1350 lines
179 KiB
XML
1350 lines
179 KiB
XML
<document id="83B539B8C3E05D86BF277380FF59335A" ID-CLB-Dataset="252076" ID-DOI="10.1643/h2021051" ID-GBIF-Dataset="a3e05a9c-2902-4857-a3ea-62a2253e135a" ID-ISSN="2766-1520" ID-Zenodo-Dep="7846895" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1681928239709" checkinUser="felipe" docAuthor="García-Vázquez, Uri Omar, Clause, Adam G., Gutiérrez-Rodríguez, Jorge, Cazares-Hernández, Erasmo & Torre-Loranca, Miguel Ángel de la" docDate="2022" docId="880587D4FFBFFFC4FFF7EEBEFDB1FA25" docLanguage="en" docName="IchthyolHerpetol.110.1.33-49.pdf" docOrigin="Ichthyology & Herpetology 110 (1)" docSource="http://dx.doi.org/10.1643/h2021051" docStyle="DocumentStyle:25DEBD253B07840CEE0E8BE8BE2C6752.2:IchthyolHerpetol.2021-.journal_article" docStyleId="25DEBD253B07840CEE0E8BE8BE2C6752" docStyleName="IchthyolHerpetol.2021-.journal_article" docStyleVersion="2" docTitle="Abronia zongolica García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca 2022, new species" docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="743CFFACFFBCFFCFFF90E90AFF95FFD9" masterDocTitle="A New Species of Abronia (Squamata: Anguidae) from the Sierra de Zongolica of Veracruz, Mexico" masterLastPageNumber="49" masterPageNumber="33" pageNumber="36" updateTime="1702000783877" updateUser="plazi">
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<mods:title id="1783E350FA5696CA1C4C2998FC3E6397">A New Species of Abronia (Squamata: Anguidae) from the Sierra de Zongolica of Veracruz, Mexico</mods:title>
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<mods:namePart id="FD9C507607B6368CA8ECFE502F6D955C">García-Vázquez, Uri Omar</mods:namePart>
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<mods:namePart id="110507C5D6AB2422EE3C1C092D54518F">Clause, Adam G.</mods:namePart>
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<mods:namePart id="435EF1974531B8E528760DC4EB71CCFA">Gutiérrez-Rodríguez, Jorge</mods:namePart>
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<mods:namePart id="ED63DA630CC268D000D2230570A27448">Cazares-Hernández, Erasmo</mods:namePart>
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<mods:namePart id="43230B6CB21D5781CB5DA1A5B7938F2D">Torre-Loranca, Miguel Ángel de la</mods:namePart>
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<mods:title id="6D3E24732213963FB9D648A20E221C86">Ichthyology & Herpetology</mods:title>
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<mods:date id="DDFEF095CE6C81B6D54E609D6CD90DB0">2022</mods:date>
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<treatment id="880587D4FFBFFFC4FFF7EEBEFDB1FA25" ID-DOI="http://doi.org/10.5281/zenodo.7852458" ID-GBIF-Taxon="209563872" ID-Zenodo-Dep="7852458" LSID="urn:lsid:plazi:treatment:880587D4FFBFFFC4FFF7EEBEFDB1FA25" httpUri="http://treatment.plazi.org/id/880587D4FFBFFFC4FFF7EEBEFDB1FA25" lastPageId="11" lastPageNumber="44" pageId="3" pageNumber="36">
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<paragraph id="001336C2FFBFFFCCFFF7EEBEFE43F812" blockId="3.[103,470,1972,1995]" box="[103,470,1972,1995]" pageId="3" pageNumber="36">
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<heading id="5B5B81AEFFBFFFCCFFF7EEBEFE43F812" bold="true" box="[103,470,1972,1995]" fontSize="10" level="2" pageId="3" pageNumber="36" reason="2">
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<emphasis id="32D8EAD0FFBFFFCCFFF7EEBEFE43F812" bold="true" box="[103,470,1972,1995]" pageId="3" pageNumber="36">
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<taxonomicName id="C7AC4D41FFBFFFCCFFF7EEBEFEA3F812" authority="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca, 2022" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[103,310,1972,1995]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="zongolica" status="sp. nov.">
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<emphasis id="32D8EAD0FFBFFFCCFFF7EEBEFEA3F812" bold="true" box="[103,310,1972,1995]" italics="true" pageId="3" pageNumber="36">Abronia zongolica</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="29EB57ABFFBFFFCCFED6EEBEFE43F812" box="[326,470,1972,1995]" pageId="3" pageNumber="36" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="001336C2FFBFFFCCFFF7EEDBFD33F831" blockId="3.[103,678,2001,2024]" box="[103,678,2001,2024]" pageId="3" pageNumber="36">urn:lsid:zoobank.org:act:0A5746B6-D2CE-4F46-AD2E-</paragraph>
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</subSubSection>
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<subSubSection id="48B66549FFBFFFCCFCA4EA8CFBC7FC2E" pageId="3" pageNumber="36" type="description">
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<paragraph id="001336C2FFBFFFCCFCA4EA8CFC71FC44" blockId="3.[820,1290,902,1015]" box="[820,996,902,925]" pageId="3" pageNumber="36">19C9CD2E6D30</paragraph>
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<paragraph id="001336C2FFBFFFCCFCA4EAAEFA9FFC62" blockId="3.[820,1290,902,1015]" box="[820,1290,932,955]" pageId="3" pageNumber="36">
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Sierra de
|
||
<taxonomicName id="C7AC4D41FFBFFFCCFC09EAAEFA9FFC62" authority="Arboreal Alligator Lizard" box="[921,1290,932,955]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="zongolica">Zongolica Arboreal Alligator Lizard</taxonomicName>
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</paragraph>
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<paragraph id="001336C2FFBFFFCCFCA4EAC8FB28FC00" blockId="3.[820,1290,902,1015]" box="[820,1213,962,985]" pageId="3" pageNumber="36">Dragoncito de la Sierra de Zongolica</paragraph>
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<paragraph id="001336C2FFBFFFCCFCA4EAEAFBC7FC2E" blockId="3.[820,1290,902,1015]" box="[820,1106,992,1015]" pageId="3" pageNumber="36">
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<figureCitation id="98972A47FFBFFFCCFCA4EAEAFC0DFC2E" box="[820,920,992,1015]" captionStart="FIG" captionStartId="4.[142,170,1536,1556]" captionTargetBox="[142,812,178,1506]" captionTargetId="figure-683@4.[142,812,178,1507]" captionTargetPageId="4" captionText="FIG. 2. Abronia zongolica, new species, head of holotype (MZFC-HE 35664, head width 25.2 mm) in dorsal view (top), right lateral view (middle; image intentionally mirrored horizontally) and ventral view (bottom), in preservative. Scale bars represent 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7846899" httpUri="https://zenodo.org/record/7846899/files/figure.png" pageId="3" pageNumber="36">Figures 2</figureCitation>
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,
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<figureCitation id="98972A47FFBFFFCCFC34EAEAFC20FC2E" box="[932,949,992,1015]" captionStart="FIG" captionStartId="5.[103,131,1898,1918]" captionTargetBox="[269,1325,178,1868]" captionTargetId="figure-1@5.[268,1325,178,1869]" captionTargetPageId="5" captionText="FIG. 3. Abronia zongolica, new species, color variation in life (photos not to scale). (A) Adult male holotype (MZFC-HE 35664, snout–vent length [SVL] 112 mm); (B) adult male paratype (MZFC-HE 35665, SVL 95.8 mm); (C) adult female paratype (MZFZ 4406, SVL 86.6 mm; image intentionally mirrored horizontally); (D) neonate paratype (MZFZ 4407, SVL 45.6 mm). Photos by Miguel Ángel de la Torre-Loranca and Antonio Esaú Valdenegro- Brito." figureDoi="http://doi.org/10.5281/zenodo.7846901" httpUri="https://zenodo.org/record/7846901/files/figure.png" pageId="3" pageNumber="36">3</figureCitation>
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,
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<figureCitation id="98972A47FFBFFFCCFC50EAEAFC44FC2E" box="[960,977,992,1015]" captionStart="FIG" captionStartId="7.[103,131,1804,1824]" captionTargetBox="[204,1388,178,1775]" captionTargetId="figure-1@7.[204,1389,178,1775]" captionTargetPageId="7" captionText="FIG. 4. Head comparison of adult Abronia zongolica and Abronia graminea, illustrating differences in (1) circumorbital area color, (2) upper and lower jaw color pattern, and (3) presence/absence of well-developed oblique row of enlarged lateral neck scales. Locality and voucher information for each image is as follows, listed from top-to-bottom. Abronia zongolica: Teopantzacoalco (paratype, MZFC-HE 35665), Ayahuatulco (holotype, MZFC-HE 35664), Huapango (referred specimen, MZFZ-IMG 310), Atiopa (paratype, MZFZ 4406), Huapango (referred specimen, MZFZ-IMG 311). Abronia graminea: El Sumidero (MZFC-HE 32991), Puerto del Aire (MZFZ-IMG 322), Acultzinapa (MZFZ-IMG 336), Puerto del Aire (Clause et al., 2016: fig. 2), Puerto del Aire (MZFZ-IMG 338). All photos taken within 1 month of capture. From top, first and third images of A. zongolica and fifth image of A. graminea intentionally mirrored horizontally. Photos by Miguel Ángel de la Torre-Loranca, Adam G. Clause, and Antonio Esaú Valdenegro- Brito." figureDoi="http://doi.org/10.5281/zenodo.7846903" httpUri="https://zenodo.org/record/7846903/files/figure.png" pageId="3" pageNumber="36">4</figureCitation>
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;
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<tableCitation id="4D2E0379FFBFFFCCFC4DEAEAFBA0FC2E" box="[989,1077,992,1015]" captionStart="Table 1" captionStartId="6.[142,190,178,198]" captionTargetPageId="6" captionText="Table 1. Character states for selected features of Abronia zongolica and all other species in the oaXacae group." httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EE9B8FB29FF1F" pageId="3" pageNumber="36" tableUuid="54D3664AFFBAFFC9FF1EE9B8FB29FF1F">Tables 1</tableCitation>
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,
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<tableCitation id="4D2E0379FFBFFFCCFBD4EAEAFBC7FC2E" box="[1092,1106,992,1015]" captionStart="Table 2" captionStartId="6.[142,190,1262,1282]" captionTargetPageId="6" captionText="Table 2. Character states for selected features of Abronia zongolica and all species in Group III of Gutiérrez-Rodríguez et al. (2021)." httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EEDE4FAEEFADB" pageId="3" pageNumber="36" tableUuid="54D3664AFFBAFFC9FF1EEDE4FAEEFADB">2</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="48B66549FFBFFFCCFCA4ED04FA2FFBFD" box="[820,1466,1038,1061]" pageId="3" pageNumber="36" type="reference_group">
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<paragraph id="001336C2FFBFFFCCFCA4ED04FA2FFBFD" blockId="3.[820,1466,1038,1061]" box="[820,1466,1038,1061]" pageId="3" pageNumber="36">
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<treatmentCitationGroup id="20BC11ECFFBFFFCCFCA4ED04FA2FFBFD" box="[820,1466,1038,1061]" pageId="3" pageNumber="36">
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<taxonomicName id="C7AC4D41FFBFFFCCFCA4ED04FC7AFBFD" baseAuthorityName="Cope" baseAuthorityYear="1864" box="[820,1007,1038,1060]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="graminea">
|
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<emphasis id="32D8EAD0FFBFFFCCFCA4ED04FC7AFBFD" box="[820,1007,1038,1060]" italics="true" pageId="3" pageNumber="36">Abronia graminea</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="810D10D3FFBFFFCCFB9EED05FADAFBFD" author="Guzman Guzman, S." box="[1038,1359,1038,1061]" page="125" pageId="3" pageNumber="36" year="2011">
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<bibRefCitation id="643D4B33FFBFFFCCFB9EED05FADAFBFD" author="Guzman Guzman, S." box="[1038,1359,1038,1061]" pageId="3" pageNumber="36" refId="ref14298" refString="Guzman Guzman, S. 2011. Anfibios y Reptiles de Veracruz: Guia Ilustrada. Consejo Veracruzano de Investigacion Cientifica y Desarrollo Tecnologico, Mexico." type="book" year="2011">Guzmán Guzmán, 2011: 125</bibRefCitation>
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</treatmentCitation>
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(in part).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="001336C2FFBFFFCCFCA4ED4EFC7CFB08" blockId="3.[820,1490,1092,1234]" pageId="3" pageNumber="36">
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<materialsCitation id="B0C43C9FFFBFFFCCFCA4ED4EFC7CFB08" ID-GBIF-Occurrence="4088553312" collectingDate="2018-12-20" collectionCode="R" collectorName="de Zongolica" country="Mexico" elevation="1600" latitude="18.608" location="Municipality of Mixtla de Altamirano" longLatPrecision="76" longitude="-97.028" municipality="Sierra de Zongolica" pageId="3" pageNumber="36" specimenCode="MZFC-HE 35664, LOR 251" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
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<emphasis id="32D8EAD0FFBFFFCCFCA4ED4EFC24FB82" bold="true" box="[820,945,1092,1115]" italics="true" pageId="3" pageNumber="36">
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<typeStatus id="DF178860FFBFFFCCFCA4ED4EFC00FB82" box="[820,917,1092,1115]" pageId="3" pageNumber="36" type="holotype">Holotype</typeStatus>
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.—
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</emphasis>
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<specimenCode id="500A9EB9FFBFFFCCFC21ED4FFBFFFB83" box="[945,1130,1092,1114]" collectionCode="MZFC-HE" pageId="3" pageNumber="36">MZFC-HE 35664</specimenCode>
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(field number
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<specimenCode id="500A9EB9FFBFFFCCFA8CED4FFA17FB82" box="[1308,1410,1092,1115]" collectionCode="LOR" pageId="3" pageNumber="36">LOR 251</specimenCode>
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), adult male, Ayahuatulco,
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,
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<collectingMunicipality id="E077ACB8FFBFFFCCFCA4ED8AFB83FB4F" box="[820,1046,1152,1174]" pageId="3" pageNumber="36">
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Sierra
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<collectorName id="AD595314FFBFFFCCFCEFED8AFB83FB4F" box="[895,1046,1152,1174]" pageId="3" pageNumber="36">de Zongolica</collectorName>
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,
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,
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<collectingCountry id="78BB7652FFBFFFCCFB0CED8AFB65FB4F" box="[1180,1264,1152,1174]" name="Mexico" pageId="3" pageNumber="36">Mexico</collectingCountry>
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,
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<geoCoordinate id="65985005FFBFFFCCFA92ED8AFAC9FB4F" box="[1282,1372,1152,1174]" degrees="18.608" direction="north" orientation="latitude" pageId="3" pageNumber="36" precision="55" value="18.608">18.608N</geoCoordinate>
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,
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<geoCoordinate id="65985005FFBFFFCCFAFFED8AFA5BFB4F" box="[1391,1486,1152,1174]" degrees="97.028" direction="west" orientation="longitude" pageId="3" pageNumber="36" precision="55" value="-97.028">97.028W</geoCoordinate>
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, datum WGS84,
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</quantity>
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||
elevation, J.
|
||
<collectionCode id="66BDAE07FFBFFFCCFB5BED95FB48FB6D" box="[1227,1245,1183,1204]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="36">R</collectionCode>
|
||
. Hernández Ginez,
|
||
<date id="74121002FFBFFFCCFA27ED94FC70FB08" pageId="3" pageNumber="36" value="2018-12-20">
|
||
<collectingDate id="6456E9EAFFBFFFCCFA27ED94FC70FB08" pageId="3" pageNumber="36" value="2018-12-20">20 December 2018</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="001336C2FFBFFFCCFCA4EDF8FA47F989" blockId="3.[820,1491,1266,1617]" pageId="3" pageNumber="36">
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFCA4EDF8FADDFAFF" ID-GBIF-Occurrence="4088553349" collectionCode="MZFC-HE" country="Mexico" location="Sierra de Zongolica" pageId="3" pageNumber="36" specimenCode="MZFC-HE 35663" specimenCount="6" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<emphasis id="32D8EAD0FFBFFFCCFCA4EDF8FC2BFAD0" bold="true" box="[820,958,1266,1289]" italics="true" pageId="3" pageNumber="36">
|
||
<typeStatus id="DF178860FFBFFFCCFCA4EDF8FC37FAD0" box="[820,930,1266,1289]" pageId="3" pageNumber="36" type="paratype">Paratypes</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
<specimenCount id="16AAFD4BFFBFFFCCFC2EEDF8FBC0FAD1" box="[958,1109,1266,1288]" pageId="3" pageNumber="36" type="generic">6 specimens</specimenCount>
|
||
, all collected in the
|
||
<location id="05736019FFBFFFCCFAF8EDF8FC36FAFF" LSID="urn:lsid:plazi:treatment:880587D4FFBFFFC4FFF7EEBEFDB1FA25:05736019FFBFFFCCFAF8EDF8FC36FAFF" country="Mexico" name="Sierra de Zongolica" pageId="3" pageNumber="36" stateProvince="Veracruz de Ignacio de la Llave">Sierra de Zongolica</location>
|
||
,
|
||
<collectingRegion id="C268F820FFBFFFCCFC24EC1BFB80FAFF" box="[948,1045,1297,1318]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="36">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry id="78BB7652FFBFFFCCFBB6EC1AFBEFFAFF" box="[1062,1146,1296,1318]" name="Mexico" pageId="3" pageNumber="36">Mexico</collectingCountry>
|
||
.
|
||
<specimenCode id="500A9EB9FFBFFFCCFB1AEC1AFADDFAFF" box="[1162,1352,1296,1318]" collectionCode="MZFC-HE" pageId="3" pageNumber="36">MZFC-HE 35663</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFAC9EC1AFCEEFABB" ID-GBIF-Occurrence="4088553315" collectingDate="2018-12-20" collectionCode="R" collectorName="de Zongolica" country="Mexico" elevation="1600" latitude="18.608" location="Municipality of Mixtla de Altamirano" longLatPrecision="76" longitude="-97.028" municipality="Sierra de Zongolica" pageId="3" pageNumber="36" specimenCode="MZFC-HE 35665" specimenCount="1" specimenCount-subadult="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<specimenCount id="16AAFD4BFFBFFFCCFAC9EC1AFA47FAFF" box="[1369,1490,1296,1318]" pageId="3" pageNumber="36" type="subadult">1 subadult</specimenCount>
|
||
female, same collection data as the holotype;
|
||
<specimenCode id="500A9EB9FFBFFFCCFAF7EC24FCEEFABB" collectionCode="MZFC-HE" pageId="3" pageNumber="36">MZFC-HE 35665</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFC19EC46FB6BFAB8" ID-GBIF-Occurrence="4088553339" box="[905,1278,1356,1378]" collectionCode="MZFC-HE" country="Mexico" pageId="3" pageNumber="36" specimenCode="MZFC-HE 35662, 1" specimenCount="1" specimenCount-adult="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<specimenCount id="16AAFD4BFFBFFFCCFC19EC46FC4DFABB" box="[905,984,1356,1378]" pageId="3" pageNumber="36" type="adult">1 adult</specimenCount>
|
||
male,
|
||
<specimenCode id="500A9EB9FFBFFFCCFBB7EC46FB76FABB" box="[1063,1251,1356,1378]" collectionCode="MZFC-HE" pageId="3" pageNumber="36">MZFC-HE 35662</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFBFFFCCFB60EC46FB6BFAB8" box="[1264,1278,1356,1377]" collectionCode="MZFC-HE" pageId="3" pageNumber="36">1</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFA97EC46FBEBFA01" ID-GBIF-Occurrence="4088553351" collectingDate="2018-11-23" collectionCode="MZFZ" collectorName="M. A. de la Torre-Loranca & J. C. Sanchez-Garcia & U. O. Garcia-Vazquez" country="Mexico" elevation="1785" latitude="18.608" location="Teopantzacoalco" longLatPrecision="76" longitude="-97.058" municipality="Municipality of San Juan Texhuacan" pageId="3" pageNumber="36" specimenCode="MZFZ 4408" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
neonate,
|
||
<location id="05736019FFBFFFCCFAE1EC46FC01FA59" LSID="urn:lsid:plazi:treatment:880587D4FFBFFFC4FFF7EEBEFDB1FA25:05736019FFBFFFCCFAE1EC46FC01FA59" country="Mexico" latitude="18.608" longLatPrecision="76" longitude="-97.058" municipality="Municipality of San Juan Texhuacan" name="Teopantzacoalco" pageId="3" pageNumber="36" stateProvince="Veracruz de Ignacio de la Llave">Teopantzacoalco</location>
|
||
,
|
||
<collectingMunicipality id="E077ACB8FFBFFFCCFC38EC60FACAFA59" box="[936,1375,1386,1408]" pageId="3" pageNumber="36">Municipality of San Juan Texhuacán</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="65985005FFBFFFCCFAE3EC60FA58FAA6" box="[1395,1485,1385,1408]" degrees="18.608" direction="north" orientation="latitude" pageId="3" pageNumber="36" precision="55" value="18.608">18.608N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="65985005FFBFFFCCFCA4EC8DFC01FA44" box="[820,916,1415,1437]" degrees="97.058" direction="west" orientation="longitude" pageId="3" pageNumber="36" precision="55" value="-97.058">97.058W</geoCoordinate>
|
||
, datum WGS84,
|
||
<quantity id="C7549B27FFBFFFCCFBDCEC8DFB0AFA44" box="[1100,1183,1415,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="3" pageNumber="36" unit="m" value="1785.0">
|
||
<elevation id="8B81D1F1FFBFFFCCFBDCEC8DFB0AFA44" box="[1100,1183,1415,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="3" pageNumber="36" unit="m" value="1785.0">1785 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<collectorName id="AD595314FFBFFFCCFA87EC82FC1BFA62" pageId="3" pageNumber="36">M. Á. de la Torre-Loranca</collectorName>
|
||
,
|
||
<collectorName id="AD595314FFBFFFCCFC0BECACFBE8FA62" box="[923,1149,1445,1467]" pageId="3" pageNumber="36">J. C. Sánchez-García</collectorName>
|
||
, and
|
||
<collectorName id="AD595314FFBFFFCCFB2AECACFA3EFA62" box="[1210,1451,1445,1467]" pageId="3" pageNumber="36">U. O. García-Vázquez</collectorName>
|
||
,
|
||
<date id="74121002FFBFFFCCFA27ECACFC7EFA01" pageId="3" pageNumber="36" value="2018-11-23">
|
||
<collectingDate id="6456E9EAFFBFFFCCFA27ECACFC7EFA01" pageId="3" pageNumber="36" value="2018-11-23">23 November 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="500A9EB9FFBFFFCCFC6BECCEFBEBFA01" box="[1019,1150,1475,1497]" collectionCode="MZFZ" pageId="3" pageNumber="36">MZFZ 4408</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFB1EECC9FA20FA00" ID-GBIF-Occurrence="4088553332" box="[1166,1461,1475,1497]" collectionCode="MZFZ" country="Mexico" pageId="3" pageNumber="36" specimenCode="MZFZ 4406" specimenCount="1" specimenCount-adult="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<specimenCount id="16AAFD4BFFBFFFCCFB1EECC9FB4AFA00" box="[1166,1247,1475,1497]" pageId="3" pageNumber="36" type="adult">1 adult</specimenCount>
|
||
male,
|
||
<specimenCode id="500A9EB9FFBFFFCCFAA1ECCEFA20FA00" box="[1329,1461,1475,1497]" collectionCode="MZFZ" pageId="3" pageNumber="36">MZFZ 4406</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFA54ECC9FBF6FA2F" ID-GBIF-Occurrence="4088553316" collectionCode="MZFZ" country="Mexico" pageId="3" pageNumber="36" specimenCode="MZFZ 4407, 1" specimenCount="1" specimenCount-adult="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
<specimenCount id="16AAFD4BFFBFFFCCFA54ECC9FCF9FA2E" pageId="3" pageNumber="36" type="adult">1 adult</specimenCount>
|
||
female,
|
||
<specimenCode id="500A9EB9FFBFFFCCFC5BECE8FBDFFA2F" box="[971,1098,1505,1527]" collectionCode="MZFZ" pageId="3" pageNumber="36">MZFZ 4407</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFBFFFCCFBC5ECEBFBF6FA2F" box="[1109,1123,1505,1526]" collectionCode="MZFZ" pageId="3" pageNumber="36">1</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="B0C43C9FFFBFFFCCFBFAECE8FA5BF989" ID-GBIF-Occurrence="4088553350" collectingDate="2019-02" collectingDateMax="2019-10" collectingDateMin="2019-02" collectionCode="R" collectorName="M. A. de la Torre-Loranca" country="Mexico" elevation="1715" latitude="18.618" location="Atiopa" longLatPrecision="76" longitude="-97.048" municipality="Municipality of San Juan Texhuacan" pageId="3" pageNumber="36" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
|
||
neonate,
|
||
<location id="05736019FFBFFFCCFB41ECEBFA89FA2E" LSID="urn:lsid:plazi:treatment:880587D4FFBFFFC4FFF7EEBEFDB1FA25:05736019FFBFFFCCFB41ECEBFA89FA2E" box="[1233,1308,1505,1527]" country="Mexico" latitude="18.618" longLatPrecision="76" longitude="-97.048" municipality="Municipality of San Juan Texhuacan" name="Atiopa" pageId="3" pageNumber="36" stateProvince="Veracruz de Ignacio de la Llave">Atiopa</location>
|
||
,
|
||
<collectingMunicipality id="E077ACB8FFBFFFCCFAB7ECEBFBBCF9CC" pageId="3" pageNumber="36">Municipality of San Juan Texhuacán</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="65985005FFBFFFCCFBADECF5FB0DF9CC" box="[1085,1176,1535,1557]" degrees="18.618" direction="north" orientation="latitude" pageId="3" pageNumber="36" precision="55" value="18.618">18.618N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="65985005FFBFFFCCFB3CECF5FA99F9CC" box="[1196,1292,1535,1557]" degrees="97.048" direction="west" orientation="longitude" pageId="3" pageNumber="36" precision="55" value="-97.048">97.048W</geoCoordinate>
|
||
, datum WGS84,
|
||
<quantity id="C7549B27FFBFFFCCFCA4EF17FC58F9EB" box="[820,973,1565,1587]" metricMagnitude="3" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.76" metricValueMin="1.67" pageId="3" pageNumber="36" unit="m" value="1715.0" valueMax="1760.0" valueMin="1670.0">
|
||
<elevation id="8B81D1F1FFBFFFCCFCA4EF17FC58F9EB" box="[820,973,1565,1587]" metricMagnitude="3" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.76" metricValueMin="1.67" pageId="3" pageNumber="36" unit="m" value="1715.0" valueMax="1760.0" valueMin="1670.0">1670–1760 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<collectorName id="AD595314FFBFFFCCFBDBEF17FAFCF9EB" box="[1099,1385,1560,1587]" pageId="3" pageNumber="36">M. Á. de la Torre-Loranca</collectorName>
|
||
and J.
|
||
<collectionCode id="66BDAE07FFBFFFCCFA2CEF17FA5BF9EB" box="[1468,1486,1565,1586]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="36">R</collectionCode>
|
||
.
|
||
<location id="05736019FFBFFFCCFCA4EF31FC63F988" LSID="urn:lsid:plazi:treatment:880587D4FFBFFFC4FFF7EEBEFDB1FA25:05736019FFBFFFCCFCA4EF31FC63F988" box="[820,1014,1595,1617]" country="Mexico" name="Hernandez Ginez" pageId="3" pageNumber="36" stateProvince="Veracruz de Ignacio de la Llave">Hernández Ginez</location>
|
||
, between
|
||
<collectingDate id="6456E9EAFFBFFFCCFBF3EF31FA5BF989" box="[1123,1486,1595,1617]" isEnumeration="true" pageId="3" pageNumber="36">
|
||
<date id="74121002FFBFFFCCFBF3EF31FB6DF989" box="[1123,1272,1595,1617]" pageId="3" pageNumber="36" value="2018-10">October 2018</date>
|
||
and
|
||
<date id="74121002FFBFFFCCFABEEF31FA5BF989" box="[1326,1486,1595,1617]" pageId="3" pageNumber="36" value="2019-02">February 2019</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFBFFFCCFCA4EF7BFC4AF881" blockId="3.[820,1490,1649,1880]" pageId="3" pageNumber="36">
|
||
<emphasis id="32D8EAD0FFBFFFCCFCA4EF7BFBB3F951" bold="true" box="[820,1062,1649,1672]" italics="true" pageId="3" pageNumber="36">Referred specimens.—</emphasis>
|
||
5 vouchers, all from the Sierra de Zongolica,
|
||
<collectingRegion id="C268F820FFBFFFCCFC21EF85FB87F97D" box="[945,1042,1679,1700]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="36">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry id="78BB7652FFBFFFCCFBB0EF85FBE1F97C" box="[1056,1140,1679,1701]" name="Mexico" pageId="3" pageNumber="36">Mexico</collectingCountry>
|
||
:
|
||
<collectionCode id="66BDAE07FFBFFFCCFB11EF85FB6FF97D" box="[1153,1274,1679,1700]" pageId="3" pageNumber="36">MZFZ-IMG</collectionCode>
|
||
309–311, Huapango, Municipality of Astacinga;
|
||
<collectionCode id="66BDAE07FFBFFFCCFB24EFA7FABAF91B" box="[1204,1327,1709,1730]" pageId="3" pageNumber="36">MZFZ-IMG</collectionCode>
|
||
312, Atiopa, Municipality of San Juan Texhuacán;
|
||
<collectionCode id="66BDAE07FFBFFFCCFA83EFC1FA1AF939" box="[1299,1423,1739,1760]" pageId="3" pageNumber="36">MZFZ-IMG</collectionCode>
|
||
313, Ayahuatulco, Municipality of Mixtla de Altamirano. All locality data in this and the preceding two paragraphs are masked; see the first paragraph of the Conservation subsection for details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="48B66549FFBFFFC9FCA4EE72FF43FB8B" lastPageId="6" lastPageNumber="39" pageId="3" pageNumber="36" type="diagnosis">
|
||
<paragraph id="001336C2FFBFFFCBFCA4EE72FE34F881" blockId="3.[820,1490,1912,2024]" lastBlockId="4.[142,812,1680,1881]" lastPageId="4" lastPageNumber="37" pageId="3" pageNumber="36">
|
||
<emphasis id="32D8EAD0FFBFFFCCFCA4EE72FBE3F857" box="[820,1142,1912,1935]" italics="true" pageId="3" pageNumber="36">
|
||
<emphasis id="32D8EAD0FFBFFFCCFCA4EE72FC2CF856" bold="true" box="[820,953,1912,1935]" italics="true" pageId="3" pageNumber="36">Diagnosis.—</emphasis>
|
||
<taxonomicName id="C7AC4D41FFBFFFCCFC29EE72FBE3F857" box="[953,1142,1912,1934]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="36" phylum="Chordata" rank="species" species="zongolica">Abronia zongolica</taxonomicName>
|
||
</emphasis>
|
||
can be distinguished from all described congeners (including members of the former genus
|
||
<taxonomicName id="C7AC4D41FFBFFFCCFCA4EEBEFC05F810" box="[820,912,1972,1993]" class="Reptilia" family="Anguidae" genus="Mesaspis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="36" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFBFFFCCFCA4EEBEFC05F810" box="[820,912,1972,1993]" italics="true" pageId="3" pageNumber="36">Mesaspis</emphasis>
|
||
</taxonomicName>
|
||
) by the following combination of characters: (1) one occipital scale; (2) two primary temporal scales contacting the postocular series; (3) posterolateral head scales moderately protuberant; (4) supra-auricular scales granular, not protuberant or spine-like; (5) 30–34 transverse dorsal scale rows; (6) dorsal scales on the flanks arranged in slightly oblique longitudinal rows relative to the ventrolateral fold; (7) lateralmost row of ventral scales unexpanded relative to the adjacent medial row.
|
||
</paragraph>
|
||
<caption id="54D3664AFFB8FFCBFF1EEF0AFD0DF9BD" ID-DOI="http://doi.org/10.5281/zenodo.7846899" ID-Zenodo-Dep="7846899" httpUri="https://zenodo.org/record/7846899/files/figure.png" pageId="4" pageNumber="37" startId="4.[142,170,1536,1556]" targetBox="[142,812,178,1506]" targetPageId="4" targetType="figure">
|
||
<paragraph id="001336C2FFB8FFCBFF1EEF0AFD0DF9BD" blockId="4.[142,811,1536,1636]" pageId="4" pageNumber="37">
|
||
FIG. 2.
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFF4BEF0AFE1FF9CD" authority="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca, 2022" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[219,394,1536,1556]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="zongolica" status="sp. nov.">
|
||
<emphasis id="32D8EAD0FFB8FFCBFF4BEF0AFE1FF9CD" bold="true" box="[219,394,1536,1556]" italics="true" pageId="4" pageNumber="37">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="29EB57ABFFB8FFCBFE08EF0AFD84F9CD" box="[408,529,1536,1556]" pageId="4" pageNumber="37" rank="species">new species</taxonomicNameLabel>
|
||
, head of holotype (MZFC-HE 35664, head width 25.2 mm) in dorsal view (top), right lateral view (middle; image intentionally mirrored horizontally) and ventral view (bottom), in preservative. Scale bars represent 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="001336C2FFB8FFCBFF1EEE72FB16FDF3" blockId="4.[142,812,1912,2024]" lastBlockId="4.[859,1530,179,2024]" pageId="4" pageNumber="37">
|
||
<emphasis id="32D8EAD0FFB8FFCBFF1EEE72FEA0F856" bold="true" box="[142,309,1912,1935]" italics="true" pageId="4" pageNumber="37">Comparisons.—</emphasis>
|
||
(
|
||
<tableCitation id="4D2E0379FFB8FFCBFEACEE73FE0CF857" box="[316,409,1913,1935]" captionStart="Table 1" captionStartId="6.[142,190,178,198]" captionTargetPageId="6" captionText="Table 1. Character states for selected features of Abronia zongolica and all other species in the oaXacae group." httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EE9B8FB29FF1F" pageId="4" pageNumber="37" tableUuid="54D3664AFFBAFFC9FF1EE9B8FB29FF1F">Tables 1</tableCitation>
|
||
,
|
||
<tableCitation id="4D2E0379FFB8FFCBFE39EE72FE22F856" box="[425,439,1912,1935]" captionStart="Table 2" captionStartId="6.[142,190,1262,1282]" captionTargetPageId="6" captionText="Table 2. Character states for selected features of Abronia zongolica and all species in Group III of Gutiérrez-Rodríguez et al. (2021)." httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EEDE4FAEEFADB" pageId="4" pageNumber="37" tableUuid="54D3664AFFBAFFC9FF1EEDE4FAEEFADB">2</tableCitation>
|
||
) Among members of the former genus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFF46EE9DFEA7F875" box="[214,306,1943,1964]" class="Reptilia" family="Anguidae" genus="Mesaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFF46EE9DFEA7F875" box="[214,306,1943,1964]" italics="true" pageId="4" pageNumber="37">Mesaspis</emphasis>
|
||
</taxonomicName>
|
||
, which was recently placed in the synonymy of
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFF3DEEBEFF6AF813" box="[173,255,1972,1994]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFF3DEEBEFF6AF813" box="[173,255,1972,1994]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
based on genomic evidence (
|
||
<bibRefCitation id="643D4B33FFB8FFCBFDDFEEBEFE83F831" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" pageId="4" pageNumber="37" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al., 2021</bibRefCitation>
|
||
),
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFEA0EEDBFE7AF83E" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[304,495,2001,2023]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB8FFCBFEA0EEDBFE7AF83E" box="[304,495,2001,2023]" italics="true" pageId="4" pageNumber="37">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
differs from all species by having dorsal scales on the flanks arranged in slightly oblique longitudinal rows relative to the ventrolateral fold (vs. parallel longitudinal rows), and by having 30–34 transverse dorsal scale rows (vs..40 rows). The new species further differs from
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBAFE823FB63FEE7" baseAuthorityName="Solano-Zavaleta, Nieto-Montes De Oca & Campbell" baseAuthorityYear="2016" box="[1087,1270,296,318]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="cuchumatanus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBAFE823FB63FEE7" box="[1087,1270,296,318]" italics="true" pageId="4" pageNumber="37">A. cuchumatanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFA96E823FAE5FEE7" baseAuthorityName="Boulenger" baseAuthorityYear="1913" box="[1286,1392,296,318]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gadovii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFA96E823FAE5FEE7" box="[1286,1392,296,318]" italics="true" pageId="4" pageNumber="37">A. gadovii</emphasis>
|
||
</taxonomicName>
|
||
, and the
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFA72E823FC21FE85" baseAuthorityName="Bocourt" baseAuthorityYear="1872" class="Reptilia" family="Anguidae" genus="Mesaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="moreletii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFA72E823FC21FE85" italics="true" pageId="4" pageNumber="37">A. moreletii</emphasis>
|
||
</taxonomicName>
|
||
complex by having 14 longitudinal dorsal scale rows (vs. 16 or 18, 16–18, and 18–22, respectively); from
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFA72E86EFC23FE4F" baseAuthorityName="Cope" baseAuthorityYear="1866" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="antauges">
|
||
<emphasis id="32D8EAD0FFB8FFCBFA72E86EFC23FE4F" italics="true" pageId="4" pageNumber="37">A. antauges</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC7FE88BFBC7FE4F" baseAuthorityName="Karges & Wright" baseAuthorityYear="1987" box="[1007,1106,385,406]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="juarezi">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC7FE88BFBC7FE4F" box="[1007,1106,385,406]" italics="true" pageId="4" pageNumber="37">A. juarezi</emphasis>
|
||
</taxonomicName>
|
||
by having vertebral and paravertebral dorsal scales distinctly keeled at midbody in adults (vs. smooth to slightly convex); from
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB41E8B6FACDFE08" baseAuthorityName="Bocourt" baseAuthorityYear="1873" box="[1233,1368,443,465]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="viridiflava">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB41E8B6FACDFE08" box="[1233,1368,443,465]" italics="true" pageId="4" pageNumber="37">A. viridiflava</emphasis>
|
||
</taxonomicName>
|
||
by having the frontonasal scale present (vs. absent); and from
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFAE4E8D3FA6CFE37" baseAuthorityName="Cope" baseAuthorityYear="1878" box="[1396,1529,472,494]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="32D8EAD0FFB8FFCBFAE4E8D3FA6CFE37" box="[1396,1529,472,494]" italics="true" pageId="4" pageNumber="37">A. monticola</emphasis>
|
||
</taxonomicName>
|
||
by having a divided postmental scale in 6/7 or 86% of specimens (vs. undivided).
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB8FFCBFCE3EB3BFADAFA42" blockId="4.[859,1530,179,2024]" pageId="4" pageNumber="37">
|
||
Among members of
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBC6EB3BFB3DFD9E" box="[1110,1192,561,583]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBC6EB3BFB3DFD9E" box="[1110,1192,561,583]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
not formerly attributed to the genus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC3BEB45FB92FDBD" box="[939,1031,591,612]" class="Reptilia" family="Anguidae" genus="Mesaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC3BEB45FB92FDBD" box="[939,1031,591,612]" italics="true" pageId="4" pageNumber="37">Mesaspis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB8CEB44FB48FDBD" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[1052,1245,590,612]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB8CEB44FB48FDBD" box="[1052,1245,590,612]" italics="true" pageId="4" pageNumber="37">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
differs from all species except the
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC4AEB66FB82FD5B" authorityName="Smith & Taylor" authorityYear="1950" box="[986,1047,620,642]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC4AEB66FB82FD5B" box="[986,1047,620,642]" italics="true" pageId="4" pageNumber="37">deppii</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<emphasis id="32D8EAD0FFB8FFCBFBE3EB67FB3EFD5B" box="[1139,1195,621,642]" italics="true" pageId="4" pageNumber="37">sensu</emphasis>
|
||
<bibRefCitation id="643D4B33FFB8FFCBFB25EB66FA3EFD5B" author="Campbell, J. A. & I. Solano-Zavaleta & O. Flores-Villela & I. W. Caviedes-Solis & D. R. Frost" box="[1205,1451,620,642]" pageId="4" pageNumber="37" pagination="149 - 156" refId="ref13073" refString="Campbell, J. A., I. Solano-Zavaleta, O. Flores-Villela, I. W. Caviedes-Solis, and D. R. Frost. 2016. A new species of Abronia (Squamata: Anguidae) from the Sierra Madre del Sur of Oaxaca, Mexico. Journal of Herpetology 50: 149 - 156." type="journal article" year="2016">Campbell et al., 2016</bibRefCitation>
|
||
;
|
||
<emphasis id="32D8EAD0FFB8FFCBFA29EB67FA6CFD5B" box="[1465,1529,621,642]" italics="true" pageId="4" pageNumber="37">contra</emphasis>
|
||
<bibRefCitation id="643D4B33FFB8FFCBFCCBEB83FB10FD79" author="Campbell, J. A. & D. R. Frost" box="[859,1157,649,672]" pageId="4" pageNumber="37" pagination="1 - 121" refId="ref12971" refString="Campbell, J. A., and D. R. Frost. 1993. Anguid lizards of the genus Abronia: revisionary notes, descriptions of four new species, a phylogenetic analysis, and key. Bulletin of the American Museum of Natural History 216: 1 - 121." type="journal article" year="1993">Campbell and Frost, 1993</bibRefCitation>
|
||
) by having dorsal scales on the flanks arranged in at least slightly oblique longitudinal rows relative to the ventrolateral fold (vs. parallel longitudinal rows in other species), and differs from all species except the subgenera
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC43EBF5FBB0FCCC" box="[979,1061,767,789]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC43EBF5FBB0FCCC" box="[979,1061,767,789]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBA4EBF5FB44FCCC" authorityName="Campbell & Frost" authorityYear="1993" box="[1076,1233,767,789]" class="Squamata" family="Anguidae" genus="Aenigmabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBA4EBF5FB44FCCC" box="[1076,1233,767,789]" italics="true" pageId="4" pageNumber="37">Aenigmabronia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFA83EBF5FA33FCCC" authorityName="Campbell & Frost" authorityYear="1993" box="[1299,1446,767,789]" class="Squamata" family="Anguidae" genus="Scopaeabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFA83EBF5FA33FCCC" box="[1299,1446,767,789]" italics="true" pageId="4" pageNumber="37">Scopaeabronia</emphasis>
|
||
</taxonomicName>
|
||
(plus a few individuals of
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBAAEA17FB59FCEB" box="[1082,1228,796,818]" class="Squamata" family="Anguidae" genus="Aenigmabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="lythrochila">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBAAEA17FB59FCEB" box="[1082,1228,796,818]" italics="true" pageId="4" pageNumber="37">A. lythrochila</emphasis>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="643D4B33FFB8FFCBFA9EEA16FC06FC89" author="Campbell, J. A. & D. R. Frost" pageId="4" pageNumber="37" pagination="1 - 121" refId="ref12971" refString="Campbell, J. A., and D. R. Frost. 1993. Anguid lizards of the genus Abronia: revisionary notes, descriptions of four new species, a phylogenetic analysis, and key. Bulletin of the American Museum of Natural History 216: 1 - 121." type="journal article" year="1993">Campbell and Frost, 1993</bibRefCitation>
|
||
) by having the lateralmost row of ventral scales unexpanded relative to the adjacent medial row (vs. expanded in other species). The new species further differs from the subgenera recognized by
|
||
<bibRefCitation id="643D4B33FFB8FFCBFB45EA98FA6CFC71" author="Campbell, J. A. & D. R. Frost" box="[1237,1529,913,936]" pageId="4" pageNumber="37" pagination="1 - 121" refId="ref12971" refString="Campbell, J. A., and D. R. Frost. 1993. Anguid lizards of the genus Abronia: revisionary notes, descriptions of four new species, a phylogenetic analysis, and key. Bulletin of the American Museum of Natural History 216: 1 - 121." type="journal article" year="1993">Campbell and Frost (1993)</bibRefCitation>
|
||
as follows. Unlike the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB7AEAA5FA15FC1C" authorityName="Campbell & Frost" authorityYear="1993" box="[1258,1408,943,965]" class="Squamata" family="Anguidae" genus="Abaculabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB7AEAA5FA15FC1C" box="[1258,1408,943,965]" italics="true" pageId="4" pageNumber="37">Abaculabronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species has a dorsum with dark, sometimes interrupted or faint crossbands in living adults (vs. dorsum with no trace of crossbands in living adults), and the supranasal scales not in contact (vs. in contact in 8/9 or 89% of specimens). Unlike the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB9BED48FB39FB81" authorityName="Campbell & Frost" authorityYear="1993" box="[1035,1196,1090,1112]" class="Squamata" family="Anguidae" genus="Aenigmabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB9BED48FB39FB81" box="[1035,1196,1090,1112]" italics="true" pageId="4" pageNumber="37">Aenigmabronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species has one occipital scale (vs. two occipitals), and protuberant head shields on the posterolateral ‘‘corners’’ of the head are present (vs. absent). Unlike the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFABDED90FA53FB69" authorityName="Campbell & Frost" authorityYear="1993" box="[1325,1478,1178,1200]" class="Squamata" family="Anguidae" genus="Auriculabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFABDED90FA53FB69" box="[1325,1478,1178,1200]" italics="true" pageId="4" pageNumber="37">Auriculabronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species has protuberant or spine-like supra-auricular scales absent (vs. present). Unlike the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFAEBEDDFFA67FB32" authorityName="Campbell & Frost" authorityYear="1993" box="[1403,1522,1237,1259]" class="Squamata" family="Anguidae" genus="Lissabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFAEBEDDFFA67FB32" box="[1403,1522,1237,1259]" italics="true" pageId="4" pageNumber="37">Lissabronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species has protuberant head shields on the posterolateral ‘‘corners’’ of the head present (vs. absent). Unlike the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBDAEC27FB48FA9A" authorityName="Campbell & Frost" authorityYear="1993" box="[1098,1245,1325,1347]" class="Squamata" family="Anguidae" genus="Scopaeabronia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBDAEC27FB48FA9A" box="[1098,1245,1325,1347]" italics="true" pageId="4" pageNumber="37">Scopaeabronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species has the lower primary temporal unexpanded (vs. expanded), six longitudinal nuchal scale rows (vs. eight rows), and 30–34 transverse dorsal scale rows (vs. 38–47 rows).
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB8FFCBFCE3ECA9FA6CF8C3" blockId="4.[859,1530,179,2024]" pageId="4" pageNumber="37">
|
||
Additionally within the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB7DECA9FAAAFA60" box="[1261,1343,1443,1465]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB7DECA9FAAAFA60" box="[1261,1343,1443,1465]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
, the new species shares with the
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB80ECCAFBD8FA0F" authorityName="Smith & Taylor" authorityYear="1950" box="[1040,1101,1472,1494]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB80ECCAFBD8FA0F" box="[1040,1101,1472,1494]" italics="true" pageId="4" pageNumber="37">deppii</emphasis>
|
||
</taxonomicName>
|
||
group the morphological synapomorphy of having dorsal scales on the flanks arranged in at least slightly oblique longitudinal rows relative to the ventrolateral fold. Because both the
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB0BEF12FB4DF9F7" authorityName="Smith & Taylor" authorityYear="1950" box="[1179,1240,1560,1582]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB0BEF12FB4DF9F7" box="[1179,1240,1560,1582]" italics="true" pageId="4" pageNumber="37">deppii</emphasis>
|
||
</taxonomicName>
|
||
group and the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFCCBEF3CFC25F995" box="[859,944,1590,1612]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFCCBEF3CFC25F995" box="[859,944,1590,1612]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
, unlike all other subgenera mentioned in the previous paragraph, are non-monophyletic based on the genomic analysis of
|
||
<bibRefCitation id="643D4B33FFB8FFCBFBDFEF7BFA5CF951" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" box="[1103,1481,1649,1672]" pageId="4" pageNumber="37" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al. (2021)</bibRefCitation>
|
||
, we here compare the new species to each species within the subgenus
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC5AEFA1FB89F918" box="[970,1052,1707,1729]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC5AEFA1FB89F918" box="[970,1052,1707,1729]" italics="true" pageId="4" pageNumber="37">Abronia</emphasis>
|
||
</taxonomicName>
|
||
. Among non-
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFB28EFA1FB60F918" authorityName="Smith & Taylor" authorityYear="1950" box="[1208,1269,1707,1729]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFB28EFA1FB60F918" box="[1208,1269,1707,1729]" italics="true" pageId="4" pageNumber="37">deppii</emphasis>
|
||
</taxonomicName>
|
||
-group members of this subgenus (
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFC40EFC3FBF8F906" baseAuthorityName="Tihen" baseAuthorityYear="1944" box="[976,1133,1737,1759]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="fuscolabialis">
|
||
<emphasis id="32D8EAD0FFB8FFCBFC40EFC3FBF8F906" box="[976,1133,1737,1759]" italics="true" pageId="4" pageNumber="37">A. fuscolabialis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFBEAEFC3FB62F906" box="[1146,1271,1737,1759]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB8FFCBFBEAEFC3FB62F906" box="[1146,1271,1737,1759]" italics="true" pageId="4" pageNumber="37">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFAA4EFC3FA30F906" box="[1332,1445,1737,1759]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="taeniata">
|
||
<emphasis id="32D8EAD0FFB8FFCBFAA4EFC3FA30F906" box="[1332,1445,1737,1759]" italics="true" pageId="4" pageNumber="37">A. taeniata</emphasis>
|
||
</taxonomicName>
|
||
; Group III of
|
||
<bibRefCitation id="643D4B33FFB8FFCBFC06EFEDFB64F924" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" box="[918,1265,1766,1789]" pageId="4" pageNumber="37" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al., 2021</bibRefCitation>
|
||
), the new species differs by having an oblique row of enlarged lateral neck scales, each
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB8FFC9FCFEEE2BFF43FB8B" blockId="4.[859,1530,179,2024]" lastBlockId="6.[142,812,751,1106]" lastPageId="6" lastPageNumber="39" pageId="4" pageNumber="37">
|
||
3 times larger than adjacent scales (vs. row of enlarged scales absent, or poorly developed and not reaching nuchal scales); flanks at least partially yellow and contrasting with darker back in adult males (vs. flanks and back similar in color); and dorsal scales on the flanks arranged in slightly oblique longitudinal rows relative to the ventrolateral fold (vs. parallel longitudinal rows). Within the
|
||
<taxonomicName id="C7AC4D41FFB8FFCBFAD0EEDBFAE8F83E" authorityName="Smith & Taylor" authorityYear="1950" box="[1344,1405,2001,2023]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFB8FFCBFAD0EEDBFAE8F83E" box="[1344,1405,2001,2023]" italics="true" pageId="4" pageNumber="37">deppii</emphasis>
|
||
</taxonomicName>
|
||
group, the new species differs from
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FE26EBFAFD82FCDC" authorityName="Smith & Taylor" authorityYear="1950" box="[438,535,751,773]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFBAFFC9FE26EBFAFD82FCDC" box="[438,535,751,773]" italics="true" pageId="6" pageNumber="39">A. deppii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDCAEBFAFCBEFCDC" box="[602,811,751,773]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="martindelcampoi">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDCAEBFAFCBEFCDC" box="[602,811,751,773]" italics="true" pageId="6" pageNumber="39">A. martindelcampoi</emphasis>
|
||
</taxonomicName>
|
||
(Group I of
|
||
<bibRefCitation id="643D4B33FFBAFFC9FE88EA04FDEEFCFD" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" box="[280,635,781,804]" pageId="6" pageNumber="39" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al., 2021</bibRefCitation>
|
||
) by having the anterior superciliary scale contacting the cantholoreal scale (vs. usually not in contact), the first postorbital supralabial scale not enlarged (vs. enlarged), two primary temporal scales contacting the postocular series (vs. one), and 30–34 transverse dorsal scale rows (vs.
|
||
<quantity id="C7549B27FFBAFFC9FE6CEAAEFDF5FC63" box="[508,608,932,954]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" metricValueMax="7.366" metricValueMin="6.858" pageId="6" pageNumber="39" unit="in" value="28.0" valueMax="29.0" valueMin="27.0">27–29 in</quantity>
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDF9EAAFFD52FC63" authorityName="Smith & Taylor" authorityYear="1950" box="[617,711,932,954]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDF9EAAFFD52FC63" box="[617,711,932,954]" italics="true" pageId="6" pageNumber="39">A. deppii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="C7549B27FFBAFFC9FC94EAAEFF5FFC00" metricMagnitude="-1" metricUnit="m" metricValue="6.604" metricValueMax="7.112" metricValueMin="6.096" pageId="6" pageNumber="39" unit="in" value="26.0" valueMax="28.0" valueMin="24.0">24– 28 in</quantity>
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FF43EAC9FE35FC01" box="[211,416,962,984]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="martindelcampoi">
|
||
<emphasis id="32D8EAD0FFBAFFC9FF43EAC9FE35FC01" box="[211,416,962,984]" italics="true" pageId="6" pageNumber="39">A. martindelcampoi</emphasis>
|
||
</taxonomicName>
|
||
). Within the
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDAFEAC9FD04FC01" box="[575,657,963,984]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="oaxacae">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDAFEAC9FD04FC01" box="[575,657,963,984]" italics="true" pageId="6" pageNumber="39">oaxacae</emphasis>
|
||
</taxonomicName>
|
||
group nested within the
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FE89EAEAFEC3FC2F" authorityName="Smith & Taylor" authorityYear="1950" box="[281,342,992,1014]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="deppii">
|
||
<emphasis id="32D8EAD0FFBAFFC9FE89EAEAFEC3FC2F" box="[281,342,992,1014]" italics="true" pageId="6" pageNumber="39">deppii</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FE2CEAEBFDA9FC2F" authorityName="Campbell, Solano-Zavaleta, Flores-Villela, Caviedes-Solis & Frost" authorityYear="2016" box="[444,572,992,1014]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="cuetzpali">
|
||
<emphasis id="32D8EAD0FFBAFFC9FE2CEAEBFDA9FC2F" box="[444,572,992,1014]" italics="true" pageId="6" pageNumber="39">A. cuetzpali</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDC2EAEBFD52FC2F" box="[594,711,993,1014]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="mixteca">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDC2EAEBFD52FC2F" box="[594,711,993,1014]" italics="true" pageId="6" pageNumber="39">A. mixteca</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FC85EAEBFF75FBCD" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="oaxacae">
|
||
<emphasis id="32D8EAD0FFBAFFC9FC85EAEBFF75FBCD" italics="true" pageId="6" pageNumber="39">A. oaxacae</emphasis>
|
||
</taxonomicName>
|
||
; Group V of
|
||
<bibRefCitation id="643D4B33FFBAFFC9FE13EAF5FD7AFBCC" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" box="[387,751,1022,1045]" pageId="6" pageNumber="39" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al., 2021</bibRefCitation>
|
||
), the new species differs by having one occipital scale (vs. two or three).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="54D3664AFFB9FFCAFFF7EE60FF0FF817" ID-DOI="http://doi.org/10.5281/zenodo.7846901" ID-Zenodo-Dep="7846901" httpUri="https://zenodo.org/record/7846901/files/figure.png" pageId="5" pageNumber="38" startId="5.[103,131,1898,1918]" targetBox="[269,1325,178,1868]" targetPageId="5" targetType="figure">
|
||
<paragraph id="001336C2FFB9FFCAFFF7EE60FF0FF817" blockId="5.[103,1490,1898,1998]" pageId="5" pageNumber="38">
|
||
FIG. 3.
|
||
<taxonomicName id="C7AC4D41FFB9FFCAFF22EE60FECBF8A7" authority="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca, 2022" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[178,350,1898,1918]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="38" phylum="Chordata" rank="species" species="zongolica" status="sp. nov.">
|
||
<emphasis id="32D8EAD0FFB9FFCAFF22EE60FECBF8A7" bold="true" box="[178,350,1898,1918]" italics="true" pageId="5" pageNumber="38">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="29EB57ABFFB9FFCAFEFAEE60FE74F8A7" box="[362,481,1898,1918]" pageId="5" pageNumber="38" rank="species">new species</taxonomicNameLabel>
|
||
, color variation in life (photos not to scale). (A) Adult male holotype (MZFC-HE 35664, snout–vent length [SVL] 112 mm); (B) adult male paratype (MZFC-HE 35665, SVL 95.8 mm); (C) adult female paratype (MZFZ 4406, SVL 86.6 mm; image intentionally mirrored horizontally); (D) neonate paratype (MZFZ 4407, SVL 45.6 mm). Photos by Miguel Ángel de la Torre-Loranca and Antonio Esaú Valdenegro- Brito.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="54D3664AFFBAFFC9FF1EE9B8FB29FF1F" ID-Table-UUID="54D3664AFFBAFFC9FF1EE9B8FB29FF1F" box="[142,1212,178,198]" httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EE9B8FB29FF1F" pageId="6" pageNumber="39" startId="6.[142,190,178,198]" targetBox="[142,1529,217,662]" targetIsTable="true" targetPageId="6" targetType="table">
|
||
<paragraph id="001336C2FFBAFFC9FF1EE9B8FB29FF1F" blockId="6.[142,1212,178,198]" box="[142,1212,178,198]" pageId="6" pageNumber="39">
|
||
Table 1. Character states for selected features of
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDFAE9B8FC82FF1F" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[618,791,178,198]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDFAE9B8FC82FF1F" bold="true" box="[618,791,178,198]" italics="true" pageId="6" pageNumber="39">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
and all other species in the
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FBB8E9B8FBEDFF1F" box="[1064,1144,178,198]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="oaxacae">
|
||
<emphasis id="32D8EAD0FFBAFFC9FBB8E9B8FBEDFF1F" bold="true" box="[1064,1144,178,198]" italics="true" pageId="6" pageNumber="39">oaXacae</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="001336C2FFBAFFC9FD47E9D3FA44FD4F" pageId="6" pageNumber="39">
|
||
<table id="72ACC462FFBA0030FF1EE9D3FA6CFD4F" box="[142,1529,217,662]" gridcols="5" gridrows="8" pageId="6" pageNumber="39">
|
||
<tr id="BE9C3480FFBA0030FF1EE9D3FA6CFF29" box="[142,1529,217,240]" gridrow="0" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<th id="FD4D5DFCFFBA0030FD47E9D3FCCCFF29" box="[727,857,217,240]" gridcol="1" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FD47E9D3FCCCFF29" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[727,857,217,240]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBAFFC9FD47E9D3FCCCFF29" bold="true" box="[727,857,217,240]" italics="true" pageId="6" pageNumber="39">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FC34E9D3FBEAFF29" box="[932,1151,217,240]" gridcol="2" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FC44E9D3FBDAFF29" authorityName="Campbell, Solano-Zavaleta, Flores-Villela, Caviedes-Solis & Frost" authorityYear="2016" box="[980,1103,217,240]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="cuetzpali">
|
||
<emphasis id="32D8EAD0FFBAFFC9FC44E9D3FBDAFF29" bold="true" box="[980,1103,217,240]" italics="true" pageId="6" pageNumber="39">A. cuetzpali</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FB5AE9D3FAADFF29" box="[1226,1336,217,240]" gridcol="3" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FB5AE9D3FAADFF29" box="[1226,1336,217,240]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="mixteca">
|
||
<emphasis id="32D8EAD0FFBAFFC9FB5AE9D3FAADFF29" bold="true" box="[1226,1336,217,240]" italics="true" pageId="6" pageNumber="39">A. miXteca</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FA13E9D3FA6CFF29" box="[1411,1529,217,240]" gridcol="4" gridrow="0" pageId="6" pageNumber="39">
|
||
<emphasis id="32D8EAD0FFBAFFC9FA13E9D3FA6CFF29" bold="true" box="[1411,1529,217,240]" italics="true" pageId="6" pageNumber="39">A. oaXacae</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EE800FA6CFEE7" box="[142,1529,266,318]" gridrow="1" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EE800FD1EFEE7" box="[142,651,266,318]" gridcol="0" gridrow="1" pageId="6" pageNumber="39">Canthal scale separate from posterior internasal scale</th>
|
||
<td id="FD4D5DFCFFBA0030FD47E800FCCCFEE7" box="[727,857,266,318]" gridcol="1" gridrow="1" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FC34E800FBEAFEE7" box="[932,1151,266,318]" gridcol="2" gridrow="1" pageId="6" pageNumber="39">yes or no</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AE800FAADFEE7" box="[1226,1336,266,318]" gridcol="3" gridrow="1" pageId="6" pageNumber="39">usually no</td>
|
||
<td id="FD4D5DFCFFBA0030FA13E800FA6CFEE7" box="[1411,1529,266,318]" gridcol="4" gridrow="1" pageId="6" pageNumber="39">no</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EE84EFA6CFEAE" box="[142,1529,324,375]" gridrow="2" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EE84EFD1EFEAE" box="[142,651,324,375]" gridcol="0" gridrow="2" pageId="6" pageNumber="39">Anterior superciliary scale contacting the cantholoreal scale</th>
|
||
<td id="FD4D5DFCFFBA0030FD47E84EFCCCFEAE" box="[727,857,324,375]" gridcol="1" gridrow="2" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FC34E84EFBEAFEAE" box="[932,1151,324,375]" gridcol="2" gridrow="2" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AE84EFAADFEAE" box="[1226,1336,324,375]" gridcol="3" gridrow="2" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FA13E84EFA6CFEAE" box="[1411,1529,324,375]" gridcol="4" gridrow="2" pageId="6" pageNumber="39">usually no</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EE877FA6CFE68" box="[142,1529,381,433]" gridrow="3" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EE877FD1EFE68" box="[142,651,381,433]" gridcol="0" gridrow="3" pageId="6" pageNumber="39">Temporal scales between posterior supralabial and posterolateral ‘‘corner’’ of head in adult males*</th>
|
||
<td id="FD4D5DFCFFBA0030FD47E877FCCCFE68" box="[727,857,381,433]" gridcol="1" gridrow="3" pageId="6" pageNumber="39">3</td>
|
||
<td id="FD4D5DFCFFBA0030FC34E877FBEAFE68" box="[932,1151,381,433]" gridcol="2" gridrow="3" pageId="6" pageNumber="39">2</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AE877FAADFE68" box="[1226,1336,381,433]" gridcol="3" gridrow="3" pageId="6" pageNumber="39">2–3</td>
|
||
<td id="FD4D5DFCFFBA0030FA13E877FA6CFE68" box="[1411,1529,381,433]" gridcol="4" gridrow="3" pageId="6" pageNumber="39">1–2</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EE8BDFA6CFDDE" box="[142,1529,439,519]" gridrow="4" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EE8BDFD1EFDDE" box="[142,651,439,519]" gridcol="0" gridrow="4" pageId="6" pageNumber="39">Occipital scale(s) Scale rows between the occipital scale(s) and first row of nuchal scales</th>
|
||
<td id="FD4D5DFCFFBA0030FD47E8BDFCCCFDDE" box="[727,857,439,519]" gridcol="1" gridrow="4" pageId="6" pageNumber="39">1 1–2</td>
|
||
<td id="FD4D5DFCFFBA0030FC34E8BDFBEAFDDE" box="[932,1151,439,519]" gridcol="2" gridrow="4" pageId="6" pageNumber="39">2–3 2</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AE8BDFAADFDDE" box="[1226,1336,439,519]" gridcol="3" gridrow="4" pageId="6" pageNumber="39">3 3</td>
|
||
<td id="FD4D5DFCFFBA0030FA13E8BDFA6CFDDE" box="[1411,1529,439,519]" gridcol="4" gridrow="4" pageId="6" pageNumber="39">3 1–2</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEB07FA6CFD99" box="[142,1529,525,576]" gridrow="5" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEB07FD1EFD99" box="[142,651,525,576]" gridcol="0" gridrow="5" pageId="6" pageNumber="39">Longitudinal nuchal scale rows</th>
|
||
<td id="FD4D5DFCFFBA0030FD47EB07FCCCFD99" box="[727,857,525,576]" gridcol="1" gridrow="5" pageId="6" pageNumber="39">6</td>
|
||
<td id="FD4D5DFCFFBA0030FC34EB07FBEAFD99" box="[932,1151,525,576]" gridcol="2" gridrow="5" pageId="6" pageNumber="39">6–8 (if 8, lateral rows reduced in size)</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AEB07FAADFD99" box="[1226,1336,525,576]" gridcol="3" gridrow="5" pageId="6" pageNumber="39">5–6</td>
|
||
<td id="FD4D5DFCFFBA0030FA13EB07FA6CFD99" box="[1411,1529,525,576]" gridcol="4" gridrow="5" pageId="6" pageNumber="39">4</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEB4CFA6CFDA3" box="[142,1529,582,634]" gridrow="6" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEB4CFD1EFDA3" box="[142,651,582,634]" gridcol="0" gridrow="6" pageId="6" pageNumber="39">Lateral neck scales between ventral scales and nuchal scales</th>
|
||
<td id="FD4D5DFCFFBA0030FD47EB4CFCCCFDA3" box="[727,857,582,634]" gridcol="1" gridrow="6" pageId="6" pageNumber="39">5–6</td>
|
||
<td id="FD4D5DFCFFBA0030FC34EB4CFBEAFDA3" box="[932,1151,582,634]" gridcol="2" gridrow="6" pageId="6" pageNumber="39">7–8</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AEB4CFAADFDA3" box="[1226,1336,582,634]" gridcol="3" gridrow="6" pageId="6" pageNumber="39">5–6</td>
|
||
<td id="FD4D5DFCFFBA0030FA13EB4CFA6CFDA3" box="[1411,1529,582,634]" gridcol="4" gridrow="6" pageId="6" pageNumber="39">3–4</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEB75FA6CFD4F" box="[142,1529,639,662]" gridrow="7" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEB75FD1EFD4F" box="[142,651,639,662]" gridcol="0" gridrow="7" pageId="6" pageNumber="39">Transverse dorsal scale rows</th>
|
||
<td id="FD4D5DFCFFBA0030FD47EB75FCCCFD4F" box="[727,857,639,662]" gridcol="1" gridrow="7" pageId="6" pageNumber="39">30–34</td>
|
||
<td id="FD4D5DFCFFBA0030FC34EB75FBEAFD4F" box="[932,1151,639,662]" gridcol="2" gridrow="7" pageId="6" pageNumber="39">32–35</td>
|
||
<td id="FD4D5DFCFFBA0030FB5AEB75FAADFD4F" box="[1226,1336,639,662]" gridcol="3" gridrow="7" pageId="6" pageNumber="39">28–31</td>
|
||
<td id="FD4D5DFCFFBA0030FA13EB75FA6CFD4F" box="[1411,1529,639,662]" gridcol="4" gridrow="7" pageId="6" pageNumber="39">27–29</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<footnote id="63B72ACCFFBAFFC9FF1EEBA7FB73FD1D" box="[142,1254,685,708]" pageId="6" pageNumber="39">
|
||
<paragraph id="001336C2FFBAFFC9FF1EEBA7FB73FD1D" blockId="6.[142,1254,685,708]" box="[142,1254,685,708]" pageId="6" pageNumber="39">
|
||
*
|
||
<emphasis id="32D8EAD0FFBAFFC9FF0EEBA7FF25FD1D" bold="true" box="[158,176,685,708]" pageId="6" pageNumber="39">¼</emphasis>
|
||
adult females sometimes also correctly segregate based on the character states presented for each species
|
||
</paragraph>
|
||
</footnote>
|
||
<subSubSection id="48B66549FFBAFFC6FF1EED7BFB17F831" lastPageId="9" lastPageNumber="42" pageId="6" pageNumber="39" type="description">
|
||
<paragraph id="001336C2FFBAFFC9FF1EED7BFCBEFB1D" blockId="6.[142,812,1137,1220]" pageId="6" pageNumber="39">
|
||
<emphasis id="32D8EAD0FFBAFFC9FF1EED7BFE52FB51" bold="true" box="[142,455,1137,1160]" italics="true" pageId="6" pageNumber="39">
|
||
Description of the
|
||
<typeStatus id="DF178860FFBAFFC9FEDEED7BFE39FB51" box="[334,428,1137,1160]" pageId="6" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="98972A47FFBAFFC9FE5FED7BFDA5FB51" box="[463,560,1137,1160]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[142,170,1536,1556]" captionStartId-1="5.[103,131,1898,1918]" captionStartId-2="7.[103,131,1804,1824]" captionTargetBox-0="[142,812,178,1506]" captionTargetBox-1="[269,1325,178,1868]" captionTargetBox-2="[204,1388,178,1775]" captionTargetId-0="figure-683@4.[142,812,178,1507]" captionTargetId-1="figure-1@5.[268,1325,178,1869]" captionTargetId-2="figure-1@7.[204,1389,178,1775]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIG. 2. Abronia zongolica, new species, head of holotype (MZFC-HE 35664, head width 25.2 mm) in dorsal view (top), right lateral view (middle; image intentionally mirrored horizontally) and ventral view (bottom), in preservative. Scale bars represent 10 mm." captionText-1="FIG. 3. Abronia zongolica, new species, color variation in life (photos not to scale). (A) Adult male holotype (MZFC-HE 35664, snout–vent length [SVL] 112 mm); (B) adult male paratype (MZFC-HE 35665, SVL 95.8 mm); (C) adult female paratype (MZFZ 4406, SVL 86.6 mm; image intentionally mirrored horizontally); (D) neonate paratype (MZFZ 4407, SVL 45.6 mm). Photos by Miguel Ángel de la Torre-Loranca and Antonio Esaú Valdenegro- Brito." captionText-2="FIG. 4. Head comparison of adult Abronia zongolica and Abronia graminea, illustrating differences in (1) circumorbital area color, (2) upper and lower jaw color pattern, and (3) presence/absence of well-developed oblique row of enlarged lateral neck scales. Locality and voucher information for each image is as follows, listed from top-to-bottom. Abronia zongolica: Teopantzacoalco (paratype, MZFC-HE 35665), Ayahuatulco (holotype, MZFC-HE 35664), Huapango (referred specimen, MZFZ-IMG 310), Atiopa (paratype, MZFZ 4406), Huapango (referred specimen, MZFZ-IMG 311). Abronia graminea: El Sumidero (MZFC-HE 32991), Puerto del Aire (MZFZ-IMG 322), Acultzinapa (MZFZ-IMG 336), Puerto del Aire (Clause et al., 2016: fig. 2), Puerto del Aire (MZFZ-IMG 338). All photos taken within 1 month of capture. From top, first and third images of A. zongolica and fifth image of A. graminea intentionally mirrored horizontally. Photos by Miguel Ángel de la Torre-Loranca, Adam G. Clause, and Antonio Esaú Valdenegro- Brito." figureDoi-0="http://doi.org/10.5281/zenodo.7846899" figureDoi-1="http://doi.org/10.5281/zenodo.7846901" figureDoi-2="http://doi.org/10.5281/zenodo.7846903" httpUri-0="https://zenodo.org/record/7846899/files/figure.png" httpUri-1="https://zenodo.org/record/7846901/files/figure.png" httpUri-2="https://zenodo.org/record/7846903/files/figure.png" pageId="6" pageNumber="39">Figs. 2–4</figureCitation>
|
||
) Adult male with both hemipenes partially everted, snout–vent length (SVL)
|
||
<quantity id="C7549B27FFBAFFC9FC93ED9AFF28FB1A" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="6" pageNumber="39" unit="mm" value="112.0">112 mm</quantity>
|
||
, head length from rostral to upper anterior edge of
|
||
</paragraph>
|
||
<paragraph id="001336C2FFBAFFC7FCCBEBE5FCB9FD78" blockId="6.[859,1529,751,1220]" lastBlockId="8.[142,812,179,1292]" lastPageId="8" lastPageNumber="41" pageId="6" pageNumber="39">
|
||
auricular opening
|
||
<quantity id="C7549B27FFBAFFC9FBBEEBE5FB0DFCDC" box="[1070,1176,751,773]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="6" pageNumber="39" unit="mm" value="25.2">25.2 mm</quantity>
|
||
, head width at broadest point
|
||
<quantity id="C7549B27FFBAFFC9FCCBEA07FC50FCFA" box="[859,965,781,803]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="6" pageNumber="39" unit="mm" value="19.6">19.6 mm</quantity>
|
||
, head width/length ratio
|
||
<emphasis id="32D8EAD0FFBAFFC9FB7DEA06FB6AFCFA" bold="true" box="[1261,1279,780,803]" pageId="6" pageNumber="39">¼</emphasis>
|
||
78.4%, tail unbroken and unregenerated, tail length (TL)
|
||
<quantity id="C7549B27FFBAFFC9FB4EEA21FAAFFC98" box="[1246,1338,811,833]" metricMagnitude="-1" metricUnit="m" metricValue="1.54" pageId="6" pageNumber="39" unit="mm" value="154.0">154 mm</quantity>
|
||
(1.37 times SVL), and 81 caudal whorls. Supranasals 1/1, somewhat expanded medially; postnasals 2/2, upper slightly smaller than lower; one pair of anterior and posterior internasals situated between rostral and frontonasal; prefrontals over twice as large as posterior internasals, contacting each other medially; canthals 1/1, preventing contact between posterior internasal and prefrontal; loreals 1/1, about one-half size of cantholoreals, and contacting both postnasals; cantholoreals 1/1, extending onto dorsum of canthus rostralis, broadly contacting anterior median supraocular, canthal, prefrontal, and fifth supralabial; median supraoculars 5/5; lateral supraoculars 3/3; superciliaries 6/5, anteriormost on left side contacting cantholoreal and similar in length to adjoining superciliary, on right side fused with adjoining superciliary; preoculars 1/1; suboculars 2/2, posteriormost barely separat- ed from lowermost primary temporal by ninth (left side) or eighth (right side) supralabial; postoculars 3/3; frontal large, separated from frontonasal and interparietal; parietal separated from median supraoculars; one large occipital, slightly larger than interparietal, abnormally divided into a small triangular anterior scale and a much larger posterior scale; one transverse row of scales separating occipital from first transverse row of nuchals; primary temporals 4/4, only lowermost two contacting postoculars on each side; secondary temporals 4/4; tertiary temporals 4/4; supralabials 10/11, antepenultimate the posteriormost to reach orbit; infralabials 9/9; postmental divided; four pairs of enlarged chinshields posterior to postmental, posteriormost ones subequal in size to adjacent ones, separated by two scales; sublabials 4/4, anteriormost contacting third infralabial but not postmental.
|
||
</paragraph>
|
||
<caption id="54D3664AFFBAFFC9FF1EEDE4FAEEFADB" ID-Table-UUID="54D3664AFFBAFFC9FF1EEDE4FAEEFADB" box="[142,1403,1261,1282]" httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EEDE4FAEEFADB" pageId="6" pageNumber="39" startId="6.[142,190,1262,1282]" targetBox="[142,1498,1300,1976]" targetIsTable="true" targetPageId="6" targetType="table">
|
||
<paragraph id="001336C2FFBAFFC9FF1EEDE4FAEEFADB" blockId="6.[142,1403,1261,1282]" box="[142,1403,1261,1282]" pageId="6" pageNumber="39">
|
||
Table 2. Character states for selected features of
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDFAEDE4FC82FADB" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[618,791,1262,1282]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDFAEDE4FC82FADB" bold="true" box="[618,791,1262,1282]" italics="true" pageId="6" pageNumber="39">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
and all species in Group III of
|
||
<bibRefCitation id="643D4B33FFBAFFC9FBADEDE7FAE2FADB" author="Gutierrez-Rodriguez, J. & A. Zaldivar-Riveron & I. Solano-Zavaleta & J. A. Campbell & R. N. Meza-Lazaro & O. Flores-Villela & A. Nieto-Montes de Oca" box="[1085,1399,1261,1282]" pageId="6" pageNumber="39" pagination="106963" refId="ref14228" refString="Gutierrez-Rodriguez, J., A. Zaldivar-Riveron, I. Solano-Zavaleta, J. A. Campbell, R. N. Meza-Lazaro, O. Flores-Villela, and A. Nieto-Montes de Oca. 2021. Phylogenomics of the Mesoamerican alligator-lizard genera Abronia and Mesaspis (Anguidae: Gerrhonotinae) reveals multiple independent clades of arboreal and terrestrial species. Molecular Phylogenetics and Evolution 154: 106963." type="journal article" year="2021">Gutiérrez-Rodríguez et al. (2021)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="001336C2FFBAFFC9FDC7EC1EFF5EF861" pageId="6" pageNumber="39">
|
||
<table id="72ACC462FFBA0030FF1EEC1EFA4FF861" box="[142,1498,1300,1976]" gridcols="5" gridrows="23" pageId="6" pageNumber="39">
|
||
<tr id="BE9C3480FFBA0030FF1EEC1EFA4FFAF2" box="[142,1498,1300,1323]" gridrow="0" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<th id="FD4D5DFCFFBA0030FDB9EC1EFC93FAF2" box="[553,774,1300,1323]" gridcol="1" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FDC7EC1EFD4CFAF2" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[599,729,1300,1323]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBAFFC9FDC7EC1EFD4CFAF2" bold="true" box="[599,729,1300,1323]" italics="true" pageId="6" pageNumber="39">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FCB4EC1EFC6BFAF2" box="[804,1022,1300,1323]" gridcol="2" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FCD3EC1EFC76FAF2" baseAuthorityName="Tihen" baseAuthorityYear="1944" box="[835,995,1300,1323]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="fuscolabialis">
|
||
<emphasis id="32D8EAD0FFBAFFC9FCD3EC1EFC76FAF2" bold="true" box="[835,995,1300,1323]" italics="true" pageId="6" pageNumber="39">A. fuscolabialis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FB8FEC1EFB6FFAF2" box="[1055,1274,1300,1323]" gridcol="3" gridrow="0" pageId="6" pageNumber="39">
|
||
<taxonomicName id="C7AC4D41FFBAFFC9FBDCEC1EFB45FAF2" box="[1100,1232,1300,1323]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="39" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFBAFFC9FBDCEC1EFB45FAF2" bold="true" box="[1100,1232,1300,1323]" italics="true" pageId="6" pageNumber="39">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="FD4D5DFCFFBA0030FA8AEC1EFA4FFAF2" box="[1306,1498,1300,1323]" gridcol="4" gridrow="0" pageId="6" pageNumber="39">
|
||
<emphasis id="32D8EAD0FFBAFFC9FADFEC1EFA51FAF2" bold="true" box="[1359,1476,1300,1323]" italics="true" pageId="6" pageNumber="39">A. taeniata</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEC4CFA4FFA87" box="[142,1498,1350,1374]" gridrow="1" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEC4CFD98FA87" box="[142,525,1350,1374]" gridcol="0" gridrow="1" pageId="6" pageNumber="39">Circumorbital area color different from</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EC4CFC93FA87" box="[553,774,1350,1374]" gridcol="1" gridrow="1" pageId="6" pageNumber="39">usually no (if yes,</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EC4CFC6BFA87" box="[804,1022,1350,1374]" gridcol="2" gridrow="1" pageId="6" pageNumber="39">no</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEC4CFB6FFA87" box="[1055,1274,1350,1374]" gridcol="3" gridrow="1" pageId="6" pageNumber="39">usually yes (bright to</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEC4CFA4FFA87" box="[1306,1498,1350,1374]" gridcol="4" gridrow="1" pageId="6" pageNumber="39">no</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEC69FA4FFAA3" box="[142,1498,1379,1402]" gridrow="2" pageId="6" pageNumber="39" rowspan-2="1" rowspan-4="1">
|
||
<th id="FD4D5DFCFFBA0030FF1EEC69FD98FAA3" box="[142,525,1379,1402]" gridcol="0" gridrow="2" pageId="6" pageNumber="39">surrounding scales in adults</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EC69FC93FAA3" box="[553,774,1379,1402]" gridcol="1" gridrow="2" pageId="6" pageNumber="39">then white or pale</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEC69FB6FFAA3" box="[1055,1274,1379,1402]" gridcol="3" gridrow="2" pageId="6" pageNumber="39">pale yellow)</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEC8AFA4FFA4E" box="[142,1498,1408,1431]" gridrow="3" pageId="6" pageNumber="39" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EC8AFC93FA4E" box="[553,774,1408,1431]" gridcol="1" gridrow="3" pageId="6" pageNumber="39">cream)</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEC96FA4FFA6D" box="[142,1498,1436,1460]" gridrow="4" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEC96FD98FA6D" box="[142,525,1436,1460]" gridcol="0" gridrow="4" pageId="6" pageNumber="39">Canthal scale separate from posterior</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EC96FC93FA6D" box="[553,774,1436,1460]" gridcol="1" gridrow="4" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EC96FC6BFA6D" box="[804,1022,1436,1460]" gridcol="2" gridrow="4" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEC96FB6FFA6D" box="[1055,1274,1436,1460]" gridcol="3" gridrow="4" pageId="6" pageNumber="39">no</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEC96FA4FFA6D" box="[1306,1498,1436,1460]" gridcol="4" gridrow="4" pageId="6" pageNumber="39">no</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EECB3FA4FFA09" box="[142,1498,1465,1488]" gridrow="5" pageId="6" pageNumber="39" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="FD4D5DFCFFBA0030FF1EECB3FD98FA09" box="[142,525,1465,1488]" gridcol="0" gridrow="5" pageId="6" pageNumber="39">internasal scale</th>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EECDCFA4FFA34" box="[142,1498,1494,1517]" gridrow="6" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EECDCFD98FA34" box="[142,525,1494,1517]" gridcol="0" gridrow="6" pageId="6" pageNumber="39">Postnasal supralabial scales and lower</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9ECDCFC93FA34" box="[553,774,1494,1517]" gridcol="1" gridrow="6" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4ECDCFC6BFA34" box="[804,1022,1494,1517]" gridcol="2" gridrow="6" pageId="6" pageNumber="39">yes</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FECDCFB6FFA34" box="[1055,1274,1494,1517]" gridcol="3" gridrow="6" pageId="6" pageNumber="39">no, lower jaw and</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AECDCFA4FFA34" box="[1306,1498,1494,1517]" gridcol="4" gridrow="6" pageId="6" pageNumber="39">yes</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EECF8FA4FF9D0" box="[142,1498,1522,1545]" gridrow="7" pageId="6" pageNumber="39" rowspan-1="1" rowspan-2="1" rowspan-4="1">
|
||
<th id="FD4D5DFCFFBA0030FF1EECF8FD98F9D0" box="[142,525,1522,1545]" gridcol="0" gridrow="7" pageId="6" pageNumber="39">temporal scales similar in color to</th>
|
||
<td id="FD4D5DFCFFBA0030FB8FECF8FB6FF9D0" box="[1055,1274,1522,1545]" gridcol="3" gridrow="7" pageId="6" pageNumber="39">throat much paler</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF05FA4FF9FF" box="[142,1498,1551,1574]" gridrow="8" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEF05FA4FF9FF" box="[142,1498,1551,1574]" colspan="5" colspanRight="4" gridcol="0" gridrow="8" pageId="6" pageNumber="39">lower jaw and throat</th>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF26FA4FF99A" box="[142,1498,1580,1603]" gridrow="9" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEF26FD98F99A" box="[142,525,1580,1603]" gridcol="0" gridrow="9" pageId="6" pageNumber="39">Lateral neck scales</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EF26FC93F99A" box="[553,774,1580,1603]" gridcol="1" gridrow="9" pageId="6" pageNumber="39">oblique row of</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EF26FC6BF99A" box="[804,1022,1580,1603]" gridcol="2" gridrow="9" pageId="6" pageNumber="39">row of distinctly</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEF26FB6FF99A" box="[1055,1274,1580,1603]" gridcol="3" gridrow="9" pageId="6" pageNumber="39">row of distinctly</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEF26FA4FF99A" box="[1306,1498,1580,1603]" gridcol="4" gridrow="9" pageId="6" pageNumber="39">row of distinctly</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF42FA4FF986" box="[142,1498,1608,1631]" gridrow="10" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EF42FC93F986" box="[553,774,1608,1631]" gridcol="1" gridrow="10" pageId="6" pageNumber="39">enlarged scales</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EF42FC6BF986" box="[804,1022,1608,1631]" gridcol="2" gridrow="10" pageId="6" pageNumber="39">enlarged scales</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEF42FB6FF986" box="[1055,1274,1608,1631]" gridcol="3" gridrow="10" pageId="6" pageNumber="39">enlarged scales</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEF42FA4FF986" box="[1306,1498,1608,1631]" gridcol="4" gridrow="10" pageId="6" pageNumber="39">enlarged scales</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF6FFA4FF9A5" box="[142,1498,1637,1660]" gridrow="11" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EF6FFC93F9A5" box="[553,774,1637,1660]" gridcol="1" gridrow="11" pageId="6" pageNumber="39">each 3 times</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EF6FFC6BF9A5" box="[804,1022,1637,1660]" gridcol="2" gridrow="11" pageId="6" pageNumber="39">absent</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEF6FFB6FF9A5" box="[1055,1274,1637,1660]" gridcol="3" gridrow="11" pageId="6" pageNumber="39">absent or poorly</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEF6FFA4FF9A5" box="[1306,1498,1637,1660]" gridcol="4" gridrow="11" pageId="6" pageNumber="39">absent</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF88FA4FF940" box="[142,1498,1666,1689]" gridrow="12" pageId="6" pageNumber="39" rowspan-0="1" rowspan-2="1" rowspan-4="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EF88FC93F940" box="[553,774,1666,1689]" gridcol="1" gridrow="12" pageId="6" pageNumber="39">larger than adjacent</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEF88FB6FF940" box="[1055,1274,1666,1689]" gridcol="3" gridrow="12" pageId="6" pageNumber="39">developed</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEF94FA4FF96C" box="[142,1498,1694,1717]" gridrow="13" pageId="6" pageNumber="39" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EF94FC93F96C" box="[553,774,1694,1717]" gridcol="1" gridrow="13" pageId="6" pageNumber="39">scales</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEFB1FA4FF90B" box="[142,1498,1723,1746]" gridrow="14" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEFB1FD98F90B" box="[142,525,1723,1746]" gridcol="0" gridrow="14" pageId="6" pageNumber="39">Longitudinal nuchal scale rows</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EFB1FC93F90B" box="[553,774,1723,1746]" gridcol="1" gridrow="14" pageId="6" pageNumber="39">6</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EFB1FC6BF90B" box="[804,1022,1723,1746]" gridcol="2" gridrow="14" pageId="6" pageNumber="39">6 (rarely 4)</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEFB1FB6FF90B" box="[1055,1274,1723,1746]" gridcol="3" gridrow="14" pageId="6" pageNumber="39">4–6</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEFB1FA4FF90B" box="[1306,1498,1723,1746]" gridcol="4" gridrow="14" pageId="6" pageNumber="39">6 (very rarely 4)</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEFD2FA4FF936" box="[142,1498,1752,1775]" gridrow="15" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEFD2FD98F936" box="[142,525,1752,1775]" gridcol="0" gridrow="15" pageId="6" pageNumber="39">Trunk coloration in adult males*</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EFD2FC93F936" box="[553,774,1752,1775]" gridcol="1" gridrow="15" pageId="6" pageNumber="39">flanks at least partially</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EFD2FC6BF936" box="[804,1022,1752,1775]" gridcol="2" gridrow="15" pageId="6" pageNumber="39">flanks and back green</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEFD2FB6FF936" box="[1055,1274,1752,1775]" gridcol="3" gridrow="15" pageId="6" pageNumber="39">flanks and back bright</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEFD2FA4FF936" box="[1306,1498,1752,1775]" gridcol="4" gridrow="15" pageId="6" pageNumber="39">flanks and back</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEFFFFA4FF8D5" box="[142,1498,1781,1804]" gridrow="16" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EFFFFC93F8D5" box="[553,774,1781,1804]" gridcol="1" gridrow="16" pageId="6" pageNumber="39">yellow, back brown</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EFFFFC6BF8D5" box="[804,1022,1781,1804]" gridcol="2" gridrow="16" pageId="6" pageNumber="39">or turquoise, with</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEFFFFB6FF8D5" box="[1055,1274,1781,1804]" gridcol="3" gridrow="16" pageId="6" pageNumber="39">green, scales often</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEFFFFA4FF8D5" box="[1306,1498,1781,1804]" gridcol="4" gridrow="16" pageId="6" pageNumber="39">yellow or cream</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEE1BFA4FF8F1" box="[142,1498,1809,1832]" gridrow="17" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EE1BFC93F8F1" box="[553,774,1809,1832]" gridcol="1" gridrow="17" pageId="6" pageNumber="39">or gray, often with</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EE1BFC6BF8F1" box="[804,1022,1809,1832]" gridcol="2" gridrow="17" pageId="6" pageNumber="39">entire or broken</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEE1BFB6FF8F1" box="[1055,1274,1809,1832]" gridcol="3" gridrow="17" pageId="6" pageNumber="39">with black bases</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEE1BFA4FF8F1" box="[1306,1498,1809,1832]" gridcol="4" gridrow="17" pageId="6" pageNumber="39">with wide dark</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEE24FA4FF89C" box="[142,1498,1838,1861]" gridrow="18" pageId="6" pageNumber="39" rowspan-0="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EE24FC93F89C" box="[553,774,1838,1861]" gridcol="1" gridrow="18" pageId="6" pageNumber="39">broken black</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EE24FC6BF89C" box="[804,1022,1838,1861]" gridcol="2" gridrow="18" pageId="6" pageNumber="39">dark or yellow</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEE24FB6FF89C" box="[1055,1274,1838,1861]" gridcol="3" gridrow="18" pageId="6" pageNumber="39">and/or yellow tips</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEE24FA4FF89C" box="[1306,1498,1838,1861]" gridcol="4" gridrow="18" pageId="6" pageNumber="39">transverse bands</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEE41FA4FF8BB" box="[142,1498,1867,1890]" gridrow="19" pageId="6" pageNumber="39" rowspan-0="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="FD4D5DFCFFBA0030FDB9EE41FC93F8BB" box="[553,774,1867,1890]" gridcol="1" gridrow="19" pageId="6" pageNumber="39">transverse bands</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EE41FC6BF8BB" box="[804,1022,1867,1890]" gridcol="2" gridrow="19" pageId="6" pageNumber="39">transverse bands</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEE6DFA4FF8A7" box="[142,1498,1895,1918]" gridrow="20" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEE6DFD98F8A7" box="[142,525,1895,1918]" gridcol="0" gridrow="20" pageId="6" pageNumber="39">Arrangement of dorsal scales on</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EE6DFC93F8A7" box="[553,774,1895,1918]" gridcol="1" gridrow="20" pageId="6" pageNumber="39">slightly oblique</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EE6DFC6BF8A7" box="[804,1022,1895,1918]" gridcol="2" gridrow="20" pageId="6" pageNumber="39">parallel longitudinal</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEE6DFB6FF8A7" box="[1055,1274,1895,1918]" gridcol="3" gridrow="20" pageId="6" pageNumber="39">parallel longitudinal</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEE6DFA4FF8A7" box="[1306,1498,1895,1918]" gridcol="4" gridrow="20" pageId="6" pageNumber="39">parallel longitudinal</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEE8EFA4FF842" box="[142,1498,1924,1947]" gridrow="21" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEE8EFD98F842" box="[142,525,1924,1947]" gridcol="0" gridrow="21" pageId="6" pageNumber="39">flanks relative to the ventrolateral</th>
|
||
<td id="FD4D5DFCFFBA0030FDB9EE8EFC93F842" box="[553,774,1924,1947]" gridcol="1" gridrow="21" pageId="6" pageNumber="39">longitudinal rows</td>
|
||
<td id="FD4D5DFCFFBA0030FCB4EE8EFC6BF842" box="[804,1022,1924,1947]" gridcol="2" gridrow="21" pageId="6" pageNumber="39">rows</td>
|
||
<td id="FD4D5DFCFFBA0030FB8FEE8EFB6FF842" box="[1055,1274,1924,1947]" gridcol="3" gridrow="21" pageId="6" pageNumber="39">rows</td>
|
||
<td id="FD4D5DFCFFBA0030FA8AEE8EFA4FF842" box="[1306,1498,1924,1947]" gridcol="4" gridrow="21" pageId="6" pageNumber="39">rows</td>
|
||
</tr>
|
||
<tr id="BE9C3480FFBA0030FF1EEEABFA4FF861" box="[142,1498,1953,1976]" gridrow="22" pageId="6" pageNumber="39">
|
||
<th id="FD4D5DFCFFBA0030FF1EEEABFA4FF861" box="[142,1498,1953,1976]" colspan="5" colspanRight="4" gridcol="0" gridrow="22" pageId="6" pageNumber="39">fold</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="001336C2FFBAFFC9FF1EEEC5FB73F83F" blockId="6.[142,1254,1998,2022]" box="[142,1254,1998,2022]" pageId="6" pageNumber="39">
|
||
<tableNote id="FD4A374CFFBAFFC9FF1EEEC5FB73F83F" box="[142,1254,1998,2022]" pageId="6" pageNumber="39" targetBox="[142,1498,1300,1976]" targetPageId="6">
|
||
*
|
||
<emphasis id="32D8EAD0FFBAFFC9FF0EEEC4FF25F83C" bold="true" box="[158,176,1998,2021]" pageId="6" pageNumber="39">¼</emphasis>
|
||
adult females sometimes also correctly segregate based on the character states presented for each species
|
||
</tableNote>
|
||
</paragraph>
|
||
<caption id="54D3664AFFBBFFC8FFF7EE06FF0FF803" ID-DOI="http://doi.org/10.5281/zenodo.7846903" ID-Zenodo-Dep="7846903" httpUri="https://zenodo.org/record/7846903/files/figure.png" pageId="7" pageNumber="40" startId="7.[103,131,1804,1824]" targetBox="[204,1388,178,1775]" targetPageId="7" targetType="figure">
|
||
<paragraph id="001336C2FFBBFFC8FFF7EE06FF0FF803" blockId="7.[103,1491,1804,2010]" pageId="7" pageNumber="40">
|
||
FIG. 4. Head comparison of adult
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FE21EE06FDCBF8F9" authorityName="García-Vázquez & Clause & Gutiérrez-Rodríguez & Cazares-Hernández & Torre-Loranca" authorityYear="2022" box="[433,606,1804,1824]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBBFFC8FE21EE06FDCBF8F9" bold="true" box="[433,606,1804,1824]" italics="true" pageId="7" pageNumber="40">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FD00EE06FCABF8F9" box="[656,830,1804,1824]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFBBFFC8FD00EE06FCABF8F9" bold="true" box="[656,830,1804,1824]" italics="true" pageId="7" pageNumber="40">Abronia graminea</emphasis>
|
||
</taxonomicName>
|
||
, illustrating differences in (1) circumorbital area color, (2) upper and lower jaw color pattern, and (3) presence/absence of well-developed oblique row of enlarged lateral neck scales. Locality and voucher information for each image is as follows, listed from top-to-bottom.
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FD11EE4BFC4CF88C" authority=": Teopantzacoalco" authorityName="Teopantzacoalco" box="[641,985,1857,1877]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBBFFC8FD11EE4BFCB8F88C" bold="true" box="[641,813,1857,1877]" italics="true" pageId="7" pageNumber="40">Abronia zongolica</emphasis>
|
||
: Teopantzacoalco
|
||
</taxonomicName>
|
||
(paratype, MZFC-HE 35665), Ayahuatulco (holotype, MZFC-HE 35664), Huapango (referred specimen, MZFZ-IMG 310), Atiopa (paratype, MZFZ 4406), Huapango (referred specimen, MZFZ-IMG 311).
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FFF7EE7CFE0DF852" authority=": El Sumidero" authorityName="El Sumidero" box="[103,408,1910,1931]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFBBFFC8FFF7EE7CFE80F853" bold="true" box="[103,277,1910,1930]" italics="true" pageId="7" pageNumber="40">Abronia graminea</emphasis>
|
||
: El Sumidero
|
||
</taxonomicName>
|
||
(MZFC-HE 32991), Puerto del Aire (MZFZ-IMG 322), Acultzinapa (MZFZ-IMG 336), Puerto del Aire (
|
||
<bibRefCitation id="643D4B33FFBBFFC8FAC4EE7DFF0EF87C" author="Clause, A. G. & G. Jimenez-Velazquez & H. A" pageId="7" pageNumber="40" pagination="142 - 145" refId="ref13465" refString="Clause, A. G., G. Jimenez-Velazquez, and H. A. Perez- Mendoza. 2016. Nature notes. Abronia graminea (Cope, 1864). Color variant. Mesoamerican Herpetology 3: 142 - 145." type="journal article" year="2016">Clause et al., 2016</bibRefCitation>
|
||
: fig. 2), Puerto del Aire (MZFZ-IMG 338). All photos taken within 1 month of capture. From top, first and third images of
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FA9AEE9BFAEBF87C" box="[1290,1406,1937,1957]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFBBFFC8FA9AEE9BFAEBF87C" bold="true" box="[1290,1406,1937,1957]" italics="true" pageId="7" pageNumber="40">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
and fifth image of
|
||
<taxonomicName id="C7AC4D41FFBBFFC8FF2DEEA1FEA5F866" box="[189,304,1963,1983]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="40" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFBBFFC8FF2DEEA1FEA5F866" bold="true" box="[189,304,1963,1983]" italics="true" pageId="7" pageNumber="40">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
intentionally mirrored horizontally. Photos by Miguel Ángel de la Torre-Loranca, Adam G. Clause, and Antonio Esaú Valdenegro- Brito.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="001336C2FFB4FFC7FF36EBA2FE75FAD2" blockId="8.[142,812,179,1292]" pageId="8" pageNumber="41">Minimum longitudinal nuchal scale rows 6; transverse dorsal scale rows 32–33; longitudinal dorsal scale rows 16, arranged in slightly oblique longitudinal rows on sides of body; eight dorsal-most longitudinal scale rows strongly keeled, others entirely smooth or with faint partial keels; transverse ventral scale rows 33; longitudinal ventral scale rows 12; lateralmost row of ventral scales unexpanded relative to the adjacent medial row of ventral scales; head and several anterior rows of nuchals with well-developed osteoderms; more posteriorly on dorsum and flanks of body, osteoderms appear very weakly developed or absent; supra-auricular scales granular and non-protruding; about seven moderately sized scales (lateral neck scales) between lateral nuchals and first large scales on ventrolateral surface of neck; lateral neck scales with enlarged oblique row reaching the lateral nuchals, each scale in row 3 times larger than adjacent lateral neck scales; antebrachial from insertion of the forelimb to wrist 12–13; ventrolateral fold moderately well developed with small scales and granules interspersed in interstitial skin throughout ventrolateral fold; subdigital lamellae on fourth toes 21/21.</paragraph>
|
||
<paragraph id="001336C2FFB4FFC7FF1EEC21FB65FE6D" blockId="8.[142,812,1323,2024]" lastBlockId="8.[859,1530,179,584]" pageId="8" pageNumber="41">
|
||
<emphasis id="32D8EAD0FFB4FFC7FF1EEC21FE46FA9B" bold="true" box="[142,467,1323,1346]" italics="true" pageId="8" pageNumber="41">
|
||
Coloration of the
|
||
<typeStatus id="DF178860FFB4FFC7FECAEC21FE2DFA9B" box="[346,440,1323,1346]" pageId="8" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="98972A47FFB4FFC7FE4BEC26FDD7FA9B" box="[475,578,1323,1346]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[142,170,1536,1556]" captionStartId-1="5.[103,131,1898,1918]" captionStartId-2="7.[103,131,1804,1824]" captionTargetBox-0="[142,812,178,1506]" captionTargetBox-1="[269,1325,178,1868]" captionTargetBox-2="[204,1388,178,1775]" captionTargetId-0="figure-683@4.[142,812,178,1507]" captionTargetId-1="figure-1@5.[268,1325,178,1869]" captionTargetId-2="figure-1@7.[204,1389,178,1775]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIG. 2. Abronia zongolica, new species, head of holotype (MZFC-HE 35664, head width 25.2 mm) in dorsal view (top), right lateral view (middle; image intentionally mirrored horizontally) and ventral view (bottom), in preservative. Scale bars represent 10 mm." captionText-1="FIG. 3. Abronia zongolica, new species, color variation in life (photos not to scale). (A) Adult male holotype (MZFC-HE 35664, snout–vent length [SVL] 112 mm); (B) adult male paratype (MZFC-HE 35665, SVL 95.8 mm); (C) adult female paratype (MZFZ 4406, SVL 86.6 mm; image intentionally mirrored horizontally); (D) neonate paratype (MZFZ 4407, SVL 45.6 mm). Photos by Miguel Ángel de la Torre-Loranca and Antonio Esaú Valdenegro- Brito." captionText-2="FIG. 4. Head comparison of adult Abronia zongolica and Abronia graminea, illustrating differences in (1) circumorbital area color, (2) upper and lower jaw color pattern, and (3) presence/absence of well-developed oblique row of enlarged lateral neck scales. Locality and voucher information for each image is as follows, listed from top-to-bottom. Abronia zongolica: Teopantzacoalco (paratype, MZFC-HE 35665), Ayahuatulco (holotype, MZFC-HE 35664), Huapango (referred specimen, MZFZ-IMG 310), Atiopa (paratype, MZFZ 4406), Huapango (referred specimen, MZFZ-IMG 311). Abronia graminea: El Sumidero (MZFC-HE 32991), Puerto del Aire (MZFZ-IMG 322), Acultzinapa (MZFZ-IMG 336), Puerto del Aire (Clause et al., 2016: fig. 2), Puerto del Aire (MZFZ-IMG 338). All photos taken within 1 month of capture. From top, first and third images of A. zongolica and fifth image of A. graminea intentionally mirrored horizontally. Photos by Miguel Ángel de la Torre-Loranca, Adam G. Clause, and Antonio Esaú Valdenegro- Brito." figureDoi-0="http://doi.org/10.5281/zenodo.7846899" figureDoi-1="http://doi.org/10.5281/zenodo.7846901" figureDoi-2="http://doi.org/10.5281/zenodo.7846903" httpUri-0="https://zenodo.org/record/7846899/files/figure.png" httpUri-1="https://zenodo.org/record/7846901/files/figure.png" httpUri-2="https://zenodo.org/record/7846903/files/figure.png" pageId="8" pageNumber="41">Figs. 2–4</figureCitation>
|
||
) In life, body gray-brown, becoming bright yellow with scattered dark speckles on paravertebral scale rows and flanks. Body with distinct but narrow and uneven black chevron-shaped crossbands, often interrupted on flanks. Many vertebral and paravertebral dorsal scales with bright yellow posterior blotch, especially at midbody. Scales in and along ventrolateral fold bright to pale yellow with scattered dark speckles, some scales with black wedge-shaped markings at anterior margin. Lateral neck scales predominantly bright to pale yellow, but some entirely black scales consolidate into one or two oblique, oblong markings. Interstitial skin between lateral neck scales often dark gray and prominent. Partial black bar borders the dorsal-most lateral neck scales. Dorsal surface of forelimbs and hindlimbs mostly yellow, but many scales flecked with medium gray-brown, sometimes heavily, especially on the hindlimbs. Digits predominantly gray-brown with yellow flecks. Head predominantly cream-colored with yellow highlights and some gray-tinged scale margins, nearly all scales moderately to faintly rugose; rugosity accentuated by heavy black vermiculations, which extend onto all nuchal scales and across 2–3 transverse dorsal scale rows posterior to the forelimb insertion. Scales between orbit and rostral on sides of head cream to pale yellow with some darker marbling; rugosity and vermiculations absent. Tail medium gray-brown with yellow anterior margins or blotches on some lateral scales, particularly anterior third of tail. Tail with roughly 22 indistinct dark brown to black crossbands that do not or only faintly extend to venter, many reduced to a single dorsal blotch. Lower jaw, chin, and throat pale yellow, many scales flecked or blotched with dark gray, sometimes heavily. Venter of body and limbs bright yellow, some scales with dark gray flecks. Iris pale gray-green.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB4FFC7FCE3E8B7FA31FD91" blockId="8.[859,1530,179,584]" pageId="8" pageNumber="41">In preservative (ethanol after formalin), overall color pattern unchanged except all yellow dramatically faded to various shades of cream, and dorsum of back and tail with slight green tinge. Manus and pes cream-colored with faint rusty tinge. Subdigital lamellae medium gray-brown.</paragraph>
|
||
<paragraph id="001336C2FFB4FFC7FCCBEB62FB54FB8C" blockId="8.[859,1531,616,2024]" pageId="8" pageNumber="41">
|
||
<emphasis id="32D8EAD0FFB4FFC7FCCBEB62FBEEFDA6" bold="true" box="[859,1147,616,639]" italics="true" pageId="8" pageNumber="41">Morphological variation.—</emphasis>
|
||
(
|
||
<figureCitation id="98972A47FFB4FFC7FB12EB62FB76FDA6" box="[1154,1251,616,639]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="5.[103,131,1898,1918]" captionStartId-1="7.[103,131,1804,1824]" captionTargetBox-0="[269,1325,178,1868]" captionTargetBox-1="[204,1388,178,1775]" captionTargetId-0="figure-1@5.[268,1325,178,1869]" captionTargetId-1="figure-1@7.[204,1389,178,1775]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIG. 3. Abronia zongolica, new species, color variation in life (photos not to scale). (A) Adult male holotype (MZFC-HE 35664, snout–vent length [SVL] 112 mm); (B) adult male paratype (MZFC-HE 35665, SVL 95.8 mm); (C) adult female paratype (MZFZ 4406, SVL 86.6 mm; image intentionally mirrored horizontally); (D) neonate paratype (MZFZ 4407, SVL 45.6 mm). Photos by Miguel Ángel de la Torre-Loranca and Antonio Esaú Valdenegro- Brito." captionText-1="FIG. 4. Head comparison of adult Abronia zongolica and Abronia graminea, illustrating differences in (1) circumorbital area color, (2) upper and lower jaw color pattern, and (3) presence/absence of well-developed oblique row of enlarged lateral neck scales. Locality and voucher information for each image is as follows, listed from top-to-bottom. Abronia zongolica: Teopantzacoalco (paratype, MZFC-HE 35665), Ayahuatulco (holotype, MZFC-HE 35664), Huapango (referred specimen, MZFZ-IMG 310), Atiopa (paratype, MZFZ 4406), Huapango (referred specimen, MZFZ-IMG 311). Abronia graminea: El Sumidero (MZFC-HE 32991), Puerto del Aire (MZFZ-IMG 322), Acultzinapa (MZFZ-IMG 336), Puerto del Aire (Clause et al., 2016: fig. 2), Puerto del Aire (MZFZ-IMG 338). All photos taken within 1 month of capture. From top, first and third images of A. zongolica and fifth image of A. graminea intentionally mirrored horizontally. Photos by Miguel Ángel de la Torre-Loranca, Adam G. Clause, and Antonio Esaú Valdenegro- Brito." figureDoi-0="http://doi.org/10.5281/zenodo.7846901" figureDoi-1="http://doi.org/10.5281/zenodo.7846903" httpUri-0="https://zenodo.org/record/7846901/files/figure.png" httpUri-1="https://zenodo.org/record/7846903/files/figure.png" pageId="8" pageNumber="41">Figs. 3–4</figureCitation>
|
||
) All
|
||
<specimenCount id="16AAFD4BFFB4FFC7FA88EB62FA32FDA4" box="[1304,1447,616,638]" pageId="8" pageNumber="41" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
|
||
similar to the
|
||
<typeStatus id="DF178860FFB4FFC7FC33EB8FFB93FD42" box="[931,1030,645,667]" pageId="8" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
in most respects. Two adult males (
|
||
<specimenCode id="500A9EB9FFB4FFC7FA1CEB8CFC37FD60" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35665</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FC3DEBA9FBBFFD63" box="[941,1066,675,698]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4408</specimenCode>
|
||
) with
|
||
<specimenCode id="500A9EB9FFB4FFC7FBE0EBA9FB45FD60" box="[1136,1232,675,697]" collectionCode="SVL" pageId="8" pageNumber="41">SVL 95.8</specimenCode>
|
||
and
|
||
<quantity id="C7549B27FFB4FFC7FA96EBA9FAFEFD61" box="[1286,1387,675,696]" metricMagnitude="-2" metricUnit="m" metricValue="9.77" pageId="8" pageNumber="41" unit="mm" value="97.7">97.7 mm</quantity>
|
||
, head length from rostral to upper anterior edge of auricular opening 20.5 and
|
||
<quantity id="C7549B27FFB4FFC7FC1FEBD4FC6DFD2A" box="[911,1016,734,755]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="8" pageNumber="41" unit="mm" value="22.4">22.4 mm</quantity>
|
||
, head width at broadest point 16.3 and
|
||
<quantity id="C7549B27FFB4FFC7FA59EBD4FC1FFCC8" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="8" pageNumber="41" unit="mm" value="16.6">16.6 mm</quantity>
|
||
, respectively, tail broken in
|
||
<specimenCode id="500A9EB9FFB4FFC7FB2EEBF1FAE1FCC8" box="[1214,1396,763,785]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35665</specimenCode>
|
||
and broken and regenerated in
|
||
<specimenCode id="500A9EB9FFB4FFC7FBD3EA13FB5CFCF7" box="[1091,1225,793,814]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4408</specimenCode>
|
||
. One adult female (
|
||
<specimenCode id="500A9EB9FFB4FFC7FA28EA13FC06FC94" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
) with SVL
|
||
<quantity id="C7549B27FFB4FFC7FB9DEA3CFBE6FC95" box="[1037,1139,822,844]" metricMagnitude="-2" metricUnit="m" metricValue="8.66" pageId="8" pageNumber="41" unit="mm" value="86.6">86.6 mm</quantity>
|
||
, head length
|
||
<quantity id="C7549B27FFB4FFC7FA9AEA3CFAE5FC95" box="[1290,1392,822,844]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="8" pageNumber="41" unit="mm" value="19.1">19.1 mm</quantity>
|
||
, head width
|
||
<quantity id="C7549B27FFB4FFC7FCCBEA5EFC50FCB0" box="[859,965,852,873]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="8" pageNumber="41" unit="mm" value="14.4">14.4 mm</quantity>
|
||
, tail unbroken and unregenerated, TL
|
||
<quantity id="C7549B27FFB4FFC7FA14EA5EFA6CFCB0" box="[1412,1529,852,873]" metricMagnitude="-1" metricUnit="m" metricValue="1.274" pageId="8" pageNumber="41" unit="mm" value="127.4">127.4 mm</quantity>
|
||
(1.47 times SVL). One subadult female (
|
||
<specimenCode id="500A9EB9FFB4FFC7FAA3EA7BFA64FC51" box="[1331,1521,881,904]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35663</specimenCode>
|
||
) with SVL
|
||
<quantity id="C7549B27FFB4FFC7FC5AEA85FBA7FC7D" box="[970,1074,911,932]" metricMagnitude="-2" metricUnit="m" metricValue="8.11" pageId="8" pageNumber="41" unit="mm" value="81.1">81.1 mm</quantity>
|
||
, head length
|
||
<quantity id="C7549B27FFB4FFC7FB5DEA85FAA1FC7D" box="[1229,1332,911,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="8" pageNumber="41" unit="mm" value="17.6">17.6 mm</quantity>
|
||
, head width
|
||
<quantity id="C7549B27FFB4FFC7FA59EA85FC1FFC1B" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="41" unit="mm" value="13.1">13.1 mm</quantity>
|
||
, tail broken and regenerated, TL
|
||
<quantity id="C7549B27FFB4FFC7FA98EAA6FAE8FC1B" box="[1288,1405,940,962]" metricMagnitude="-1" metricUnit="m" metricValue="1.042" pageId="8" pageNumber="41" unit="mm" value="104.2">104.2 mm</quantity>
|
||
(including
|
||
<quantity id="C7549B27FFB4FFC7FCCBEAC0FC5FFC06" box="[859,970,970,992]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="8" pageNumber="41" unit="mm" value="26.7">26.7 mm</quantity>
|
||
regenerated tissue). Two neonates (
|
||
<specimenCode id="500A9EB9FFB4FFC7FA1AEAC0FC37FC24" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FC25EAE2FBAEFC27" box="[949,1083,999,1022]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
) with
|
||
<specimenCode id="500A9EB9FFB4FFC7FB02EAEDFB69FC25" box="[1170,1276,999,1020]" collectionCode="SVL" pageId="8" pageNumber="41">SVL 44.2</specimenCode>
|
||
and
|
||
<quantity id="C7549B27FFB4FFC7FAD3EAEDFA25FC24" box="[1347,1456,999,1021]" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="8" pageNumber="41" unit="mm" value="45.6">45.6 mm</quantity>
|
||
, head length 10.5 and
|
||
<quantity id="C7549B27FFB4FFC7FBB3ED0FFB1AFBC3" box="[1059,1167,1029,1050]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="41" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, head width 7.1 and
|
||
<quantity id="C7549B27FFB4FFC7FA06ED0FFA61FBC3" box="[1430,1524,1029,1051]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="41" unit="mm" value="7.5">7.5 mm</quantity>
|
||
, respectively, tail broken in
|
||
<specimenCode id="500A9EB9FFB4FFC7FB32ED29FABCFBEE" box="[1186,1321,1058,1080]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
, and broken and regenerated in
|
||
<specimenCode id="500A9EB9FFB4FFC7FB92ED4AFB28FB8C" box="[1026,1213,1087,1109]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB4FFC7FCE3ED57FC18F831" blockId="8.[859,1531,616,2024]" pageId="8" pageNumber="41">
|
||
The
|
||
<specimenCount id="16AAFD4BFFB4FFC7FC3FED57FBD8FBAA" box="[943,1101,1117,1139]" pageId="8" pageNumber="41" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
|
||
closely resemble the
|
||
<typeStatus id="DF178860FFB4FFC7FAFAED57FA44FBAA" box="[1386,1489,1117,1139]" pageId="8" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
in scalation, with exceptions as follows. Supranasal markedly expanded medially on left side and contacting right posterior internasal in
|
||
<specimenCode id="500A9EB9FFB4FFC7FC67EDBCFB27FB15" box="[1015,1202,1206,1228]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
and
|
||
<specimenCode id="500A9EB9FFB4FFC7FB64EDBCFAE2FB12" box="[1268,1399,1206,1227]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
; postnasals subequal in size in
|
||
<specimenCode id="500A9EB9FFB4FFC7FBA4EDD9FB7BFB30" box="[1076,1262,1235,1257]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35665</specimenCode>
|
||
; postnasal on right side absent in
|
||
<specimenCode id="500A9EB9FFB4FFC7FC42EDFBFBC3FADE" box="[978,1110,1265,1287]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
; prefrontals in narrow contact with each other in
|
||
<specimenCode id="500A9EB9FFB4FFC7FB81EC04FB46FAFD" box="[1041,1235,1294,1316]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
; cantholoreal in narrow contact with supralabials on right side and separated from supralabials on left side by expanded preocular in MZFZ 4408; superciliaries 6/
|
||
<quantity id="C7549B27FFB4FFC7FBC7EC6DFB1DFAA4" box="[1111,1160,1383,1405]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="41" unit="in" value="4.0">4 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FB05EC6DFAC1FAA5" box="[1173,1364,1382,1404]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FAF5EC6CFAE7FAA5" box="[1381,1394,1382,1404]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">5</specimenCode>
|
||
/
|
||
<quantity id="C7549B27FFB4FFC7FAECEC6CFA38FAA4" box="[1404,1453,1382,1405]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="41" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FA2AEC6DFC01FA40" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4408</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FC35EC8EFC26FA43" box="[933,947,1412,1434]" collectionCode="MZFZ" pageId="8" pageNumber="41">6</specimenCode>
|
||
/
|
||
<quantity id="C7549B27FFB4FFC7FC2CEC8EFC79FA43" box="[956,1004,1412,1434]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="41" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FC68EC8FFBE9FA40" box="[1016,1148,1412,1434]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
, and 6/6 on all other
|
||
<typeStatus id="DF178860FFB4FFC7FA19EC8FFA60FA43" box="[1417,1525,1413,1434]" pageId="8" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
; anteriormost superciliary markedly elongate on right side on
|
||
<specimenCode id="500A9EB9FFB4FFC7FCCBECCAFC48FA0D" box="[859,989,1471,1493]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4408</specimenCode>
|
||
, due to aberrant fusion with adjacent superciliary; postoculars 2–4 on left side and 3–4 on right side across all
|
||
<typeStatus id="DF178860FFB4FFC7FC16ECF0FC65F9D6" box="[902,1008,1530,1551]" pageId="8" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
(mean
|
||
<emphasis id="32D8EAD0FFB4FFC7FBC5ECF3FBF2F9C9" bold="true" box="[1109,1127,1529,1552]" pageId="8" pageNumber="41">¼</emphasis>
|
||
3.1); frontal–interparietal contact absent in
|
||
<specimenCode id="500A9EB9FFB4FFC7FC5AEF12FBDCF9F4" box="[970,1097,1560,1581]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
, narrow contact in all other
|
||
<typeStatus id="DF178860FFB4FFC7FA19EF12FA60F9F4" box="[1417,1525,1560,1581]" pageId="8" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
; parietal–median supraocular contact broad in
|
||
<specimenCode id="500A9EB9FFB4FFC7FAE0EF3FFA61F992" box="[1392,1524,1589,1611]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
, narrow in
|
||
<specimenCode id="500A9EB9FFB4FFC7FC5DEF59FB11F9B1" box="[973,1156,1618,1640]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35663</specimenCode>
|
||
, absent in all other
|
||
<typeStatus id="DF178860FFB4FFC7FACAEF59FA53F9B1" box="[1370,1478,1619,1640]" pageId="8" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
; one large occipital entire and not aberrantly divided in all
|
||
<typeStatus id="DF178860FFB4FFC7FCCBEF84FC51F97A" box="[859,964,1678,1699]" pageId="8" pageNumber="41" type="paratype">paratypes</typeStatus>
|
||
except
|
||
<specimenCode id="500A9EB9FFB4FFC7FB8BEF84FB40F97A" box="[1051,1237,1677,1699]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35663</specimenCode>
|
||
, in which a small triangular scale separates the occipital from the interparietal (same as
|
||
<typeStatus id="DF178860FFB4FFC7FCCBEFC2FC54F906" box="[859,961,1736,1759]" pageId="8" pageNumber="41" type="holotype">holotype</typeStatus>
|
||
); two transverse rows of scales separating occipital from first transverse row of nuchals in
|
||
<specimenCode id="500A9EB9FFB4FFC7FAA5EFECFA60F925" box="[1333,1525,1766,1788]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35665</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FCCBEE0EFC75F8C0" box="[859,992,1795,1817]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
, and
|
||
<specimenCode id="500A9EB9FFB4FFC7FBB7EE09FB73F8C0" box="[1063,1254,1795,1817]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35662</specimenCode>
|
||
; primary temporals 4/2 with uppermost on left side very small in
|
||
<specimenCode id="500A9EB9FFB4FFC7FAB0EE2BFA4DF8EE" box="[1312,1496,1825,1847]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35663</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FA72EE2BFA65F8EF" box="[1506,1520,1825,1846]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">4</specimenCode>
|
||
/ 3 with medial temporals presumably fused on right side in
|
||
<specimenCode id="500A9EB9FFB4FFC7FCCBEE56FC75F8A8" box="[859,992,1884,1905]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FC61EE56FC6BF8A8" box="[1009,1022,1884,1905]" collectionCode="MZFZ" pageId="8" pageNumber="41">3</specimenCode>
|
||
/
|
||
<quantity id="C7549B27FFB4FFC7FB97EE56FBACF8AB" box="[1031,1081,1884,1906]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="41" unit="in" value="2.0">2 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FBD5EE56FB5FF8AB" box="[1093,1226,1884,1906]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
; tertiary temporals 4/
|
||
<quantity id="C7549B27FFB4FFC7FA59EE56FA6CF8AB" box="[1481,1529,1884,1906]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="41" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FCCBEE70FC71F856" box="[859,996,1913,1935]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4406</specimenCode>
|
||
and
|
||
<specimenCode id="500A9EB9FFB4FFC7FBA0EE70FB29F857" box="[1072,1212,1913,1935]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB4FFC7FB40EE73FB4BF856" box="[1232,1246,1913,1935]" collectionCode="MZFZ" pageId="8" pageNumber="41">5</specimenCode>
|
||
/
|
||
<quantity id="C7549B27FFB4FFC7FB78EE73FA88F856" box="[1256,1309,1913,1935]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="41" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="500A9EB9FFB4FFC7FABEEE73FA60F856" box="[1326,1525,1913,1935]" collectionCode="MZFC-HE" pageId="8" pageNumber="41">MZFC-HE 35665</specimenCode>
|
||
; supralabials 10–12 (mean
|
||
<emphasis id="32D8EAD0FFB4FFC7FBEDEE9FFB1AF875" bold="true" box="[1149,1167,1941,1964]" pageId="8" pageNumber="41">¼</emphasis>
|
||
10.8); infralabials 9–11 (mean
|
||
<emphasis id="32D8EAD0FFB4FFC7FA79EE9FFA6EF875" bold="true" box="[1513,1531,1941,1964]" pageId="8" pageNumber="41">¼</emphasis>
|
||
9.1); postmental single in
|
||
<specimenCode id="500A9EB9FFB4FFC7FBE8EEBFFB63F810" box="[1144,1270,1972,1994]" collectionCode="MZFZ" pageId="8" pageNumber="41">MZFZ 4407</specimenCode>
|
||
; sublabials 4–5 (mean
|
||
<emphasis id="32D8EAD0FFB4FFC7FA79EEB9FA6EF813" bold="true" box="[1513,1531,1971,1994]" pageId="8" pageNumber="41">¼</emphasis>
|
||
4.5).
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FFEFE9B9FDD0FE4E" blockId="9.[103,773,177,407]" pageId="9" pageNumber="42">
|
||
Transverse dorsal scale rows 30–34 (mean
|
||
<emphasis id="32D8EAD0FFB5FFC6FDF2E9BBFDE1FF11" bold="true" box="[610,628,177,200]" pageId="9" pageNumber="42">¼</emphasis>
|
||
32.3); transverse ventral scale rows
|
||
<quantity id="C7549B27FFB5FFC6FE11E9DAFE2AFF3F" box="[385,447,208,230]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="9" pageNumber="42" unit="in" value="32.0">32 in</quantity>
|
||
MZFC-HE 35663 and MZFZ 4406; lateralmost row of ventral scales weakly enlarged overall in MZFC-HE 35662; osteoderms appear weakly developed in MZFC-HE 35662 and MZFZ 4407; lateral neck scales as few as
|
||
<quantity id="C7549B27FFB5FFC6FE86E84CFED6FE85" box="[278,323,326,348]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="42" unit="in" value="5.0">5 in</quantity>
|
||
MZFC-HE 35663; subdigital lamellae on fourth toes 17–22 (mean
|
||
<emphasis id="32D8EAD0FFB5FFC6FEEFE868FE04FEA0" bold="true" box="[383,401,354,377]" pageId="9" pageNumber="42">¼</emphasis>
|
||
19.8), excluding the counts from MZFZ 4408 which is missing both toe tips.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FFF7E8BCFEF5FAFD" blockId="9.[103,773,438,2024]" pageId="9" pageNumber="42">
|
||
<emphasis id="32D8EAD0FFB5FFC6FFF7E8BCFEECFE14" bold="true" box="[103,377,438,461]" italics="true" pageId="9" pageNumber="42">Color pattern variation.—</emphasis>
|
||
In life, the
|
||
<specimenCount id="16AAFD4BFFB5FFC6FE61E8BDFDCCFE14" box="[497,601,439,461]" pageId="9" pageNumber="42" type="adult">two adult</specimenCount>
|
||
male
|
||
<typeStatus id="DF178860FFB5FFC6FD0BE8BDFC91FE15" box="[667,772,439,460]" pageId="9" pageNumber="42" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<specimenCode id="500A9EB9FFB5FFC6FFFDE8DEFF7BFE30" box="[109,238,468,489]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
and
|
||
<specimenCode id="500A9EB9FFB5FFC6FEBAE8DEFE77FE32" box="[298,482,468,491]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
) differ from the
|
||
<typeStatus id="DF178860FFB5FFC6FD32E8DEFC90FE33" box="[674,773,468,490]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
as follows: dorsum medium gray; black chevron-shaped crossbands 0.5–2 scales wide, often extend to ventrolateral fold in
|
||
<specimenCode id="500A9EB9FFB5FFC6FF2DEB27FEABFD9B" box="[189,318,557,578]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
, reduced to narrow interrupted paravertebral dashes in
|
||
<specimenCode id="500A9EB9FFB5FFC6FE8FEB40FE4FFDB9" box="[287,474,586,608]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; many scales in and along ventrolateral fold with black wedge-shaped anterior marking in
|
||
<specimenCode id="500A9EB9FFB5FFC6FF1DEB8CFE86FD43" box="[141,275,645,667]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
; lateral neck scales mostly pale cream to whitish; partial black bar bordering dorsal-most lateral neck scales broken and indistinct in
|
||
<specimenCode id="500A9EB9FFB5FFC6FE59EBCAFD10FD0F" box="[457,645,704,726]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; dorsum of head with indistinct, arrowhead-shaped dark marking centered at occipital scale in
|
||
<specimenCode id="500A9EB9FFB5FFC6FE3AEBF6FDA7FCC9" box="[426,562,763,785]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
; dorsum of head medium gray with cream highlights in
|
||
<specimenCode id="500A9EB9FFB5FFC6FDA0EA13FD26FCF7" box="[560,691,793,814]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
, cream with light gray margins on some scales in
|
||
<specimenCode id="500A9EB9FFB5FFC6FDD6EA3CFC95FC95" box="[582,768,822,844]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; black vermiculations extend across all nuchal scales and onto 3–4 transverse dorsal scale rows posterior to forelimb insertion in
|
||
<specimenCode id="500A9EB9FFB5FFC6FF64EA85FEE3FC7D" box="[244,374,911,932]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
; sides of head from rostral to orbit medium gray with pale cream marbling in
|
||
<specimenCode id="500A9EB9FFB5FFC6FDE9EAA7FC95FC18" box="[633,768,940,962]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
, uniform pale gray in
|
||
<specimenCode id="500A9EB9FFB5FFC6FEFEEAC0FDBAFC06" box="[366,559,969,991]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; supralabial scales posterior to orbit, and lowermost temporal scales, medium-gray with cream marbling and light to moderate black vermiculations in
|
||
<specimenCode id="500A9EB9FFB5FFC6FEACED28FE54FBEE" box="[316,449,1058,1079]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
, these scales cream to pale gray with vermiculations only on temporals in
|
||
<specimenCode id="500A9EB9FFB5FFC6FD09ED4AFF3BFBAA" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; anterior to autotomy point, tail with roughly 13 indistinct dark gray-brown crossbands in
|
||
<specimenCode id="500A9EB9FFB5FFC6FDCBED71FD76FB49" box="[603,739,1147,1168]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
,
|
||
<specimenCode id="500A9EB9FFB5FFC6FD67ED71FC90FB49" box="[759,773,1147,1168]" collectionCode="MZFZ" pageId="9" pageNumber="42">3</specimenCode>
|
||
heavily obscured dark gray crossbands in
|
||
<specimenCode id="500A9EB9FFB5FFC6FDD4ED92FC95FB77" box="[580,768,1176,1198]" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; ventral scales on head and body with hairline dark margins in
|
||
<specimenCode id="500A9EB9FFB5FFC6FF15EDD9FE91FB31" box="[133,260,1235,1256]" collectionCode="MZFZ" pageId="9" pageNumber="42">MZFZ 4408</specimenCode>
|
||
, only some scales with dark margins in
|
||
<specimenCode id="500A9EB9FFB5FFC6FD2AEDD9FF4DFADF" collectionCode="MZFC-HE" pageId="9" pageNumber="42">MZFC-HE 35665</specimenCode>
|
||
; venter of head, body, and tail, and limbs mostly pale cream to whitish.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FFEFEC26FB2BFEFB" blockId="9.[103,773,438,2024]" lastBlockId="9.[820,1491,179,2024]" pageId="9" pageNumber="42">
|
||
In life, the adult female
|
||
<typeStatus id="DF178860FFB5FFC6FE19EC26FE7DFA98" box="[393,488,1324,1345]" pageId="9" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
(MZFZ 4406) differs from the
|
||
<typeStatus id="DF178860FFB5FFC6FF0AEC43FF68FA86" box="[154,253,1353,1375]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
as follows: body medium brown-gray with strong green tinge, becoming pale gray on paravertebral scale rows and flanks; chevron-shaped crossbands dark gray, 1–2 scales wide, often offset and partially fused along dorsal midline, terminate on flanks; posterolateral margins of some dark crossbands cream or pale yellow; scales in and along ventrolateral fold pale gray or cream, some with dark speckling; lateral neck scales mostly pale cream, lack adjacent dark dorsal bar; dorsum of all limbs pale gray with green tinge, hindlimbs blotched with medium gray-brown; digits tan to brown with green tinge; dorsum of head pale yellow with green tinge, most scales faintly rugose; rugosity accentuated by faint to moderate dark gray vermiculations, which extend onto most vertebral and paravertebral nuchal scales; sides of head from rostral to orbit to lowermost secondary temporal scale pale yellow with green tinge, with hairline black margins on many scales and some dark marks; temporal scales with moderate to weak black vermiculations, absent on supralabial scales; tail medium gray-brown becoming pale gray on sides with green tinge throughout; tail with roughly 24 broken, often offset crossbands, none reduced to a single dorsal blotch; lower jaw, chin, and throat cream to pale yellow, lack green tinge, lateralmost scales often with black to dark gray margins; venter of body, limbs, and tail pale yellow to cream, becoming pale gray laterally; dark crossbands on tail weakly visible on venter, most prominent on posterior half of tail.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FCDCE821FABDFBF2" blockId="9.[820,1491,179,2024]" pageId="9" pageNumber="42">
|
||
In preservative, the subadult female
|
||
<typeStatus id="DF178860FFB5FFC6FB62E821FAC6FE99" box="[1266,1363,299,320]" pageId="9" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
(MZFC-HE 35663) differs from the
|
||
<typeStatus id="DF178860FFB5FFC6FBD2E843FB32FE86" box="[1090,1191,329,351]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
as follows: body medium-brown, becoming slightly rusty brown on flanks; black chevron-shaped crossbands 0.5–1 scales wide, some broken at dorsal midline, usually with pale yellow or cream blotch or dash at posterolateral margin; scales in and along ventrolateral fold rusty brown with pale yellow to cream highlights; lateral neck scales mostly pale brown or cream, lack adjacent dark dorsal bar; interstitial skin on lateral neck inconspicuous, medium-gray, not forming oblique dark markings; limbs rusty brown, becoming dark gray posterolaterally; dorsum of head medium-brown with some gray highlights; black vermiculations weak on lateralmost nuchal scales, terminate at forelimb insertion; sides of head from rostral to orbit to lowermost secondary temporal scale cream with brown tinge, many scales with hairline dark gray margins; temporal scales with weak black vermiculations, absent on supralabial scales; tail medium-brown with rusty and gray highlights; anterior to autotomy point, tail with roughly 13 broken, often offset dark gray to black crossbands, none reduced to a single dorsal blotch, some with pale yellow to cream spot at posterior or posterolateral margin; regenerated portion of tail uniform medium-brown; lower jaw, chin, and throat pale cream to very pale brown with dark flecking; venter cream to pale gray or pale brown, often with moderate to heavy dark flecking.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FCDCED3EFABFF8CC" blockId="9.[820,1491,179,2024]" pageId="9" pageNumber="42">
|
||
In preservative, the two neonate
|
||
<typeStatus id="DF178860FFB5FFC6FB43ED3EFAA9FB90" box="[1235,1340,1076,1097]" pageId="9" pageNumber="42" type="paratype">paratypes</typeStatus>
|
||
(MZFZ 4407 and MZFC-HE 35662) differ from the
|
||
<typeStatus id="DF178860FFB5FFC6FB7DED5BFAC5FBBE" box="[1261,1360,1105,1127]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
as follows: body pale gray to medium brown-gray lacking green tinge; dorsal crossbands black to dark gray, partly bordered posteriorly by pale gray to white; vertebral and paravertebral dorsal scales lack pale posterior blotches; lateral neck scales mostly pale gray; dorsum of all limbs pale gray to medium brown-gray, moderately to heavily flecked or blotched with black or dark gray; fingers and toes dark gray to medium brown-gray; dorsum of head same color as body, scales flat, most scales with hairline black margins, few scales rugose and only faintly so; rugosity accentuated with limited black vermiculations, not extending onto nuchal scales or transverse dorsal scales; sides of head from rostral to orbit to lowermost secondary temporal scale pale gray to pale brown-gray with hairline black margins on most scales, otherwise unmarked; tail same color as body with black to dark gray crossbands; regenerated portion of tail in MZFC-HE 35662 uniform dark gray; lower jaw, chin, and throat pale gray lacking green tinge; cream tinge on anterior half of jaw in MZFC-HE 35662; venter of body and limbs pale gray, nearly all scales moderately to heavily flecked with dark gray; venter of tail same as body except for uniform dark gray regenerated portion in MZFC-HE 35662; venter of tail mostly brown-gray with heavy dark gray flecking in MZFZ 4407.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB5FFC6FCDCEE15FB17F831" blockId="9.[820,1491,179,2024]" pageId="9" pageNumber="42">
|
||
In preservative (ethanol after formalin), all other
|
||
<typeStatus id="DF178860FFB5FFC6FAF9EE15FA47F8ED" box="[1385,1490,1823,1844]" pageId="9" pageNumber="42" type="paratype">paratypes</typeStatus>
|
||
maintain their general color pattern as described above, but like the
|
||
<typeStatus id="DF178860FFB5FFC6FC1CEE50FC7AF8A9" box="[908,1007,1882,1904]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
all yellow or cream is faded to pale cream or whitish, and brown areas are more gray and occasionally have a green tinge. Also, as in the
|
||
<typeStatus id="DF178860FFB5FFC6FB28EE9CFA88F875" box="[1208,1309,1942,1964]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
, manus and pes cream or whitish with faint rusty tinge, and subdigital lamellae medium gray-brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="54D3664AFFB6FFC5FF1EEBC3FCB9FCF4" ID-DOI="http://doi.org/10.5281/zenodo.7846905" ID-Zenodo-Dep="7846905" httpUri="https://zenodo.org/record/7846905/files/figure.png" pageId="10" pageNumber="43" startId="10.[142,170,713,733]" targetBox="[142,812,178,684]" targetPageId="10" targetType="figure">
|
||
<paragraph id="001336C2FFB6FFC5FF1EEBC3FCB9FCF4" blockId="10.[142,812,713,813]" pageId="10" pageNumber="43">
|
||
FIG. 5. Geographic distribution of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FE75EBC3FD01FD04" box="[485,660,713,733]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FE75EBC3FD01FD04" bold="true" box="[485,660,713,733]" italics="true" pageId="10" pageNumber="43">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
and proximate populations of the only known sympatric congener,
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FD07EBEEFC9AFD21" box="[663,783,740,760]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB6FFC5FD07EBEEFC9AFD21" bold="true" box="[663,783,740,760]" italics="true" pageId="10" pageNumber="43">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
in Mexico. The star represents the type locality of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FDEEEBF4FD62FCCB" box="[638,759,766,786]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FDEEEBF4FD62FCCB" bold="true" box="[638,759,766,786]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
. See Appendix 1 for locality information of mapped records of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FD22EA13FCB3FCF4" box="[690,806,793,813]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB6FFC5FD22EA13FCB3FCF4" bold="true" box="[690,806,793,813]" italics="true" pageId="10" pageNumber="43">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="48B66549FFB6FFC5FF1EEA53FEA8FB9F" pageId="10" pageNumber="43" type="etymology">
|
||
<paragraph id="001336C2FFB6FFC5FF1EEA53FEA8FB9F" blockId="10.[142,812,857,1094]" pageId="10" pageNumber="43">
|
||
<emphasis id="32D8EAD0FFB6FFC5FF1EEA53FE89FCA9" bold="true" box="[142,284,857,880]" italics="true" pageId="10" pageNumber="43">Etymology.—</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FD88EA53FDECFCB6" box="[536,633,857,879]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FD88EA53FDECFCB6" box="[536,633,857,879]" italics="true" pageId="10" pageNumber="43">zongolica</emphasis>
|
||
</taxonomicName>
|
||
is a feminine singular adjective in the nominative case and refers to the Sierra de Zongolica of
|
||
<collectingRegion id="C268F820FFB6FFC5FEEEEA9DFE75FC75" box="[382,480,919,940]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="43">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry id="78BB7652FFB6FFC5FE79EA9DFDA8FC74" box="[489,573,919,941]" name="Mexico" pageId="10" pageNumber="43">Mexico</collectingCountry>
|
||
. This mountain range supports the only confirmed populations of the new species. The name ‘‘zongolica’’ appears to be derived from the words ‘‘tzoncolican’’ or ‘‘tzoncolihucan’’ in the Nahuatl language, which roughly translate as ‘‘where hair is braided’’ (Melgarejo Vivanco, 1950).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="48B66549FFB6FFC4FF1EED6CFDB1FA25" lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43" type="distribution">
|
||
<paragraph id="001336C2FFB6FFC5FF1EED6CFC45FE83" blockId="10.[142,812,1126,1485]" lastBlockId="10.[859,1530,179,1256]" pageId="10" pageNumber="43">
|
||
<emphasis id="32D8EAD0FFB6FFC5FF1EED6CFE21FBA4" bold="true" box="[142,436,1126,1149]" italics="true" pageId="10" pageNumber="43">Distribution and ecology.—</emphasis>
|
||
Based on vouchered material, the distribution of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FEA5ED8CFE26FB42" box="[309,435,1157,1179]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FEA5ED8CFE26FB42" box="[309,435,1157,1179]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
extends across less than
|
||
<quantity id="C7549B27FFB6FFC5FD58ED8FFC98FB42" box="[712,781,1157,1179]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="43" unit="km" value="10.0">10 km</quantity>
|
||
in the Sierra de Zongolica from near Ayahuatulco to Huapango (
|
||
<figureCitation id="98972A47FFB6FFC5FF05EDC8FF42FB00" box="[149,215,1218,1241]" captionStart="FIG" captionStartId="10.[142,170,713,733]" captionTargetBox="[142,812,178,684]" captionTargetId="figure-608@10.[142,812,178,684]" captionTargetPageId="10" captionText="FIG. 5. Geographic distribution of Abronia zongolica and proximate populations of the only known sympatric congener, A. graminea in Mexico. The star represents the type locality of A. zongolica. See Appendix 1 for locality information of mapped records of A. graminea." figureDoi="http://doi.org/10.5281/zenodo.7846905" httpUri="https://zenodo.org/record/7846905/files/figure.png" pageId="10" pageNumber="43">Fig. 5</figureCitation>
|
||
). All confirmed localities lie within the Río Alpatlahuaya drainage, and span an elevational range of
|
||
<quantity id="C7549B27FFB6FFC5FD72EDEBFF60FACC" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="10" pageNumber="43" unit="m" value="1850.0" valueMax="2200.0" valueMin="1500.0">
|
||
<geoCoordinate id="65985005FFB6FFC5FD72EDEBFC88FB2E" box="[738,797,1249,1271]" degrees="1,500" direction="north" orientation="latitude" pageId="10" pageNumber="43" precision="55" value="1.5">1,500</geoCoordinate>
|
||
<geoCoordinate id="65985005FFB6FFC5FC8EEDEBFF5EFACC" degrees="2,200" direction="west" orientation="longitude" pageId="10" pageNumber="43" precision="55" value="-2.2">– 2,200</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
. Suitable elevations above
|
||
<quantity id="C7549B27FFB6FFC5FDD0EDF5FD36FACC" box="[576,675,1279,1301]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="43" unit="m" value="1500.0">1,500 m</quantity>
|
||
and mesic tropical forests are continuous between the known localities, and thus
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FF6FEC37FE14FA8B" box="[255,385,1340,1362]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FF6FEC37FE14FA8B" box="[255,385,1340,1362]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
likely occurs in other regions of the Sierra de Zongolica. We are aware of records of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FD0CEC51FD7BFAA8" box="[668,750,1371,1393]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB6FFC5FD0CEC51FD7BFAA8" box="[668,750,1371,1393]" italics="true" pageId="10" pageNumber="43">Abronia</emphasis>
|
||
</taxonomicName>
|
||
from nearby forested areas of west-central
|
||
<collectingRegion id="C268F820FFB6FFC5FDB0EC70FDEAFA56" box="[544,639,1402,1423]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="43">Veracruz</collectingRegion>
|
||
that also might represent
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FE92EC93FE12FA77" box="[258,391,1432,1454]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FE92EC93FE12FA77" box="[258,391,1432,1454]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
. Pending detailed analysis of these populations and deposition of museum vouchers, we have considered their taxonomic allocation uncertain at this time. Additionally,
|
||
<bibRefCitation id="643D4B33FFB6FFC5FC65E9DAFA87FF3E" author="Guzman Guzman, S." box="[1013,1298,208,231]" pageId="10" pageNumber="43" refId="ref14298" refString="Guzman Guzman, S. 2011. Anfibios y Reptiles de Veracruz: Guia Ilustrada. Consejo Veracruzano de Investigacion Cientifica y Desarrollo Tecnologico, Mexico." type="book" year="2011">Guzmán Guzmán (2011)</bibRefCitation>
|
||
published a photograph from an unspecified locality of what is identified as a female
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FC3BE801FBB2FEF9" box="[939,1063,267,288]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB6FFC5FC3BE801FBB2FEF9" box="[939,1063,267,288]" italics="true" pageId="10" pageNumber="43">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
, but which is actually an adult
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAECE801FA6CFEF9" box="[1404,1529,266,288]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAECE801FA6CFEF9" box="[1404,1529,266,288]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
(likely a male) based on multiple diagnostic features as given in
|
||
<tableCitation id="4D2E0379FFB6FFC5FCEAE84FFC59FE83" box="[890,972,325,347]" captionStart="Table 2" captionStartId="6.[142,190,1262,1282]" captionTargetPageId="6" captionText="Table 2. Character states for selected features of Abronia zongolica and all species in Group III of Gutiérrez-Rodríguez et al. (2021)." httpUri="http://table.plazi.org/id/54D3664AFFBAFFC9FF1EEDE4FAEEFADB" pageId="10" pageNumber="43" tableUuid="54D3664AFFBAFFC9FF1EEDE4FAEEFADB">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB6FFC5FCE3E868FB21FC25" blockId="10.[859,1530,179,1256]" pageId="10" pageNumber="43">
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FCE3E868FBB8FEA1" box="[883,1069,354,376]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FCE3E868FBB8FEA1" box="[883,1069,354,376]" italics="true" pageId="10" pageNumber="43">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
is found in mature pine-oak forest (
|
||
<emphasis id="32D8EAD0FFB6FFC5FA51E869FA6CFEA1" box="[1473,1529,355,376]" italics="true" pageId="10" pageNumber="43">sensu</emphasis>
|
||
<bibRefCitation id="643D4B33FFB6FFC5FCCBE88AFB8AFE4E" author="Rzedowski, J." box="[859,1055,384,407]" pageId="10" pageNumber="43" refId="ref15828" refString="Rzedowski, J. 2006. Vegetacion de Mexico. 1 ra Edicion digital, Comision Nacional para el Conocimiento y Uso de la Biodiversidad, Mexico." type="book" year="2006">Rzedowski, 2006</bibRefCitation>
|
||
), with dominant tree species including
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FCCBE894FB82FE6A" box="[859,1047,414,435]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="conspersa">
|
||
<emphasis id="32D8EAD0FFB6FFC5FCCBE894FB82FE6A" box="[859,1047,414,435]" italics="true" pageId="10" pageNumber="43">Quercus conspersa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FBB5E894FB39FE6A" box="[1061,1196,413,435]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="crassifolia">
|
||
<emphasis id="32D8EAD0FFB6FFC5FBB5E894FB39FE6A" box="[1061,1196,413,435]" italics="true" pageId="10" pageNumber="43">Q. crassifolia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FB2AE894FAB1FE6A" box="[1210,1316,413,435]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="laurina">
|
||
<emphasis id="32D8EAD0FFB6FFC5FB2AE894FAB1FE6A" box="[1210,1316,413,435]" italics="true" pageId="10" pageNumber="43">Q. laurina</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAF4E894FA53FE6A" box="[1380,1478,414,435]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="rugosa">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAF4E894FA53FE6A" box="[1380,1478,414,435]" italics="true" pageId="10" pageNumber="43">Q. rugosa</emphasis>
|
||
</taxonomicName>
|
||
; the arboreal stratum also regularly contains
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAAAE8B0FA67FE09" box="[1338,1522,442,464]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="acuminata">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAAAE8B0FA67FE09" box="[1338,1522,442,464]" italics="true" pageId="10" pageNumber="43">Alnus acuminata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FCCBE8D2FBACFE37" box="[859,1081,472,494]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="benthamii">
|
||
<emphasis id="32D8EAD0FFB6FFC5FCCBE8D2FBACFE37" box="[859,1081,472,494]" italics="true" pageId="10" pageNumber="43">Cupressus benthamii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FBDDE8D2FB2BFE37" box="[1101,1214,472,494]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="lindleyi">
|
||
<emphasis id="32D8EAD0FFB6FFC5FBDDE8D2FB2BFE37" box="[1101,1214,472,494]" italics="true" pageId="10" pageNumber="43">C. lindleyi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FA9AE8D3FA04FE37" box="[1290,1425,472,494]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="patula">
|
||
<emphasis id="32D8EAD0FFB6FFC5FA9AE8D3FA04FE37" box="[1290,1425,472,494]" italics="true" pageId="10" pageNumber="43">Pinus patula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="98972A47FFB6FFC5FA37E8D2FA79FE36" box="[1447,1516,472,495]" captionStart="FIG" captionStartId="10.[1178,1206,1952,1972]" captionTargetBox="[142,1129,1528,2020]" captionTargetId="figure-594@10.[142,1129,1528,2020]" captionTargetPageId="10" captionText="FIG. 6. Habitat of Abronia zongolica at the type locality in Veracruz, Mexico." figureDoi="http://doi.org/10.5281/zenodo.7846907" httpUri="https://zenodo.org/record/7846907/files/figure.png" pageId="10" pageNumber="43">Fig. 6</figureCitation>
|
||
). Epiphytes are typically abundant and primarily consist of bryophytes, but also include the bromeliad
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FA1BEB18FC55FD9C" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="imperialis">
|
||
<emphasis id="32D8EAD0FFB6FFC5FA1BEB18FC55FD9C" italics="true" pageId="10" pageNumber="43">Tillandsia imperialis</emphasis>
|
||
</taxonomicName>
|
||
and fern
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FBB9EB25FAA1FD9C" box="[1065,1332,559,581]" class="Polypodiopsida" family="Dryopteridaceae" genus="Elaphoglossum" kingdom="Plantae" order="Polypodiales" pageId="10" pageNumber="43" phylum="Tracheophyta" rank="species" species="paleaceum">
|
||
<emphasis id="32D8EAD0FFB6FFC5FBB9EB25FAA1FD9C" box="[1065,1332,559,581]" italics="true" pageId="10" pageNumber="43">Elaphoglossum paleaceum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAD1EB25FA6CFD9C" box="[1345,1529,559,581]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAD1EB25FA6CFD9C" box="[1345,1529,559,581]" italics="true" pageId="10" pageNumber="43">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
seems to be both arboreal and diurnal. Most individuals were found hidden behind or within bromeliads on tree trunks or branches, ranging from the root or buttress level up to
|
||
<quantity id="C7549B27FFB6FFC5FA22EB82FA4EFD44" box="[1458,1499,648,669]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="43" unit="m" value="4.0">4 m</quantity>
|
||
in height. Individuals were also directly observed while active by day amongst the branches of trees and shrubs. An analysis of fecal material from two individuals of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAB4EBEAFA37FD2C" box="[1316,1442,735,757]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAB4EBEAFA37FD2C" box="[1316,1442,735,757]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
(MZFZ-IMG 312 and 313) indicates that the species feeds on insects (Orthoptera, Coleoptera, Lepidoptera, and Hemiptera) along with other unidentified small invertebrates. To our knowledge, this is the first published wild dietary data for any arboreal species of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FBA8EA78FB1FFC51" box="[1080,1162,882,904]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB6FFC5FBA8EA78FB1FFC51" box="[1080,1162,882,904]" italics="true" pageId="10" pageNumber="43">Abronia</emphasis>
|
||
</taxonomicName>
|
||
. Like all arboreal congeners for which data exist (
|
||
<bibRefCitation id="643D4B33FFB6FFC5FBB1EA85FAF2FC7F" author="Schmidt-Ballardo, W. & I. Solano-Zavaleta & A. G. Clause" box="[1057,1383,911,934]" pageId="10" pageNumber="43" pagination="192 - 194" refId="ref15984" refString="Schmidt-Ballardo, W., I. Solano-Zavaleta, and A. G. Clause. 2015. Nature notes. Abronia deppii. Reproduction. Mesoamerican Herpetology 2: 192 - 194." type="journal article" year="2015">Schmidt-Ballardo et al., 2015</bibRefCitation>
|
||
),
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAEAEA9AFA6CFC7C" box="[1402,1529,911,933]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAEAEA9AFA6CFC7C" box="[1402,1529,911,933]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
is a viviparous species that gives birth in the spring: a captive female (MZFZ-IMG 309) birthed four neonates in early May that measured
|
||
<quantity id="C7549B27FFB6FFC5FB92EAEDFBECFC24" box="[1026,1145,999,1021]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="10" pageNumber="43" unit="mm" value="31.5" valueMax="33.0" valueMin="30.0">30–33 mm</quantity>
|
||
SVL.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB6FFC5FCE3ED0EFA31FB3E" blockId="10.[859,1530,179,1256]" pageId="10" pageNumber="43">
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FCE3ED0EFBA1FBC3" box="[883,1076,1028,1050]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FCE3ED0EFBA1FBC3" box="[883,1076,1028,1050]" italics="true" pageId="10" pageNumber="43">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
is sympatric with
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FA8CED0FFA37FBC3" box="[1308,1442,1029,1050]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB6FFC5FA8CED0FFA37FBC3" box="[1308,1442,1029,1050]" italics="true" pageId="10" pageNumber="43">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
in the municipalities of Astacinga and
|
||
<collectingRegion id="C268F820FFB6FFC5FB2FED28FAB5FBEE" box="[1215,1312,1058,1079]" country="Nicaragua" name="Rio San Juan" pageId="10" pageNumber="43">San Juan</collectingRegion>
|
||
Texhuacán (
|
||
<figureCitation id="98972A47FFB6FFC5FA3DED28FA7EFBE0" box="[1453,1515,1058,1081]" captionStart="FIG" captionStartId="10.[142,170,713,733]" captionTargetBox="[142,812,178,684]" captionTargetId="figure-608@10.[142,812,178,684]" captionTargetPageId="10" captionText="FIG. 5. Geographic distribution of Abronia zongolica and proximate populations of the only known sympatric congener, A. graminea in Mexico. The star represents the type locality of A. zongolica. See Appendix 1 for locality information of mapped records of A. graminea." figureDoi="http://doi.org/10.5281/zenodo.7846905" httpUri="https://zenodo.org/record/7846905/files/figure.png" pageId="10" pageNumber="43">Fig. 5</figureCitation>
|
||
). Both species occupy similar forest microenvironments, but
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FA72ED4AFC2EFBAB" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="graminea">
|
||
<emphasis id="32D8EAD0FFB6FFC5FA72ED4AFC2EFBAB" italics="true" pageId="10" pageNumber="43">A. graminea</emphasis>
|
||
</taxonomicName>
|
||
is usually more abundant and is also distributed at higher elevations in coniferous forests, where
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAE6ED70FA6CFB56" box="[1398,1529,1145,1167]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAE6ED70FA6CFB56" box="[1398,1529,1145,1167]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
appears to be absent. The only other gerrhonotine anguid lizard species that shares habitat with
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FAB1EDBFFA30FB13" box="[1313,1445,1204,1226]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FAB1EDBFFA30FB13" box="[1313,1445,1204,1226]" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
in the Sierra de Zongolica is the terrestrial
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FB7BEDD8FA08FB3E" baseAuthorityName="Wiegmann" baseAuthorityYear="1828" box="[1259,1437,1233,1255]" class="Reptilia" family="Anguidae" genus="Barisia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="imbricata">
|
||
<emphasis id="32D8EAD0FFB6FFC5FB7BEDD8FA08FB3E" box="[1259,1437,1233,1255]" italics="true" pageId="10" pageNumber="43">Barisia imbricata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="001336C2FFB6FFC4FCCBEC0DFDA6FCEF" blockId="10.[859,1530,1287,1485]" lastBlockId="11.[103,773,179,1532]" lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43">
|
||
<emphasis id="32D8EAD0FFB6FFC5FCCBEC0DFB91FAC7" bold="true" box="[859,1028,1287,1310]" italics="true" pageId="10" pageNumber="43">Conservation.—</emphasis>
|
||
None of the vouchered populations of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FA72EC0DFC2EFAE3" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FA72EC0DFC2EFAE3" italics="true" pageId="10" pageNumber="43">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
occur in designated protected areas. However, the species might eventually be documented in the Parque Nacional Cañón del Río Blanco, the Reserva Ecológica Natural en la Cuenca Alta del Río Atoyac, or even the Reserva del Bicentenario (
|
||
<bibRefCitation id="643D4B33FFB6FFC5FB02EC93FC78FA15" author="Castillo-Hernandez, L. A. & H. Flores-Olvera" pageId="10" pageNumber="43" pagination="539 - 563" refId="ref13287" refString="Castillo-Hernandez, L. A., and H. Flores-Olvera. 2017. Floristic composition of the cloud forest of the Bicentenario Reserve, Zongolica, Veracruz, Mexico. Botanical Science 95: 539 - 563." type="journal article" year="2017">Castillo-Hernández and Flores-Olvera, 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB6FFC5FC6BECBDFB63FA14" author="Perez-Sato, J. A. & N. M. Ceron de la Luz & R. Serna-Lagunes & J. E. Rivera-Hernandez & N. Mora-Collado & J. Salazar-Ortiz" box="[1019,1270,1462,1485]" pageId="10" pageNumber="43" pagination="38 - 44" refId="ref15453" refString="Perez-Sato, J. A., N. M. Ceron de la Luz, R. Serna-Lagunes, J. E. Rivera-Hernandez, N. Mora-Collado, and J. Salazar-Ortiz. 2018. Herpetofauna de tres localidades del Municipio de Amatlan de Los Reyes, Veracruz, Mexico. Agro Productividad 11: 38 - 44." type="journal article" year="2018">Pérez-Sato et al., 2018</bibRefCitation>
|
||
). Regionally, deforestation has caused extensive habitat loss and fragmentation (
|
||
<bibRefCitation id="643D4B33FFB7FFC4FFFEE9DBFE15FF3E" author="Gomez-Diaz, J. A. & K. Brast & J. Degener & T. Kromer & E. Ellis & F. Heitkamp & G. Gerold" box="[110,384,208,231]" pageId="11" pageNumber="44" pagination="1 - 12" refId="ref14100" refString="Gomez-Diaz, J. A., K. Brast, J. Degener, T. Kromer, E. Ellis, F. Heitkamp, and G. Gerold. 2018. Long-term changes in forest cover in central Veracruz, Mexico (1993 - 2014). Tropical Conservation Science 11: 1 - 12." type="journal article" year="2018">Gómez-Díaz et al., 2018</bibRefCitation>
|
||
), including to the surroundings of all vouchered localities for
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FE0DE9E4FD8BFEDD" box="[413,542,238,260]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FE0DE9E4FD8BFEDD" box="[413,542,238,260]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
. Due to agricultural demands and other livelihood requirements for the growing human population in the Sierra de Zongolica, we expect that this deforestation will continue. Climate change is also forecasted to dramatically reduce the area of cloud forests in this region (
|
||
<bibRefCitation id="643D4B33FFB7FFC4FF7BE888FE62FE4E" author="Ponce-Reyes, R. & V. - H. Reynoso-Rosales & J. E. M. Watson & J. VanDerWal & R. A. Fuller & R. L. Pressey & H. H. Possingham" box="[235,503,386,408]" pageId="11" pageNumber="44" pagination="448 - 452" refId="ref15650" refString="Ponce-Reyes, R., V. - H. Reynoso-Rosales, J. E. M. Watson, J. VanDerWal, R. A. Fuller, R. L. Pressey, and H. H. Possingham. 2012. Vulnerability of cloud forest reserves in Mexico to climate change. Nature Climate Change 2: 448 - 452." type="journal article" year="2012">Ponce-Reyes et al., 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FD93E888FD6FFE40" author="Rojas-Soto, O. R. & V. Sosa & J. F. Ornelas" box="[515,762,386,409]" pageId="11" pageNumber="44" pagination="2671 - 2690" refId="ref15782" refString="Rojas-Soto, O. R., V. Sosa, and J. F. Ornelas. 2012. Forecasting cloud forest in eastern and southern Mexico: conservation insights under future climate change scenarios. Biodiversity and Conservation 21: 2671 - 2690." type="journal article" year="2012">Rojas-Soto et al., 2012</bibRefCitation>
|
||
). Because
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FF59E8AAFED9FE6C" box="[201,332,415,437]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FF59E8AAFED9FE6C" box="[201,332,415,437]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
also occurs in more widespread mid-elevation forests, the threat posed by climate change remains difficult to quantify (but see
|
||
<bibRefCitation id="643D4B33FFB7FFC4FE35E8D1FDF0FE28" author="Elsen, P. R. & W. B. Monahan & A. M. Merenlender" box="[421,613,474,497]" pageId="11" pageNumber="44" pagination="1974" refId="ref13818" refString="Elsen, P. R., W. B. Monahan, and A. M. Merenlender. 2020. Topography and human pressure in mountain ranges alter expected species responses to climate change. Nature Communications 11: 1974." type="journal article" year="2020">Elsen et al., 2020</bibRefCitation>
|
||
). Additionally, the species could be threatened by direct killing due to mistaken beliefs that it is dangerously venomous (
|
||
<bibRefCitation id="643D4B33FFB7FFC4FD0CEB1CFEA0FD93" author="Penguilly Macias, M. A. & A. Moreno Fuentes, I. & G. Espinoza Pineda" pageId="11" pageNumber="44" pagination="99 - 105" refId="ref15271" refString="Penguilly Macias, M. A., A. Moreno Fuentes, I. G. Mayer- Goyenechea, and G. Espinoza Pineda. 2010. Percepcion acerca de las lagartijas consideradas nocivas por algunos Otomies, Nahuas, Tepehuas y Mestizos en el Estado de Hidalgo, Mexico, p. 99 - 105. In: Etnobiologia y Sistemas Biocognitivos Tradicionales: Paradigmas en la Conservacion Biologica y el Fortalecimiento Cultural. A. Moreno, R. Valadez, M. Teresa Pulido, R. Mariaca, P. Mejia, and T. V. Gutierrez Santillan (eds.). Asociacion Etnobiologica Mexicana y Sociedad Latinoamericana de Etnobiologia, Universidad Autonoma del Estado de Hidalgo, Mexico." type="book chapter" year="2010">Penguilly Macías et al., 2010</bibRefCitation>
|
||
) and by harvest for the pet trade (
|
||
<bibRefCitation id="643D4B33FFB7FFC4FD20EB39FF35FDBF" author="Altherr, S." pageId="11" pageNumber="44" refId="ref12577" refString="Altherr, S. 2014. Stolen wildlife - why the EU needs to tackle smuggling of nationally protected species. Report by Pro Wildlife, Munich, Germany." type="book" year="2014">Altherr, 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FF3CEB5BFEE2FDBE" author="Auliya, M. & S. Altherr & D. Ariano-Sanchez & E. H. Baard & C. Brown & Brown, J. - C. & Cantu, G. & Gentile, P. & Gildenhuys, E. & Henningheim, J. & Hintzmann, K. & Kanari, M. & Krvavac, M. & Lettink & T. Ziegler" box="[172,375,593,615]" pageId="11" pageNumber="44" pagination="103 - 119" refId="ref12639" refString="Auliya, M., S. Altherr, D. Ariano-Sanchez, E. H. Baard, C. Brown, R. M. Brown, J. - C. Cantu, G. Gentile, P. Gildenhuys, E. Henningheim, J. Hintzmann, K. Kanari, M. Krvavac, M. Lettink... T. Ziegler. 2016. Trade in live reptiles, its impact on wild populations, and the role of the European market. Biological Conservation 204: 103 - 119." type="journal article" year="2016">Auliya et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FE14EB5BFD18FDB1" author="Hernandez Mares, P." box="[388,653,593,616]" pageId="11" pageNumber="44" refId="ref14325" refString="Hernandez Mares, P. 2018. ' Dragoncitos': Trafico Ilegal Pone en Peligro a Lagartijas Mexicanas. Mongabay web article: https: // es. mongabay. com / 2018 / 03 / trafico-ilegal-lagartijasmexicanas / (accessed 24 March 2018)." type="url" year="2018">Hernández Mares, 2018</bibRefCitation>
|
||
), although we consider these factors less important compared to deforestation and climate change. Nonetheless, based on the risk of hobbyist commercialization, we have preemptively masked the locality data reported herein by (1) rounding all GPS coordinates to the nearest hundredth of a decimal degree, and (2) not giving distances or cardinal directions from the nearest settlement or landmark.
|
||
</paragraph>
|
||
<caption id="54D3664AFFB6FFC5FB0AEEAAFB76F830" ID-DOI="http://doi.org/10.5281/zenodo.7846907" ID-Zenodo-Dep="7846907" httpUri="https://zenodo.org/record/7846907/files/figure.png" pageId="10" pageNumber="43" startId="10.[1178,1206,1952,1972]" targetBox="[142,1129,1528,2020]" targetPageId="10" targetType="figure">
|
||
<paragraph id="001336C2FFB6FFC5FB0AEEAAFB76F830" blockId="10.[1178,1530,1951,2025]" pageId="10" pageNumber="43">
|
||
FIG. 6. Habitat of
|
||
<taxonomicName id="C7AC4D41FFB6FFC5FADCEE95FA6DF86A" box="[1356,1528,1951,1971]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="43" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB6FFC5FADCEE95FA6DF86A" bold="true" box="[1356,1528,1951,1971]" italics="true" pageId="10" pageNumber="43">Abronia zongolica</emphasis>
|
||
</taxonomicName>
|
||
at the type locality in Veracruz, Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="001336C2FFB7FFC4FFEFEA34FDB1FA25" blockId="11.[103,773,179,1532]" pageId="11" pageNumber="44">
|
||
Given this available threat data, together with the aforementioned distribution and ecology of
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FDC8EA56FD42FCA8" box="[600,727,859,881]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FDC8EA56FD42FCA8" box="[600,727,859,881]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
, we propose its categorization under three relevant speciesimperilment frameworks. First, we suggest that
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FD14EA9DFC90FC75" box="[644,773,918,940]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FD14EA9DFC90FC75" box="[644,773,918,940]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
be provisionally assessed as Endangered (B1ab[iii]
|
||
<emphasis id="32D8EAD0FFB7FFC4FD30EAB9FD25FC13" bold="true" box="[672,688,947,970]" pageId="11" pageNumber="44">þ</emphasis>
|
||
2ab[iii]) on the International Union for Conservation of Nature’s (IUCN) Red List of Threatened Species. Second, we evaluate
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FFF7ED07FF73FBFB" box="[103,230,1036,1058]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FFF7ED07FF73FBFB" box="[103,230,1036,1058]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
as having an Environmental Vulnerability Score (EVS;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FF3AED20FE15FB98" author="Wilson, L. D. & V. Mata-Silva & J. D. Johnson" box="[170,384,1066,1089]" pageId="11" pageNumber="44" refId="ref16799" refString="Wilson, L. D., V. Mata-Silva, and J. D. Johnson. 2013. A conservation reassessment of the reptiles of Mexico based" type="book" year="2013">Wilson et al., 2013</bibRefCitation>
|
||
) of 18 out of 20, placing it in the High Vulnerability category. Contributory geographic distribution, ecological distribution, and human persecution scores of 6, 8, and 4, respectively, substantiate this EVS, but if the species is shown to have a broader distribution and/or becomes commercially exploited for the pet trade, the latter two scores would warrant being updated to 7 and 6, respectively. Lastly, we recommend a categorization of Amenazada (Threatened) for
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FE51EC1DFDDDFAF5" box="[449,584,1302,1324]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="species" species="zongolica">
|
||
<emphasis id="32D8EAD0FFB7FFC4FE51EC1DFDDDFAF5" box="[449,584,1302,1324]" italics="true" pageId="11" pageNumber="44">A. zongolica</emphasis>
|
||
</taxonomicName>
|
||
on the Norma Oficial Mexicana list (
|
||
<bibRefCitation id="643D4B33FFB7FFC4FECDEC3FFDBDFA92" author="SEMARNAT & Secretaria de Medio Ambiente & Recursos Naturales" box="[349,552,1332,1355]" pageId="11" pageNumber="44" refId="ref16018" refString="SEMARNAT (Secretaria de Medio Ambiente y Recursos Naturales). 2010. Norma Oficial Mexicana NOM- 059 - SEMARNAT- 2010. Proteccion Ambiental de Especies Nativas de Mexico de Flora y Fauna Silvestres. Categorias de Riesgo y Especificaciones para su Inclusion, Exclusion o Cambio. Lista de Especies en Riesgo. Diario Oficial de la Federacion, Mexico." type="book" year="2010">SEMARNAT, 2010</bibRefCitation>
|
||
), which is the only legally binding imperiled-species list in
|
||
<collectingCountry id="78BB7652FFB7FFC4FD8CEC58FDE5FAB1" box="[540,624,1362,1384]" name="Mexico" pageId="11" pageNumber="44">Mexico</collectingCountry>
|
||
. We base this recommendation on the species meeting Criteria A.(I)
|
||
<emphasis id="32D8EAD0FFB7FFC4FF04EC86FF31FA7A" bold="true" box="[148,164,1420,1443]" pageId="11" pageNumber="44">þ</emphasis>
|
||
B.(II)
|
||
<emphasis id="32D8EAD0FFB7FFC4FF48EC86FF7DFA7A" bold="true" box="[216,232,1420,1443]" pageId="11" pageNumber="44">þ</emphasis>
|
||
C.(II)
|
||
<emphasis id="32D8EAD0FFB7FFC4FE8FEC86FEBAFA7A" bold="true" box="[287,303,1420,1443]" pageId="11" pageNumber="44">þ</emphasis>
|
||
D.(II) as defined by the Anexo Normativo I. All three proposed categorizations are consistent with those of many other arboreal species of
|
||
<taxonomicName id="C7AC4D41FFB7FFC4FE4AECC2FDB9FA07" box="[474,556,1480,1502]" class="Reptilia" family="Anguidae" genus="Abronia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="32D8EAD0FFB7FFC4FE4AECC2FDB9FA07" box="[474,556,1480,1502]" italics="true" pageId="11" pageNumber="44">Abronia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="643D4B33FFB7FFC4FDA9ECC3FC95FA07" author="SEMARNAT & Secretaria de Medio Ambiente & Recursos Naturales" box="[569,768,1481,1502]" pageId="11" pageNumber="44" refId="ref16018" refString="SEMARNAT (Secretaria de Medio Ambiente y Recursos Naturales). 2010. Norma Oficial Mexicana NOM- 059 - SEMARNAT- 2010. Proteccion Ambiental de Especies Nativas de Mexico de Flora y Fauna Silvestres. Categorias de Riesgo y Especificaciones para su Inclusion, Exclusion o Cambio. Lista de Especies en Riesgo. Diario Oficial de la Federacion, Mexico." type="book" year="2010">SEMARNAT, 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FFF7ECECFEAEFA22" author="Wilson, L. D. & V. Mata-Silva & J. D. Johnson" box="[103,315,1510,1532]" pageId="11" pageNumber="44" refId="ref16799" refString="Wilson, L. D., V. Mata-Silva, and J. D. Johnson. 2013. A conservation reassessment of the reptiles of Mexico based" type="book" year="2013">Wilson et al., 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="643D4B33FFB7FFC4FED8ECECFD8CFA25" author="Clause, A. G. & R. Luna-Reyes & A. Nieto-Montes de Oca" box="[328,537,1509,1532]" pageId="11" pageNumber="44" pagination="330 - 343" refId="ref13508" refString="Clause, A. G., R. Luna-Reyes, and A. Nieto-Montes de Oca. 2020. A new species of Abronia (Squamata: Anguidae) from a protected area in Chiapas, Mexico. Herpetologica 76: 330 - 343." type="journal article" year="2020">Clause et al., 2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |