265 lines
27 KiB
XML
265 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1166.96193" ID-Pensoft-Pub="1313-2970-1166-91" ID-Pensoft-UUID="4632CBCBEB0659009F53366E2553A21C" ID-ZooBank="4B15C8CDDFF94ACEAD2A749D3AB5C5AD" ModsDocID="1313-2970-1166-91" checkinTime="1686086550896" checkinUser="pensoft" docAuthor="Dvorak, Libor, Fogasova, Katarina, Obona, Jozef, Toeroek, Edina & Manko, Peter" docDate="2023" docId="4ABF56F3B29C529A85B5F704180E8F15" docLanguage="en" docName="ZooKeys 1166: 91-102" docOrigin="ZooKeys 1166" docPubDate="2023-06-06" docSource="http://dx.doi.org/10.3897/zookeys.1166.96193" docTitle="Ptychoptera xanthopleura Dvorak, Obona & Manko 2023, sp. nov." docType="treatment" docUuid="CD413B9C-177D-4A4C-ACB9-3565C502C27B" docUuidSource="ZooBank" docVersion="2" id="4632CBCBEB0659009F53366E2553A21C" lastPageNumber="91" masterDocId="4632CBCBEB0659009F53366E2553A21C" masterDocTitle="Two new Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from the Western Palaearctic" masterLastPageNumber="102" masterPageNumber="91" pageNumber="91" updateTime="1686086854992" updateUser="ExternalLinkService">
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<mods:title id="33B5734CFEFDB259E160F71E40BC5688">Two new Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from the Western Palaearctic</mods:title>
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<mods:name id="3C024154E483C89A19CBEA72E3163A44" type="personal">
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<mods:namePart id="A2FE343D4469BD4369C6603BD08557A3">Dvorak, Libor</mods:namePart>
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<mods:nameIdentifier id="2EC68B18713CBB7F507AB928A11126A2" type="ORCID">https://orcid.org/0000-0002-4712-3679</mods:nameIdentifier>
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<mods:affiliation id="AD5EB95DAF423E2DA757C23C385FA598">Tri Sekery 21, CZ- 35301 Marianske Lazne, Czech Republic</mods:affiliation>
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<mods:roleTerm id="B2686559E6F472D6E8041F4A0F2793EA">Author</mods:roleTerm>
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<mods:namePart id="AD4FF92BE3B782C459C16FDE1102ABA1">Fogasova, Katarina</mods:namePart>
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<mods:nameIdentifier id="EE964B759DB0183C497ADE3DB4CB350F" type="ORCID">https://orcid.org/0009-0003-9604-1885</mods:nameIdentifier>
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<mods:affiliation id="2A1C43D51C43F4BBF9F82C0FFE01F104">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
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<mods:nameIdentifier id="672B6DDBBD17AEFAACF30C1F4FC7BA6E" type="email">katarina.fogasova@smail.unipo.sk</mods:nameIdentifier>
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<mods:roleTerm id="B5746EDA4F1D817D12009FFD17DF0246">Author</mods:roleTerm>
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<mods:namePart id="A03FDFC2A541DA58B02B38E49F77AF15">Obona, Jozef</mods:namePart>
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<mods:nameIdentifier id="CAEF5A2DB140F4FDBEB2ACCF5183914F" type="ORCID">https://orcid.org/0000-0002-1185-658X</mods:nameIdentifier>
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<mods:affiliation id="01D72E8747B6A4A241A7402E9199E07B">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name id="8C4B3F124F71C73CF9CE5B6881E6D067" type="personal">
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<mods:roleTerm id="0352C1FBDDEFF8FC801AD8C9971B18A3">Author</mods:roleTerm>
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<mods:namePart id="E03968A86139AF2C0881901CBC4E933E">Toeroek, Edina</mods:namePart>
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<mods:nameIdentifier id="A2B0B7A411A27AABA0077C998CA78471" type="ORCID">https://orcid.org/0000-0001-5982-7078</mods:nameIdentifier>
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<mods:affiliation id="CAB4DE9224D67C89FD800431211DE8FD">" Lenduelet " Landscape and Conservation Ecology Research Group, Institute of Ecology and Botany, Centre for Ecological Research, Alkotmany str. 2 - 4, Vacratot, H- 2163, Hungary</mods:affiliation>
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<mods:namePart id="8C84F94B619E4EC27EB35D4FEEE395EA">Manko, Peter</mods:namePart>
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<mods:nameIdentifier id="382B2AC380C429DED89DC7C2C7D87EF4" type="ORCID">https://orcid.org/0000-0003-1862-9117</mods:nameIdentifier>
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<mods:affiliation id="B37176F5648158E9807C5C7042C27FA6">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Presov, 17. novembra 1, SK- 08116 Presov, Slovakia</mods:affiliation>
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<mods:title id="EB7E0FB052F3B9C724E5C1F9F6BD3423">ZooKeys</mods:title>
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<mods:date id="98C47B6640C665ED2DF752B63AE2EE3B">2023</mods:date>
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<mods:number id="54F76C155DA052DAC708BC64E42D2A83">2023-06-06</mods:number>
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<mods:number id="FA6345F10E3D0E75AEC8EF5B4D6B5895">1166</mods:number>
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<mods:start id="E96D272AF72A7996BD967762A005C04D">91</mods:start>
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<mods:url id="C21E0704ECDE1FE1AB31AB8B5C6A8DB2">http://dx.doi.org/10.3897/zookeys.1166.96193</mods:url>
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<mods:classification id="3668934B5E48A36C2E12C7464A2031C5">journal article</mods:classification>
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<mods:identifier id="8E392EEEAD6027AEEA7DA801465D18FD" type="DOI">http://dx.doi.org/10.3897/zookeys.1166.96193</mods:identifier>
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<mods:identifier id="B8EAC6BF305B5E95CE0408925BA94D2A" type="Pensoft-Pub">1313-2970-1166-91</mods:identifier>
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<mods:identifier id="DE84348BC56FE7DF929B97E511CA7D21" type="ZooBank">4B15C8CDDFF94ACEAD2A749D3AB5C5AD</mods:identifier>
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<treatment id="4ABF56F3B29C529A85B5F704180E8F15" LSID="urn:lsid:zoobank.org:act:CD413B9C-177D-4A4C-ACB9-3565C502C27B" httpUri="http://treatment.plazi.org/id/4ABF56F3B29C529A85B5F704180E8F15" lastPageNumber="91" pageId="0" pageNumber="91">
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<subSubSection id="E9CA13382B098E866A4DBC452996B9EF" pageId="0" pageNumber="91" type="nomenclature">
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<paragraph id="9345ABDF2AD976033068CE5B0A3CD5A5" pageId="0" pageNumber="91">
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<taxonomicName id="CF595C008C151CECBAEB67E4246953A2" LSID="https://zoobank.org/CD413B9C-177D-4A4C-ACB9-3565C502C27B" authority="Dvořák, Oboňa & Manko" authorityName="Dvorak, Obona & Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura" status="sp. nov.">
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Ptychoptera xanthopleura
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<normalizedToken id="4158862F3B12A8DCA8DBC2E60A0340F2" originalValue="Dvořák">Dvorak</normalizedToken>
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,
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<normalizedToken id="7BEE7F3B1DADDBF928AD81A53CFA4298" originalValue="Oboňa">Obona</normalizedToken>
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& Manko
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</taxonomicName>
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<taxonomicNameLabel id="E6696C222EF66598B57D7A347B4855AE" pageId="0" pageNumber="91">sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4F778EC72C86C5256CBD392079EB17DA" pageId="0" pageNumber="91" type="description">
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<paragraph id="BF82C0EF0AE4661D66F72A8EA49354D3" pageId="0" pageNumber="91">
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||
<figureCitation id="4E22844503B4C414A0EB97E13D9E6499" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ptychoptera xanthopleura sp. nov. a habitus of male, lateral view b pleurae, lateral view c wing." figureDoi="10.3897/zookeys.1166.96193.figure1" httpUri="https://binary.pensoft.net/fig/860365" pageId="0" pageNumber="91">Figs 1</figureCitation>
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<figureCitation id="BFA4FBA165637DD6B7DA0621607D53E8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ptychoptera xanthopleura sp. nov. terminalia a epandrium b paramere c aedeagal complex d gonostylus e hypandrium. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GAS = apical stylus of gonostylus, ALP = lateral ejaculatory process." figureDoi="10.3897/zookeys.1166.96193.figure2" httpUri="https://binary.pensoft.net/fig/860366" pageId="0" pageNumber="91">, 2</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A2AACD83AAF8EFB324FE633F026198B9" pageId="0" pageNumber="91" type="type material">
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||
<paragraph id="E048764EB1E5C2C04E54119D81FB354A" pageId="0" pageNumber="91">Type material.</paragraph>
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<paragraph id="C618EFC4DC7C1A0614C65BFBE8758AEF" pageId="0" pageNumber="91">
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<emphasis id="4A375A134CF722A8B1D08F492A9B46BE" bold="true" italics="true" pageId="0" pageNumber="91">Holotype</emphasis>
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: 1 ♂: Azerbaijan, Qum, sidebrook/small tributary of the Ardavacaj (Ardavachay) River + wetland, 845 m a. s. l.,
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<geoCoordinate id="934C5CC48EEFE08C507A146EC7949C4D" degrees="41" direction="north" minutes="28" orientation="latitude" precision="1" seconds="10.3" value="41.46953">41°28'10.3"N</geoCoordinate>
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,
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<geoCoordinate id="CD4F263781B7B59E45A2893FFB1DE2E3" degrees="46" direction="east" minutes="55" orientation="longitude" precision="1" seconds="57.2" value="46.932556">46°55'57.2"E</geoCoordinate>
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, 8.V.2019, leg. J.
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<normalizedToken id="292F2B65A61D34669F7AD7AA6D1DCFC4" originalValue="Oboňa">Obona</normalizedToken>
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& P. Manko.
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<emphasis id="D82A22A463036615E7E736A15149078F" bold="true" italics="true" pageId="0" pageNumber="91">Paratypes</emphasis>
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: 2 ♂♂: Georgia, border of Imereti and Samtskhe-Javakheti regions, brook and spring, south slope of Zekari pass, 2 050 m a. s. l.,
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<geoCoordinate id="5B0ECE83BDF63E7B0EE8D2BF6E28D681" degrees="41" direction="north" minutes="49" orientation="latitude" precision="15" seconds="23" value="41.823055">41°49'23"N</geoCoordinate>
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,
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<geoCoordinate id="662EBC50F1419C29E14A8CC2C1E27283" degrees="42" direction="east" minutes="51" orientation="longitude" precision="15" seconds="09" value="42.852497">42°51'09"E</geoCoordinate>
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, 17.VII.2019, leg. G.
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<normalizedToken id="D8A5B4312DA85C4A8401C1F5BC23768A" originalValue="Vinçon">Vincon</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="8C05879903783F5AD86B569FC018DB24" pageId="0" pageNumber="91" type="description">
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<paragraph id="B5775ABAF0DB3BFD84D78D0728E61C5A" pageId="0" pageNumber="91">Description.</paragraph>
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<paragraph id="F030C4A3581CC4EAAF72085C80EC2BA1" pageId="0" pageNumber="91">
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<emphasis id="47762FFC98A92682ECCDA9A28456CC01" bold="true" pageId="0" pageNumber="91">
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Male.
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<emphasis id="63DCB6DB263D649C589C3BE47C5ED79B" italics="true" pageId="0" pageNumber="91">Head</emphasis>
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</emphasis>
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: Frons, vertex, and occiput black with metallic blue shine, mouthparts including palpi pale yellow, scape and pedicel yellowish orange, antennal flagellomeres a somewhat darker, tending to pale brown.
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</paragraph>
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<paragraph id="70E70870D298307C852EC833983B8146" pageId="0" pageNumber="91">
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<emphasis id="7061F10AF517D897C97AA89760D26E4C" bold="true" italics="true" pageId="0" pageNumber="91">Thorax</emphasis>
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: Scutum, paratergite, and mediotergite blackish with metallic blue shine; scutellum, pleurotergite, katepisternum, and katepimeron brownish black with lighter metallic blue shine; other parts yellow. Halteres yellow with light brown knob. Legs yellow except brown extreme apex of femora and tibiae, tarsi somewhat darkened.
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</paragraph>
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<paragraph id="BCC4123DCCD055BE70221D5FB9FDD249" pageId="0" pageNumber="91">
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Wing length 10 mm (holotype, Fig.
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<figureCitation id="06DD018C63CD922D4269AA71DA7D90DB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ptychoptera xanthopleura sp. nov. a habitus of male, lateral view b pleurae, lateral view c wing." figureDoi="10.3897/zookeys.1166.96193.figure1" httpUri="https://binary.pensoft.net/fig/860365" pageId="0" pageNumber="91">1c</figureCitation>
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). Wing almost hyaline, veins yellowish brown, distinct spots brownish black, forming more or less three stripes, at base of wing at the level of crossvein h, from C to Cu. Middle stripe touching vein C, running through cross-veins up to middle part of vein Cu, isolated spot before end of vein Cu. Isolated spot on around middle of R1. Third stripe consist of three large, almost touching spots: at the tip of R1 and fork of R2+3, one on fork vein R4+5 and one on fork vein M1+2. Small spot at the end of vein R3.
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</paragraph>
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<caption id="C11F15FC5F84922D85844B9C870F5136" doi="10.3897/zookeys.1166.96193.figure1" httpUri="https://binary.pensoft.net/fig/860365" pageId="0" pageNumber="91" start="Figure 1" startId="F1">
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||
<paragraph id="BEDF4BD880264051236BF39C6F44637E" pageId="0" pageNumber="91">
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||
<emphasis id="4A11D0E1F9428EE2C4D74A78D37DE8C6" bold="true" pageId="0" pageNumber="91">Figure 1.</emphasis>
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<taxonomicName id="646EE3739AA818A68D5B0FC06713460F" authorityName="Dvorak, Obona & Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura">
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<emphasis id="25E7E0FC5417976137BAA1B3904439AD" italics="true" pageId="0" pageNumber="91">Ptychoptera xanthopleura</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="3876139F951380FBB4D897014BB434D8" bold="true" pageId="0" pageNumber="91">a</emphasis>
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habitus of male, lateral view
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<emphasis id="D146916AD4EF0AB6AC43E0D0B7862EC4" bold="true" pageId="0" pageNumber="91">b</emphasis>
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pleurae, lateral view
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<emphasis id="AC8CFB0E81675B4058B38A76AE6A8191" bold="true" pageId="0" pageNumber="91">c</emphasis>
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wing.
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</paragraph>
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</caption>
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<paragraph id="7B6900AA2337A7BB66F080E141EB6EF4" pageId="0" pageNumber="91">
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<emphasis id="2F871210716C324761D7BAE4D59B0176" bold="true" italics="true" pageId="0" pageNumber="91">Abdomen</emphasis>
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: Tergum 1 dark shiny brown with yellow apex, sternum 1 yellow. Tergum 2 brown basally and apically, yellow in middle, sternum 2 yellow. Tergum 3 yellow basally, brown apically, sternum 3 yellow. Remaining terga and sterna brown, sternum 4 yellow basally. Auxiliar copulatory organ yellow.
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</paragraph>
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<paragraph id="4517D58C65825214149E1DB5B53C230F" pageId="0" pageNumber="91">
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<emphasis id="DEAA30E0674C0E85057475354337D8E9" bold="true" italics="true" pageId="0" pageNumber="91">Male genitalia</emphasis>
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(Fig.
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<figureCitation id="C19D51D09D410AE4BEBE1F332B24D4A6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ptychoptera xanthopleura sp. nov. terminalia a epandrium b paramere c aedeagal complex d gonostylus e hypandrium. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GAS = apical stylus of gonostylus, ALP = lateral ejaculatory process." figureDoi="10.3897/zookeys.1166.96193.figure2" httpUri="https://binary.pensoft.net/fig/860366" pageId="0" pageNumber="91">2</figureCitation>
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): Hypopygium almost 2
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<normalizedToken id="789E72C28AD0113FD59B02E0632DD649" originalValue="×">x</normalizedToken>
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as wide as long, widest in basal quarter, medially with very deep emargination. Epandrial claspers simple and long (length/width ratio ca. 5.8) and covered by long pale hairs, the longest hair up to 1.75 longer than width of epandrium. Apical stylus of gonostylus robust and rounded apically, secondary lobe long, reaching almost 0.75 of apical stylus length. Media lobe of basal lobe of gonostylus long and sharply pointed (saber-like); anterior lobe of basal lobe of gonostylus bulbous apically with several setae at extreme apex.
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</paragraph>
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<caption id="34A24E90DAEA87AF5D8FC48759E81EAB" doi="10.3897/zookeys.1166.96193.figure2" httpUri="https://binary.pensoft.net/fig/860366" pageId="0" pageNumber="91" start="Figure 2" startId="F2">
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||
<paragraph id="8CCE50672B9FFAB07D5A3EA7D33A4F50" pageId="0" pageNumber="91">
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<emphasis id="372D252F8D0B675A73606B7A3722314B" bold="true" pageId="0" pageNumber="91">Figure 2.</emphasis>
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<taxonomicName id="E97504B371B6FD952F20CA2F295F9472" authorityName="Dvorak, Obona & Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura">
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<emphasis id="364F419642E854F5A656C56E32B0C885" italics="true" pageId="0" pageNumber="91">Ptychoptera xanthopleura</emphasis>
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</taxonomicName>
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sp. nov. terminalia
|
||
<emphasis id="9791557E8F100BD8381682B527650A5E" bold="true" pageId="0" pageNumber="91">a</emphasis>
|
||
epandrium
|
||
<emphasis id="74AFDBE3DA2D2A8A71BCF083AA2B5E20" bold="true" pageId="0" pageNumber="91">b</emphasis>
|
||
paramere
|
||
<emphasis id="6581CD39EE695A493E6812D9D299A648" bold="true" pageId="0" pageNumber="91">c</emphasis>
|
||
aedeagal complex
|
||
<emphasis id="DD8E70CF722B56AF9403684E0AA80917" bold="true" pageId="0" pageNumber="91">d</emphasis>
|
||
gonostylus
|
||
<emphasis id="2F0B8A8C0F9353FDEA757D20E1A74779" bold="true" pageId="0" pageNumber="91">e</emphasis>
|
||
hypandrium. Abbreviations: ECP = epandrial clasper, EL = epandrial lobe, EPI = epiproct, GBM = medial lobe of basal lobe of gonostylus, GAS = apical stylus of gonostylus, ALP = lateral ejaculatory process.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DB33040F8B598EF498E9E414CF40D789" pageId="0" pageNumber="91">
|
||
<emphasis id="26B158BC908BFB1EEA5B0430D47A1322" bold="true" pageId="0" pageNumber="91">Female.</emphasis>
|
||
The authors have an immature female which was sampled in Lesser Caucasus (Georgia, Kakheti region, Ilto river, above (N of) the
|
||
<normalizedToken id="648CCBF25F176B29B0FCA2B1DBA30D4F" originalValue="Chart’ala">Chart'ala</normalizedToken>
|
||
village, 790 m a. s. l.,
|
||
<geoCoordinate id="CF06E3CB98A1A1B8441CCBEFCF681A2F" degrees="42" direction="north" minutes="8" orientation="latitude" precision="15" seconds="18" value="42.138336">42°8'18"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BA016E67FF3FC2F406B34717C44D97C" degrees="45" direction="east" minutes="7" orientation="longitude" precision="15" seconds="32" value="45.125553">45°7'32"E</geoCoordinate>
|
||
, 8.VII.2019, leg. P. Manko & G.
|
||
<normalizedToken id="D15A13B976760CCB74146F5430DC5BF5" originalValue="Vinçon">Vincon</normalizedToken>
|
||
). The characters correspond to the above-described new species. However, its identity cannot be confirmed in this stage of ontogenesis/development and could be solved after collecting more specimens of the genus
|
||
<taxonomicName id="BCF7D1459806C133F59B4579C805525A" authorityName="Meigen" authorityYear="1803" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F682A22A47C46FFF303D6937519F9A68" italics="true" pageId="0" pageNumber="91">Ptychoptera</emphasis>
|
||
</taxonomicName>
|
||
from the Transcaucasia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="80EB01A0250C05858D7A00AAC1244F3F" pageId="0" pageNumber="91" type="etymology">
|
||
<paragraph id="0CCC59371BB2B7BE0A96E7A21C9F99BE" pageId="0" pageNumber="91">Etymology.</paragraph>
|
||
<paragraph id="0E6D0BCED0F4E0FBD855CC59A14AB2BB" pageId="0" pageNumber="91">
|
||
The name reflects predominantly yellow pleurae (Fig.
|
||
<figureCitation id="7D20D1E6C4F6B9A76E2593C2C3B2CD3F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ptychoptera xanthopleura sp. nov. a habitus of male, lateral view b pleurae, lateral view c wing." figureDoi="10.3897/zookeys.1166.96193.figure1" httpUri="https://binary.pensoft.net/fig/860365" pageId="0" pageNumber="91">1b</figureCitation>
|
||
), which are unique for the Western Palaearctic species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEFA3ADC7FBF0AE238F773ED6AA484DD" pageId="0" pageNumber="91" type="differential diagnosis">
|
||
<paragraph id="8BC4EC09046E24780C1EFFCD937F4BCA" pageId="0" pageNumber="91">Differential diagnosis.</paragraph>
|
||
<paragraph id="FAAA2B797F5ACCAC247AAD53E5E86045" pageId="0" pageNumber="91">
|
||
According to the presence of auxiliary sexual organ and shining pleurae,
|
||
<taxonomicName id="4350C3CAF332C983F5D39B80BAF62FE8" class="Insecta" kingdom="Animalia" lsidName="P. xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura">
|
||
<emphasis id="21D02282C6AF781FF63A7CC38494E34E" italics="true" pageId="0" pageNumber="91">P. xanthopleura</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the subgenus
|
||
<taxonomicName id="17D33F478DA0F3A9FD6F5F9354BA2241" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera (Paraptychoptera)" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="subGenus" subGenus="Paraptychoptera">Ptychoptera Paraptychoptera</taxonomicName>
|
||
and according to male genitalia and the maximum parsimonious tree based on 53 morphological characters (see Fig.
|
||
<figureCitation id="354645D179CCEDCCCFDA4F75658F1B54" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The maximum parsimonious tree based on 53 morphological characters. The characteristics are shown above branches. The parentheses show the maximum parsimony phylogeny bootstrap support value expressed as percentages. Support values under 40 are not shown." figureDoi="10.3897/zookeys.1166.96193.figure3" httpUri="https://binary.pensoft.net/fig/860367" pageId="0" pageNumber="91">3</figureCitation>
|
||
and Table
|
||
<tableCitation id="7C1C9D3EA60616B68001CA87D2D2C7DB" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Matrix of the 53 morphological data (based on the Keresztes et al. 2021) used in the phylogenetic analyses. For description of characters and character states, see Keresztes et al. (2021). '? ' - missing data." httpUri="http://table.plazi.org/id/CF8E29BB06282969F5BAF729122E839C" pageId="0" pageNumber="91" tableUuid="CF8E29BB06282969F5BAF729122E839C">1</tableCitation>
|
||
), the nearest species is
|
||
<taxonomicName id="5D4BDFB9F66143B7D8326EC326F26797" class="Insecta" kingdom="Animalia" lsidName="P. lacustris" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="lacustris">
|
||
<emphasis id="50D6EC28C3A8F3FB99AA74AD5C2C5C34" italics="true" pageId="0" pageNumber="91">P. lacustris</emphasis>
|
||
</taxonomicName>
|
||
and belongs to the highly divergent monophyletic unit, the
|
||
<taxonomicName id="95D4182DADDA5BD1B1155D0D03EF48BA" authorityName="Meigen" authorityYear="1830" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera lacustris" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="lacustris">
|
||
<emphasis id="1E06B5AAEDA08132785714D511BC584F" italics="true" pageId="0" pageNumber="91">Ptychoptera lacustris</emphasis>
|
||
</taxonomicName>
|
||
group, including five species,
|
||
<taxonomicName id="3393EDC6AD61D1426573C329337A7897" class="Insecta" kingdom="Animalia" lsidName="P. xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura">
|
||
<emphasis id="65E2A192A741BD26266D9A3B60BA6980" italics="true" pageId="0" pageNumber="91">P. xanthopleura</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="F75046962D697BA5E9197920B9AFB22A" class="Insecta" kingdom="Animalia" lsidName="P. lacustris" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="lacustris">
|
||
<emphasis id="030846CECD58B80A1B272B833AB5A43D" italics="true" pageId="0" pageNumber="91">P. lacustris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="127390736533F4AA32C5446CBEDFD63F" class="Insecta" kingdom="Animalia" lsidName="P. castor" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="castor">
|
||
<emphasis id="9565E612A8D46F681498299E70C270A7" italics="true" pageId="0" pageNumber="91">P. castor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="04FAACFC070BCC31C7373A9C4DF30732" class="Insecta" kingdom="Animalia" lsidName="P. helena" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="helena">
|
||
<emphasis id="9BE346A76CCC40FDDA78CF2DCF9D1107" italics="true" pageId="0" pageNumber="91">P. helena</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="5C64AAA80592D1643546C101EBD329A3" class="Insecta" kingdom="Animalia" lsidName="P. pollux" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="pollux">
|
||
<emphasis id="4F1EC02EDD7FD28ED1F9CCF5F4CDED44" italics="true" pageId="0" pageNumber="91">P. pollux</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="BD13F16EA055EEC6E29863415CC9235F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The maximum parsimonious tree based on 53 morphological characters. The characteristics are shown above branches. The parentheses show the maximum parsimony phylogeny bootstrap support value expressed as percentages. Support values under 40 are not shown." figureDoi="10.3897/zookeys.1166.96193.figure3" httpUri="https://binary.pensoft.net/fig/860367" pageId="0" pageNumber="91">3</figureCitation>
|
||
).
|
||
<taxonomicName id="1BD60718CEB1E57B58AF32F2D391A939" authorityName="Dvorak, Obona & Manko" authorityYear="2023" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera xanthopleura" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthopleura">
|
||
<emphasis id="D3ACCACF21F9F0CD02F1A95787C14A32" italics="true" pageId="0" pageNumber="91">Ptychoptera xanthopleura</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is close to but differs from the most similar species
|
||
<taxonomicName id="8A64EE4D783242FAD8CBB256CC319FF1" class="Insecta" kingdom="Animalia" lsidName="P. lacustris" order="Diptera" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="lacustris">
|
||
<emphasis id="543154AFC34827758B6FEB6E44BF683A" italics="true" pageId="0" pageNumber="91">P. lacustris</emphasis>
|
||
</taxonomicName>
|
||
mainly by having an almost completely yellow pleurae, the shape of the hypopygium (epandrial claspers, secondary lobe of gonostylus, and medial lobe of basal lobe of gonostylus).
|
||
</paragraph>
|
||
<caption id="B3FFD18E47FA66C7C8CCC125AC9F9878" doi="10.3897/zookeys.1166.96193.figure3" httpUri="https://binary.pensoft.net/fig/860367" pageId="0" pageNumber="91" start="Figure 3" startId="F3">
|
||
<paragraph id="57B3B57F969C0AFC4E447BD89BF0258B" pageId="0" pageNumber="91">
|
||
<emphasis id="7CA384D284E484FAC79882E47C1DF5A3" bold="true" pageId="0" pageNumber="91">Figure 3.</emphasis>
|
||
The maximum parsimonious tree based on 53 morphological characters. The characteristics are shown above branches. The parentheses show the maximum parsimony phylogeny bootstrap support value expressed as percentages. Support values under 40 are not shown.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |