.70 - 0.73</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.68-0.73</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.66-0.69 (n=5)</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.69-0.73</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.67-0.70</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">0.67-0.72 (n=7)</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.71 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.70 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.67 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.71 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.69 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01
</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">
0.71 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02
</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">Maximun SVL males</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">81</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">95</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">103</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">96</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">82</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">89</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">99</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">95</td>
</tr>
<tr pageId="0" pageNumber="79">
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">Maximun SVL females</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">unknown</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">unknown</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">91</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">82</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">75</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">unknown</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">75</td>
<td colspan="1" pageId="0" pageNumber="79" rowspan="1">74</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="distribution">
<paragraph pageId="0" pageNumber="79">Distribution and natural history observations.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Venegas &amp; García-Ayachi &amp; Chávez-Arribasplata &amp; Chávez &amp; Wong &amp; García-Bravo" authorityYear="2020" class="Reptilia" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus catherineae" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">Stenocercus catherineae</emphasis>
</taxonomicName>
sp. nov. is only known from three proximate localities at Huembo and Cuispes in the northern extreme of the Cordillera Central at the 
<normalizedToken originalValue="Río">Rio</normalizedToken>
Utcubamba basin in the Department of Amazonas, at elevations between 1466 to 2085 m a.s.l. (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of S. catherineae sp. nov. (red triangles), S. dracopennatus sp. nov. (yellow star), and S. philmayi sp. nov. (blue squares), and the similar species: S. aculeatus (fuchsia diamonds), S. angulifer (blue diamonds), S. huancabambae (orange diamonds), S. prionotus (turquoise diamonds), and S. scapularis (red diamonds). Symbols with a dot in the middle correspond to type localities. White circles with a dot are important cities for reference." httpUri="https://binary.pensoft.net/fig/476352" pageId="0" pageNumber="79">5</figureCitation>
). According to the terrestrial ecoregions of the world by 
<bibRefCitation DOI="https://doi.org/10.1641/0006-3568(2001)051[0933:TEOTWA]2.0.CO;2" author="Olson, DM" journalOrPublisher="The Annals and magazine of natural history" pageId="0" pageNumber="79" refId="B69" refString="Olson, DM, Dinerstein, E, Wikramanayake, ED, Burgess, ND, Powell, GVN, Underwood, EC, D'Amico, JA, Itoua, I, Strand, HE, Morrison, JC, Loucks, CJ, Allnutt, TF, Ricketts, TH, Kura, Y, Lamoreux, JF, Wettengel, WW, Hedao, P, Kassem, KR, 2001. Terrestrial Ecoregions of the World: A new Map of Life on Earth. BioScience 51: 933-938. https://doi.org/10.1641/0006-3568(2001)051[0933:TEOTWA]2.0.CO;2" title="Terrestrial Ecoregions of the World: A new Map of Life on Earth. BioScience 51: 933 - 938." url="https://doi.org/10.1641/0006-3568(2001)051[0933:TEOTWA]2.0.CO;2" year="2001">Olson et al. (2001)</bibRefCitation>
, this species occurs in the Peruvian Yungas ecoregion. The habitat at the type locality of 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. is a steep area located on the sides of the 
<normalizedToken originalValue="Río">Rio</normalizedToken>
Chido with presence of corn, coffee and fruit plantations, and some patches of secondary forest and shrub vegetation. Several individuals were observed basking on fallen tree trunks or at the base of bushes close to trails between 1000 and 1200 hours. When the lizards were disturbed, they ran to hide inside fern patches. No other lizards or reptiles were observed in sympatry.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/476352" pageId="0" pageNumber="79" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 5.</emphasis>
Distribution map of 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. (red triangles), 
<taxonomicName genus="S." lsidName="S. dracopennatus" pageId="0" pageNumber="79" rank="species" species="dracopennatus">
<emphasis italics="true" pageId="0" pageNumber="79">S. dracopennatus</emphasis>
</taxonomicName>
sp. nov. (yellow star), and 
<taxonomicName genus="S." lsidName="S. philmayi" pageId="0" pageNumber="79" rank="species" species="philmayi">
<emphasis italics="true" pageId="0" pageNumber="79">S. philmayi</emphasis>
</taxonomicName>
sp. nov. (blue squares), and the similar species: 
<taxonomicName genus="S." lsidName="S. aculeatus" pageId="0" pageNumber="79" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="79">S. aculeatus</emphasis>
</taxonomicName>
(fuchsia diamonds), 
<taxonomicName genus="S." lsidName="S. angulifer" pageId="0" pageNumber="79" rank="species" species="angulifer">
<emphasis italics="true" pageId="0" pageNumber="79">S. angulifer</emphasis>
</taxonomicName>
(blue diamonds), 
<taxonomicName genus="S." lsidName="S. huancabambae" pageId="0" pageNumber="79" rank="species" species="huancabambae">
<emphasis italics="true" pageId="0" pageNumber="79">S. huancabambae</emphasis>
</taxonomicName>
(orange diamonds), 
<taxonomicName genus="S." lsidName="S. prionotus" pageId="0" pageNumber="79" rank="species" species="prionotus">
<emphasis italics="true" pageId="0" pageNumber="79">S. prionotus</emphasis>
</taxonomicName>
(turquoise diamonds), and 
<taxonomicName genus="S." lsidName="S. scapularis" pageId="0" pageNumber="79" rank="species" species="scapularis">
<emphasis italics="true" pageId="0" pageNumber="79">S. scapularis</emphasis>
</taxonomicName>
(red diamonds). Symbols with a dot in the middle correspond to type localities. White circles with a dot are important cities for reference.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="79">
At Cuispes only one individual was collected in the croplands close to the village. Other squamate reptiles collected with 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. at Cuispes were 
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Atractus</emphasis>
</taxonomicName>
sp. Wagler, 1828, 
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Dipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dipsas palmeri" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="palmeri">
<emphasis italics="true" pageId="0" pageNumber="79">Dipsas palmeri</emphasis>
</taxonomicName>
Boulenger, 1912, and 
<taxonomicName genus="S." lsidName="S. flagracanthus" pageId="0" pageNumber="79" rank="species" species="flagracanthus">
<emphasis italics="true" pageId="0" pageNumber="79">S. flagracanthus</emphasis>
</taxonomicName>
sp. nov. At Cocachimba, the two specimens collected were found basking at 1000 hours on the base of rocky fences with bushes along a trail. An uncollected adult female was observed basking at 900 hours on a fallen wall of an abandoned house. The general landscape at Cocachimba is composed of croplands of corn, fruit and sugar cane bordered by rocky fences and some streams with narrow fringes of riverine forest or shrub vegetation. Other species of squamate reptiles collected with 
<taxonomicName genus="S." lsidName="S. catherineae" pageId="0" pageNumber="79" rank="species" species="catherineae">
<emphasis italics="true" pageId="0" pageNumber="79">S. catherineae</emphasis>
</taxonomicName>
sp. nov. at Cocachimba were: 
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Atractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="79">Atractus</emphasis>
</taxonomicName>
sp., 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Colubridae" genus="Chironius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chironius exoletus" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="exoletus">
<emphasis italics="true" pageId="0" pageNumber="79">Chironius exoletus</emphasis>
</taxonomicName>
Linnaeus, 1758, 
<taxonomicName baseAuthorityName="Sentzen" baseAuthorityYear="1796" class="Reptilia" family="Colubridae" genus="Mastigodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastigodryas boddaerti" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="boddaerti">
<emphasis italics="true" pageId="0" pageNumber="79">Mastigodryas boddaerti</emphasis>
</taxonomicName>
Sentzen, 1796, and 
<taxonomicName class="Reptilia" family="Gymnophthalmidae" genus="Petracola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Petracola angustisoma" order="Squamata" pageId="0" pageNumber="79" phylum="Chordata" rank="species" species="angustisoma">
<emphasis italics="true" pageId="0" pageNumber="79">Petracola angustisoma</emphasis>
</taxonomicName>
<normalizedToken originalValue="Echevarría">Echevarria</normalizedToken>
&amp; Venegas, 2015.
</paragraph>
<paragraph pageId="0" pageNumber="79">
The female paratype (CORBIDI 21367) collected during the rainy season (December 2019) had 2 well developed follicles, one in the left and the other in the right ovary. The sizes of these follicles are 10.83 
<normalizedToken originalValue="×">x</normalizedToken>
9.67 mm and 10.99 
<normalizedToken originalValue="×">x</normalizedToken>
9.53 mm; their volumes were 530.2 mm³ and 522.6 mm³, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The specific name is a noun in the genitive case and is a patronym for Catherine Dupont, a Peruvian veterinary specialist in One Health, who is actively working searching and monitoring viruses and other zoonotic pathogens. The specific name of this lizard is in recognition of her passion for the natural world and its creatures, and her selfless support of the herpetological division of CORBIDI.</paragraph>
</subSubSection>
</treatment>
</document>