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<mods:title id="6810A8D40AA2E3BE3843C0D4F30C89D4">Notes on the taxonomic status and distribution of some Cylindrotomidae (Diptera, Tipuloidea), with emphasis on Japanese species</mods:title>
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<treatment id="F9511990009B516C993E0AF41571BD3C" ID-GBIF-Taxon="192586634" LSID="urn:lsid:plazi:treatment:F9511990009B516C993E0AF41571BD3C" httpUri="http://treatment.plazi.org/id/F9511990009B516C993E0AF41571BD3C" lastPageNumber="13" pageId="0" pageNumber="13" scope_family="Cylindrotomidae" scope_order="Diptera">
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<taxonomicName id="373FD7CEC4501F7131C9800873B10AFB" ID-CoL="3V67C" LSID="F9511990-009B-516C-993E-0AF41571BD3C" authority="(Alexander, 1919)" baseAuthorityName="Alexander" baseAuthorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">Liogma mikado (Alexander, 1919)</taxonomicName>
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<figureCitation id="40AC416CCCD85984075DBBFCD34E232E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">Figs 4D</figureCitation>
<figureCitation id="7437F8BF11E0565F1BE82DA22C57042A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of &quot; Triogma kuwanai &quot; form H Triogma kuwanai (Alexander, 1913) of &quot; Triogma limbinervis &quot; form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">, 5D</figureCitation>
<figureCitation id="BBE3B85C000B42794BC0C7C404E4291C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Occurrence data in Japan and surrounding areas of A Liogma brevipecten Alexander, 1932 B Liogma mikado (Alexander, 1919). Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data." figureDoi="10.3897/zookeys.1083.75624.figure22" httpUri="https://binary.pensoft.net/fig/637495" pageId="0" pageNumber="13">, 22B</figureCitation>
<figureCitation id="DABB0A26C24F26210134E627F3E1C1C8" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">, 23</figureCitation>
<figureCitation id="8DCB75FD30F57AB3D3DA82F04DE7DDC3" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">, 24</figureCitation>
<figureCitation id="CC6BB7F11D18CB2BF7276F316781A776" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">, 25</figureCitation>
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<taxonomicName id="B0DE5F4F349D43A50086F42A1D3E6D5B" authorityName="Alexander" authorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">Phalacrocera mikado</taxonomicName>
in
<bibRefCitation id="AA44A2D00F508E1C46CC01652A9BEB3E" DOI="https://doi.org/10.1093/aesa/12.4.327" author="Alexander, CP" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="13" pagination="327 - 348" refId="B2" refString="Alexander, CP, 1919. Undescribed species of Japanese crane-flies (Tipulidae, Diptera). Annals of the Entomological Society of America 12: 327 - 348, DOI: https://doi.org/10.1093/aesa/12.4.327" title="Undescribed species of Japanese crane-flies (Tipulidae, Diptera)." url="https://doi.org/10.1093/aesa/12.4.327" volume="12" year="1919">Alexander 1919</bibRefCitation>
: 346: original description;
<bibRefCitation id="9A18EE461CBAB5904A00C789712B70C2" author="Alexander, CP" journalOrPublisher="Genera Insectorum" pageId="0" pageNumber="13" pagination="1 - 16" refId="B6" refString="Alexander, CP, 1928. Diptera. Family Tipulidae, Subfam. Cylindrotominae. Genera Insectorum 187: 1 - 16" title="Diptera. Family Tipulidae, Subfam. Cylindrotominae." volume="187" year="1928">Alexander 1928</bibRefCitation>
: 10: distribution, illustration;
<bibRefCitation id="F4B5D1BB8ED42622AB377677A1387BCD" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander 1953a</bibRefCitation>
: 57: faunistic record;
<bibRefCitation id="89AF42A26135AEA30BEA0A8E7E0D6F74" author="Ishida, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="13" refId="B27" refString="Ishida, H, 1955. The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera. II. Tipula (part) and Cylindrotominae. Annual Report of the Hyogo Agricultural College (Kenkyu Shuroku, 1955) 5: 59-77." title="The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera. II. Tipula (part) and Cylindrotominae. Annual Report of the Hyogo Agricultural College (Kenkyu Shuroku, 1955) 5: 59 - 77." year="1955">Ishida 1955</bibRefCitation>
: 77: distribution.
</paragraph>
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<taxonomicName id="159A714E2E5707BB453EE79F3223D1A2" baseAuthorityName="Alexander" baseAuthorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">Liogma mikado</taxonomicName>
in
<bibRefCitation id="83722E71147F425A7921D33C62997FA1" author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
: 85-90: new combination, distribution, faunistic records, larva and pupa description, illustrations;
<bibRefCitation id="3D7D1E9FD8BAF089A8391E1172F44CBD" author="Sidorenko, VS" editor="Ler, PA" journalOrPublisher="Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="68 - 70" refId="B58" refString="Sidorenko, VS, 1999. Cylindrotomidae. In: Ler, PA, Ed., Key to the insects of Russian Far East. Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok: 68 - 70" title="Cylindrotomidae." volumeTitle="Key to the insects of Russian Far East." year="1999">Sidorenko 1999</bibRefCitation>
: 68-70: identification key, illustration, distribution;
<bibRefCitation id="10D913C2D14D9A3014AC231996A3CBFE" author="Nakamura, T" journalOrPublisher="Bulletin of the Tochigi Prefectural Museum" pageId="0" pageNumber="13" pagination="23 - 30" refId="B35" refString="Nakamura, T, 2001. Cylindrotomidae of Tochigi Prefecture, Japan (Diptera, Tipulidae). Bulletin of the Tochigi Prefectural Museum 18: 23 - 30" title="Cylindrotomidae of Tochigi Prefecture, Japan (Diptera, Tipulidae)." volume="18" year="2001">Nakamura 2001</bibRefCitation>
: 23-29: identification key, illustration, distribution, faunistic records;
<bibRefCitation id="124C5F63781193B9423F53C118F88E01" author="Paramonov, NM" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" refId="B41" refString="Paramonov, NM, 2004b. To the fauna Cylindrotomidae (Diptera, Nematocera) of the Asian part of Russia. Tomsk State University Journal of Biology 11: 69." title="To the fauna Cylindrotomidae (Diptera, Nematocera) of the Asian part of Russia. Tomsk State University Journal of Biology 11: 69." year="2004 b">Paramonov 2004b</bibRefCitation>
: 69: faunistic record;
<bibRefCitation id="4092ECBED4DD341BF558AAC464205DAF" author="Paramonov, NM" editor="Leley, AS" journalOrPublisher="6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="887 - 889" refId="B43" refString="Paramonov, NM, 2006. Cylindrotomidae. In: Leley, AS, Ed., Key to the Insects of Russian Far East. 6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok: 887 - 889" title="Cylindrotomidae." volumeTitle="Key to the Insects of Russian Far East." year="2006">Paramonov 2006</bibRefCitation>
: 888-889: identification key, distribution;
<bibRefCitation id="B8DDE77CF47EDF912F9898FF477FFB16" author="Nakamura, T" journalOrPublisher="Tokai University Press, Kanawaga" pageId="0" pageNumber="13" refId="B37" refString="Nakamura, T, 2014. Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." title="Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." year="2014">Nakamura 2014</bibRefCitation>
: 54: distribution;
<bibRefCitation id="F8859B1EB0D5E508E9D402EB3A44F829" author="Kato, D" journalOrPublisher="Makunagi / Acta Dipterologica" pageId="0" pageNumber="13" pagination="7 - 24" refId="B29" refString="Kato, D, Suzuki, Y, 2017. A revised list of the crane flies of Kanagawa, Japan, with newly recorded species (Diptera, Tipuloidea). Makunagi/Acta Dipterologica 28: 7 - 24" title="A revised list of the crane flies of Kanagawa, Japan, with newly recorded species (Diptera, Tipuloidea)." volume="28" year="2017">Kato and Suzuki 2017</bibRefCitation>
: 16: distribution;
<bibRefCitation id="3334218EAB76FDA67D748B5CF71BE9D5" DOI="https://doi.org/10.33910/2686-9519-2019-11-2-119-125" author="Paramonov, NM" journalOrPublisher="Amurian Zoological Journal" pageId="0" pageNumber="13" pagination="119 - 125" refId="B44" refString="Paramonov, NM, 2019. Superfamily Tipuloidea (Diptera), new taxon for Jewish Autonomous Region of Russia. Amurian Zoological Journal 11: 119 - 125, DOI: https://doi.org/10.33910/2686-9519-2019-11-2-119-125" title="Superfamily Tipuloidea (Diptera), new taxon for Jewish Autonomous Region of Russia." url="https://doi.org/10.33910/2686-9519-2019-11-2-119-125" volume="11" year="2019">Paramonov 2019</bibRefCitation>
: 120: faunistic data;
<bibRefCitation id="A419059F6D6215FC42499A5DB3423C1B" DOI="https://doi.org/10.1093/zoolinnean/zlaa177" author="Imada, Y" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="13" pagination="1156 - 1204" refId="B26" refString="Imada, Y, 2020. Moss mimesis par excellence: integrating previous and new data on the life history and larval ecomorphology of long-bodied craneflies (Diptera: Cylindrotomidae: Cylindrotominae). Zoological Journal of the Linnean Society 193 (4): 1156 - 1204, DOI: https://doi.org/10.1093/zoolinnean/zlaa177" title="Moss mimesis par excellence: integrating previous and new data on the life history and larval ecomorphology of long-bodied craneflies (Diptera: Cylindrotomidae: Cylindrotominae)." url="https://doi.org/10.1093/zoolinnean/zlaa177" volume="193" year="2020">Imada 2020</bibRefCitation>
: biology and ecology of larvae;
<bibRefCitation id="47E0147797ADA8ADA159C95E806F5C2C" author="Kim, J" journalOrPublisher="Entomological Research Bulletin" pageId="0" pageNumber="13" pagination="89 - 91" refId="B30" refString="Kim, J, Bae, YJ, 2020. First Record of the Genus Liogma (Diptera: Cylindrotomidae) in Korea. Entomological Research Bulletin 36 (2): 89 - 91" title="First Record of the Genus Liogma (Diptera: Cylindrotomidae) in Korea." volume="36" year="2020">Kim and Bae 2020</bibRefCitation>
: distribution.
</paragraph>
</subSubSection>
<subSubSection id="9C913731F23A5A9EC031A0051E5097AA" pageId="0" pageNumber="13" type="type material">
<paragraph id="DB89542AEEDBB909F3CED74C9D8F0486" pageId="0" pageNumber="13">Type material examined.</paragraph>
<paragraph id="A11FE5C8E86DF1F85D16AACEBBAA0136" pageId="0" pageNumber="13">
<taxonomicName id="B3A191D4F2054A8A8D4D606B57BD259E" authorityName="Alexander" authorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">
<emphasis id="1CC32AB6FEDA200D6C7B1A63CBF96126" italics="true" pageId="0" pageNumber="13">Phalacrocera mikado</emphasis>
</taxonomicName>
Alexander:
<emphasis id="81E49A2E0B3B274665A901DBBB8EF905" bold="true" italics="true" pageId="0" pageNumber="13">ALLOTYPE</emphasis>
♂: •
<emphasis id="616DECDCDC7A3F43468F37E6C696D8D0" bold="true" pageId="0" pageNumber="13">Japan</emphasis>
, Tokyo, Tokyo metropolis, 1919.04.?, leg. R. Takahashi (USNM).
</paragraph>
</subSubSection>
<subSubSection id="8DD2D2D7B62BE8B2CB19CC430B2AD848" pageId="0" pageNumber="13" type="type material">
<paragraph id="4CCAB4EAD5C4AB6920E2E9952E9D9DDB" pageId="0" pageNumber="13">Non-type material examined.</paragraph>
<paragraph id="3AF20FA4BB7B44AACC9EB0F057EAF722" pageId="0" pageNumber="13">
<emphasis id="300D051988A14DFFDEA02EC741F37817" bold="true" pageId="0" pageNumber="13">Japan</emphasis>
• 2 ♀; Aichi, Toyota, Kawashimo, triburary of Yahagi River;
<geoCoordinate id="2F568D173EE1DBF44B85402C1EF26384" degrees="35.20376" direction="north" orientation="latitude" precision="1" value="35.20376">35.20376°N</geoCoordinate>
,
<geoCoordinate id="C57A776FC24190D93E73A7975B721455" degrees="137.3012" direction="east" orientation="longitude" precision="5" value="137.3012">137.3012°E</geoCoordinate>
; alt. 140 m; 4 May. 2014; D. Kato leg.; BLKU. • 1 ♀; Aichi, Seto, Iwaya-cho, near Iwayada Park;
<geoCoordinate id="F04BDE92C2D7081AE020696E08875CF8" degrees="35.23957" direction="north" orientation="latitude" precision="1" value="35.23957">35.23957°N</geoCoordinate>
,
<geoCoordinate id="10E7DE1D70018FFFE6AE2A4DC06A9A6E" degrees="137.15084" direction="east" orientation="longitude" precision="1" value="137.15083">137.15084°E</geoCoordinate>
; alt. 300 m; 4 May. 2016; D. Kato leg.; BLKU. • 1 ♀; Aichi, Seto, Minamiazuma;
<geoCoordinate id="95A06563CDB933A446536309F97F0B8A" degrees="35.223213" direction="north" orientation="latitude" precision="1" value="35.223213">35.223213°N</geoCoordinate>
,
<geoCoordinate id="1A9DE3166724E3E5B6F5DDF3D448EBF0" degrees="137.1131" direction="east" orientation="longitude" precision="5" value="137.1131">137.1131°E</geoCoordinate>
; alt. 150 m; 5 May. 2014; D. Kato leg.; BLKU. • 3 ♀; Aomori, Hirosaki, Koguriyama, Inekari River;
<geoCoordinate id="9AC8D7A89BFB8C67D1EB81E8DFC4BE72" degrees="40.53658" direction="north" orientation="latitude" precision="1" value="40.53658">40.53658°N</geoCoordinate>
,
<geoCoordinate id="4194D52E3557EE5A97135C207083678F" degrees="140.48701" direction="east" orientation="longitude" precision="1" value="140.48701">140.48701°E</geoCoordinate>
; alt. 170 m; 28 May 2013; • 1 ♂; same locality; 31 May. 2013; D. Kato leg.; BLKU. • 1 ♂; Aomori, Fukaura, Mt. Takanio;
<geoCoordinate id="355DC9B827C4FAC923A153CCB3DB5CF0" degrees="40.68993" direction="north" orientation="latitude" precision="1" value="40.68993">40.68993°N</geoCoordinate>
,
<geoCoordinate id="21704F7589AD0E7D885B74F59E041B2F" degrees="140.10285" direction="east" orientation="longitude" precision="1" value="140.10284">140.10285°E</geoCoordinate>
; alt. 140 m; 11 May. 2014; D. Kato leg.; BLKU. • 1 ♂; Aomori, Hirosaki, Soma Path;
<geoCoordinate id="5E0B2412CE85E1D5F8194A2F2F11473E" degrees="40.49479" direction="north" orientation="latitude" precision="1" value="40.49479">40.49479°N</geoCoordinate>
,
<geoCoordinate id="53D8D71D483D3718FAC74F124FFD5D37" degrees="140.40231" direction="east" orientation="longitude" precision="1" value="140.40231">140.40231°E</geoCoordinate>
; alt. 392 m; 31 May. 2013; D. Kato leg.; BLKU. • Ehime, Kumakogen, River Myogadani springs, 1275 m,
<geoCoordinate id="1FAB3574C9E75218EC4B1F89B649927F" degrees="33.55808" direction="north" orientation="latitude" precision="1" value="33.55808">33.55808°N</geoCoordinate>
,
<geoCoordinate id="8A94F5F9079B0C3DD1F7AA6ED1E39D65" degrees="132.93805" direction="east" orientation="longitude" precision="1" value="132.93805">132.93805°E</geoCoordinate>
, 2019.05.19, 2 ♂ 1 ♀, L.-P.
<normalizedToken id="182F969C0541F789963FE346EFF2B67E" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 5 ♂, 11 ♀; Ehime, Wakayama, Mount Ishizuchi;
<geoCoordinate id="218DB211F904E83E7C6F57FBC65468BE" degrees="33.76491" direction="north" orientation="latitude" precision="1" value="33.76491">33.76491°N</geoCoordinate>
,
<geoCoordinate id="974B42AE151BE126FF37C711D56C5627" degrees="133.12948" direction="east" orientation="longitude" precision="1" value="133.12949">133.12948°E</geoCoordinate>
; alt. 1600 m; 5 Jul. 2019; L.-P.
<normalizedToken id="D8597B7F45BFEDA99287AA7480ED33EB" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 1 ♂; Ehime, Wakayama, small waterfall and stream;
<geoCoordinate id="A1FE30EDB6FFD073C24C7E586EE295E1" degrees="33.74519" direction="north" orientation="latitude" precision="1" value="33.74519">33.74519°N</geoCoordinate>
,
<geoCoordinate id="3608077D5CF32F5AD56CB0B34DAE4779" degrees="133.13714" direction="east" orientation="longitude" precision="1" value="133.13715">133.13714°E</geoCoordinate>
; alt. 1305 m; 18 May. 2019; L.-P.
<normalizedToken id="9794040ED1EA0B5607E1D0EF84F9304C" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 2 ♀; Ehime, Wakayama, small waterfall;
<geoCoordinate id="55FDF85FBBE8A78FB809FE99A9068AAF" degrees="33.71591" direction="north" orientation="latitude" precision="1" value="33.71591">33.71591°N</geoCoordinate>
,
<geoCoordinate id="AF6678B6D32860157BF9F040844A6449" degrees="133.10839" direction="east" orientation="longitude" precision="1" value="133.10838">133.10839°E</geoCoordinate>
; alt. 930 m; 18 May. 2019; L.-P.
<normalizedToken id="E64343C458BA8B5398C6E39C4F3533DB" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 2 ♂; Fukuoka, Fukuoka, Sawara-ku, Itaya, Mt. Sefuri;
<geoCoordinate id="E3262AB244F932F90468BEB2964D5ED0" degrees="33.43811" direction="north" orientation="latitude" precision="1" value="33.43811">33.43811°N</geoCoordinate>
,
<geoCoordinate id="A776ABCE3A5800F4259AE43D15B11B59" degrees="130.36673" direction="east" orientation="longitude" precision="1" value="130.36673">130.36673°E</geoCoordinate>
; alt. 970 m; 2 May. 2015; • 1 ♂; same locality; 13 May. 2015; D. Kato leg.; BLKU. • 1 ♂; Fukuoka, Miyako, Saigawa-Hobashira, Notoge Pass;
<geoCoordinate id="E8D0A3EF0123285C957ED6CAF56A9AD8" degrees="33.49565" direction="north" orientation="latitude" precision="1" value="33.49565">33.49565°N</geoCoordinate>
,
<geoCoordinate id="AE53C01B654BBCE7F0304115C176E4AA" degrees="130.96156" direction="east" orientation="longitude" precision="1" value="130.96156">130.96156°E</geoCoordinate>
; alt. 740 m; 22 Apr. 2016; D. Kato leg.; BLKU. • 1 ♀; Fukuoka, Soeda, rocky streem and moss covered cliff;
<geoCoordinate id="CD5E19CB7EEF7E178ED609D75977D34D" degrees="33.48309" direction="north" orientation="latitude" precision="1" value="33.48309">33.48309°N</geoCoordinate>
,
<geoCoordinate id="6776EA2D5069B26592857A493BB93C6B" degrees="130.93289" direction="east" orientation="longitude" precision="1" value="130.93289">130.93289°E</geoCoordinate>
; alt. 900 m; 21 May. 2019; L.-P.
<normalizedToken id="9AE3A8D99AC8E2A544F7B0E38B940D37" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 2 ♂; Ishikawa, Hakusan, near to Hakusan National Park;
<geoCoordinate id="C4C64EB318BBFD735010DFA4F0C85DBC" degrees="36.25869" direction="north" orientation="latitude" precision="1" value="36.25869">36.25869°N</geoCoordinate>
,
<geoCoordinate id="5CB052D7AFF0FFE33CFC157F6CA87466" degrees="136.72558" direction="east" orientation="longitude" precision="1" value="136.72559">136.72558°E</geoCoordinate>
; alt. 678 m; 27 May. 2015; M. Kato leg.; CYI. • 1 ♀; Iwate, Nishiwaga, Mahirudake;
<geoCoordinate id="C54F69019C3F0F9D98EC812EE852A8CB" degrees="39.46511" direction="north" orientation="latitude" precision="1" value="39.46511">39.46511°N</geoCoordinate>
,
<geoCoordinate id="D1E1FAD0D0169F955242287DCB0B04FE" degrees="140.69365" direction="east" orientation="longitude" precision="1" value="140.69365">140.69365°E</geoCoordinate>
; alt. 900 m; 19 Jun. 2015; Y. Imada leg.; CYI. • 1 ♂; Niigata, Echigo, Sugatani, Kitakanbara;
<geoCoordinate id="5F78BF562A6CFE690FD2B875B8A8004F" degrees="37.84" direction="north" orientation="latitude" precision="555" value="37.84">37.84°N</geoCoordinate>
,
<geoCoordinate id="2E6013FDE3E4EB69DCE28302BEEF469E" degrees="139" direction="east" orientation="longitude" precision="55555" value="139.0">139°E</geoCoordinate>
; 8 May. 1955; H. Koike leg.; USNM. • 1 ♀; Saitama, Ogano, Mt. Futago;
<geoCoordinate id="AE21AB170CA421CFAFC6B4A4E98BC668" degrees="36.06994" direction="north" orientation="latitude" precision="1" value="36.06994">36.06994°N</geoCoordinate>
,
<geoCoordinate id="D11581641FFE624EE6DF4587395DBCBF" degrees="138.86753" direction="east" orientation="longitude" precision="1" value="138.86752">138.86753°E</geoCoordinate>
; alt. 942 m; larva collected: 28 Nov. 2014, emerged 15 Dec. 2014; M. Kato leg.; CYI. • 1 ♀; Shizuoka, Aoi-ku, Umegashima, Akamizu;
<geoCoordinate id="D3DE32087C3BB334E2D373A6190D6AA2" degrees="35.27455" direction="north" orientation="latitude" precision="1" value="35.27455">35.27455°N</geoCoordinate>
,
<geoCoordinate id="644D6EB63E2D87BF1833E5E0EE2A5B66" degrees="138.32731" direction="east" orientation="longitude" precision="1" value="138.32732">138.32731°E</geoCoordinate>
; alt. 680 m; larva collected: 8 Jan. 2007, emerged: 22 Feb. 2007; leg. Y. Sato EUMJ. • 9 ♂; Shizuoka, Shizuoka, Hatanagi;
<geoCoordinate id="BA81981499563F744DEFF2165347A819" degrees="35.2976" direction="north" orientation="latitude" precision="5" value="35.2976">35.2976°N</geoCoordinate>
,
<geoCoordinate id="063E308F46C8F68A414EE2C3DCD57029" degrees="138.21557" direction="east" orientation="longitude" precision="1" value="138.21558">138.21557°E</geoCoordinate>
; alt. 828 m; 12 May. 2013; M. Kato leg.; CYI. • 1 ♂; Shizuoka, Shizuoka, Abenoootaki;
<geoCoordinate id="09DB2A360E16F0B49349829D07AAD39D" degrees="35.30031" direction="north" orientation="latitude" precision="1" value="35.30031">35.30031°N</geoCoordinate>
,
<geoCoordinate id="DB3BDE47BCE9FFE8B67DA736158117BA" degrees="138.35084" direction="east" orientation="longitude" precision="1" value="138.35085">138.35084°E</geoCoordinate>
; alt. 930 m; larva collected: 15 Jan. 2014, emerged: 19 Apr. 2014; M. Kato leg.; CYI. • 8 ♂, 1 ♀; Shizuoka, Ikawa-touge;
<geoCoordinate id="8AABD3811B1D28EE9012EDAC28D942E1" degrees="35.24094" direction="north" orientation="latitude" precision="1" value="35.24094">35.24094°N</geoCoordinate>
,
<geoCoordinate id="DEA656748A7C6FC9A14F5ED9E6FD68F6" degrees="138.28156" direction="east" orientation="longitude" precision="1" value="138.28156">138.28156°E</geoCoordinate>
; alt. 1471 m; 10 May. 2015; M. Kato / Y. Imada leg.; CYI. • 1 ♂; Tokushima, Awa, Mt. Tsurugi;
<geoCoordinate id="6AC84D540A67264DDAB93A9D442115AA" degrees="33.87" direction="north" orientation="latitude" precision="555" value="33.87">33.87°N</geoCoordinate>
,
<geoCoordinate id="078B9F0E49204F74ACECBC1FE2E1BE71" degrees="134.11" direction="east" orientation="longitude" precision="555" value="134.11">134.11°E</geoCoordinate>
; 30 May. 1950; Issiki-Ito leg.; USNM. • 1 ♂; Tokushima, Mima, Koyadaira;
<geoCoordinate id="1777FBAB4570AA45869B433B9D80EA01" degrees="33.87543" direction="north" orientation="latitude" precision="1" value="33.87543">33.87543°N</geoCoordinate>
,
<geoCoordinate id="15F9D232C4DCBF5FB25AF16164C1C794" degrees="134.09571" direction="east" orientation="longitude" precision="1" value="134.0957">134.09571°E</geoCoordinate>
; alt. 1340 m; 30 Apr. 2016; D. Kato leg.; BLKU. • 1 ♂; Tokushima, Miyoshi, Higashiiya-Sugeoi, near Nagoro Dam;
<geoCoordinate id="A2FDC69B8BB6DC74722372A89D3D446A" degrees="33.85182" direction="north" orientation="latitude" precision="1" value="33.85182">33.85182°N</geoCoordinate>
,
<geoCoordinate id="9E07088DD5A7415149BC5DD989744A2C" degrees="134.0234" direction="east" orientation="longitude" precision="5" value="134.0234">134.0234°E</geoCoordinate>
; 29 Apr. 2016; D. Kato leg.; BLKU. • 2 ♂; Tokyo, Mt. Mitake;
<geoCoordinate id="EA6081E9068CE240A46650AC1C1EB526" degrees="35.78" direction="north" orientation="latitude" precision="555" value="35.78">35.78°N</geoCoordinate>
,
<geoCoordinate id="2592F268ECC30BC96586AF84C369BD93" degrees="139.14" direction="east" orientation="longitude" precision="555" value="139.14">139.14°E</geoCoordinate>
; 10 May. 1931; B. Oda leg.; USNM. • 1 sex unknown; Tokyo, Mt. Takao;
<geoCoordinate id="F05BB3CD373EE4427F2BBA645B270369" degrees="35.62" direction="north" orientation="latitude" precision="555" value="35.62">35.62°N</geoCoordinate>
,
<geoCoordinate id="69AE1061F2F33FA05C9F7AE3791C2849" degrees="139.24" direction="east" orientation="longitude" precision="555" value="139.24">139.24°E</geoCoordinate>
; alt. 300-600 m; 7 May. 1922; Esaki leg.; USNM. • 1 ♀; Tokyo, Tokyo, Akiruno, rocky river and stream;
<geoCoordinate id="228DCBD6FC8F7A70CA6BE95AB3F4E7A3" degrees="35.74766" direction="north" orientation="latitude" precision="1" value="35.74766">35.74766°N</geoCoordinate>
,
<geoCoordinate id="20C6BCB7547918FE087A89A05A90E91D" degrees="139.18466" direction="east" orientation="longitude" precision="1" value="139.18466">139.18466°E</geoCoordinate>
; alt. 288 m; 11 May. 2019; L.-P.
<normalizedToken id="381CC53DB18C11E699D1800278F3092D" originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 1 ♂; Tottori, Mt. Daisen;
<geoCoordinate id="7DD41AF32C86B808DE44A9FB4EA51DCA" degrees="35.38" direction="north" orientation="latitude" precision="555" value="35.38">35.38°N</geoCoordinate>
,
<geoCoordinate id="CE7B764C57F67483DC27611F5AED54D6" degrees="133.54" direction="east" orientation="longitude" precision="555" value="133.54">133.54°E</geoCoordinate>
; 7 Jun. 1930; Hibi leg.; USNM. • 1 ♂; Yamagata, Iide, Mt. Iide;
<geoCoordinate id="C6E01D9C7F935048706262756A5DD89E" degrees="37.85122" direction="north" orientation="latitude" precision="1" value="37.85122">37.85122°N</geoCoordinate>
,
<geoCoordinate id="7E2B1E411D1041C11950C1E87B215160" degrees="139.78064" direction="east" orientation="longitude" precision="1" value="139.78064">139.78064°E</geoCoordinate>
, alt. 600 m; 23 May. 2015; Y. Imada leg.; CYI. • 2 ♂, 1 ♀; Yamagata, Oguni, Nukumidaira;
<geoCoordinate id="4BFAE926CA018CDFC9D8CAFB0554DEFF" degrees="37.92293" direction="north" orientation="latitude" precision="1" value="37.92293">37.92293°N</geoCoordinate>
,
<geoCoordinate id="79331935DFF9BBF1883120FAD2A65781" degrees="139.67546" direction="east" orientation="longitude" precision="1" value="139.67546">139.67546°E</geoCoordinate>
; alt. 433 m; larvae collected: 9 Nov. 2014, emerged: 22 Apr. 2014; Y. Imada leg.; CYI.
<emphasis id="E55EE37F47D2264A8E52E02D1D8C2473" bold="true" pageId="0" pageNumber="13">Russia</emphasis>
• Primorsky Krai, Khasansky District, Primorsky Settlement, Zolotistyy [Golden] Stream;
<geoCoordinate id="E4FCCD31C0C60F2B6A717866F182DC40" degrees="43.10075" direction="north" orientation="latitude" precision="1" value="43.10075">43.10075°N</geoCoordinate>
,
<geoCoordinate id="4DE385305EBE81F257825C06834F8407" degrees="131.54862" direction="east" orientation="longitude" precision="1" value="131.54861">131.54862°E</geoCoordinate>
; alt. 62 m; 10 Jun. 2007 - 11 Jun. 2007; N.M. Paramonov leg.; CKLP. • 2 ♂; Sakhalin Oblast, Yuzhno-Kurilsk Urban Settlement, Kuril/Kunashir Island, near Lagunnoe Lake;
<geoCoordinate id="26837A49C69F95395E749A3EA2DCAC7A" degrees="44.0623" direction="north" orientation="latitude" precision="5" value="44.0623">44.0623°N</geoCoordinate>
,
<geoCoordinate id="BC29F5287D76E68909502986852EA18C" degrees="145.759" direction="east" orientation="longitude" precision="55" value="145.759">145.759°E</geoCoordinate>
; alt. 20 m; 11 Jul. 1954 - 12 Jul. 1954; leg. N.A. Violovich ZIN. • 1 ♂; Sakhalin Oblast, Kunashir Island, Mendeleevo Settlement;
<geoCoordinate id="FD4E3E987E853B82CBD78D2C441EADED" degrees="43.971" direction="north" orientation="latitude" precision="55" value="43.971">43.971°N</geoCoordinate>
,
<geoCoordinate id="5CE86334863711E491D305BF779053A9" degrees="145.694" direction="east" orientation="longitude" precision="55" value="145.694">145.694°E</geoCoordinate>
; alt. 220 m; 28 Jun. 1973; I.M. Kerzhner leg.; ZIN. • 1 ♀; Sakhalin Oblast, Kunashir Island, Alekhino Settlement [uninhabited];
<geoCoordinate id="26D729D12071F4777BF022C5DF33605C" degrees="43.91" direction="north" orientation="latitude" precision="555" value="43.91">43.91°N</geoCoordinate>
,
<geoCoordinate id="190D4A5AA1EFCD630D339DA1E409CCBF" degrees="145.52" direction="east" orientation="longitude" precision="555" value="145.52">145.52°E</geoCoordinate>
; alt. 5 m, 29 Jun. 1962; G.O. Krivoluckaja leg.; ZIN. • 2 ♂; Sakhalin Oblast, Kunashir Island, the mouth of the Tjatina River;
<geoCoordinate id="3C1C2928CD782EC86691BFC4A62DDB72" degrees="44.2711" direction="north" orientation="latitude" precision="5" value="44.2711">44.2711°N</geoCoordinate>
,
<geoCoordinate id="C5521907670512A8B13CF892A097B739" degrees="146.1583" direction="east" orientation="longitude" precision="5" value="146.1583">146.1583°E</geoCoordinate>
; alt. 15 m; 21 Jul. 2014; Y.N. Sundukov leg.; ZIN.
</paragraph>
</subSubSection>
<subSubSection id="D5CD6E68037F2E4FB6EACA55D42CC440" pageId="0" pageNumber="13" type="redescription">
<paragraph id="D2585580AA28174860B9119FFC27DADE" pageId="0" pageNumber="13">Redescription.</paragraph>
<paragraph id="61CC1A260362FDCC0BA2D9A547F238CC" pageId="0" pageNumber="13">
<emphasis id="9F711B4198E3468A0300F09C2786BC50" bold="true" pageId="0" pageNumber="13">Head.</emphasis>
Dark brown to black, with greyish pubescence (Fig.
<figureCitation id="AB724BBB13137465777BBF0FB3A1436F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23B-D</figureCitation>
). Rostrum short without nasus. Mouth parts pale brown to brown. Palpus brown to dark brown, five segmented; last segment 1.2-1.4
<normalizedToken id="AEFDC013239C2DC39D36AC7D46384D71" originalValue="×">x</normalizedToken>
longer than penultimate (Fig.
<figureCitation id="1373AA95DC0768D3A444EDAFE59FB372" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23B, D</figureCitation>
). Scape cylindrical, 1.6-1.8
<normalizedToken id="A2FE67728F6D51362735178F1BDADFF2" originalValue="×">x</normalizedToken>
longer than pedicel; pedicel ovate; pedicel brown, scape yellow to brown (Figs
<figureCitation id="6E2C209C093B98B45D052031828DA780" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4D</figureCitation>
,
<figureCitation id="145B0142886283F3099418EDE17F67D0" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23B, D</figureCitation>
); flagellum 14-segmented, pale brown to brown, monochrome or gradually darkening from base to tip. Flagellar segments simple, cylindrical in both sexes, not expanded ventrally; all male flagellomeres and 2-8 female flagellomeres covered with sparse whitish setae/sensilla; sensilla slightly denser in ventral side; verticels less prominent, 4-6 verticels not showing clear arrangement (Figs
<figureCitation id="2092DA56FCEB7687076EAB19FA53C711" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4D</figureCitation>
,
<figureCitation id="C6A734AF150919AB8484FBC6D770C80D" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23B, D</figureCitation>
).
</paragraph>
<caption id="D8029A04CBF68A315872F092BEB74BBB" doi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13" start="Figure 23" startId="F23">
<paragraph id="A2DD65F05E6EA45229EDB3FA9A2679A8" pageId="0" pageNumber="13">
<emphasis id="B311C8FA2EF660F74C1FEEC732393B76" bold="true" pageId="0" pageNumber="13">Figure 23.</emphasis>
<taxonomicName id="FBD72537B414E2C8485805B3E036DE17" baseAuthorityName="Alexander" baseAuthorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">
<emphasis id="1229645FDFFB5371373AAE7B19DD9A37" italics="true" pageId="0" pageNumber="13">Liogma mikado</emphasis>
</taxonomicName>
(Alexander, 1919)
<emphasis id="06E702E7A7CF093DFDCA30C5864D5846" bold="true" pageId="0" pageNumber="13">A</emphasis>
habitus of male, lateral view (colouration of wings is artefact)
<emphasis id="FD14D9EF07FE1F5CB15C192F4E50EE63" bold="true" pageId="0" pageNumber="13">B</emphasis>
head and thorax of female, lateral view
<emphasis id="E21CAD43A1A106384096506B07FEDD76" bold="true" pageId="0" pageNumber="13">C</emphasis>
head and thorax, dorsal view
<emphasis id="09B2F50ED0D148D580966C53787AEA38" bold="true" pageId="0" pageNumber="13">D</emphasis>
head of male, dorsal view
<emphasis id="168C707B85BF14ADB1E10653DD097986" bold="true" pageId="0" pageNumber="13">E</emphasis>
female terminalia lateral view.
</paragraph>
</caption>
<paragraph id="1FD744825C0E1BC939080FFA1922DF4E" pageId="0" pageNumber="13">
<emphasis id="72AA67FB24EEC0C38DA2BC5C494D4F31" bold="true" pageId="0" pageNumber="13">Thorax.</emphasis>
General colour shiny dark brown to black, with yellowish area in lateral side. Pronotum dark brown to black. Anterior part of mesonotum brown with black stripes or patches, usually forming three longitudinal, black markings on presutural area of scutum, and two drop-shaped black markings on postsutural area of scutum (Fig.
<figureCitation id="4CEE2CB3A57C574BCBB9C836BC29107A" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23C</figureCitation>
) or one large marking; black parts bare and shiny; paler parts with pubescence and with several long yellow hairs, forming longitudinal lines (Fig.
<figureCitation id="568E2B86245E43FBEF2084F87FF45B75" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23C</figureCitation>
). Dorsal pleural area, base of wing, anepimeron, and base of halter yellowish. Coxa black, ventrally paler (Fig.
<figureCitation id="ACF9D7C722938E8D8355AD044866523F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23B</figureCitation>
); trochanter yellow; femur gradually darkening distally, basal part yellowish, apical part dark brown to black; tibia and tarsus dark brown. Wing hyaline; veins brown; pterostigma pale; three branches of M reaching wing margin, M1 at same level as M1+2, cell a2 narrow,&gt; 8
<normalizedToken id="BC7A50678F8E2C59F9C57BB8882DA8C7" originalValue="×">x</normalizedToken>
longer than wide (Fig.
<figureCitation id="0C3F846D44ED09CC7DD8B1DB6DCEDB8F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of &quot; Triogma kuwanai &quot; form H Triogma kuwanai (Alexander, 1913) of &quot; Triogma limbinervis &quot; form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5D</figureCitation>
); membrane with interference patterns, visible with dark background (Fig.
<figureCitation id="0052B3CD04C58CC30C28E70B2BACC22B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23A</figureCitation>
). Halter stem pale brown, knob brown.
</paragraph>
<paragraph id="103820E937B8D90A59CCA70512445FF0" pageId="0" pageNumber="13">
<emphasis id="84300F86DF9EB242B60E513BD3E547B0" bold="true" pageId="0" pageNumber="13">Abdomen.</emphasis>
Black, without any distinct patterns (Fig.
<figureCitation id="BF20169DFD015B050C59CB3133D9D541" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23A</figureCitation>
).
</paragraph>
<paragraph id="22CFC6DC4FB323EF555E004FE23E478A" pageId="0" pageNumber="13">
<emphasis id="1DB2453C0A6AF97C826FB6AF7D400C65" bold="true" pageId="0" pageNumber="13">Male terminalia</emphasis>
: Relatively small, uniformly black, directed caudally (Fig.
<figureCitation id="505C2FF67F65E5E767BAF133C56A32A1" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23A</figureCitation>
). Tergite 9 fused with gonocoxite and sternite 9 (Fig.
<figureCitation id="AFDAEA4BB5874C4555460198500E8A81" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24C</figureCitation>
); tergite 9 with median lobe, with notch at middle (Fig.
<figureCitation id="2985B9F1A67B3A7461E65F53A7FAD976" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24A</figureCitation>
); lateral lobes of tergite 9 not prominent. Sternite 9 reduced to narrow band (Fig.
<figureCitation id="940CF7149FBB8FC2E6875270D9F5AA06" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24B, C</figureCitation>
). Gonocoxite relatively large, 1.2-1.4
<normalizedToken id="A5CA71883C5F2AD6A96450CA9A3CC8DC" originalValue="×">x</normalizedToken>
longer than tergite 9, in lateral view (Fig.
<figureCitation id="A8CDEE42E192DE3C4B8383C5DFC1AD0C" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24C</figureCitation>
); without any distinct lobes (Fig.
<figureCitation id="57C4F9B22076DD1A14E68E22399755C4" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24B, C</figureCitation>
); inner side of gonocoxite membranous; small round sclerotised patch on membranous area between gonocoxites present (Fig.
<figureCitation id="0789C5B3FE58FABE0DC8BE5F4B58565E" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24B</figureCitation>
), triangular in lateral view (Fig.
<figureCitation id="3A47AA44EAB91D2F8E304A6ABFF2E6FB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24C, F</figureCitation>
); holding base of aedeagal complex if it moved dorsally. Gonostylus with a strongly sclerotised, claw-like outgrowth; tip of gonostylus finger-like (Fig.
<figureCitation id="9F94820C19A7E80D1A2669B04AAC4D53" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24A, C</figureCitation>
). Aedeagus complex as long as gonocoxite and sternite 9 together; sperm pump and ejaculatory apodeme, partly covered by parameres (Fig.
<figureCitation id="D28D6D496AF9EF6003D3E3999B5AEF71" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24F</figureCitation>
); interbase simple L-shaped, both in lateral and dorsal/ventral views (Fig.
<figureCitation id="E2B57D5D2C6716CE4DEEDAD63FA75836" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24D-F</figureCitation>
); posterior part blade-like, with a small notch on dorsal side, in lateral view (Fig.
<figureCitation id="4D625C1A9A50FD09C6CED6C67905F893" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24F</figureCitation>
); aedeagus widened and curved dorsally at right angle in midlength, covered with prominent spines on ventral and lateral sides; membranous area on ventral side behind ventral spines, make flexible the aedeagus and able to straighten, probably during copulation (Fig.
<figureCitation id="3CC29FECE451BB8437D84E40C3E41CB5" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24D, F</figureCitation>
); aedeagus with apical branches short, directed caudally; median branch slightly longer and wider than lateral ones (Fig.
<figureCitation id="B8C4A489440472CC90234160EA37808E" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13">24D, F</figureCitation>
).
</paragraph>
<caption id="116250A8FA64989B7AB4A71C99F1BE77" doi="10.3897/zookeys.1083.75624.figure24" httpUri="https://binary.pensoft.net/fig/637497" pageId="0" pageNumber="13" start="Figure 24" startId="F24">
<paragraph id="F56B6DC5657D140F2492C25100842B65" pageId="0" pageNumber="13">
<emphasis id="66BFDCF4D1C6607F973200E9D50A2EC1" bold="true" pageId="0" pageNumber="13">Figure 24.</emphasis>
Male genital structures of
<taxonomicName id="1D65655AB305E82512A758CBBA5626BB" baseAuthorityName="Alexander" baseAuthorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">
<emphasis id="0764C7AD37F2C18573945F57A042F673" italics="true" pageId="0" pageNumber="13">Liogma mikado</emphasis>
</taxonomicName>
(Alexander, 1919)
<emphasis id="945C6272E120ADAFA6B1B2BD9CFC177F" bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis id="977BE526DCD866C4864297B8B7CAE461" bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, ventral view
<emphasis id="E075E281E611BD9F675B726C348613B9" bold="true" pageId="0" pageNumber="13">C</emphasis>
terminalia, lateral view
<emphasis id="BF039F2B68A69DE3A9AFAC06482E103F" bold="true" pageId="0" pageNumber="13">D</emphasis>
aedeagus complex, dorsal view
<emphasis id="901D87DE6D82C0033102CC102A8ED8C5" bold="true" pageId="0" pageNumber="13">E</emphasis>
aedeagus complex, ventral view
<emphasis id="686E3AD4481A007EC60506321448DBCB" bold="true" pageId="0" pageNumber="13">F</emphasis>
aedeagus complex, lateral view.
</paragraph>
</caption>
<paragraph id="E6080F052A3D626630758A1B331000EC" pageId="0" pageNumber="13">
<emphasis id="3242EE6FB74CC9625A313CF1E4DD5B55" bold="true" pageId="0" pageNumber="13">Female terminalia</emphasis>
: Black, tips of cercus and hypopygial valve yellowish (Fig.
<figureCitation id="941BFEFBF941275E56314DF0280174C4" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Liogma mikado (Alexander, 1919) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax, dorsal view D head of male, dorsal view E female terminalia lateral view." figureDoi="10.3897/zookeys.1083.75624.figure23" httpUri="https://binary.pensoft.net/fig/637496" pageId="0" pageNumber="13">23E</figureCitation>
). Tergite 8, 2
<normalizedToken id="06312A8A58ABBD4E545D991EF42BF3D5" originalValue="×">x</normalizedToken>
larger than tergite 9 in lateral view (Fig.
<figureCitation id="F2B4C3AF4B0E5294D989EA133372DCAB" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25B</figureCitation>
); not divided at middle (Fig.
<figureCitation id="4E931636E53CF8B48FDB2CF197A5AB7A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25A</figureCitation>
). Caudal margin of tergite 9 straight in lateral view (Fig.
<figureCitation id="87CE9707BAE72C3AB7EB4F2A1DD0F642" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25B</figureCitation>
). Lateral lobe of tergite 10 finger-like, 3
<normalizedToken id="2D2FB9A7C7BAA94439F4A2CC835117DB" originalValue="×">x</normalizedToken>
longer than wide, well separated from tergite 10 (Fig.
<figureCitation id="BE63F8A03E57D953A038FD01D87AC313" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25A</figureCitation>
); triangular sclerite large, separated from tergite 10 (Fig.
<figureCitation id="3941D8A275F507CDEEC7B2187D99EAE0" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25A</figureCitation>
). Cercus and hypogynial valve blade-like, relative narrow compared to other cylindrotomines (Fig.
<figureCitation id="07CCAFC883592FFC1A8FF1592FBC79BC" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25B</figureCitation>
); rough surface on dorsal tip of cercus hardly recognisable, only a few small pyramid teeth present. Genital fork large, heavily sclerotised plate; common sperm duct after genital opening relatively short, hardly recognisable; sperm ducts carrot-shaped; wall of sperm wrinkled (Fig.
<figureCitation id="80F6C0025F9610902021327D50F28AC8" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25C</figureCitation>
); three round spermathecae present, diameter ~ 1/3-1/2
<normalizedToken id="2802B5D664B863687DE3820F598E8244" originalValue="×">x</normalizedToken>
width of genital fork (Fig.
<figureCitation id="5A1CCE2F192263BD468868CED7735118" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genital structures of Liogma mikado (Alexander, 1919) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13">25D</figureCitation>
).
</paragraph>
<caption id="3DAE46BC0D70C327F17D0E75F1216491" doi="10.3897/zookeys.1083.75624.figure25" httpUri="https://binary.pensoft.net/fig/637498" pageId="0" pageNumber="13" start="Figure 25" startId="F25">
<paragraph id="4F4DE33F50E88A44BC841908E13EFB6C" pageId="0" pageNumber="13">
<emphasis id="201362F33C1FCF28D0F1E1B154F40EA4" bold="true" pageId="0" pageNumber="13">Figure 25.</emphasis>
Female genital structures of
<taxonomicName id="75261BBE0EAF063961B881D199841265" baseAuthorityName="Alexander" baseAuthorityYear="1919" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma mikado" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="mikado">
<emphasis id="951449566203EF537236ADABB1F6BBB5" italics="true" pageId="0" pageNumber="13">Liogma mikado</emphasis>
</taxonomicName>
(Alexander, 1919)
<emphasis id="C24AF4D69BFAF338DF21CF94A4D7C9FF" bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis id="61F16FFEB49D3DE941FA0CACB9B041E6" bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, lateral view
<emphasis id="DBD74BD6E588D563305B22351BF64E30" bold="true" pageId="0" pageNumber="13">C</emphasis>
sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view
<emphasis id="5F8D11B1B221BC31F9C4D273A12E4EE5" bold="true" pageId="0" pageNumber="13">D</emphasis>
spermathecae.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A979F641F83A198C5A1E2E11FFEAC8ED" pageId="0" pageNumber="13" type="distribution">
<paragraph id="3A7B13893768AE575EEBE53EFD9F09FF" pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph id="E0B59BCF52185C1ADEC15923ED211C16" pageId="0" pageNumber="13">
South Korea, Japan (Honshu I and Shikoku I), and Russia (Jewish Autonomous Oblast, Sakhalin Oblast (Kuril Is: Kunashir I) (
<bibRefCitation id="E87610D3DF7FF02100A2AB15AC7F09AA" author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B39" refString="Oosterbroek, P, 2021. Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea). https://ccw.naturalis.nl/index.php" title="Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea)." url="https://ccw.naturalis.nl/index.php" year="2021">Oosterbroek 2021</bibRefCitation>
). First records from Japan: Kyushu I (Fig.
<figureCitation id="63A0F0BB2EC7D5039C25911BAE44A80C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Occurrence data in Japan and surrounding areas of A Liogma brevipecten Alexander, 1932 B Liogma mikado (Alexander, 1919). Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data." figureDoi="10.3897/zookeys.1083.75624.figure22" httpUri="https://binary.pensoft.net/fig/637495" pageId="0" pageNumber="13">22B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B50138960D098528AE9E49BB5F4599AB" pageId="0" pageNumber="13" type="comments">
<paragraph id="F47D925538F39926BCA4A31F6784FE27" pageId="0" pageNumber="13">Comments.</paragraph>
<paragraph id="CFFF0299303614F8BAA5AC02D9DB5268" pageId="0" pageNumber="13">
As with other
<taxonomicName id="448D70C0AC0F276F02764EF86370572D" lsidName="" pageId="0" pageNumber="13" rank="subFamily" subFamily="Cylindrotominae">Cylindrotominae</taxonomicName>
species that have simple antennae and three M vein branches, this species was also originally described as
<taxonomicName id="B31796D1EC83FA5C5F03FC547687176F" authorityName="Schiner" authorityYear="1863" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="930A56C7A366BAB3F288B69D0E6BD6CC" italics="true" pageId="0" pageNumber="13">Phalacrocera</emphasis>
</taxonomicName>
(
<bibRefCitation id="51DA3E7BACF40979764F45DC7DBAE2E4" DOI="https://doi.org/10.1093/aesa/12.4.327" author="Alexander, CP" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="13" pagination="327 - 348" refId="B2" refString="Alexander, CP, 1919. Undescribed species of Japanese crane-flies (Tipulidae, Diptera). Annals of the Entomological Society of America 12: 327 - 348, DOI: https://doi.org/10.1093/aesa/12.4.327" title="Undescribed species of Japanese crane-flies (Tipulidae, Diptera)." url="https://doi.org/10.1093/aesa/12.4.327" volume="12" year="1919">Alexander 1919</bibRefCitation>
). Later
<bibRefCitation id="81B2745B2AEFDBCA37DB98149ED12720" author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi (1960)</bibRefCitation>
moved this species to the
<taxonomicName id="C43807901737629442668E282D1ADF50" authorityName="Osten Sacken" authorityYear="1869" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="267405D85262A181EC4DF8E8A96A78B3" italics="true" pageId="0" pageNumber="13">Liogma</emphasis>
</taxonomicName>
genus based on the morphological similarity of the immature stages with
<taxonomicName id="A7DE492270E97E74A45B33026A0E8C42" baseAuthorityName="Osten Sacken" baseAuthorityYear="1865" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma nodicornis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="nodicornis">
<emphasis id="4954CC656B58F565A3A02E614919BF21" italics="true" pageId="0" pageNumber="13">Liogma nodicornis</emphasis>
</taxonomicName>
(Osten Sacken, 1865). However,
<taxonomicName id="FC7CBCB4B8A2CA72C53467EC7C9443B3" lsidName="L. mikado" pageId="0" pageNumber="13" rank="species" species="mikado">
<emphasis id="7D62D19489CDC07F1A15B0274FC86EE4" italics="true" pageId="0" pageNumber="13">L. mikado</emphasis>
</taxonomicName>
is a morphologically and genetically quite distinct species from the other
<taxonomicName id="7C18A9AEC0E36735FEF7F19BF2B0F299" authorityName="Osten Sacken" authorityYear="1869" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="4FEA8C3750FD0927D7606A348B4038C3" italics="true" pageId="0" pageNumber="13">Liogma</emphasis>
</taxonomicName>
species, and the exact phylogenetic position remains unclear.
</paragraph>
</subSubSection>
</treatment>
</document>