treatments-xml/data/80/0A/7F/800A7F3A42BB5B2D966ED916A855AADC.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1075.67459" ID-GBIF-Dataset="e83d47ed-a6b8-4706-9df8-e1686d1be39f" ID-PMC="PMC8671714" ID-Pensoft-Pub="1313-2970-1075-33" ID-Pensoft-UUID="DBA502BCE85F5F3C922FDD3123C6365B" ID-PubMed="35046751" ID-ZooBank="BB034E5C10CB49179FCDEF6A72816F6C" ModsDocID="1313-2970-1075-33" checkinTime="1638937131209" checkinUser="pensoft" docAuthor="Moreno-Gonzalez 1, Jairo A., Pinto-da-Rocha 1, Ricardo &amp; Gallao 2, Jonas E." docDate="2021" docId="800A7F3A42BB5B2D966ED916A855AADC" docLanguage="en" docName="ZooKeys 1075: 33-75" docOrigin="ZooKeys 1075" docPubDate="2021-12-07" docSource="http://dx.doi.org/10.3897/zookeys.1075.67459" docTitle="Tityus (Tityus) C. L. Koch 1836" docType="treatment" docVersion="4" id="DBA502BCE85F5F3C922FDD3123C6365B" lastPageNumber="33" masterDocId="DBA502BCE85F5F3C922FDD3123C6365B" masterDocTitle="Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species" masterLastPageNumber="75" masterPageNumber="33" pageNumber="33" updateTime="1668151205184" updateUser="ExternalLinkService">
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<mods:title>Bringing order to a complex system: phenotypic and genotypic evidence contribute to the taxonomy of Tityus (Scorpiones, Buthidae) and support the description of a new species</mods:title>
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<mods:namePart>Moreno-Gonzalez 1, Jairo A.</mods:namePart>
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
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<mods:namePart>Pinto-da-Rocha 1, Ricardo</mods:namePart>
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gallao 2, Jonas E.</mods:namePart>
<mods:affiliation>Instituto de Biociencias - Universidade de Sao Paulo, Departamento de Zoologia. Rua do Matao, travessa 14, 321, 005508 - 900, Sao Paulo, Brazil</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-12-07</mods:number>
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<mods:number>1075</mods:number>
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<mods:start>33</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1075.67459</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1075.67459</mods:identifier>
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<treatment ID-GBIF-Taxon="191254654" LSID="urn:lsid:plazi:treatment:800A7F3A42BB5B2D966ED916A855AADC" httpUri="http://treatment.plazi.org/id/800A7F3A42BB5B2D966ED916A855AADC" lastPageNumber="33" pageId="0" pageNumber="33">
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Subgenus 
<taxonomicName LSID="800A7F3A-42BB-5B2D-966E-D916A855AADC" authority="(Tityus) C. L. Koch, 1836" authorityName="C. L. Koch" authorityYear="1836" baseAuthorityName="Tityus" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus)" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">Tityus (Tityus) C. L. Koch, 1836</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="33">
<taxonomicName authority="(Tityus)" authorityName="C. L. Koch" authorityYear="1836" baseAuthorityName="Tityus" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus)" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">Tityus (Tityus)</taxonomicName>
: 
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="0" pageNumber="33" pagination="55 - 67" refId="B34" refString="Lourenco, WR, 2006. Une nouvelle proposition de decoupage sous-generique du genre &quot; Tityus &quot; C.L. Koch, 1836 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 39: 55 - 67" title="Une nouvelle proposition de decoupage sous-generique du genre &quot; Tityus &quot; C. L. Koch, 1836 (Scorpiones, Buthidae)." volume="39" year="2006">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
(2006)
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: 57, 58, 60, figures 3-6, 10-13, 22.
</paragraph>
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<paragraph pageId="0" pageNumber="33">Type species.</paragraph>
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Perty" authorityYear="1833" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" lsidName="Scorpio bahiensis" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="bahiensis">
<emphasis italics="true" pageId="0" pageNumber="33">Scorpio bahiensis</emphasis>
</taxonomicName>
Perty, 1833 by monotypy.
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<subSubSection pageId="0" pageNumber="33" type="comments">
<paragraph pageId="0" pageNumber="33">Comments.</paragraph>
<paragraph pageId="0" pageNumber="33">
This subgenus currently includes, among others, all species assigned to the 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. bahiensis" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="bahiensis">
<emphasis italics="true" pageId="0" pageNumber="33">T. bahiensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1945; 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. bolivianus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis italics="true" pageId="0" pageNumber="33">T. bolivianus</emphasis>
</taxonomicName>
Kraepelin, 1895; 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. stigmurus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="stigmurus">
<emphasis italics="true" pageId="0" pageNumber="33">T. stigmurus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1945, and 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. trivittatus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="trivittatus">
<emphasis italics="true" pageId="0" pageNumber="33">T. trivittatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
, 1945 species-groups, according to the classification proposal of 
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="0" pageNumber="33" pagination="55 - 67" refId="B34" refString="Lourenco, WR, 2006. Une nouvelle proposition de decoupage sous-generique du genre &quot; Tityus &quot; C.L. Koch, 1836 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 39: 55 - 67" title="Une nouvelle proposition de decoupage sous-generique du genre &quot; Tityus &quot; C. L. Koch, 1836 (Scorpiones, Buthidae)." volume="39" year="2006">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
(2006)
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. In addition to 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. brazilae" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="brazilae">
<emphasis italics="true" pageId="0" pageNumber="33">T. brazilae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
&amp; Eickstedt, 1984, here transferred to this subgenus (see Discussion). On the other hand, 
<taxonomicName authorityName="Lourenco &amp; Florez" authorityYear="1990" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus sastrei" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="sastrei">
<emphasis italics="true" pageId="0" pageNumber="33">Tityus sastrei</emphasis>
</taxonomicName>
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
&amp; 
<normalizedToken originalValue="Flórez">Florez</normalizedToken>
, 1990 belongs to 
<taxonomicName authority="" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Atreus)" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">Tityus (Atreus)</taxonomicName>
and is excluded from 
<taxonomicName authority="(Tityus)" authorityName="C. L. Koch" authorityYear="1836" baseAuthorityName="Tityus" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus)" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">Tityus (Tityus)</taxonomicName>
(see Discussion). Finally, according to previous hypotheses and our data, the 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. bolivianus" order="Orthoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis italics="true" pageId="0" pageNumber="33">T. bolivianus</emphasis>
</taxonomicName>
Kraepelin, 1895 species-group forms an independent clade outside 
<taxonomicName authority="(Tityus)" authorityName="C. L. Koch" authorityYear="1836" baseAuthorityName="Tityus" class="Insecta" family="Buthidae" genus="Tityus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tityus (Tityus)" order="Scorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">Tityus (Tityus)</taxonomicName>
, but additional studies, including the study of the type species of this group, are required to propose a formal taxonomic decision.
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