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<mods:title id="A535DB6B16CCBED3306DF849F3DADAA3">Phylogenetic evidence and morphological diagnosis for a new species of Parasesarma De Man, 1895 (Decapoda, Brachyura, Sesarmidae) from Sulawesi, Indonesia</mods:title>
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<mods:namePart id="E9B136ECFEB2EC5F308A66C029F7FC5E">Shahdadi, Adnan</mods:namePart>
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<mods:affiliation id="16BAC9F424AF86A27C0CE4A4D6879DF8">Department of Marine Biology, Faculty of Marine Sciences and Technology, University of Hormozgan, Bandar Abbas, Iran. adnan 1361 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5131 - 7700</mods:affiliation>
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<mods:namePart id="FAB3B70D7145565DAA682087A0C57227">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="DB2D14E7FCB223613E9B85AA435A6293">Mendoza, Jose Christopher E.</mods:namePart>
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<mods:affiliation id="6FE0EAE3FA31A3EE028ABEDD7780642F">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377, Singapore. jcmendoza @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 9729 - 5703</mods:affiliation>
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<emphasis id="3A9CEAF3C472831E1BB80B72FE46F9BA" bold="true" box="[151,504,1590,1617]" pageId="1" pageNumber="568">
<taxonomicName id="CFE84D62C472831E1BB80B72FE11F9BA" ID-CoL="c13f4821-166b-461e-ba47-4420faf15783" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[151,431,1590,1617]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C472831E1BB80B72FE11F9BA" bold="true" box="[151,431,1590,1617]" italics="true" pageId="1" pageNumber="568">Parasesarma rahayuae</emphasis>
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<paragraph id="085736E1C472831E1BB80B1EFE67F99E" blockId="1.[151,504,1590,1653]" box="[151,473,1626,1653]" pageId="1" pageNumber="568">
(
<figureCitation id="90D32A64C472831E1BB00B1EFEB2F99E" box="[159,268,1627,1653]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1847,1871]" captionStartId-1="5.[151,250,1688,1712]" captionStartId-2="6.[151,250,1016,1040]" captionTargetBox-0="[194,1394,194,1808]" captionTargetBox-1="[201,1386,181,1665]" captionTargetBox-2="[151,1436,181,992]" captionTargetId-0="figure-20@4.[181,1407,181,1823]" captionTargetId-1="figure-15@5.[188,1399,181,1665]" captionTargetId-2="figure-20@6.[151,1436,181,992]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." captionText-1="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." captionText-2="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi-0="http://doi.org/10.5281/zenodo.8282243" figureDoi-1="http://doi.org/10.5281/zenodo.8282247" figureDoi-2="http://doi.org/10.5281/zenodo.8282249" httpUri-0="https://zenodo.org/record/8282243/files/figure.png" httpUri-1="https://zenodo.org/record/8282247/files/figure.png" httpUri-2="https://zenodo.org/record/8282249/files/figure.png" pageId="1" pageNumber="568">Figs. 2‒4</figureCitation>
, 
<figureCitation id="90D32A64C472831E1A380B1FFEE0F99E" box="[279,350,1626,1653]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="1" pageNumber="568">5A, D</figureCitation>
, 
<figureCitation id="90D32A64C472831E1A460B1EFE6FF99E" box="[361,465,1627,1653]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="1" pageNumber="568">6A, D, G</figureCitation>
)
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<emphasis id="3A9CEAF3C472831E1BB80BE7FE3FF957" bold="true" box="[151,385,1698,1724]" pageId="1" pageNumber="568">Material examined</emphasis>
. 
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<typeStatus id="D7538843C472831E1AB80BE6FDBAF956" box="[407,516,1699,1725]" pageId="1" pageNumber="568" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="584E9E9AC472831E193A0BE6FD27F956" box="[533,665,1698,1725]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="568" type="Museum">MZB 5709</specimenCode>
(ex. 
<specimenCode id="584E9E9AC472831E19F60BE6FC24F956" box="[729,922,1698,1725]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="568">ZRC 2019.1055</specimenCode>
), male (11.47×10.11), 
<collectingCountry id="70FF7671C472831E1F810BE6FAA0F956" box="[1198,1310,1699,1725]" name="Indonesia" pageId="1" pageNumber="568">Indonesia</collectingCountry>
, 
<collectingRegion id="CA2CF803C472831E1E1F0BE6FA26F956" box="[1328,1432,1699,1725]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="568">Sulawesi</collectingRegion>
, Manado, Bitung, Pulau Lembeh, 
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, mangrove near NAD Resort, coll. 
<collectorName id="A51D5337C472831E1F0A0B82FB31F90A" box="[1061,1167,1735,1761]" pageId="1" pageNumber="568">H.H. Tan</collectorName>
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<date id="7C561021C472831E1FB40B82FAA3F90A" box="[1179,1309,1735,1761]" pageId="1" pageNumber="568" value="2018-01-12">
<collectingDate id="6C12E9C9C472831E1FB40B82FAA3F90A" box="[1179,1309,1735,1761]" pageId="1" pageNumber="568" value="2018-01-12">01.12.2018</collectingDate>
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; 
<materialsCitation id="B8803CBCC472831E1E070B82FBDBF8C2" ID-GBIF-Occurrence="4400641306" collectingDate="2018-01-12" collectionCode="ZRC" collectorName="H. H. Tan" country="Indonesia" location="Mawali Bay" pageId="1" pageNumber="568" specimenCode="ZRC 2019.1055" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="D7538843C472831E1E070B82FA27F90A" box="[1320,1433,1735,1761]" pageId="1" pageNumber="568" type="paratype">Paratypes</typeStatus>
: 
<specimenCode id="584E9E9AC472831E1BB80BAEFEF3F8EE" box="[151,333,1770,1797]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="568">ZRC 2019.1055</specimenCode>
, 
<specimenCount id="1EEEFD68C472831E1A780BAEFE12F8EE" box="[343,428,1771,1797]" pageId="1" pageNumber="568" type="male">3 males</specimenCount>
(11.32 × 9.6; 10.87 × 9.4; 10.86 × 9.42), 
<specimenCount id="1EEEFD68C472831E185D0BAEFC63F8EE" box="[882,989,1771,1797]" pageId="1" pageNumber="568" type="female">6 females</specimenCount>
((ovig. 11.90 × 9.77; ovig. 11.47 × 9.35; ovig. 10.93 × 9.12; ovig. 9.88 ×8.71; 9.1 ×7.7; 7.67 × 6.54), same data as the holotype.
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</paragraph>
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<emphasis id="3A9CEAF3C472831E1BE80A77FE6EF8A7" bold="true" box="[199,464,1842,1868]" pageId="1" pageNumber="568">Comparative material</emphasis>
. 
<taxonomicName id="CFE84D62C472831E1AF10A76FBBAF8A6" authority="Yeo, Rahayu &amp; Ng, 2004" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[478,1028,1843,1869]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C472831E1AF10A76FD61F8A6" box="[478,735,1843,1869]" italics="true" pageId="1" pageNumber="568">Parasesarma anambas</emphasis>
<materialsCitation id="B8803CBCC472831E19CA0A76FBBAF8A6" ID-GBIF-Occurrence="4400641307" box="[741,1028,1843,1869]" collectionCode="ZRC" country="Indonesia" county="Air Terjun" location="eastern Pulau Siantan" municipality="Sungei Temburun" pageId="1" pageNumber="568" specimenCode="ZRC 2003.0726" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<bibRefCitation id="6C794B10C472831E19CA0A76FBBAF8A6" author="Yeo, D. C. J. &amp; Rahayu, D. L. &amp; Ng, P. K. L." box="[741,1028,1843,1869]" pageId="1" pageNumber="568" pagination="79 - 88" refId="ref5226" refString="Yeo, D. C. J., Rahayu, D. L. &amp; Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas expedition 2002. Raffles Bulletin of Zoology, Supplement No. 11, 79 - 88." type="book chapter" year="2004">Yeo, Rahayu &amp; Ng, 2004</bibRefCitation>
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</taxonomicName>
: 
<specimenCode id="584E9E9AC472831E1F200A76FB78F8A6" box="[1039,1222,1843,1869]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="568">ZRC 2003.0726</specimenCode>
, 
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, 
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, 
<specimenCount id="1EEEFD68C472831E1BB80A12FF45F89A" box="[151,251,1879,1905]" pageId="1" pageNumber="568" type="female">1 female</specimenCount>
, 
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, 
<taxonomicName id="CFE84D62C472831E1A500A12FDFCF89A" authority="Islands" authorityName="Islands" box="[383,578,1879,1905]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="anambas">Anambas Islands</taxonomicName>
, 
<location id="0D37603AC472831E19620A12FCFFF89A" LSID="urn:lsid:plazi:treatment:804187F7C47283181BB80B72FC97FDF2:0D37603AC472831E19620A12FCFFF89A" box="[589,833,1879,1905]" country="Indonesia" county="Air Terjun" municipality="Sungei Temburun" name="eastern Pulau Siantan" pageId="1" pageNumber="568">eastern Pulau Siantan</location>
, 
<location id="0D37603AC472831E18640A12FC66F89A" LSID="urn:lsid:plazi:treatment:804187F7C47283181BB80B72FC97FDF2:0D37603AC472831E18640A12FC66F89A" box="[843,984,1879,1905]" country="Indonesia" county="Air Terjun" municipality="Sungei Temburun" name="Teluk Baruk" pageId="1" pageNumber="568">Teluk Baruk</location>
, 
<collectingMunicipality id="E833AC9BC472831E18CC0A12FB10F89A" box="[995,1198,1879,1905]" pageId="1" pageNumber="568">Sungei Temburun</collectingMunicipality>
, 
<collectingCounty id="E1364E6DC472831E1F980A12FA95F89A" box="[1207,1323,1879,1905]" pageId="1" pageNumber="568">Air Terjun</collectingCounty>
waterfall, stn. EA-JL05.
</paragraph>
<paragraph id="085736E1C472831E1BE80ADAFBDFF836" blockId="1.[151,1437,1698,2013]" pageId="1" pageNumber="568">
<taxonomicName id="CFE84D62C472831E1BE80ADAFD60F852" authority="Li, Shih &amp; Ng, 2019" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[199,734,1951,1977]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C472831E1BE80ADAFE5FF853" box="[199,481,1951,1976]" italics="true" pageId="1" pageNumber="568">Parasesarma gemmatum</emphasis>
<bibRefCitation id="6C794B10C472831E1AC50ADAFD60F852" author="Li, J. - J. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[490,734,1951,1977]" pageId="1" pageNumber="568" pagination="40" refId="ref4223" refString="Li, J. - J., Shih, H. - T. &amp; Ng, P. K. L. (2019) Three new species and two new records of Parasesarma De Man, 1895 (Crustacea: Brachyura: Sesarmidae) from Taiwan and the Philippines from morphological and molecular evidence. Zoological Studies, 58, e 40." type="journal article" year="2019">Li, Shih &amp; Ng, 2019</bibRefCitation>
</taxonomicName>
, 
<materialsCitation id="B8803CBCC472831E19C40ADAFB9FF852" ID-GBIF-Occurrence="4400641305" box="[747,1057,1951,1977]" collectionCode="ZRC" pageId="1" pageNumber="568" specimenCode="ZRC 2019.1076" specimenCount="1" typeStatus="paratype">
<specimenCode id="584E9E9AC472831E19C40ADAFC1BF852" box="[747,933,1951,1977]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="568">ZRC 2019.1076</specimenCode>
, 
<typeStatus id="D7538843C472831E189C0ADAFB9FF852" box="[947,1057,1951,1977]" pageId="1" pageNumber="568" type="paratype">paratypes</typeStatus>
</materialsCitation>
, 
<specimenCount id="1EEEFD68C472831E1F000ADAFBC1F852" box="[1071,1151,1951,1977]" pageId="1" pageNumber="568" type="male">1 male</specimenCount>
, 
<materialsCitation id="B8803CBCC472831E1FA20ADAFD9EF836" ID-GBIF-Occurrence="4400641302" country="Taiwan" location="Pingtung" pageId="1" pageNumber="568" specimenCount="1" stateProvince="Pingtung" typeStatus="paratype">
1 ovig. female, 
<collectingCountry id="70FF7671C472831E1E6C0ADAFA26F852" box="[1347,1432,1951,1977]" name="Taiwan" pageId="1" pageNumber="568">Taiwan</collectingCountry>
, 
<collectingRegion id="CA2CF803C472831E1BB80A86FF41F836" box="[151,255,1987,2013]" country="Taiwan" name="Pingtung" pageId="1" pageNumber="568">Pingtung</collectingRegion>
, Kenting, Tanzih fishport
</materialsCitation>
; 
<materialsCitation id="B8803CBCC472831E19040A86FC85F836" ID-GBIF-Occurrence="4400641303" box="[555,827,1986,2013]" collectionCode="ZRC" country="Taiwan" pageId="1" pageNumber="568" specimenCode="ZRC 2023.0256" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCode id="584E9E9AC472831E19040A86FD5CF836" box="[555,738,1986,2013]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="568">ZRC 2023.0256</specimenCode>
, 
<specimenCount id="1EEEFD68C472831E19C20A86FC85F836" box="[749,827,1987,2013]" pageId="1" pageNumber="568" type="male">1 male</specimenCount>
</materialsCitation>
, 
<materialsCitation id="B8803CBCC472831E18680A86FBE3F836" ID-GBIF-Occurrence="4400641304" box="[839,1117,1987,2013]" country="Philippines" location="Cebu" pageId="1" pageNumber="568" specimenCount="1" stateProvince="Cebu" typeStatus="paratype">
<collectingCountry id="70FF7671C472831E18680A86FC76F836" box="[839,968,1987,2013]" name="Philippines" pageId="1" pageNumber="568">Philippines</collectingCountry>
, 
<collectingRegion id="CA2CF803C472831E18FC0A86FBB0F836" box="[979,1038,1987,2013]" country="Philippines" name="Cebu" pageId="1" pageNumber="568">Cebu</collectingRegion>
Island
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="5C976669C471831D1BB80A17FB1BF810" ID-DOI="http://doi.org/10.5281/zenodo.8282239" ID-Zenodo-Dep="8282239" httpUri="https://zenodo.org/record/8282239/files/figure.png" pageId="2" pageNumber="569" startId="2.[151,250,1874,1898]" targetBox="[221,1367,189,1843]" targetPageId="2" targetType="figure">
<paragraph id="085736E1C471831D1BB80A17FB1BF810" blockId="2.[151,1437,1874,2043]" pageId="2" pageNumber="569">
<emphasis id="3A9CEAF3C471831D1BB80A17FEAFF881" bold="true" box="[151,273,1874,1898]" pageId="2" pageNumber="569">FIGURE 1.</emphasis>
A, maximum likelihood (ML) phylogenetic consensus tree of 
<taxonomicName id="CFE84D62C471831D18550A16FBBFF881" authorityName="De Man" authorityYear="1895" box="[890,1025,1875,1898]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="3A9CEAF3C471831D18550A16FBBFF881" box="[890,1025,1875,1898]" italics="true" pageId="2" pageNumber="569">Parasesarma</emphasis>
</taxonomicName>
based on the COX1 gene and constructed in raxml GUI. 
<taxonomicName id="CFE84D62C471831D1A1E0A32FD04F865" authority="(De Haan, 1835)" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[305,698,1910,1935]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="3A9CEAF3C471831D1A1E0A32FDBBF865" box="[305,517,1911,1934]" italics="true" pageId="2" pageNumber="569">Parasesarma pictum</emphasis>
(
<bibRefCitation id="6C794B10C471831D193B0A32FD0DF865" author="De Haan, W." box="[532,691,1910,1935]" pageId="2" pageNumber="569" pagination="9 - 15" refId="ref3798" refString="De Haan, W. (1835) Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive Descriptio Animalium, quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suspecto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Apud Auctorem, Lugduni-Batavorum, pp. 25 - 64, pls. 9 - 15, 17, C, D." type="journal article" year="1835">De Haan, 1835</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="CFE84D62C471831D19C00A32FC42F864" authority="(Shen, 1940)" baseAuthorityName="Shen" baseAuthorityYear="1940" box="[751,1020,1911,1935]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="tripectinis">
<emphasis id="3A9CEAF3C471831D19C00A32FCCEF865" box="[751,880,1911,1934]" italics="true" pageId="2" pageNumber="569">P. tripectinis</emphasis>
(
<bibRefCitation id="6C794B10C471831D18500A32FC4BF864" author="Shen, C. - J." box="[895,1013,1911,1935]" pageId="2" pageNumber="569" pagination="255 - 262" refId="ref4952" refString="Shen, C. - J. (1940) Four new species of Brachyura from China Seas. Journal of the Hong Kong Fisheries Research Station, 1 (2), 255 - 262." type="journal article" year="1940">Shen, 1940</bibRefCitation>
)
</taxonomicName>
were used as outgroup (numbers on the branches correspond to bootstrap value), black arrow indicates clade of 
<taxonomicName id="CFE84D62C471831D185B0ADEFC53F858" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[884,1005,1947,1971]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C471831D185B0ADEFC53F858" box="[884,1005,1947,1971]" italics="true" pageId="2" pageNumber="569">P. rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C471831D18DB0ADEFB93F858" bold="true" box="[1012,1069,1947,1971]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="21AF5788C471831D18DB0ADEFB93F858" box="[1012,1069,1947,1971]" pageId="2" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; B, maximum parsimony haplotype network (COX1 gene, 628 bp) of close congeners, 
<taxonomicName id="CFE84D62C471831D19BB0AFAFBB4F83C" authority="Yeo, Rahayu &amp; Ng, 2004" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[660,1034,1983,2007]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C471831D19BB0AFAFCB8F83C" box="[660,774,1983,2007]" italics="true" pageId="2" pageNumber="569">P. anambas</emphasis>
<bibRefCitation id="6C794B10C471831D18240AFAFBB4F83C" author="Yeo, D. C. J. &amp; Rahayu, D. L. &amp; Ng, P. K. L." box="[779,1034,1983,2007]" pageId="2" pageNumber="569" pagination="79 - 88" refId="ref5226" refString="Yeo, D. C. J., Rahayu, D. L. &amp; Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas expedition 2002. Raffles Bulletin of Zoology, Supplement No. 11, 79 - 88." type="book chapter" year="2004">Yeo, Rahayu &amp; Ng, 2004</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CFE84D62C471831D1F3C0AFAFAD0F83C" authority="Li, Shih &amp; Ng, 2019" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[1043,1390,1983,2007]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C471831D1F3C0AFAFB24F83D" box="[1043,1178,1983,2006]" italics="true" pageId="2" pageNumber="569">P. gemmatum</emphasis>
<bibRefCitation id="6C794B10C471831D1F8F0AFAFAD0F83C" author="Li, J. - J. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1184,1390,1983,2007]" pageId="2" pageNumber="569" pagination="40" refId="ref4223" refString="Li, J. - J., Shih, H. - T. &amp; Ng, P. K. L. (2019) Three new species and two new records of Parasesarma De Man, 1895 (Crustacea: Brachyura: Sesarmidae) from Taiwan and the Philippines from morphological and molecular evidence. Zoological Studies, 58, e 40." type="journal article" year="2019">Li, Shih &amp; Ng, 2019</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="CFE84D62C471831D1BB80AA6FEAEF810" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[151,272,2019,2043]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C471831D1BB80AA6FEAEF810" box="[151,272,2019,2043]" italics="true" pageId="2" pageNumber="569">P. rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C471831D1A390AA6FEF1F810" bold="true" box="[278,335,2019,2043]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="21AF5788C471831D1A390AA6FEF1F810" box="[278,335,2019,2043]" pageId="2" pageNumber="569" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, constructed with PopART (the numbers and hatch marks refer to mutational steps).
</paragraph>
</caption>
<subSubSection id="40F2656AC470831C1BE80DD3FE9DFE62" pageId="3" pageNumber="570" type="diagnosis">
<paragraph id="085736E1C470831C1BE80DD3FE9DFE62" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
<emphasis id="3A9CEAF3C470831C1BE80DD3FE84FF5B" bold="true" box="[199,314,150,176]" pageId="3" pageNumber="570">Diagnosis</emphasis>
. Relatively long ambulatory legs. Carapace subquadrate, slightly broader than long, dorsal surface entirely covered with small granules, front moderately deflexed, sinuous in dorsal view, median postfrontal lobes slightly broader than lateral ones. Eyestalk longer than wide, cornea wider than eyestalk. Chelipeds without subdistal spine on dorsal border of merus; male chela with two transverse pectinated crests on the upper surface of palm, dactylus with 17‒19 transverse, broadly ovate tubercles. Male pleon triangular, somite 2 medially as long as lateral edges. G1 stout, straight, apical corneous process relatively long, bent at an angle of ~80° to vertical axis, aperture subterminal.
</paragraph>
</subSubSection>
<subSubSection id="40F2656AC470831C1BE80CD7FD37F956" pageId="3" pageNumber="570" type="description">
<paragraph id="085736E1C470831C1BE80CD7FD2AFD42" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
<emphasis id="3A9CEAF3C470831C1BE80CD7FEEFFE47" bold="true" box="[199,337,402,428]" pageId="3" pageNumber="570">Description</emphasis>
. Maximum cw of material examined = 
<quantity id="CF109B04C470831C18050CD6FC1CFE46" box="[810,930,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="3" pageNumber="570" unit="mm" value="11.9">11.90 mm</quantity>
. Carapace (
<figureCitation id="90D32A64C470831C1F030CD6FB2AFE46" box="[1068,1172,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Figs. 2A</figureCitation>
, 
<figureCitation id="90D32A64C470831C1F8C0CD6FB76FE46" box="[1187,1224,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">3A</figureCitation>
, 
<figureCitation id="90D32A64C470831C1FF70CD6FB45FE46" box="[1240,1275,403,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">4A</figureCitation>
) subquadrate, slightly broader than long, greatest width between exorbital angles (
<typeStatus id="D7538843C470831C18A60CF2FC53FE3A" box="[905,1005,439,465]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
cw/cl = 1.1); dorsal surface with small granules scattered all over (
<figureCitation id="90D32A64C470831C1AF70C9EFD8DFE1E" box="[472,563,475,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2C</figureCitation>
); front in 
<typeStatus id="D7538843C470831C19840C9EFCB3FE1E" box="[683,781,475,501]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
~0.53× carapace width, moderately deflexed, sinuous in dorsal view; postfrontal lobes (
<figureCitation id="90D32A64C470831C1ADD0CBAFDF7FDF2" box="[498,585,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2D</figureCitation>
) low but distinct, median lobes slightly broader than lateral lobes, separated by shallow furrow; dorsal regions moderately well indicated, gastric region demarcated, cardiac region separated from intestinal region; lateral branchial ridges prominent; anterolateral margin with sharp exorbital angle directed obliquely; lateral margins straight with no indication of epibranchial tooth, edged with row of short setae. Eyestalk longer than wide, cornea wider than eyestalk.
</paragraph>
<paragraph id="085736E1C470831C1BE80FF6FE3EFB2E" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
Chelipeds homochelous (
<figureCitation id="90D32A64C470831C1AC20FF6FDCFFD26" box="[493,625,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2A, B</figureCitation>
, 
<figureCitation id="90D32A64C470831C19AD0FF6FD19FD26" box="[642,679,691,717]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">3A</figureCitation>
); chelae (
<figureCitation id="90D32A64C470831C18320FF6FCC7FD26" box="[797,889,691,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2B</figureCitation>
) large (palm length ~0.75× carapace width in 
<typeStatus id="D7538843C470831C1BB80F92FF43FD1A" box="[151,253,727,753]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
), robust (length ~1.6× palm width in 
<typeStatus id="D7538843C470831C19B70F92FD41FD1A" box="[664,767,727,753]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
). Merus with granulate dorsal border, but no subdistal spine; ventral border granulate; anterior border granulate, with large subdistal spine; inner face smooth with a longitudinal rows of setae (
<figureCitation id="90D32A64C470831C1A6F0E5AFE24FCD2" box="[320,410,799,825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">Fig. 3B</figureCitation>
). Upper surface of palm with two transverse pectinated crests and a third crest consisting of granules (
<figureCitation id="90D32A64C470831C1A090E06FE1FFCB6" box="[294,417,835,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2D, E</figureCitation>
, 
<figureCitation id="90D32A64C470831C1A810E06FE6CFCB6" box="[430,466,835,861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">3D</figureCitation>
), distal (primary) crest composed of 10 or 11 tall teeth (varying on opposite claws of 
<typeStatus id="D7538843C470831C1BB80E22FF43FC6A" box="[151,253,871,897]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
), secondary crest well-developed, with 7 teeth; both crests ending on inner side in short swollen, tubercular ridge (
<figureCitation id="90D32A64C470831C1BCE0ECEFE84FC4E" box="[225,314,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">Fig. 3D</figureCitation>
) and several small granules at outer side; upper margin of palm with scattered setae; outer surface of palm with few scattered setae, granular except for smooth, punctate fixed finger (
<figureCitation id="90D32A64C470831C1F6B0EEAFB27FC22" box="[1092,1177,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2F</figureCitation>
); inner surface of palm generally smooth, except for some granules forming a low vertical ridge; ventral border of chela sinuous; length of cutting margin of fixed finger of 
<typeStatus id="D7538843C470831C193D0EB2FDCAFBFA" box="[530,628,1015,1041]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
~0.4× length of entire propodus. Dactylus (
<figureCitation id="90D32A64C470831C1F450EB2FB5AFBFA" box="[1130,1252,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2E‒H</figureCitation>
) slightly curved downward and inward, ~0.6× propodus length in 
<typeStatus id="D7538843C470831C19CE095EFCF9FBDE" box="[737,839,1051,1077]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
; dorsal surface bearing 17–19 transversely broadly ovate tubercles (
<figureCitation id="90D32A64C470831C1A60097AFE6DFBB2" box="[335,467,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Figs. 2E‒H</figureCitation>
, 
<figureCitation id="90D32A64C470831C1AF1097AFDBFFBB2" box="[478,513,1087,1113]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">3C</figureCitation>
), distinct to tip, proximal tubercles with transverse keel and longitudinal wrinkles, proximal tubercles positioned at inner part of upper dactylar face, row of ~6 rounded tubercles on proximal onethird of inner edge of dorsal surface; fingers with chitinous tips, cutting edge of both fingers with a series of variably sized teeth (
<figureCitation id="90D32A64C470831C1A3209EEFECDFB2E" box="[285,371,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="085736E1C470831C1BE8098AFCEAFADA" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
Ambulatory legs (
<figureCitation id="90D32A64C470831C1ABE098AFD94FB02" box="[401,554,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Figs. 2A, B, I</figureCitation>
, 
<figureCitation id="90D32A64C470831C191A098AFDE7FB02" box="[565,601,1231,1257]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">3A</figureCitation>
, 
<figureCitation id="90D32A64C470831C194A098AFD36FB02" box="[613,648,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">4A</figureCitation>
) relatively long; P4 longest, length (ischium–dactylus) ~1.8× carapace width, merus with anterior margin crenulated, ~2.3× as long as wide, propodus ~3.3× as long as wide, dactylus length ~0.9× length of propodus (all measures from 
<typeStatus id="D7538843C470831C19CE0852FCF6FADA" box="[737,840,1303,1329]" pageId="3" pageNumber="570" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="085736E1C470831C1BE8087EFE95FA76" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
Male pleon (
<figureCitation id="90D32A64C470831C1A7A087EFE7DFABE" box="[341,451,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[151,250,1847,1871]" captionTargetBox="[194,1394,194,1808]" captionTargetId="figure-20@4.[181,1407,181,1823]" captionTargetPageId="4" captionText="FIGURE 2. Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view." figureDoi="http://doi.org/10.5281/zenodo.8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="3" pageNumber="570">Fig. 2B, J</figureCitation>
) triangular; telson slightly longer than basal width, slightly longer than somite 6; somite 6 longer than others; somite 5 and 4 trapezoidal, somite 3 widest, laterally convex, somite 2 medially as long as lateral edges.
</paragraph>
<paragraph id="085736E1C470831C1BE808E2FBD2F9E2" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
G1 (
<figureCitation id="90D32A64C470831C1BD708E2FEC5FA2A" box="[248,379,1447,1473]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="3" pageNumber="570">Figs. 3E‒H</figureCitation>
, 
<figureCitation id="90D32A64C470831C1AA808E2FE14FA2A" box="[391,426,1447,1473]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="3" pageNumber="570">6A</figureCitation>
) stout, straight, stem triangular with blunt angles in cross-section; apical corneous process relatively long, bent at an angle of ~80° to vertical axis, arched in cross-section, distinctly wider than the rest of the apical corneous process when seen from the dorsal (sternal) view, aperture subterminal.
</paragraph>
<paragraph id="085736E1C470831C1BE80B56FD37F956" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
Females (
<figureCitation id="90D32A64C470831C1A190B56FEC5F9C6" box="[310,379,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">Fig. 4</figureCitation>
) with smaller chelipeds than in males; pectinated crest absent on palm, replaced by rows of granules; dactylus with 5 or 6 small, round tubercles on proximal half (
<figureCitation id="90D32A64C470831C18FD0B72FB8EF9BA" box="[978,1072,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">Fig. 4D</figureCitation>
). Pleon (
<figureCitation id="90D32A64C470831C1FB80B72FB4EF9BA" box="[1175,1264,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">Fig. 4B</figureCitation>
) broad, evenly ovate, broadest at somites 4 and 5, fringed with long setae, touching coxae of ambulatory legs. Vulva (
<figureCitation id="90D32A64C470831C1E300B1EFAC8F99E" box="[1311,1398,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,1016,1040]" captionTargetBox="[151,1436,181,992]" captionTargetId="figure-20@6.[151,1436,181,992]" captionTargetPageId="6" captionText="FIGURE 4. Parasesarma rahayuae n. sp., paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="3" pageNumber="570">Fig. 4C</figureCitation>
) in depression on anterior edge of sternite 6, operculum touching line of sternite 5, oval operculum in anterior part with a sternal cap (cover) posterior to operculum.
</paragraph>
</subSubSection>
<subSubSection id="40F2656AC470831C1BE80B82FA31F8C2" pageId="3" pageNumber="570" type="etymology">
<paragraph id="085736E1C470831C1BE80B82FA31F8C2" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
<emphasis id="3A9CEAF3C470831C1BE80B82FEF6F90A" bold="true" box="[199,328,1735,1761]" pageId="3" pageNumber="570">Etymology</emphasis>
. The new species is warmly dedicated to and named after Professor Dwi Listyo Rahayu (National Research and Innovation Agency, formerly the Indonesian Institute of Sciences), for her work on the genera 
<taxonomicName id="CFE84D62C470831C1BB80A4AFE92F8C3" authorityName="De Man" authorityYear="1895" box="[151,300,1807,1832]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="3A9CEAF3C470831C1BB80A4AFE92F8C3" box="[151,300,1807,1832]" italics="true" pageId="3" pageNumber="570">Parasesarma</emphasis>
</taxonomicName>
and 
<taxonomicName id="CFE84D62C470831C1A4C0A4AFE4FF8C3" authorityName="De Man" authorityYear="1895" box="[355,497,1807,1832]" class="Malacostraca" family="Sesarmidae" genus="Perisesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="3A9CEAF3C470831C1A4C0A4AFE4FF8C3" box="[355,497,1807,1832]" italics="true" pageId="3" pageNumber="570">Perisesarma</emphasis>
</taxonomicName>
, making significant contributions to our taxonomic understanding of these genera.
</paragraph>
</subSubSection>
<subSubSection id="40F2656AC47083181BE80A76FC97FDF2" lastPageId="7" lastPageNumber="574" pageId="3" pageNumber="570" type="discussion">
<paragraph id="085736E1C470831C1BE80A76FA2DF836" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="570">
<emphasis id="3A9CEAF3C470831C1BE80A76FE8BF8A6" bold="true" box="[199,309,1843,1869]" pageId="3" pageNumber="570">Remarks</emphasis>
. Corresponding to the molecular phylogenetic relationships, 
<taxonomicName id="CFE84D62C470831C18DE0A76FB45F8A6" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[1009,1275,1843,1869]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C470831C18DE0A76FB45F8A6" box="[1009,1275,1843,1869]" italics="true" pageId="3" pageNumber="570">Parasesarma rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C470831C1E2C0A76FAFAF8A6" bold="true" box="[1283,1348,1843,1869]" pageId="3" pageNumber="570">
<taxonomicNameLabel id="21AF5788C470831C1E2C0A76FAFAF8A6" box="[1283,1348,1843,1869]" pageId="3" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, is also morphologically similar to 
<taxonomicName id="CFE84D62C470831C1AE90A12FDE2F89B" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[454,604,1879,1904]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C470831C1AE90A12FDE2F89B" box="[454,604,1879,1904]" italics="true" pageId="3" pageNumber="570">P. gemmatum</emphasis>
</taxonomicName>
and 
<taxonomicName id="CFE84D62C470831C19A00A12FCB0F89A" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[655,782,1879,1905]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C470831C19A00A12FCB0F89A" box="[655,782,1879,1905]" italics="true" pageId="3" pageNumber="570">P. anambas</emphasis>
</taxonomicName>
. These three species are small crabs (all known individuals less than 
<quantity id="CF109B04C470831C1BD00A3FFEF3F87E" box="[255,333,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="570" unit="mm" value="15.0">15 mm</quantity>
in carapace width) with a subquadrate carapace (
<figureCitation id="90D32A64C470831C185B0A3EFC53F87E" box="[884,1005,1914,1941]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="3" pageNumber="570">Fig. 5A‒C</figureCitation>
) and a large number (more than 15) of transversely oval tubercles on the dactylus of the male chela (
<figureCitation id="90D32A64C470831C18670ADAFC02F852" box="[840,956,1950,1977]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="3" pageNumber="570">Fig. 5E‒F</figureCitation>
; 
<bibRefCitation id="6C794B10C470831C18E90ADAFBCAF852" author="Yeo, D. C. J. &amp; Rahayu, D. L. &amp; Ng, P. K. L." box="[966,1140,1951,1977]" pageId="3" pageNumber="570" pagination="79 - 88" refId="ref5226" refString="Yeo, D. C. J., Rahayu, D. L. &amp; Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas expedition 2002. Raffles Bulletin of Zoology, Supplement No. 11, 79 - 88." type="book chapter" year="2004">
Yeo 
<emphasis id="3A9CEAF3C470831C18D60ADAFB97F852" box="[1017,1065,1951,1977]" italics="true" pageId="3" pageNumber="570">et al</emphasis>
. 2004
</bibRefCitation>
: fig. 2; 
<bibRefCitation id="6C794B10C470831C1FE80ADAFADFF852" author="Li, J. - J. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[1223,1377,1951,1977]" pageId="3" pageNumber="570" pagination="40" refId="ref4223" refString="Li, J. - J., Shih, H. - T. &amp; Ng, P. K. L. (2019) Three new species and two new records of Parasesarma De Man, 1895 (Crustacea: Brachyura: Sesarmidae) from Taiwan and the Philippines from morphological and molecular evidence. Zoological Studies, 58, e 40." type="journal article" year="2019">
Li 
<emphasis id="3A9CEAF3C470831C1FC90ADAFAA9F852" box="[1254,1303,1951,1977]" italics="true" pageId="3" pageNumber="570">et al</emphasis>
. 2019
</bibRefCitation>
: figs. 9, 10). The new species, however, differs from these two species in a number of morphological characteristics.
</paragraph>
<caption id="5C976669C477831B1BB80A72FD08F850" ID-DOI="http://doi.org/10.5281/zenodo.8282243" ID-Zenodo-Dep="8282243" httpUri="https://zenodo.org/record/8282243/files/figure.png" pageId="4" pageNumber="571" startId="4.[151,250,1847,1871]" targetBox="[194,1394,194,1808]" targetPageId="4" targetType="figure">
<paragraph id="085736E1C477831B1BB80A72FD08F850" blockId="4.[151,1437,1847,1979]" pageId="4" pageNumber="571">
<emphasis id="3A9CEAF3C477831B1BB80A72FEADF8A4" bold="true" box="[151,275,1847,1871]" pageId="4" pageNumber="571">FIGURE 2.</emphasis>
<taxonomicName id="CFE84D62C477831B1A360A7DFDB8F8A4" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[281,518,1847,1871]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C477831B1A360A7DFDB8F8A4" box="[281,518,1847,1871]" italics="true" pageId="4" pageNumber="571">Parasesarma rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C477831B19240A72FDFAF8A4" bold="true" box="[523,580,1847,1871]" pageId="4" pageNumber="571">
<taxonomicNameLabel id="21AF5788C477831B19240A72FDFAF8A4" box="[523,580,1847,1871]" pageId="4" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: A, dorsal habitus; B, ventral habitus; C, carapace, dorsal view; D, frontal view; E, left chela, dorsal view; F, left chela, outer view; G, cheliped dactylar tubercles, dorsal view; H, cheliped dactylar tubercles, outer view; I, ambulatory leg, right P4, dorsal view; J, pleonal somites 1 and 2, posteroventral view.
</paragraph>
</caption>
<caption id="5C976669C476831A1BB80BDDFE03F8F7" ID-DOI="http://doi.org/10.5281/zenodo.8282247" ID-Zenodo-Dep="8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="5" pageNumber="572" startId="5.[151,250,1688,1712]" targetBox="[201,1386,181,1665]" targetPageId="5" targetType="figure">
<paragraph id="085736E1C476831A1BB80BDDFE03F8F7" blockId="5.[151,1436,1688,1820]" pageId="5" pageNumber="572">
<emphasis id="3A9CEAF3C476831A1BB80BDDFEADF95B" bold="true" box="[151,275,1688,1712]" pageId="5" pageNumber="572">FIGURE 3.</emphasis>
<taxonomicName id="CFE84D62C476831A1A360BDCFDB9F95B" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[281,519,1688,1712]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C476831A1A360BDCFDB9F95B" box="[281,519,1688,1712]" italics="true" pageId="5" pageNumber="572">Parasesarma rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C476831A19220BDCFDF8F95A" bold="true" box="[525,582,1689,1713]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="21AF5788C476831A19220BDCFDF8F95A" box="[525,582,1689,1713]" pageId="5" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part.
</paragraph>
</caption>
<caption id="5C976669C47583191BB80EBDFC73FBDF" ID-DOI="http://doi.org/10.5281/zenodo.8282249" ID-Zenodo-Dep="8282249" httpUri="https://zenodo.org/record/8282249/files/figure.png" pageId="6" pageNumber="573" startId="6.[151,250,1016,1040]" targetBox="[151,1436,181,992]" targetPageId="6" targetType="figure">
<paragraph id="085736E1C47583191BB80EBDFC73FBDF" blockId="6.[151,1436,1016,1077]" pageId="6" pageNumber="573">
<emphasis id="3A9CEAF3C47583191BB80EBDFEAAFBFB" bold="true" box="[151,276,1016,1040]" pageId="6" pageNumber="573">FIGURE 4.</emphasis>
<taxonomicName id="CFE84D62C47583191A340EBCFDB4FBFB" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[283,522,1016,1040]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C47583191A340EBCFDB4FBFB" box="[283,522,1016,1040]" italics="true" pageId="6" pageNumber="573">Parasesarma rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C4758319193F0EBCFDF5FBFA" bold="true" box="[528,587,1017,1041]" pageId="6" pageNumber="573">
<taxonomicNameLabel id="21AF5788C4758319193F0EBCFDF5FBFA" box="[528,587,1017,1041]" pageId="6" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, ovig. female (11.9 mm × 9.77 mm) ZRC 2019.1055, Manado. A, dorsal habitus; B, pleon, ventral view, C, vulvae, ventral view; D, left chela, outer view.
</paragraph>
</caption>
<caption id="5C976669C47583191BB80A41FD56F862" ID-DOI="http://doi.org/10.5281/zenodo.8282253" ID-Zenodo-Dep="8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="6" pageNumber="573" startId="6.[151,250,1796,1820]" targetBox="[151,1436,1137,1773]" targetPageId="6" targetType="figure">
<paragraph id="085736E1C47583191BB80A41FD56F862" blockId="6.[151,1436,1796,1929]" pageId="6" pageNumber="573">
<emphasis id="3A9CEAF3C47583191BB80A41FEAAF8F6" bold="true" box="[151,276,1796,1821]" pageId="6" pageNumber="573">FIGURE 5.</emphasis>
A, D, 
<taxonomicName id="CFE84D62C47583191A710A40FDF3F8F6" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[350,589,1797,1821]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C47583191A710A40FDF3F8F6" box="[350,589,1797,1821]" italics="true" pageId="6" pageNumber="573">Parasesarma rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C4758319197A0A40FD31F8F6" bold="true" box="[597,655,1797,1821]" pageId="6" pageNumber="573">
<taxonomicNameLabel id="21AF5788C4758319197A0A40FD31F8F6" box="[597,655,1797,1821]" pageId="6" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, 
<taxonomicName id="CFE84D62C47583191A1D0A6CFD0FF8AA" authority="Yeo, Rahayu &amp; Ng, 2004" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[306,689,1833,1857]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C47583191A1D0A6CFE19F8AA" box="[306,423,1833,1857]" italics="true" pageId="6" pageNumber="573">P. anambas</emphasis>
<bibRefCitation id="6C794B10C47583191A820A6CFD0FF8AA" author="Yeo, D. C. J. &amp; Rahayu, D. L. &amp; Ng, P. K. L." box="[429,689,1833,1857]" pageId="6" pageNumber="573" pagination="79 - 88" refId="ref5226" refString="Yeo, D. C. J., Rahayu, D. L. &amp; Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas expedition 2002. Raffles Bulletin of Zoology, Supplement No. 11, 79 - 88." type="book chapter" year="2004">Yeo, Rahayu &amp; Ng, 2004</bibRefCitation>
</taxonomicName>
, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, 
<taxonomicName id="CFE84D62C47583191BB80A08FE46F88E" authority="Li, Shih &amp; Ng, 2019" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[151,504,1869,1893]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C47583191BB80A08FEA1F88F" box="[151,287,1869,1892]" italics="true" pageId="6" pageNumber="573">P. gemmatum</emphasis>
<bibRefCitation id="6C794B10C47583191A0A0A08FE46F88E" author="Li, J. - J. &amp; Shih, H. - T. &amp; Ng, P. K. L." box="[293,504,1869,1893]" pageId="6" pageNumber="573" pagination="40" refId="ref4223" refString="Li, J. - J., Shih, H. - T. &amp; Ng, P. K. L. (2019) Three new species and two new records of Parasesarma De Man, 1895 (Crustacea: Brachyura: Sesarmidae) from Taiwan and the Philippines from morphological and molecular evidence. Zoological Studies, 58, e 40." type="journal article" year="2019">Li, Shih &amp; Ng, 2019</bibRefCitation>
</taxonomicName>
, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles.
</paragraph>
</caption>
<paragraph id="085736E1C47483181BB80DD2FC97FDF2" blockId="7.[151,1437,150,537]" pageId="7" pageNumber="574">
<taxonomicName id="CFE84D62C47483181BB80DD2FE0CFF5B" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[151,434,151,176]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C47483181BB80DD2FE0CFF5B" box="[151,434,151,176]" italics="true" pageId="7" pageNumber="574">Parasesarma gemmatum</emphasis>
</taxonomicName>
lacks the pectinated crests on the dorsal surface of the male palm (
<figureCitation id="90D32A64C47483181FE40DD2FA9DFF5A" box="[1227,1315,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="7" pageNumber="574">Fig. 5F</figureCitation>
), while 
<taxonomicName id="CFE84D62C47483181EAB0DD2FEBFFF3E" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C47483181EAB0DD2FA2BFF5B" box="[1412,1429,151,176]" italics="true" pageId="7" pageNumber="574">P</emphasis>
. 
<emphasis id="3A9CEAF3C47483181BB80DFEFEBFFF3E" box="[151,257,187,213]" italics="true" pageId="7" pageNumber="574">rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C47483181A260DFEFEF7FF3E" bold="true" box="[265,329,187,213]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="21AF5788C47483181A260DFEFEF7FF3E" box="[265,329,187,213]" pageId="7" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has two pectinated crests on the dorsal face of the palm (
<figureCitation id="90D32A64C474831818F30DFEFB89FF3E" box="[988,1079,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="7" pageNumber="574">Fig. 5D</figureCitation>
), like most other 
<taxonomicName id="CFE84D62C47483181E2F0DFEFA2BFF3F" authorityName="De Man" authorityYear="1895" box="[1280,1429,187,212]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="3A9CEAF3C47483181E2F0DFEFA2BFF3F" box="[1280,1429,187,212]" italics="true" pageId="7" pageNumber="574">Parasesarma</emphasis>
</taxonomicName>
. The G1 of 
<taxonomicName id="CFE84D62C47483181A3C0D9AFE14FF13" authorityName="Li, Shih &amp; Ng" authorityYear="2019" box="[275,426,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="gemmatum">
<emphasis id="3A9CEAF3C47483181A3C0D9AFE14FF13" box="[275,426,223,248]" italics="true" pageId="7" pageNumber="574">P. gemmatum</emphasis>
</taxonomicName>
has a proportionally shorter and wider apical corneous process in a more upright position (approximately 45 degrees from the vertical axis; 
<figureCitation id="90D32A64C474831819FC0C46FC97FEF6" box="[723,809,258,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="7" pageNumber="574">Fig. 5C</figureCitation>
) in comparison to the longer, narrower, and more bent process in 
<taxonomicName id="CFE84D62C47483181A3D0C62FE27FEAA" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[274,409,295,321]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C47483181A3D0C62FE27FEAA" box="[274,409,295,321]" italics="true" pageId="7" pageNumber="574">P. rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C47483181A8E0C62FE5FFEAA" bold="true" box="[417,481,295,321]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="21AF5788C47483181A8E0C62FE5FFEAA" box="[417,481,295,321]" pageId="7" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(almost 90° from the vertical axis; 
<figureCitation id="90D32A64C474831818560C62FC6AFEAA" box="[889,980,294,321]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="7" pageNumber="574">Fig. 5A</figureCitation>
). Compared to 
<taxonomicName id="CFE84D62C47483181FAB0C62FAB8FEAA" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[1156,1286,295,321]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C47483181FAB0C62FAB8FEAA" box="[1156,1286,295,321]" italics="true" pageId="7" pageNumber="574">P. anambas</emphasis>
</taxonomicName>
, 
<taxonomicName id="CFE84D62C47483181E390C62FA22FEAA" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[1302,1436,295,321]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C47483181E390C62FA22FEAA" box="[1302,1436,295,321]" italics="true" pageId="7" pageNumber="574">P. rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C47483181BB80C0EFF69FE8E" bold="true" box="[151,215,331,357]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="21AF5788C47483181BB80C0EFF69FE8E" box="[151,215,331,357]" pageId="7" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has relatively broader tubercles on the dorsal face of the male cheliped dactylus (
<figureCitation id="90D32A64C47483181F5A0C0EFB4FFE8E" box="[1141,1265,330,357]" captionStart="FIGURE 5" captionStartId="6.[151,250,1796,1820]" captionTargetBox="[151,1436,1137,1773]" captionTargetId="figure-57@6.[151,1436,1137,1773]" captionTargetPageId="6" captionText="FIGURE 5. A, D, Parasesarma rahayuae n. sp., holotype, male (11.47 mm ×10.11 mm), MZB 5709 (ex. ZRC 2019.1055), Manado: B, E, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm× 10.1 mm) ZRC 2023.0256, Cebu. A‒C, carapace dorsal view; D‒F, dorsal face of left cheliped palm and proximal dactylar tubercles." figureDoi="http://doi.org/10.5281/zenodo.8282253" httpUri="https://zenodo.org/record/8282253/files/figure.png" pageId="7" pageNumber="574">Fig. 5D, C</figureCitation>
). It also differs in the morphology of the G1, with the apical corneous process proportionally shorter (
<figureCitation id="90D32A64C47483181F4C0C2AFB61FE62" box="[1123,1247,367,393]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">Fig. 6A, D</figureCitation>
), semicircular in cross-section (
<figureCitation id="90D32A64C47483181A170CD6FE2FFE46" box="[312,401,403,429]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">Fig. 6G</figureCitation>
), distinctly wider than the rest of the apical corneous process when seen from the dorsal view (
<figureCitation id="90D32A64C47483181BB00CF2FF45FE3A" box="[159,251,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1688,1712]" captionTargetBox="[201,1386,181,1665]" captionTargetId="figure-15@5.[188,1399,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm), ZRC 2019.1055, Manado: A, dorsal habitus; B, merus, inner surface; C, right cheliped dactylus, dorso-external view; D, right palm secondary pectinated crest, frontal view; E, right G1, full dorsal view; F, right G1 distal tip, dorsal view; G, right G1 front view of apical corneous part, H, right G1 upper view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.8282247" httpUri="https://zenodo.org/record/8282247/files/figure.png" pageId="7" pageNumber="574">Figs. 3F</figureCitation>
, 
<figureCitation id="90D32A64C47483181A290CF2FE94FE3A" box="[262,298,439,465]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">6A</figureCitation>
) and with a subterminal aperture in 
<taxonomicName id="CFE84D62C474831819970CF2FC82FE3A" authority="Shahdadi &amp; Schubart &amp; Mendoza, 2023" authorityName="Shahdadi &amp; Schubart &amp; Mendoza" authorityYear="2023" box="[696,828,439,465]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="rahayuae" status="sp. nov.">
<emphasis id="3A9CEAF3C474831819970CF2FC82FE3A" box="[696,828,439,465]" italics="true" pageId="7" pageNumber="574">P. rahayuae</emphasis>
</taxonomicName>
<emphasis id="3A9CEAF3C4748318186D0CF2FC3EFE3A" bold="true" box="[834,896,439,465]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="21AF5788C4748318186D0CF2FC3EFE3A" box="[834,896,439,465]" pageId="7" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, while it is longer (
<figureCitation id="90D32A64C47483181F7A0CF2FB12FE3A" box="[1109,1196,439,465]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">Fig. 6B</figureCitation>
), only slightly arched in cross-section (
<figureCitation id="90D32A64C47483181A780C9EFE0CFE1E" box="[343,434,475,501]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">Fig. 6H</figureCitation>
), not much wider than the rest of the apical corneous process when seen from the dorsal view (
<figureCitation id="90D32A64C47483181BF20CBAFE8DFDF2" box="[221,307,511,537]" captionStart="FIGURE 6" captionStartId="7.[151,250,1815,1839]" captionTargetBox="[151,1436,561,1792]" captionTargetId="figure-226@7.[151,1436,561,1792]" captionTargetPageId="7" captionText="FIGURE 6. A, D, G, Parasesarma rahayuae n. sp., paratype, male (11.32 mm × 9.6 mm) ZRC 2019.1055, Manado; B, E, H, P. anambas Yeo, Rahayu &amp; Ng, 2004, paratype male (7.9 mm × 6.3 mm) ZRC 2003.0726, Pulau Siantan; C, F, I, P. gemmatum Li, Shih &amp; Ng, 2019, male (11.9 mm × 10.1 mm) ZRC 2023.0256, Cebu. A‒C, dorsal view; B‒F, apical corneous part, upper view; G‒I, apical corneous part, front view." figureDoi="http://doi.org/10.5281/zenodo.8282255" httpUri="https://zenodo.org/record/8282255/files/figure.png" pageId="7" pageNumber="574">Fig. 6B</figureCitation>
) and with a terminal aperture in 
<taxonomicName id="CFE84D62C4748318198D0CBAFC9CFDF2" authorityName="Yeo, Rahayu &amp; Ng" authorityYear="2004" box="[674,802,511,537]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="anambas">
<emphasis id="3A9CEAF3C4748318198D0CBAFC9CFDF2" box="[674,802,511,537]" italics="true" pageId="7" pageNumber="574">P. anambas</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>