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<document id="C6CD08BC5FDB4C0617FA276737502716" ID-DOI="10.5281/zenodo.8400309" ID-ISSN="1942-1354" ID-Zenodo-Dep="8400309" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630209611898" checkinUser="felipe" docAuthor="Santos-Silva, Antonio &amp; Heffern, Daniel" docDate="2012" docId="DE2B87B87C49FF9EFF7A1AB9FD79FA47" docLanguage="en" docName="0220.pdf" docOrigin="Insecta Mundi 2012 (220)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Malukandra heterostyla" docType="treatment" docVersion="6" lastPageNumber="3" masterDocId="2212FFC07C4BFF9AFFBA1C3FFF8BFFD9" masterDocTitle="Description of the female of Malukandra heterostyla (Lameere, 1902) (Coleoptera, Cerambycidae, Parandrinae) Antonio Santos-Silva" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1699186649247" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="E301D8D9BF1F914482D84B2C18B4D888">Description of the female of Malukandra heterostyla (Lameere, 1902) (Coleoptera, Cerambycidae, Parandrinae) Antonio Santos-Silva</mods:title>
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<taxonomicName id="91824D2D7C49FF98FF7A1AB9FD7BF947" ID-CoL="3XQTZ" authority="(Lameere, 1902)" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[192,752,1670,1694]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C49FF98FF7A1AB9FD7BF947" bold="true" box="[192,752,1670,1694]" pageId="2" pageNumber="1">
<emphasis id="64F6EABC7C49FF98FF7A1AB9FD85F947" bold="true" box="[192,526,1670,1694]" italics="true" pageId="2" pageNumber="1">Malukandra heterostyla</emphasis>
(
<bibRefCitation id="32134B5F7C49FF98FDA41AB9FD6CF947" author="Lameere, A. A." box="[542,743,1670,1694]" pageId="2" pageNumber="1" pagination="59 - 111" refId="ref2559" refString="Lameere, A. A. 1902. Revision des Prionides. Premier memoire. - Parandrines. Annales de la Societe Entomologique de Belgique 46 (3): 59 - 111." type="journal article" year="1902">Lameere, 1902</bibRefCitation>
)
</emphasis>
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<paragraph id="563D36AE7C49FF98FF7A1A99FEBDF967" blockId="2.[192,752,1670,1726]" box="[192,310,1702,1726]" pageId="2" pageNumber="1">
(
<figureCitation id="CEB92A2B7C49FF98FF731A99FEA0F967" box="[201,299,1702,1726]" captionStart="Figures 1–3" captionStartId="4.[192,288,1154,1176]" captionTargetBox="[197,1432,200,1119]" captionTargetId="figure-229@4.[192,1440,192,1124]" captionTargetPageId="4" captionText="Figures 1–3. Malukandra heterostyla, female, habitus. 1) Dorsal. 2) Ventral. 3) Lateral." figureDoi="http://doi.org/10.5281/zenodo.8400311" httpUri="https://zenodo.org/record/8400311/files/figure.png" pageId="2" pageNumber="1">Fig. 1–3</figureCitation>
)
</paragraph>
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<paragraph id="563D36AE7C49FF98FF7A1AD9FBE0F927" blockId="2.[192,1440,1766,1918]" box="[192,1131,1766,1790]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="769211807C49FF98FF7A1AD9FBE0F927" box="[192,1131,1766,1790]" pageId="2" pageNumber="1">
<treatmentCitation id="D72310BF7C49FF98FF7A1AD9FD4BF927" author="Lameere, A. A." box="[192,704,1766,1790]" page="100" pageId="2" pageNumber="1" year="1902">
<taxonomicName id="91824D2D7C49FF98FF7A1AD9FD4BF927" ID-CoL="4D5GJ" authority="Lameere, 1902: 100" authorityName="Lameere" authorityPageNumber="100" authorityYear="1902" box="[192,704,1766,1790]" class="Insecta" family="Cerambycidae" genus="Parandra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C49FF98FF7A1AD9FE4FF927" box="[192,452,1766,1790]" italics="true" pageId="2" pageNumber="1">Parandra heterostyla</emphasis>
<bibRefCitation id="32134B5F7C49FF98FE761AD8FD4BF927" author="Lameere, A. A." box="[460,704,1766,1790]" pageId="2" pageNumber="1" pagination="59 - 111" refId="ref2559" refString="Lameere, A. A. 1902. Revision des Prionides. Premier memoire. - Parandrines. Annales de la Societe Entomologique de Belgique 46 (3): 59 - 111." type="journal article" year="1902">Lameere, 1902: 100</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="D72310BF7C49FF98FD761AD9FCCDF927" author="Lameere, A. A." box="[716,838,1766,1790]" page="116" pageId="2" pageNumber="1" year="1912">
<bibRefCitation id="32134B5F7C49FF98FD761AD9FCCDF927" author="Lameere, A. A." box="[716,838,1766,1790]" pageId="2" pageNumber="1" pagination="113 - 188" refId="ref2593" refString="Lameere, A. A. 1912. Revision des Prionides. Vingt-deuxieme Memoire. - Addenda et Corrigenda. Memoires de la Societe Entomologique de Belgique 21: 113 - 188." type="journal article" year="1912">1912: 116</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="D72310BF7C49FF98FCE81AD9FB9EF927" author="Webb, G. A." box="[850,1045,1766,1790]" page="327" pageId="2" pageNumber="1" year="1994">
<bibRefCitation id="32134B5F7C49FF98FCE81AD9FB9EF927" author="Webb, G. A." box="[850,1045,1766,1790]" pageId="2" pageNumber="1" pagination="325 - 328" refId="ref2797" refString="Webb, G. A. 1994. Parandra araucariae Gressitt (Coleoptera: Cerambycidae: Parandrinae): a new Record for Norfolk Island. Memoirs of the Queensland Museum 37 (1): 325 - 328." type="journal article" year="1994">Webb 1994: 327</bibRefCitation>
</treatmentCitation>
(note).
</treatmentCitationGroup>
</paragraph>
<paragraph id="563D36AE7C49FF98FF7A1B39FC69F8E7" blockId="2.[192,1440,1766,1918]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="769211807C49FF98FF7A1B39FC69F8E7" pageId="2" pageNumber="1">
<taxonomicName id="91824D2D7C49FF98FF7A1B39FDD2F8C7" ID-CoL="4D5GJ" authorityName="Lameere" authorityYear="1902" box="[192,601,1798,1822]" class="Insecta" family="Cerambycidae" genus="Parandra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="heterostyla" subGenus="Parandra">
<emphasis id="64F6EABC7C49FF98FF7A1B39FEB3F8C7" box="[192,312,1798,1822]" italics="true" pageId="2" pageNumber="1">Parandra</emphasis>
(
<emphasis id="64F6EABC7C49FF98FEF01B39FE49F8C7" box="[330,450,1798,1822]" italics="true" pageId="2" pageNumber="1">Parandra</emphasis>
)
<emphasis id="64F6EABC7C49FF98FE6E1B39FDD2F8C7" box="[468,601,1798,1822]" italics="true" pageId="2" pageNumber="1">heterostyla</emphasis>
</taxonomicName>
;
<treatmentCitation id="D72310BF7C49FF98FDD01B38FCB1F8C7" author="Lameere, A. A." box="[618,826,1798,1822]" page="7" pageId="2" pageNumber="1" year="1913">
<bibRefCitation id="32134B5F7C49FF98FDD01B38FCB1F8C7" author="Lameere, A. A." box="[618,826,1798,1822]" pageId="2" pageNumber="1" pagination="1 - 108" refId="ref2626" refString="Lameere, A. A. 1913. Cerambycidae: Prioninae. Coleopterum Catalogus 22 (pars 52): 1 - 108." type="journal article" year="1913">Lameere 1913: 7</bibRefCitation>
</treatmentCitation>
(cat.); 1919: 18;
<treatmentCitation id="D72310BF7C49FF98FBB61B39FB69F8C7" author="Arigony, T. H. A." box="[1036,1250,1798,1822]" page="89" pageId="2" pageNumber="1" year="1984">
<bibRefCitation id="32134B5F7C49FF98FBB61B39FB69F8C7" author="Arigony, T. H. A." box="[1036,1250,1798,1822]" pageId="2" pageNumber="1" pagination="87 - 125" refId="ref2513" refString="Arigony, T. H. A. 1984. O subgenero Parandra Latreille, 1804 (Coleoptera, Cerambycidae, Parandrinae): Estudo fenetico e cladistico de 12 especies. Iheringia (Serie Zoologia) 64: 87 - 125, 66 fig." type="journal article" year="1984">Arigony 1984: 89</bibRefCitation>
</treatmentCitation>
, 90, 95, 96, 97, 109, 111, 125, fig. 16, 60, 64, 65;
<treatmentCitation id="D72310BF7C49FF98FD3B1B19FC07F8E7" author="Santos-Silva, A." box="[641,908,1830,1854]" page="32" pageId="2" pageNumber="1" year="2002">
<bibRefCitation id="32134B5F7C49FF98FD3B1B19FC07F8E7" author="Santos-Silva, A." box="[641,908,1830,1854]" pageId="2" pageNumber="1" pagination="29 - 52" refId="ref2679" refString="Santos-Silva, A. 2002. Notas e descriCOes em Parandrini (Coleoptera, Cerambycidae, Parandrinae). Iheringia (Serie Zoologia) 92 (2): 29 - 52, 56 fig." type="journal article" year="2002">Santos-Silva 2002: 32</bibRefCitation>
</treatmentCitation>
(note).
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</paragraph>
<paragraph id="563D36AE7C49FF98FF7A1B79FC78F887" blockId="2.[192,1440,1766,1918]" box="[192,1011,1862,1886]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="769211807C49FF98FF7A1B79FC78F887" box="[192,1011,1862,1886]" pageId="2" pageNumber="1">
<taxonomicName id="91824D2D7C49FF98FF7A1B79FDC7F887" ID-CoL="LVVS" authorityName="Santos-Silva &amp; Shute" authorityYear="2009" box="[192,588,1862,1886]" class="Insecta" family="Cerambycidae" genus="Birandra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="heterostyla" subGenus="Birandra">
<emphasis id="64F6EABC7C49FF98FF7A1B79FEB8F887" box="[192,307,1862,1886]" italics="true" pageId="2" pageNumber="1">Birandra</emphasis>
(
<emphasis id="64F6EABC7C49FF98FEF91B79FE3DF887" box="[323,438,1862,1886]" italics="true" pageId="2" pageNumber="1">Birandra</emphasis>
)
<emphasis id="64F6EABC7C49FF98FE7D1B79FDC7F887" box="[455,588,1862,1886]" italics="true" pageId="2" pageNumber="1">heterostyla</emphasis>
</taxonomicName>
;
<treatmentCitation id="D72310BF7C49FF98FDE11B79FC64F887" author="Santos-Silva, A. &amp; S. Shute" box="[603,1007,1862,1886]" page="32" pageId="2" pageNumber="1" year="2009">
<bibRefCitation id="32134B5F7C49FF98FDE11B79FC64F887" author="Santos-Silva, A. &amp; S. Shute" box="[603,1007,1862,1886]" pageId="2" pageNumber="1" pagination="19 - 35" refId="ref2715" refString="Santos-Silva, A., and S. Shute. 2009. The identity of Parandra laevis Latreille, 1804 and nomenclatural changes in the Parandrinae (Coleoptera: Cerambycidae). ZooKeys 25: 19 - 35, 9 fig." type="journal article" year="2009">Santos-Silva and Shute 2009: 32</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="563D36AE7C49FF98FF7A1B59FCC5F8A7" blockId="2.[192,1440,1766,1918]" box="[192,846,1894,1918]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="769211807C49FF98FF7A1B59FCC5F8A7" box="[192,846,1894,1918]" pageId="2" pageNumber="1">
<taxonomicName id="91824D2D7C49FF98FF7A1B59FE63F8A7" ID-CoL="3XQTZ" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[192,488,1894,1918]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C49FF98FF7A1B59FE63F8A7" box="[192,488,1894,1918]" italics="true" pageId="2" pageNumber="1">Malukandra heterostyla</emphasis>
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;
<treatmentCitation id="D72310BF7C49FF98FE4D1B59FCC1F8A7" ID-DOI="http://doi.org/10.5281/zenodo.8400189" ID-Zenodo-Dep="8400189" author="Santos-Silva, A. &amp; D. Heffern &amp; K. Matsuda" box="[503,842,1894,1918]" httpUri="http://treatment.plazi.org/id/975887B7FFEBFFD466D0FBF8168D34B6" page="59" pageId="2" pageNumber="1" year="2010">
<bibRefCitation id="32134B5F7C49FF98FE4D1B59FCC1F8A7" author="Santos-Silva, A. &amp; D. Heffern &amp; K. Matsuda" box="[503,842,1894,1918]" pageId="2" pageNumber="1" pagination="1 - 120" refId="ref2757" refString="Santos-Silva, A., D. Heffern and K. Matsuda. 2010. Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae). Insecta Mundi 0130: 1 - 120." type="book chapter" year="2010">Santos-Silva et al. 2010: 59</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="1E9865257C48FF99FF791CFEFE1BFA03" pageId="3" pageNumber="2" type="description">
<paragraph id="563D36AE7C48FF99FF791CFEFD2CFC03" blockId="3.[195,1443,193,1370]" pageId="3" pageNumber="2">
<emphasis id="64F6EABC7C48FF99FF791CFEFEEDFF00" bold="true" box="[195,358,193,217]" pageId="3" pageNumber="2">Description</emphasis>
. Female: General coloration of integument brown; dorsal surface darker than ventral; distal portion of head, parts of mandibles, margins of pro- meso- and metathorax, of scutellum, of elytra (suture and epipleura), of ventrite V, and parts of legs from dark-brown to blackish; extreme distal of ventrites slightly darker than remaining surface. Body not flattened dorsoventrally (
<figureCitation id="CEB92A2B7C48FF99FABA1D1DFAC2FEE3" box="[1280,1353,290,314]" captionStart="Figures 1–3" captionStartId="4.[192,288,1154,1176]" captionTargetBox="[197,1432,200,1119]" captionTargetId="figure-229@4.[192,1440,192,1124]" captionTargetPageId="4" captionText="Figures 1–3. Malukandra heterostyla, female, habitus. 1) Dorsal. 2) Ventral. 3) Lateral." figureDoi="http://doi.org/10.5281/zenodo.8400311" httpUri="https://zenodo.org/record/8400311/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
). Head elongated behind eyes. Width of head plus eyes equal to 0.9 times that of pronotum at anterolateral angles. Length of mandible equal to 0.7 times that of head; moderately coarsely, sparsely punctate on dorsal surface (sometimes, distinctly coarsely, abundantly punctate on basal half near inner edge), slightly finer towards apex; outer surface vertical, triangularly excavated between base and ventral tooth, moderately coarsely, sparsely punctate on basal half, finer and closer on apical half; pilosity very short, sparse on outer and dorsal surface, somewhat longer near inner edge. Dorsal surface of head coarsely, abundantly punctate, slightly finer towards clypeus and coarser laterally between the eyes and prothorax; sulcus between the gibbosities moderately shallow, enlarged toward clypeus. Clypeus medially in the same level as the area of the sulcus between the gibbosities of the head; laterally slightly lower than centrally; moderately coarsely, sparsely punctate (mainly laterally); pilosity very short, sparse. Clypeolabral suture distinct throughout. Median projection of labrum wide, short, rounded at apex; punctures as in clypeus; pilosity very short, moderately longer on centro-apical region. Ocular carina distinct. Largest width of eyes about 0.6 times length; posterior edge of eyes strongly elevated in relation to the area behind it. Area behind eyes
<emphasis id="64F6EABC7C48FF99FC871EDDFCC7FD23" bold="true" box="[829,844,738,762]" pageId="3" pageNumber="2">c</emphasis>
oarsely punctate near apex of upper ocular lobe, gradually sparser, finer towards gula, mainly after apex of lower ocular lobe. Submentum distinctly separated from genae and gula, slightly tumid between gula and anterior third, then distinctly oblique towards anterior edge, somewhat depressed laterally in this region; surface coarsely, moderately sparsely punctate; anterior edge elevated; pilosity short, sparse, with some long hairs in middle area near anterior edge. Antennae not reaching apex of prothorax; antennomere XI 1.7 times longer than X; ventral sensorial area of antennomeres III-X not divided by carinae and not visible from side; XI divided and distinctly visible from side.
</paragraph>
<paragraph id="563D36AE7C48FF99FF491FDDFC81FA83" blockId="3.[195,1443,193,1370]" pageId="3" pageNumber="2">
Sides of prothorax (
<figureCitation id="CEB92A2B7C48FF99FE5B1FDDFDCFFC23" box="[481,580,994,1018]" captionStart="Figures 1–3" captionStartId="4.[192,288,1154,1176]" captionTargetBox="[197,1432,200,1119]" captionTargetId="figure-229@4.[192,1440,192,1124]" captionTargetPageId="4" captionText="Figures 1–3. Malukandra heterostyla, female, habitus. 1) Dorsal. 2) Ventral. 3) Lateral." figureDoi="http://doi.org/10.5281/zenodo.8400311" httpUri="https://zenodo.org/record/8400311/files/figure.png" pageId="3" pageNumber="2">Fig. 1–2</figureCitation>
) nearly parallel or slightly rounded from the area of the middle lateral angles to anterolateral angles, convergent from the former towards posterolateral angles; anterolateral angles projecting forward, acute (not as pronounced and rounded in
<specimenCount id="4084FD277C48FF99FBBF181DFB06FBE3" box="[1029,1165,1058,1082]" pageId="3" pageNumber="2" type="female">one female</specimenCount>
); posterolateral angles acute, projecting; lateral angles absent. Disc of pronotum almost flat centrally, somewhat coarsely punctate centrally, gradually coarser, more abundantly punctate laterally;
<materialsCitation id="E6EA3CF37C48FF99FB39185DFE75FB00" collectorName="Metasternum" location="Metepisterna" municipality="Elytra" pageId="3" pageNumber="2" specimenCount="1">
anterior edge straight, except near anterolateral angles.
<collectorName id="FB7753787C48FF99FDC818BDFC90FB40" box="[626,795,1154,1177]" pageId="3" pageNumber="2">Metasternum</collectorName>
coarsely, abundantly punctate laterally, gradually finer, sparser towards central area.
<location id="535D60757C48FF99FD38189DFCACFB63" LSID="urn:lsid:plazi:treatment:DE2B87B87C49FF9EFF7A1AB9FD79FA47:535D60757C48FF99FD38189DFCACFB63" box="[642,807,1186,1210]" municipality="Elytra" name="Metepisterna" pageId="3" pageNumber="2">Metepisterna</location>
coarsely, abundantly punctate.
<collectingMunicipality id="B659ACD47C48FF99FB0C189DFA8EFB63" box="[1206,1285,1186,1210]" pageId="3" pageNumber="2">Elytra</collectingMunicipality>
abundantly, coarsely, deeply punctate
</materialsCitation>
; dorsal carinae distinct or almost indistinct. Ventrites I–IV finely, sparsely punctate centrally, gradually coarser, more abundant laterally; ventrite V moderately coarsely punctate throughout, mainly laterally;
<materialsCitation id="E6EA3CF37C48FF99FD90193DFBC2FAE3" collectionCode="V" location="Metatarsomere" pageId="3" pageNumber="2" specimenCount="1">
pilosity very short and sparse, longer and more abundant on centro-apical region of
<collectionCode id="3093AE6B7C48FF99FE8D191DFEC5FAE0" box="[311,334,1314,1337]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V</collectionCode>
.
<location id="535D60757C48FF99FEE0191DFD9CFAE0" LSID="urn:lsid:plazi:treatment:DE2B87B87C49FF9EFF7A1AB9FD79FA47:535D60757C48FF99FEE0191DFD9CFAE0" box="[346,535,1314,1337]" name="Metatarsomere" pageId="3" pageNumber="2">Metatarsomere</location>
I about 1.2 times longer than II–III together
</materialsCitation>
; length of metatarsomere V (without claws) about 0.9 that of I–III together.
</paragraph>
<paragraph id="563D36AE7C48FF99FF7919BEFE1BFA03" blockId="3.[195,1442,1409,1498]" pageId="3" pageNumber="2">
<emphasis id="64F6EABC7C48FF99FF7919BEFD75FA40" bold="true" box="[195,766,1409,1434]" pageId="3" pageNumber="2">
Dimensions in mm (female) (
<specimenCount id="4084FD277C48FF99FDF619BEFD7EFA40" box="[588,757,1409,1433]" pageId="3" pageNumber="2" type="generic">3 specimens</specimenCount>
)
</emphasis>
. Total length (including mandibles), 12.4–16.9; prothorax: length 2.5-3.4; anterior width, 3.1–4.1; posterior width, 2.8–3.8; humeral width, 3.2–4.3; elytral length, 7.7–10.6.
</paragraph>
</subSubSection>
<subSubSection id="1E9865257C48FF99FF791A3EFAF6F943" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="563D36AE7C48FF99FF791A3EFAF6F943" blockId="3.[195,1443,1537,1690]" pageId="3" pageNumber="2">
<emphasis id="64F6EABC7C48FF99FF791A3EFEC0F9C0" bold="true" box="[195,331,1537,1561]" pageId="3" pageNumber="2">Diagnosis</emphasis>
. Females of
<taxonomicName id="91824D2D7C48FF99FE531A3DFC99F9C3" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[489,786,1538,1562]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C48FF99FE531A3DFC99F9C3" box="[489,786,1538,1562]" italics="true" pageId="3" pageNumber="2">Malukandra heterostyla</emphasis>
</taxonomicName>
differ from females of
<taxonomicName id="91824D2D7C48FF99FB951A3DFB65F9C3" authorityName="Santos-Silva, Heffern and Matsuda" authorityYear="2010" box="[1071,1262,1538,1562]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hornabrooki">
<emphasis id="64F6EABC7C48FF99FB951A3DFB65F9C3" box="[1071,1262,1538,1562]" italics="true" pageId="3" pageNumber="2">M. hornabrooki</emphasis>
</taxonomicName>
mainly by the sculpture on the dorsal surface of head, between the eyes, formed by coarse and abundant punctures (fine and very sparse in the latter). Females of
<taxonomicName id="91824D2D7C48FF99FCB01A7DFC52F980" authorityName="Santos-Silva, Heffern and Matsuda" authorityYear="2010" box="[778,985,1602,1625]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jayawijayana">
<emphasis id="64F6EABC7C48FF99FCB01A7DFC52F980" box="[778,985,1602,1625]" italics="true" pageId="3" pageNumber="2">M. jayawijayana</emphasis>
</taxonomicName>
remain unknown, but since males of this species have metatarsomere V slightly longer than I–III together, we believe this is also the case in females. In males and females of
<taxonomicName id="91824D2D7C48FF99FDC71ABDFCA1F943" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[637,810,1666,1690]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C48FF99FDC71ABDFCA1F943" box="[637,810,1666,1690]" italics="true" pageId="3" pageNumber="2">M. heterostyla</emphasis>
</taxonomicName>
metatarsomere V is shorter than I–III together.
</paragraph>
</subSubSection>
<subSubSection id="1E9865257C48FF9EFF791AFEFE64FAC4" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="discussion">
<paragraph id="563D36AE7C48FF9EFF791AFEFE64FAC4" blockId="3.[195,1443,1729,1850]" lastBlockId="4.[192,1440,1221,1309]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<emphasis id="64F6EABC7C48FF99FF791AFEFEDDF900" bold="true" box="[195,342,1729,1753]" pageId="3" pageNumber="2">Comments</emphasis>
. With the specimens examined in this study,
<taxonomicName id="91824D2D7C48FF99FC301AFDFB39F903" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[906,1202,1730,1754]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C48FF99FC301AFDFB39F903" box="[906,1202,1730,1754]" italics="true" pageId="3" pageNumber="2">Malukandra heterostyla</emphasis>
</taxonomicName>
is currently known from
<specimenCount id="4084FD277C48FF99FEB91ADDFE3EF923" box="[259,437,1762,1786]" pageId="3" pageNumber="2" type="generic">five specimens</specimenCount>
(
<specimenCount id="4084FD277C48FF99FE7E1ADDFDCAF923" box="[452,577,1762,1786]" pageId="3" pageNumber="2" type="male">two males</specimenCount>
and
<specimenCount id="4084FD277C48FF99FDC61ADDFCACF923" box="[636,807,1762,1786]" pageId="3" pageNumber="2" type="female">three females</specimenCount>
). At the time of the transfer of
<taxonomicName id="91824D2D7C48FF99FB251ADDFA29F923" authorityName="Lameere" authorityYear="1902" box="[1183,1442,1762,1786]" class="Insecta" family="Cerambycidae" genus="Parandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C48FF99FB251ADDFA29F923" box="[1183,1442,1762,1786]" italics="true" pageId="3" pageNumber="2">Parandra heterostyla</emphasis>
</taxonomicName>
to
<taxonomicName id="91824D2D7C48FF99FF591B3DFEF4F8C3" authorityName="Santos-Silva, Heffern &amp; Matsuda" authorityYear="2010" box="[227,383,1794,1818]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="64F6EABC7C48FF99FF591B3DFEF4F8C3" box="[227,383,1794,1818]" italics="true" pageId="3" pageNumber="2">Malukandra</emphasis>
</taxonomicName>
, only
<specimenCount id="4084FD277C48FF99FE761B3DFDC0F8C3" box="[460,587,1794,1818]" pageId="3" pageNumber="2" type="male">two males</specimenCount>
were known. Now we had the opportunity to examine
<specimenCount id="4084FD277C48FF99FB401B3DFA28F8C3" box="[1274,1443,1794,1818]" pageId="3" pageNumber="2" type="female">three females</specimenCount>
that belong to the species. Although we have examined a great number of specimens of
<taxonomicName id="91824D2D7C48FF99FABD1B1DFA28F8E3" box="[1287,1443,1826,1850]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subFamily" subFamily="Parandrinae">Parandrinae</taxonomicName>
from the Oriental Region in recent years, few specimens were from
<collectingRegion id="9446F84C7C4FFF9EFC4E18FAFBECFB04" box="[1012,1127,1221,1245]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="3">Sulawesi</collectingRegion>
. Thus, it is not possible to exclude the possibility of the occurrence of
<taxonomicName id="91824D2D7C4FFF9EFD7018D9FCFDFB24" authorityName="Lameere" authorityYear="1902" box="[714,886,1253,1277]" class="Insecta" family="Cerambycidae" genus="Parandra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C4FFF9EFD7018D9FCFDFB24" box="[714,886,1253,1277]" italics="true" pageId="4" pageNumber="3">M. heterostyla</emphasis>
</taxonomicName>
on this island, mainly because of the proximity of Halmahera Island.
</paragraph>
</subSubSection>
<caption id="02FD66267C4FFF9EFF7A18BDFB17FB41" ID-DOI="http://doi.org/10.5281/zenodo.8400311" ID-Zenodo-Dep="8400311" box="[192,1180,1150,1180]" httpUri="https://zenodo.org/record/8400311/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,288,1154,1176]" targetBox="[197,1432,200,1119]" targetPageId="4">
<paragraph id="563D36AE7C4FFF9EFF7A18BDFB17FB41" blockId="4.[192,1180,1150,1180]" box="[192,1180,1150,1180]" pageId="4" pageNumber="3">
<emphasis id="64F6EABC7C4FFF9EFF7A18BDFED2FB4E" bold="true" box="[192,345,1150,1180]" pageId="4" pageNumber="3">Figures 1–3.</emphasis>
<taxonomicName id="91824D2D7C4FFF9EFEE518BDFDE1FB4E" baseAuthorityName="Lameere" baseAuthorityYear="1902" box="[351,618,1154,1175]" class="Insecta" family="Cerambycidae" genus="Malukandra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="heterostyla">
<emphasis id="64F6EABC7C4FFF9EFEE518BDFDE1FB4E" box="[351,618,1154,1175]" italics="true" pageId="4" pageNumber="3">Malukandra heterostyla</emphasis>
</taxonomicName>
, female, habitus.
<emphasis id="64F6EABC7C4FFF9EFC8A18BDFCCCFB41" bold="true" box="[816,839,1154,1176]" pageId="4" pageNumber="3">1)</emphasis>
Dorsal.
<emphasis id="64F6EABC7C4FFF9EFC1F18BDFC30FB41" bold="true" box="[933,955,1154,1176]" pageId="4" pageNumber="3">2)</emphasis>
Ventral.
<emphasis id="64F6EABC7C4FFF9EFB9F18BDFBB0FB41" bold="true" box="[1061,1083,1154,1176]" pageId="4" pageNumber="3">3)</emphasis>
Lateral.
</paragraph>
</caption>
<subSubSection id="1E9865257C4FFF9EFF7A197AFD79FA47" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="563D36AE7C4FFF9EFF7A197AFD79FA47" blockId="4.[192,1440,1349,1438]" pageId="4" pageNumber="3">
<emphasis id="64F6EABC7C4FFF9EFF7A197AFE4DFA84" bold="true" box="[192,454,1349,1373]" pageId="4" pageNumber="3">Material examined</emphasis>
.
<materialsCitation id="E6EA3CF37C4FFF9EFE6C197AFD66FA47" collectingDate="1981-02-01" collectingDateMax="1981-02-14" collectingDateMin="1981-02-01" collectionCode="USNM, MZSP" collectorName="Tg. Loleo &amp; A. C. Messer &amp; P. M. Taylor" country="Indonesia" county="Pan-Tunggal Lumber Co." location="Camp" municipality="Halmahera Island" pageId="4" pageNumber="3" specimenCount="3" specimenCount-female="3" stateProvince="Maluku Utara">
<emphasis id="64F6EABC7C4FFF9EFE6C197AFD0AFA84" bold="true" box="[470,641,1349,1373]" pageId="4" pageNumber="3">
<collectingCountry id="2E95763E7C4FFF9EFE6C197AFD0AFA84" box="[470,641,1349,1373]" name="Indonesia" pageId="4" pageNumber="3">INDONESIA</collectingCountry>
</emphasis>
,
<emphasis id="64F6EABC7C4FFF9EFD2B197AFCCFFA84" box="[657,836,1349,1373]" italics="true" pageId="4" pageNumber="3">
<collectingRegion id="9446F84C7C4FFF9EFD2B197AFCCFFA84" box="[657,836,1349,1373]" country="Indonesia" name="Maluku Utara" pageId="4" pageNumber="3">North Maluku</collectingRegion>
</emphasis>
:
<collectingMunicipality id="B659ACD47C4FFF9EFCE9197AFBBDFA84" box="[851,1078,1349,1373]" pageId="4" pageNumber="3">Halmahera Island</collectingMunicipality>
(“kao Dist.,
<location id="535D60757C4FFF9EFB691979FA97FA87" LSID="urn:lsid:plazi:treatment:DE2B87B87C49FF9EFF7A1AB9FD79FA47:535D60757C4FFF9EFB691979FA97FA87" box="[1235,1308,1350,1374]" country="Indonesia" county="Pan-Tunggal Lumber Co." municipality="Halmahera Island" name="Camp" pageId="4" pageNumber="3" stateProvince="Maluku Utara">Camp</location>
34, 34 km inland from
<collectingCounty id="BF5C4E227C4FFF9EFEE0195AFD1DFAA7" box="[346,662,1381,1406]" pageId="4" pageNumber="3">Pan-Tunggal Lumber Co.</collectingCounty>
; base camp at
<collectorName id="FB7753787C4FFF9EFCF41959FC4FFAA4" box="[846,964,1381,1406]" pageId="4" pageNumber="3">Tg. Loleo</collectorName>
”),
<specimenCount id="4084FD277C4FFF9EFC591959FB84FAA4" box="[995,1039,1382,1406]" pageId="4" pageNumber="3" type="female">3 ♀</specimenCount>
,
<date id="223C106E7C4FFF9EFBA41959FB3EFAA7" box="[1054,1205,1382,1406]" pageId="4" pageNumber="3" value="1981-02-01" valueMax="1981-02-14" valueMin="1981-02-01">
<collectingDate id="3278E9867C4FFF9EFBA41959FB3EFAA7" box="[1054,1205,1382,1406]" pageId="4" pageNumber="3" value="1981-02-01" valueMax="1981-02-14" valueMin="1981-02-01">1-14.II.1981</collectingDate>
</date>
,
<collectorName id="FB7753787C4FFF9EFB791959FAEBFAA4" box="[1219,1376,1382,1406]" pageId="4" pageNumber="3">A. C. Messer</collectorName>
&amp;
<collectorName id="FB7753787C4FFF9EFA3D1959FEB2FA44" pageId="4" pageNumber="3">P. M. Taylor</collectorName>
coll. (2 from
<collectionCode id="3093AE6B7C4FFF9EFE6319B9FDBAFA47" box="[473,561,1414,1438]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="3" type="Museum">USNM</collectionCode>
; 1 from
<collectionCode id="3093AE6B7C4FFF9EFD2F19B9FD6DFA47" box="[661,742,1414,1438]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="3" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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