treatments-xml/data/CC/6D/87/CC6D87D9FFADFFCAA8F5FBEAE8F7FB26.xml

<document id="1F84483CED5294AB81455E21F6F1FFEA" ID-DOI="10.1649/0010-065X-77.3.319" ID-ISSN="1938-4394" ID-Zenodo-Dep="13243120" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722967910837" checkinUser="felipe" docAuthor="Martínez-Luque, Erick Omar &amp; Johnson, Paul J." docDate="2023" docId="CC6D87D9FFADFFCAA8F5FBEAE8F7FB26" docLanguage="en" docName="TheColeopBull.77.3.319-327.pdf" docOrigin="The Coleopterists Bulletin 77 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-77.3.319" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Scaptolenus acrognathus Champion 1896" docType="treatment" docVersion="2" lastPageNumber="326" masterDocId="3054FFA1FFAFFFC2A842FF97E904FF97" masterDocTitle="Description of the Female and Redescription of the Male of Scaptolenus acrognathus Champion (Coleoptera: Elateridae)" masterLastPageNumber="327" masterPageNumber="319" pageNumber="320" updateTime="1722970792159" updateUser="ExternalLinkService">
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<mods:title id="5A8A3A77962B1770BB05269B4FCE09F6">Description of the Female and Redescription of the Male of Scaptolenus acrognathus Champion (Coleoptera: Elateridae)</mods:title>
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<mods:namePart id="608F56CB38524135F8C1C2351848D207">Martínez-Luque, Erick Omar</mods:namePart>
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<mods:title id="0ECAEC41F272E67AED16078EE2F3418B">The Coleopterists Bulletin</mods:title>
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<heading id="1F3381A3FFADFFC0A8F5FBEAEB5BFB06" box="[183,607,1148,1170]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="320" reason="2">
<taxonomicName id="83C44D4CFFADFFC0A8F5FBEAEB5BFB06" authority="Champion, 1896" authorityName="Champion" authorityYear="1896" box="[183,607,1148,1170]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFADFFC0A8F5FBEAEB5BFB06" bold="true" box="[183,607,1148,1170]" pageId="2" pageNumber="320">
<emphasis id="76B0EADDFFADFFC0A8F5FBEAE8B5FB05" bold="true" box="[183,433,1149,1170]" italics="true" pageId="2" pageNumber="320">SCAPTOLENUs ACROGNATHUs</emphasis>
<bibRefCitation id="20554B3EFFADFFC0A9F5FBEBEB5BFB06" author="Champion, G. C." box="[439,607,1148,1170]" pageId="2" pageNumber="320" pagination="557 - 573" refId="ref4964" refString="Champion, G. C. 1896. Fam. Cebrionidae [pp. 557 - 573, pl. 25]. In: Biologia Centrali-Americana, Insecta, Coleoptera, Vol. 3, part 1, Serricornia (F. D. Godman and O. Salvin, editors). R. H. Porter, London. xv + [1] + 690 pp., 27 pl." type="book chapter" year="1896">Champion, 1896</bibRefCitation>
</emphasis>
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<emphasis id="76B0EADDFFADFFC0A8DAFB30E894FB2B" box="[152,400,1191,1212]" italics="true" pageId="2" pageNumber="320">Scaptolenus acrognathus</emphasis>
<bibRefCitation id="20554B3EFFADFFC0A9DCFB30EB7DFB2A" author="Champion, G. C." box="[414,633,1191,1213]" pageId="2" pageNumber="320" pagination="557 - 573" refId="ref4964" refString="Champion, G. C. 1896. Fam. Cebrionidae [pp. 557 - 573, pl. 25]. In: Biologia Centrali-Americana, Insecta, Coleoptera, Vol. 3, part 1, Serricornia (F. D. Godman and O. Salvin, editors). R. H. Porter, London. xv + [1] + 690 pp., 27 pl." type="book chapter" year="1896">Champion 1896: 564</bibRefCitation>
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, table XXV, fig. 8, 8a, male (drawing). References: 
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<bibRefCitation id="20554B3EFFADFFC0A8F1FB4AE889FB65" author="Dalla Torre, K. W. von" box="[179,397,1245,1267]" pageId="2" pageNumber="320" refId="ref5135" refString="Dalla Torre, K. W. von. 1911. Coleopterorum Catalo - gus, Vol. 11, Pars 25, Cebrionidae [pp. 11 - 13]. W. Junk, Berlin." type="book" year="1911">Dalla Torre 1911: 11</bibRefCitation>
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; 
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<bibRefCitation id="20554B3EFFADFFC0A9DCFB4AEB7CFB65" author="Dalla Torre, K. W. von" box="[414,632,1245,1267]" pageId="2" pageNumber="320" refId="ref5173" refString="Dalla Torre, K. W. von. 1913. Coleoptera Fam. Cebrionidae. Genera Insectorum, fasc. 127 (P. Wytsman, editor). V. Verteneuil &amp; L. Desmet, Bruxelles, 17 pp. + 1 pl." type="book" year="1913">Dalla Torre 1913: 11</bibRefCitation>
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; 
<treatmentCitation id="C56510DEFFADFFC0A8F1FB60E892FA9A" author="Blackwelder, R. H." box="[179,406,1271,1293]" page="275" pageId="2" pageNumber="320" year="1944">
<bibRefCitation id="20554B3EFFADFFC0A8F1FB60E892FA9A" author="Blackwelder, R. H." box="[179,406,1271,1293]" pageId="2" pageNumber="320" pagination="189 - 341" refId="ref4921" refString="Blackwelder, R. H. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, Part 2. Bulletin of the United States National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944: 275</bibRefCitation>
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.
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<emphasis id="76B0EADDFFADFFC0A8F2FAB5EB0FFAAF" bold="true" box="[176,523,1312,1336]" pageId="2" pageNumber="320">
Material Examined (
<specimenCount id="52C2FD46FFADFFC0A9D1FAB5E8C4FAAF" box="[403,448,1312,1336]" count="1" pageId="2" pageNumber="320" type="female">1 ♀</specimenCount>
; 
<specimenCount id="52C2FD46FFADFFC0A990FAB5E8F9FAAF" box="[466,509,1312,1336]" count="3" pageId="2" pageNumber="320" type="male">3 ♂</specimenCount>
).
</emphasis>
<materialsCitation id="F4AC3C92FFADFFC0AA5AFAB5EB4BFA35" collectingDate="1997-08-14" collectorName="de Capula &amp; Col. J. L. Cozar" country="Mexico" latitude="20.341972" location="Cerro de Capula" longLatPrecision="1" longitude="-100.29381" municipality="Huimilpan" pageId="2" pageNumber="320" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Queretaro de Arteaga">
<collectingCountry id="3CD3765FFFADFFC0AA5AFAB5EB7FFAA0" box="[536,635,1314,1335]" name="Mexico" pageId="2" pageNumber="320">MEXICO</collectingCountry>
: 
<collectingRegion id="8600F82DFFADFFC0A8DAFAAAE9FEFAC5" box="[152,250,1341,1362]" country="Mexico" name="Queretaro de Arteaga" pageId="2" pageNumber="320">Querétaro</collectingRegion>
, 
<collectingMunicipality id="A41FACB5FFADFFC0A95FFAAAE882FAC4" box="[285,390,1341,1363]" pageId="2" pageNumber="320">Huimilpan</collectingMunicipality>
, 
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Cerro 
<collectorName id="E9315319FFADFFC0A9BCFAAAEB7DFAC4" box="[510,633,1341,1363]" pageId="2" pageNumber="320">de Capula</collectorName>
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, (
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, 
<geoCoordinate id="21F05008FFADFFC0A974FAC1E8CBFAFA" box="[310,463,1366,1389]" degrees="100" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="320" precision="1" seconds="37.7" value="-100.29381">100°17′37.7′′W</geoCoordinate>
), oak forest, leaf litter, 2,536 msnm, 
<date id="307A100FFFADFFC0A91BFAE5E8D0FA1F" box="[345,468,1394,1416]" pageId="2" pageNumber="320" value="1997-08-14">
<collectingDate id="203EE9E7FFADFFC0A91BFAE5E8D0FA1F" box="[345,468,1394,1416]" pageId="2" pageNumber="320" value="1997-08-14">14-viii-1997</collectingDate>
</date>
, 
<collectorName id="E9315319FFADFFC0A99DFAE5EB79FA10" box="[479,637,1394,1416]" pageId="2" pageNumber="320">Col. J. L. Cozar</collectorName>
(UAQE-22753: 
<specimenCount id="52C2FD46FFADFFC0A971FA1AE85DFA35" box="[307,345,1421,1442]" count="1" pageId="2" pageNumber="320" type="female">1 ♀</specimenCount>
; and UAQE-22754: 
<specimenCount id="52C2FD46FFADFFC0AA62FA1AEB43FA35" box="[544,583,1421,1442]" count="1" pageId="2" pageNumber="320" type="male">1 ♂</specimenCount>
)
</materialsCitation>
; 
<materialsCitation id="F4AC3C92FFADFFC0AA1BFA1AE85AFA40" collectingDate="1971-07-05" collectionCode="FSCA" collectorName="Col. B. K. Dozier" country="Mexico" location="Matehuala" pageId="2" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="San Luis Potosi">
<collectingRegion id="8600F82DFFADFFC0AA1BFA1AE80AFA28" country="Mexico" name="San Luis Potosi" pageId="2" pageNumber="320">San Luis PotosÍ</collectingRegion>
, 
<location id="411B6014FFADFFC0A95CFA30E882FA2A" LSID="urn:lsid:plazi:treatment:CC6D87D9FFADFFCAA8F5FBEAE8F7FB26:411B6014FFADFFC0A95CFA30E882FA2A" box="[286,390,1447,1469]" country="Mexico" name="Matehuala" pageId="2" pageNumber="320" stateProvince="San Luis Potosi">Matehuala</location>
, 
<date id="307A100FFFADFFC0A9D4FA30E8F9FA2A" box="[406,509,1447,1469]" pageId="2" pageNumber="320" value="1971-07-05">
<collectingDate id="203EE9E7FFADFFC0A9D4FA30E8F9FA2A" box="[406,509,1447,1469]" pageId="2" pageNumber="320" value="1971-07-05">5-vii-1971</collectingDate>
</date>
, 
<collectorName id="E9315319FFADFFC0AA4CFA30E9DDFA4F" pageId="2" pageNumber="320">Col. B. K. Dozier</collectorName>
(
<collectionCode id="22D5AE0AFFADFFC0A8ABFA55E82EFA40" box="[233,298,1474,1495]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="2" pageNumber="320" type="Museum">FSCA</collectionCode>
: 
<specimenCount id="52C2FD46FFADFFC0A974FA55E850FA40" box="[310,340,1474,1495]" count="1" pageId="2" pageNumber="320" type="male">1♂</specimenCount>
)
</materialsCitation>
; 
<materialsCitation id="F4AC3C92FFADFFC0A929FA55E82AF99B" collectingDate="1961-06-17" collectionCode="CNIN-IBUNAM" collectorName="de Bravo &amp; Col. J. Hendrichs S. &amp; Col. J. Hendrichs" country="Mexico" location="Valle de Bravo" pageId="2" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Mexico">
<collectingRegion id="8600F82DFFADFFC0A929FA55EB25FA4F" box="[363,545,1474,1496]" country="Mexico" name="Mexico" pageId="2" pageNumber="320">Estado de México</collectingRegion>
, 
<location id="411B6014FFADFFC0AA6FFA55E9D1FA65" LSID="urn:lsid:plazi:treatment:CC6D87D9FFADFFCAA8F5FBEAE8F7FB26:411B6014FFADFFC0AA6FFA55E9D1FA65" country="Mexico" name="Valle de Bravo" pageId="2" pageNumber="320" stateProvince="Mexico">
Valle 
<collectorName id="E9315319FFADFFC0AA25FA55E9D1FA65" pageId="2" pageNumber="320">de Bravo</collectorName>
</location>
, 
<date id="307A100FFFADFFC0A8A1FA4AE854FA64" box="[227,336,1501,1523]" pageId="2" pageNumber="320" value="1961-06-17">
<collectingDate id="203EE9E7FFADFFC0A8A1FA4AE854FA64" box="[227,336,1501,1523]" pageId="2" pageNumber="320" value="1961-06-17">17-vi-1961</collectingDate>
</date>
, 
<collectorName id="E9315319FFADFFC0A91FFA4AEB2DFA65" box="[349,553,1501,1523]" pageId="2" pageNumber="320">Col. J. Hendrichs S.</collectorName>
(
<collectionCode id="22D5AE0AFFADFFC0AA7BFA4AE9F9F99B" pageId="2" pageNumber="320">CNIN-IBUNAM</collectionCode>
: 
<specimenCount id="52C2FD46FFADFFC0A944FA60E822F99B" box="[262,294,1527,1548]" count="1" pageId="2" pageNumber="320" type="male">1♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0CDE6544FFADFFC4A8F2F985EA6AFA43" lastPageId="6" lastPageNumber="324" pageId="2" pageNumber="320" type="description">
<paragraph id="447B36CFFFADFFC0A8F2F985EAA6FDA4" blockId="2.[152,639,1312,1709]" lastBlockId="2.[684,1172,194,1709]" pageId="2" pageNumber="320">
<emphasis id="76B0EADDFFADFFC0A8F2F985E894F9B0" bold="true" box="[176,400,1554,1576]" pageId="2" pageNumber="320">Description of Female</emphasis>
(
<figureCitation id="DCFF2A4AFFADFFC0A9DEF985EB0BF9B0" box="[412,527,1554,1576]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Figs. 11–18</figureCitation>
, 
<figureCitation id="DCFF2A4AFFADFFC0AA5BF985E9B7F9D4" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="2" pageNumber="320">20, 22, 24, 26</figureCitation>
). Body 
<quantity id="833C9B2AFFADFFC0A8BEF9BAE854F9D5" box="[252,336,1581,1602]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="320" unit="mm" value="13.8">13.8 mm</quantity>
long, 
<quantity id="833C9B2AFFADFFC0A9CEF9BAE8D0F9D5" box="[396,468,1581,1602]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="320" unit="mm" value="4.7">4.7 mm</quantity>
wide; head, mandibles, antennae, pronotum, abdomen, and elytra fusco-rufous, basal part fulvous (
<figureCitation id="DCFF2A4AFFADFFC0A989F9F5EB38F9E0" box="[459,572,1634,1656]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Figs. 11–13</figureCitation>
). Head with setae similar in length to length of penultimate maxillary palp, thin, fulvous-colored, scattered on frons and grouped apically, denser at base of mandibles and antennae. Eye black, small, similar in length to antennomere 1. Antenna with setae similar in length to antennomere 3 and fulvous in color. Pronotal margin setae similar in length to setae on head, setae only on margin of pronotum, densest at anterior and posterior angles. Legs with small setae half the length of setae present on head and fulvous in color. Elytra with setae similar in length to setae present on legs, with thin, fulvous setae on outer marginal part, abundant at base and apically. Abdomen bright fusco-rufous (
<figureCitation id="DCFF2A4AFFADFFC0AB9FFE70ED50FE6B" box="[989,1108,487,509]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Figs. 12, 13</figureCitation>
), with scattered testaceous setae, thin and similar in length to setae present on elytra.
</paragraph>
<paragraph id="447B36CFFFADFFC1AA87FDA0EB2EFD8F" blockId="2.[684,1172,194,1709]" lastBlockId="3.[173,660,194,1736]" lastPageId="3" lastPageNumber="321" pageId="2" pageNumber="320">
Head 
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long, 
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wide, with dense, deep punctation. Front irregularly shaped (not smooth, but distinctive or tubercled). Maxillary palp (
<figureCitation id="DCFF2A4AFFADFFC0AAF6FD10EBFEFD0A" box="[692,762,647,669]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="2" pageNumber="320">Fig. 26</figureCitation>
) with apical part of palpomere cylindrical, 2.6 times as long as wide, wider at apex than at base; penultimate palpomere cylindrical, 4 times as long as wide, gradually widening apically. Antenna clavate with short, seemingly fused antennomeres (
<figureCitation id="DCFF2A4AFFADFFC0AC00FD65ED8CFC90" box="[1090,1160,754,776]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="2" pageNumber="320">Fig. 20</figureCitation>
), length ratio of antennomeres 2–11: 0.2:0.2:0.1:0.1: 0.1:0.1:0.08:0.09:0.05:0.1. Pronotum 1.6 times as wide as long; width of pronotum 
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, length of pronotum 
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; anterior margin convergent medially, subacute, lateral margins rounded, with carina extending from the posterior angle to more than half of lateral margin, not reaching anterior margin. Pronotum with scattered, shallow punctation; posterior angles divergent and not very acute, exceeding apex of scutellar shield. Scutellar shield subtriangular with rounded edges. Elytra together 
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long and 
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wide (across humeri), punctures very similar to punctures of pronotum, gradually sulcate (
<figureCitation id="DCFF2A4AFFADFFC0AB1CFBF0EAA1FBEB" box="[862,933,1127,1149]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Fig. 11</figureCitation>
). Abdomen smooth with a shiny appearance (
<figureCitation id="DCFF2A4AFFADFFC0AB37FB15EAB9FB00" box="[885,957,1154,1176]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Fig. 12</figureCitation>
), with 5 ventrites and with fine, sparse punctuation, each puncture with very thin, decumbent, fulvous seta. Hind wing length 
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, 2 times as long as wide (
<figureCitation id="DCFF2A4AFFADFFC0AC7DFB45ED8CFB70" box="[1087,1160,1234,1256]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="2" pageNumber="320">Fig. 14</figureCitation>
). R cell open, r3, r4, RML, MSP, and SS present; basal portion of RP measuring 
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, MP 
<subScript id="D840348AFFADFFC0AC05FA84ED5BFAB6" attach="left" box="[1095,1119,1299,1313]" fontSize="6" pageId="2" pageNumber="320">3+4</subScript>
short measuring 
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; CuA 
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and CAS directly connected with Cu; CuA 
<subScript id="D840348AFFADFFC0AB3DFADFEA83FAC1" attach="left" box="[895,903,1352,1366]" fontSize="6" pageId="2" pageNumber="320">1</subScript>
, CuA 
<subScript id="D840348AFFADFFC0AB81FADFEADFFAC1" attach="left" box="[963,987,1352,1366]" fontSize="6" pageId="2" pageNumber="320">3+4</subScript>
, W cell, AA 
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, and AA 
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absent. Prosternum wider than long, length 
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, width of prosternum 
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, not exceeding coxae, abruptly truncated in anterior part, discal part gradually curved, punctation similar to that of abdomen. Mesoventrite and metaventrite with fine, shallow punctures, each puncture with fulvous seta, similar in length to setae present on head. Metacoxal plate broadly arcuate anteriorly (
<figureCitation id="DCFF2A4AFFADFFC0ABA8F985ED36F9B0" box="[1002,1074,1554,1576]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="2" pageNumber="320">Fig. 24</figureCitation>
), strongly narrowing laterally, with 2 acute teeth separated by an emargination over femoral attachment; posterior margin carinate from ectal tooth to lateral apex; coxal plate finely and shallow punctate, each puncture with seta similar in size to setae present on head, setae fulvous. Metanepisternum triangular anteriorly, narrowing to metacoxal angle. Protibia with ectal border with subacute, triangular central tooth (
<figureCitation id="DCFF2A4AFFACFFC1A8B2FE85E838FEB0" box="[240,316,274,296]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">Fig. 22</figureCitation>
), tibial apex with subacute lateral tooth, tibia with fine, shallow punctures, each puncture with seta. Tibia with setae densely clustered at apical margin; length of the ectal tooth 
<quantity id="833C9B2AFFACFFC1AA02FEF6EB8BFEE0" box="[576,655,353,375]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="321" unit="mm" value="0.5">0.5 mm</quantity>
. Protibial spurs long, equal, 
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long; nearly straight, gradually curved, apex subtruncate. Average lengths of tarsomeres 1–5, basal to distal: 1.0 mm, 
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, 
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, 
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, 
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; each robust, subcylindrical, with fine, rigid setae of similar length to setae present on protibiae.
</paragraph>
<paragraph id="447B36CFFFACFFC1A887FD8BE8C4FBBA" blockId="3.[173,660,194,1736]" pageId="3" pageNumber="321">
<emphasis id="76B0EADDFFACFFC1A887FD8BE888FDA5" bold="true" box="[197,396,540,562]" pageId="3" pageNumber="321">Female Terminalia.</emphasis>
Tergite VIII (
<figureCitation id="DCFF2A4AFFACFFC1AA51FD8BEB5CFDA5" box="[531,600,540,562]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="3" pageNumber="321">Fig. 15</figureCitation>
) widening apically in basal third, then gradually narrowing apically; apex broadly rounded with long, thick setae, similar in length to setae present on head; base with deep median emargination, each basolateral lobe extended, punctation dense and deep along entire length of tergite. Sternite VIII (
<figureCitation id="DCFF2A4AFFACFFC1AA7DFD2BEB88FD45" box="[575,652,700,722]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="3" pageNumber="321">Fig. 16</figureCitation>
) broadly subtriangular, punctate with thin setae, similar in length to setae present on tergite VIII, located on lateral margin, with 2 groups of setae in apical fifth; basal angles slightly extended, with carinate margin running medially from lateral margin of basal fifth of sternite and merging with base of spiculum ventrale; basodiscal area of sternite translucent, difficult to delimit; spiculum ventrale (broken in figure) very long, 3.16 times as long as sternite. Ovipositor (
<figureCitation id="DCFF2A4AFFACFFC1A92AFC50E8D0FC4B" box="[360,468,967,989]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="3" pageNumber="321">Figs. 17, 18</figureCitation>
) baculum 4.04 times length of gonocoxites; base of ovipositor 
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wide, tapering to dorsoventrally compressed gonocoxites, each 
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wide.
</paragraph>
<paragraph id="447B36CFFFACFFC1A887FBA6EB96F9CB" blockId="3.[173,660,194,1736]" pageId="3" pageNumber="321">
<emphasis id="76B0EADDFFACFFC1A887FBA6E8AEFBD0" bold="true" box="[197,426,1073,1095]" pageId="3" pageNumber="321">Redescription of Male</emphasis>
(
<figureCitation id="DCFF2A4AFFACFFC1A9F5FBA5EB24FBD0" box="[439,544,1074,1096]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Figs. 1–10</figureCitation>
, 
<figureCitation id="DCFF2A4AFFACFFC1AA69FBA5E9CCFBF5" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">19, 21, 23, 25</figureCitation>
). Body 13.2–16.0 mm (
<emphasis id="76B0EADDFFACFFC1A9FAFBDCE8C7FBF5" box="[440,451,1099,1122]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 14.8, 
<emphasis id="76B0EADDFFACFFC1AA58FBDAEB22FBF5" box="[538,550,1101,1122]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) long, 
<quantity id="833C9B2AFFACFFC1A8EFFBF0E825FBEB" box="[173,289,1127,1149]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.2" metricValueMin="5.6" pageId="3" pageNumber="321" unit="mm" value="5.9" valueMax="6.2" valueMin="5.6">5.6–6.2 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1A973FBF1E838FBEA" box="[305,316,1126,1149]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 6.0, 
<emphasis id="76B0EADDFFACFFC1A9C0FBFFE88AFBEA" box="[386,398,1128,1149]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) wide; Dorsum brown (
<figureCitation id="DCFF2A4AFFACFFC1A8F6FB15E9EEFB00" box="[180,234,1154,1176]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Fig. 1</figureCitation>
), thoracic venter and femora fulvous; abdominal ventrites, femora, tibiae, tarsomeres, and claws fuscous in color (
<figureCitation id="DCFF2A4AFFACFFC1A911FB20E8AAFB5B" box="[339,430,1207,1229]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Figs. 2, 3</figureCitation>
). Head black with anterior setae similar in length to antennomere 3, decumbent and fulvous. Eyes black, longer than half the length of antennomere 1. Antennae brown with setae fulvous, setae similar in length to antennomere 3. Pronotum dark brown with long, fine, fulvous-colored setae similar in length to antennomere 6 (
<figureCitation id="DCFF2A4AFFACFFC1A8F6FAE5E9E8FA10" box="[180,236,1394,1416]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Fig. 1</figureCitation>
). Legs with coxae and femora fulvous, with long, thin decumbent setae and similar in length to antennomere 6. Tibiae, tarsomeres, and claws fuscous with decumbent setae similar in length to antennomere 6 (
<figureCitation id="DCFF2A4AFFACFFC1A96CFA4BE881FA66" box="[302,389,1500,1522]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Figs. 2, 3</figureCitation>
). Elytra fuscous with shorter fulvous setae similar in length to antennomere 3, concentrated on the lateral margin (
<figureCitation id="DCFF2A4AFFACFFC1A9B2F985EB21F9B0" box="[496,549,1554,1576]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Fig. 1</figureCitation>
). Abdomen fuscous on all segments, with testaceous, decumbent, evenly spaced, short, scattered setae (
<figureCitation id="DCFF2A4AFFACFFC1AA08F9D0EB81F9CB" box="[586,645,1607,1629]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="447B36CFFFACFFC1A887F9F5EDA7FA35" blockId="3.[173,660,194,1736]" lastBlockId="3.[706,1193,193,1736]" pageId="3" pageNumber="321">
Head 
<quantity id="833C9B2AFFACFFC1A8B8F9F6E86CF9E0" box="[250,360,1633,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="3" pageNumber="321" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1A931F9F7E87AF9E0" box="[371,382,1632,1655]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.5, 
<emphasis id="76B0EADDFFACFFC1A9FBF9F5E8C1F9E0" box="[441,453,1634,1655]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) long, 
<quantity id="833C9B2AFFACFFC1AA67F9F5EB97F9E0" box="[549,659,1634,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="3" pageNumber="321" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1A8F7F9ECE9C4F905" box="[181,192,1659,1682]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.5, 
<emphasis id="76B0EADDFFACFFC1A945F9EAE817F905" box="[263,275,1661,1682]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) wide, with dense, deep punctation with setae. Maxillary palp (
<figureCitation id="DCFF2A4AFFACFFC1A9F5F900E8FAF93A" box="[439,510,1687,1709]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">Fig. 25</figureCitation>
) with length of penultimate palpomere 0.4–1.0 mm (
<emphasis id="76B0EADDFFACFFC1AA4BF927EB10F950" box="[521,532,1712,1735]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 0.6, 
<emphasis id="76B0EADDFFACFFC1AA17F925EB65F950" box="[597,609,1714,1735]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3), length of last palpomere 0.4–1.0 mm (
<emphasis id="76B0EADDFFACFFC1AC15FF56ED66FF4F" box="[1111,1122,193,216]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 0.7, 
<emphasis id="76B0EADDFFACFFC1AA81FF4AEBCBFF65" box="[707,719,221,242]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3). Antennomeres 4–10 serrate (
<figureCitation id="DCFF2A4AFFACFFC1AC55FF4BED58FF66" box="[1047,1116,220,242]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">Fig. 19</figureCitation>
), length ratios of antennomeres 2–11: 0.1:0.3:0.7:0.7:0.6: 0.6:0.6:0.6:0.6:0.6:0.7. Pronotum 1.6× wider than long; width 
<quantity id="833C9B2AFFACFFC1AB7FFEBBEAB7FED5" box="[829,947,300,322]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="3" pageNumber="321" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.5–3.8 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AB87FEBCEAD4FED5" box="[965,976,299,322]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 3.6, 
<emphasis id="76B0EADDFFACFFC1AC5FFEBAED2DFED5" box="[1053,1065,301,322]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3), length 
<quantity id="833C9B2AFFACFFC1AA81FED0EA30FECB" box="[707,820,327,348]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="3" pageNumber="321" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1–2.2 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AB02FED1EA4FFECA" box="[832,843,326,349]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 2.2, 
<emphasis id="76B0EADDFFACFFC1ABC8FEDFEA92FECA" box="[906,918,328,349]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3); discal punctation dense and deep, slightly sparser and shallower than on head; lateral margins convergent anteriorly from mid-length, anterior angles subacute, lateral margins rounded, without carina; posterior angles widely divergent, elongate, apically acute. Scutellar shield subtriangular. Elytra together 9.5–12.0 mm (
<emphasis id="76B0EADDFFACFFC1ACC7FE71ED94FE6A" box="[1157,1168,486,509]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 11.0, 
<emphasis id="76B0EADDFFACFFC1AABBFD95EA01FD80" box="[761,773,514,535]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) long and 
<quantity id="833C9B2AFFACFFC1ABD5FD96ED0FFD80" box="[919,1035,513,535]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.2" metricValueMin="5.6" pageId="3" pageNumber="321" unit="mm" value="5.9" valueMax="6.2" valueMin="5.6">5.6–6.2 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AC59FD97ED22FD80" box="[1051,1062,512,535]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 6.0, 
<emphasis id="76B0EADDFFACFFC1AC2EFD95ED7CFD80" box="[1132,1144,514,535]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) wide across humeri, with punctures similar to those of the head, striae shallowly sulciform. Abdomen with 5 ventrites, punctation fine, sparse, each puncture with a fine, decumbent seta. Hind wing length 
<quantity id="833C9B2AFFACFFC1AA81FD10EA2CFD0B" box="[707,808,647,668]" metricMagnitude="-2" metricUnit="m" metricValue="1.273" pageId="3" pageNumber="321" unit="mm" value="12.73">12.73 mm</quantity>
, 2.1× as long as wide (
<figureCitation id="DCFF2A4AFFACFFC1AC4EFD10ED4DFD0B" box="[1036,1097,647,669]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Fig. 4</figureCitation>
). Support sclerites (SS) V-shaped; R cell 4.56 times as long as wide; r3, r4, RML, MSP, MP 
<subScript id="D840348AFFACFFC1ABB4FD5FED0BFD41" attach="left" box="[1014,1039,712,726]" fontSize="6" pageId="3" pageNumber="321">1+2</subScript>
, MP 
<subScript id="D840348AFFACFFC1AC7FFD5FED51FD41" attach="left" box="[1085,1109,712,726]" fontSize="6" pageId="3" pageNumber="321">3+4</subScript>
present; MP 
<subScript id="D840348AFFACFFC1AAA7FD75EBF9FD67" attach="left" box="[741,765,738,752]" fontSize="6" pageId="3" pageNumber="321">3+4</subScript>
short, 
<quantity id="833C9B2AFFACFFC1AB79FD40EA95FD7B" box="[827,913,727,749]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="3" pageNumber="321" unit="mm" value="1.53">1.53 mm</quantity>
; basal portion of RP 
<quantity id="833C9B2AFFACFFC1AC0DFD40EDA1FD7B" box="[1103,1189,727,749]" metricMagnitude="-3" metricUnit="m" metricValue="3.76" pageId="3" pageNumber="321" unit="mm" value="3.76">3.76 mm</quantity>
; CuA 
<subScript id="D840348AFFACFFC1AAB2FD6AEBFCFC9C" attach="left" box="[752,760,765,779]" fontSize="6" pageId="3" pageNumber="321">1</subScript>
joining CuA 
<subScript id="D840348AFFACFFC1AB31FD6AEA7FFC9C" attach="left" box="[883,891,765,779]" fontSize="6" pageId="3" pageNumber="321">2</subScript>
; W cell present, 2.6× as long as wide; CuA 
<subScript id="D840348AFFACFFC1AB6CFC8FEA42FCB1" attach="left" box="[814,838,792,806]" fontSize="6" pageId="3" pageNumber="321">3+4</subScript>
, CAS, AA 
<subScript id="D840348AFFACFFC1ABF7FC8FEAB9FCB1" attach="left" box="[949,957,792,806]" fontSize="6" pageId="3" pageNumber="321">4</subScript>
, AA 
<subScript id="D840348AFFACFFC1ABAFFC8FEAF1FCB1" attach="left" box="[1005,1013,792,806]" fontSize="6" pageId="3" pageNumber="321">3</subScript>
, AA 
<subScript id="D840348AFFACFFC1AC67FC8FED39FCB1" attach="left" box="[1061,1085,792,806]" fontSize="6" pageId="3" pageNumber="321">3+4</subScript>
and AP 
<subScript id="D840348AFFACFFC1ACD2FC8FEDACFCB1" attach="left" box="[1168,1192,792,806]" fontSize="6" pageId="3" pageNumber="321">3+4</subScript>
present. Prosternum 
<quantity id="833C9B2AFFACFFC1ABC9FCB0EAD1FCAB" box="[907,981,807,828]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="321" unit="mm" value="1.1">1.1 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1ABA6FCB1EAEBFCAA" box="[996,1007,806,829]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.1, 
<emphasis id="76B0EADDFFACFFC1AC77FCBFED45FCAA" box="[1077,1089,808,829]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) long, 
<quantity id="833C9B2AFFACFFC1AA81FCD5EA0EFCC0" box="[707,778,834,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="321" unit="mm" value="1.3">1.3 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AB55FCD7EA26FCC0" box="[791,802,832,855]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.3, 
<emphasis id="76B0EADDFFACFFC1AB20FCD5EA6AFCC0" box="[866,878,834,855]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3) wide, short and not reaching coxae, anterior margin truncate, with disc convex, with fine, shallow and very dispersed setate punctation. Mesoventrite and metaventrite with fine, shallow punctures, each puncture with a long, fine seta similar in length to the setae of the pronotum. Metacoxal ventral plate broad medially, with 3 subacute to acute teeth at coxal base covering metafemoral insertion, then gradually tapering laterally (
<figureCitation id="DCFF2A4AFFACFFC1AC15FB80ED99FBBB" box="[1111,1181,1047,1069]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">Fig. 23</figureCitation>
), with fine, shallow, setate punctation, seta longer than length of antennomere 6. Metanepisternum triangular, broad anteriorly, narrowing to metacoxal angle. Protibial dorsal angle with subacute triangular tooth near midlength (
<figureCitation id="DCFF2A4AFFACFFC1ABF4FB0BEAF9FB26" box="[950,1021,1180,1202]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="3" pageNumber="321">Fig. 21</figureCitation>
), apex with large, subacute lateral tooth, protibial surface with fine, shallow, setate punctures, setae dense on the upper and lower faces; length of the ectal tooth 
<quantity id="833C9B2AFFACFFC1AC7BFB7BEDADFA95" box="[1081,1193,1260,1282]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="321" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AA89FA91EBD2FA8A" box="[715,726,1286,1309]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 0.5, 
<emphasis id="76B0EADDFFACFFC1AB57FA9FEA25FA8A" box="[789,801,1288,1309]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3). Protibial spurs long, subacute, shallowly curved inner spur 
<quantity id="833C9B2AFFACFFC1AB85FAB5ED47FAA0" box="[967,1091,1314,1336]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="321" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AC17FAB7ED64FAA0" box="[1109,1120,1312,1335]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 0.6, 
<emphasis id="76B0EADDFFACFFC1AA81FAAAEBCBFAC5" box="[707,719,1341,1362]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3); length of outer spur 
<quantity id="833C9B2AFFACFFC1AB84FAABED0BFAC5" box="[966,1039,1340,1362]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="321" unit="mm" value="0.7">0.7 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AC5EFAACED23FAC5" box="[1052,1063,1339,1362]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 0.7, 
<emphasis id="76B0EADDFFACFFC1AC2BFAAAED71FAC5" box="[1129,1141,1341,1362]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3); apices semi-acute and curved. Length ratios (in mm) of tarsomeres 1–5: 1.3:0.8:0.7:0.6:0.9; tarsomeres slender, subcylindrical, with stiff, fine, long setae.
</paragraph>
<paragraph id="447B36CFFFACFFC1AA99FA30EDACF950" blockId="3.[706,1193,193,1736]" pageId="3" pageNumber="321">
<emphasis id="76B0EADDFFACFFC1AA99FA30EA81FA2A" bold="true" box="[731,901,1447,1469]" pageId="3" pageNumber="321">Male Terminalia.</emphasis>
Trilobed aedeagus (
<figureCitation id="DCFF2A4AFFACFFC1AC00FA30ED98FA2B" box="[1090,1180,1447,1469]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="3" pageNumber="321">Figs. 6, 7</figureCitation>
): phallobase 1.1 times longer than wide, curved laterally, with a semi-rectangular emagination at its base and a posterior internal emargination narrower than that at base. Median lobe fusiform in dorsal aspect, wide at base, narrowing, then widening after midlength, apically tapering to a narrowly obtuse apex gradually curved ventrally, length 
<quantity id="833C9B2AFFACFFC1ABE3F9F6ED15F9E0" box="[929,1041,1633,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="3" pageNumber="321" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AC5CF9F7ED2DF9E0" box="[1054,1065,1632,1655]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.6, 
<emphasis id="76B0EADDFFACFFC1AC28F9F5ED72F9E0" box="[1130,1142,1634,1655]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3); parameres divergent basally, gently curved at midlength, subparallel apically, with apices obliquely subtruncate, length 
<quantity id="833C9B2AFFACFFC1ABC7F925EAFFF950" box="[901,1019,1714,1735]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="3" pageNumber="321" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 mm</quantity>
(
<emphasis id="76B0EADDFFACFFC1AC4EF927ED13F950" box="[1036,1047,1712,1735]" italics="true" pageId="3" pageNumber="321">x</emphasis>
= 1.5, 
<emphasis id="76B0EADDFFACFFC1AC21F925ED6BF950" box="[1123,1135,1714,1735]" italics="true" pageId="3" pageNumber="321">n</emphasis>
= 3),
</paragraph>
<caption id="10BB6647FFABFFC6A8F2F91FED97F90B" ID-DOI="http://doi.org/10.5281/zenodo.13243122" ID-Zenodo-Dep="13243122" box="[176,1171,1672,1692]" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="4" pageNumber="322" startId="4.[176,215,1672,1691]" targetBox="[156,1169,193,1652]" targetPageId="4" targetType="figure">
<paragraph id="447B36CFFFABFFC6A8F2F91FED97F90B" blockId="4.[152,1171,1672,1716]" box="[176,1171,1672,1692]" pageId="4" pageNumber="322">
<emphasis id="76B0EADDFFABFFC6A8F2F91FE814F90B" bold="true" box="[176,272,1672,1692]" pageId="4" pageNumber="322">Figs. 1–10.</emphasis>
<taxonomicName id="83C44D4CFFABFFC6A967F91EE8FFF90B" authorityName="Champion" authorityYear="1896" box="[293,507,1673,1692]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="322" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFABFFC6A967F91EE8FFF90B" box="[293,507,1673,1692]" italics="true" pageId="4" pageNumber="322">Scaptolenus acrognathus</emphasis>
</taxonomicName>
(male). 
<emphasis id="76B0EADDFFABFFC6AA03F91FEB56F90C" bold="true" box="[577,594,1672,1691]" pageId="4" pageNumber="322">1)</emphasis>
Dorsal habitus; 
<emphasis id="76B0EADDFFABFFC6AA9FF91FEBEAF90C" bold="true" box="[733,750,1672,1691]" pageId="4" pageNumber="322">2)</emphasis>
Ventral habitus; 
<emphasis id="76B0EADDFFABFFC6AB3FF91FEA8AF90C" bold="true" box="[893,910,1672,1691]" pageId="4" pageNumber="322">3)</emphasis>
Lateral habitus; 
<emphasis id="76B0EADDFFABFFC6AC5EF91FED2AF90C" bold="true" box="[1052,1070,1672,1691]" pageId="4" pageNumber="322">4)</emphasis>
Hind wing;
</paragraph>
</caption>
<paragraph id="447B36CFFFABFFC6A8DAF937ED5FF923" blockId="4.[152,1171,1672,1716]" box="[152,1115,1696,1716]" pageId="4" pageNumber="322">
<emphasis id="76B0EADDFFABFFC6A8DAF937E9AEF924" bold="true" box="[152,170,1696,1715]" pageId="4" pageNumber="322">5)</emphasis>
Sternite VIII. Aedeagus: 
<emphasis id="76B0EADDFFABFFC6A9C6F937E892F924" bold="true" box="[388,406,1696,1715]" pageId="4" pageNumber="322">6)</emphasis>
Ventral view; 
<emphasis id="76B0EADDFFABFFC6AA50F937EB20F924" bold="true" box="[530,548,1696,1715]" pageId="4" pageNumber="322">7)</emphasis>
Dorsal view. 
<emphasis id="76B0EADDFFABFFC6AAD8F937EBA9F924" bold="true" box="[666,685,1696,1715]" pageId="4" pageNumber="322">8)</emphasis>
Sternite IX; 
<emphasis id="76B0EADDFFABFFC6AB59F937EA29F924" bold="true" box="[795,813,1696,1715]" pageId="4" pageNumber="322">9)</emphasis>
Tergite IX and X; 
<emphasis id="76B0EADDFFABFFC6AB8FF937EAEEF924" bold="true" box="[973,1002,1696,1715]" pageId="4" pageNumber="322">10)</emphasis>
Tergite VIII.
</paragraph>
<caption id="10BB6647FFAAFFC7A887F902EDACF93F" ID-DOI="http://doi.org/10.5281/zenodo.13243124" ID-Zenodo-Dep="13243124" box="[197,1192,1685,1704]" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="5" pageNumber="323" startId="5.[197,238,1685,1704]" targetBox="[173,1185,203,1656]" targetPageId="5" targetType="figure">
<paragraph id="447B36CFFFAAFFC7A887F902EDACF93F" blockId="5.[173,1192,1685,1728]" box="[197,1192,1685,1704]" pageId="5" pageNumber="323">
<emphasis id="76B0EADDFFAAFFC7A887F902E831F93F" bold="true" box="[197,309,1685,1704]" pageId="5" pageNumber="323">Figs. 11–18.</emphasis>
<taxonomicName id="83C44D4CFFAAFFC7A909F902EB28F93F" authorityName="Champion" authorityYear="1896" box="[331,556,1685,1704]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="323" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFAAFFC7A909F902EB28F93F" box="[331,556,1685,1704]" italics="true" pageId="5" pageNumber="323">Scaptolenus acrognathus</emphasis>
</taxonomicName>
(female). 
<emphasis id="76B0EADDFFAAFFC7AAC9F902EBA3F93F" bold="true" box="[651,679,1685,1704]" pageId="5" pageNumber="323">11)</emphasis>
Dorsal habitus; 
<emphasis id="76B0EADDFFAAFFC7AB7CF902EA5FF93F" bold="true" box="[830,859,1685,1704]" pageId="5" pageNumber="323">12)</emphasis>
Ventral habitus; 
<emphasis id="76B0EADDFFAAFFC7ABB5F902ED11F93F" bold="true" box="[1015,1045,1685,1704]" pageId="5" pageNumber="323">13)</emphasis>
Lateral habitus;
</paragraph>
</caption>
<paragraph id="447B36CFFFAAFFC7A8EFF93AED42F957" blockId="5.[173,1192,1685,1728]" box="[173,1094,1709,1728]" pageId="5" pageNumber="323">
<emphasis id="76B0EADDFFAAFFC7A8EFF93AE9CDF957" bold="true" box="[173,201,1709,1728]" pageId="5" pageNumber="323">14)</emphasis>
Hind wing; 
<emphasis id="76B0EADDFFAAFFC7A977F93AE855F957" bold="true" box="[309,337,1709,1728]" pageId="5" pageNumber="323">15)</emphasis>
Tergite VIII with spiculum ventrale; 
<emphasis id="76B0EADDFFAAFFC7AAD2F93AEBA8F957" bold="true" box="[656,684,1709,1728]" pageId="5" pageNumber="323">16)</emphasis>
Sternite VIII; 
<emphasis id="76B0EADDFFAAFFC7AB6BF93AEA41F957" bold="true" box="[809,837,1709,1728]" pageId="5" pageNumber="323">17)</emphasis>
Ovipositor; 
<emphasis id="76B0EADDFFAAFFC7ABF3F93AEAC9F957" bold="true" box="[945,973,1709,1728]" pageId="5" pageNumber="323">18)</emphasis>
Gonocoxites.
</paragraph>
<caption id="10BB6647FFA9FFC4A8F2FB75EBDFFA99" ID-DOI="http://doi.org/10.5281/zenodo.13243126" ID-Zenodo-Dep="13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="6" pageNumber="324" startId="6.[176,215,1250,1269]" targetBox="[177,1152,223,1182]" targetPageId="6" targetType="figure">
<paragraph id="447B36CFFFA9FFC4A8F2FB75EBDFFA99" blockId="6.[152,1171,1250,1294]" pageId="6" pageNumber="324">
<emphasis id="76B0EADDFFA9FFC4A8F2FB75E81FFB61" bold="true" box="[176,283,1250,1270]" pageId="6" pageNumber="324">Figs. 19–26.</emphasis>
<taxonomicName id="83C44D4CFFA9FFC4A972FB74EB03FB61" authorityName="Champion" authorityYear="1896" box="[304,519,1251,1270]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="324" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFA9FFC4A972FB74EB03FB61" box="[304,519,1251,1270]" italics="true" pageId="6" pageNumber="324">Scaptolenus acrognathus</emphasis>
</taxonomicName>
. Antenna: 
<emphasis id="76B0EADDFFA9FFC4AA21FB75EB7BFB62" bold="true" box="[611,639,1250,1269]" pageId="6" pageNumber="324">19)</emphasis>
Male; 
<emphasis id="76B0EADDFFA9FFC4AAFEFB75EBDCFB62" bold="true" box="[700,728,1250,1269]" pageId="6" pageNumber="324">20)</emphasis>
Female. Protibia: 
<emphasis id="76B0EADDFFA9FFC4AB37FB75EA95FB62" bold="true" box="[885,913,1250,1269]" pageId="6" pageNumber="324">21)</emphasis>
Male; 
<emphasis id="76B0EADDFFA9FFC4AB8CFB75EAEEFB62" bold="true" box="[974,1002,1250,1269]" pageId="6" pageNumber="324">22)</emphasis>
Female. Metacoxal plate: 
<emphasis id="76B0EADDFFA9FFC4A88FFB6DE9EDFA9A" bold="true" box="[205,233,1274,1293]" pageId="6" pageNumber="324">23)</emphasis>
Male; 
<emphasis id="76B0EADDFFA9FFC4A964FB6DE846FA9A" bold="true" box="[294,322,1274,1293]" pageId="6" pageNumber="324">24)</emphasis>
Female. Maxillary palp: 
<emphasis id="76B0EADDFFA9FFC4AA58FB6DEB33FA9A" bold="true" box="[538,567,1274,1293]" pageId="6" pageNumber="324">25)</emphasis>
Male; 
<emphasis id="76B0EADDFFA9FFC4AA31FB6DEB94FA9A" bold="true" box="[627,656,1274,1293]" pageId="6" pageNumber="324">26)</emphasis>
Female.
</paragraph>
</caption>
<paragraph id="447B36CFFFA9FFC4A8DAFAAEEA6AFA43" blockId="6.[151,638,1337,1733]" lastBlockId="6.[684,1172,1337,1733]" pageId="6" pageNumber="324">
central elevation in apical third of paramere with thin, decumbent setae. Sternite VIII (
<figureCitation id="DCFF2A4AFFA9FFC4A986FAC3E8F9FAFD" box="[452,509,1364,1386]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="6" pageNumber="324">Fig. 5</figureCitation>
) 1.7 times as wide as long, broader at base than at apex; basal margin straight, with rounded corners; medio-basal part gradually translucent, difficult to delimit; setae thin, longer than antennomere 6, concentrated at margins; distal margin sinuately bilobed.Sternite IX (
<figureCitation id="DCFF2A4AFFA9FFC4AA60FA4EEB52FA78" box="[546,598,1497,1519]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="6" pageNumber="324">Fig.8</figureCitation>
) 2.5 times as long as wide, straight from base to two-thirds length; distal third narrowing, with broadly rounded apex, with translucent discal area in basal third, with rounded angles, setae thin and longer in length than antennomere 6, grouped in apical third. Tergite VIII (
<figureCitation id="DCFF2A4AFFA9FFC4A8DDF9EEE9E6F918" box="[159,226,1657,1679]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="6" pageNumber="324">Fig. 10</figureCitation>
) triangular, 1.07 times as wide as long; base with tubercles on margin, stippled on apical two-thirds, setae long, slender, decumbent. Tergite IX (
<figureCitation id="DCFF2A4AFFA9FFC4AA79F938EB72F953" box="[571,630,1711,1733]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="6" pageNumber="324">Fig. 9</figureCitation>
) gradually narrowing distally, posterior margin strongly notched at middle, posterolateral angles rounded, sparsely punctate, and with setae thin and longer in length than antennomere 6, located on distal and apical margins. Tergite X reduced (
<figureCitation id="DCFF2A4AFFA9FFC4ABECFA33EAE1FA2E" box="[942,997,1444,1466]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="6" pageNumber="324">Fig. 9</figureCitation>
), no setae or punctation, apex truncate.
</paragraph>
</subSubSection>
<subSubSection id="0CDE6544FFA9FFCAAA87FA4EE8F7FB26" lastPageId="8" lastPageNumber="326" pageId="6" pageNumber="324" type="distribution">
<paragraph id="447B36CFFFA9FFC4AA87FA4EED48F9CD" blockId="6.[684,1172,1337,1733]" pageId="6" pageNumber="324">
<emphasis id="76B0EADDFFA9FFC4AA87FA4EEA46FA78" bold="true" box="[709,834,1497,1519]" pageId="6" pageNumber="324">Distribution</emphasis>
(
<figureCitation id="DCFF2A4AFFA9FFC4AB17FA4EEAA7FA78" box="[853,931,1497,1519]" captionStart="Fig" captionStartId="7.[197,228,932,951]" captionTargetBox="[200,1168,232,879]" captionTargetId="figure-423@7.[173,1193,193,914]" captionTargetPageId="7" captionText="Fig. 27. Distribution of Scaptolenus acrognathus." figureDoi="http://doi.org/10.5281/zenodo.13243130" httpUri="https://zenodo.org/record/13243130/files/figure.png" pageId="6" pageNumber="324">Fig. 27</figureCitation>
). 
<bibRefCitation id="20554B3EFFA9FFC4ABFBFA4EED6DFA78" author="Champion, G. C." box="[953,1129,1497,1519]" pageId="6" pageNumber="324" pagination="557 - 573" refId="ref4964" refString="Champion, G. C. 1896. Fam. Cebrionidae [pp. 557 - 573, pl. 25]. In: Biologia Centrali-Americana, Insecta, Coleoptera, Vol. 3, part 1, Serricornia (F. D. Godman and O. Salvin, editors). R. H. Porter, London. xv + [1] + 690 pp., 27 pl." type="book chapter" year="1896">Champion (1896)</bibRefCitation>
described this species from the state of 
<collectingRegion id="8600F82DFFA9FFC4AC74FA63ED8BF99E" box="[1078,1167,1524,1545]" country="Mexico" name="Guerrero" pageId="6" pageNumber="324">Guerrero</collectingRegion>
, 
<collectingCountry id="3CD3765FFFA9FFC4AAEFF998EBFDF9B2" box="[685,761,1551,1573]" name="Mexico" pageId="6" pageNumber="324">Mexico</collectingCountry>
. Here we report this species for the states of Estado de 
<collectingCountry id="3CD3765FFFA9FFC4AB7FF9BEEA8DF9A8" box="[829,905,1577,1599]" name="Mexico" pageId="6" pageNumber="324">México</collectingCountry>
, 
<collectingRegion id="8600F82DFFA9FFC4ABDAF9BDEAFEF9A8" box="[920,1018,1578,1599]" country="Mexico" name="Queretaro de Arteaga" pageId="6" pageNumber="324">Querétaro</collectingRegion>
, and 
<collectingRegion id="8600F82DFFA9FFC4AC7AF9BDEBE8F9CB" country="Mexico" name="San Luis Potosi" pageId="6" pageNumber="324">San Luis PotosÍ</collectingRegion>
, which represent 
<emphasis id="76B0EADDFFA9FFC4ABD6F9D3ED42F9CE" bold="true" box="[916,1094,1604,1625]" pageId="6" pageNumber="324">new state records</emphasis>
.
</paragraph>
<paragraph id="447B36CFFFA9FFC5AA87F9C8EB97FADC" blockId="6.[684,1172,1337,1733]" lastBlockId="7.[173,660,1013,1729]" lastPageId="7" lastPageNumber="325" pageId="6" pageNumber="324">
<emphasis id="76B0EADDFFA9FFC4AA87F9C8EA9CF9E2" bold="true" box="[709,920,1631,1653]" pageId="6" pageNumber="324">Sexual Dimorphism.</emphasis>
<taxonomicName id="83C44D4CFFA9FFC4ABE2F9C8ED97F9E3" authorityName="Champion" authorityYear="1896" box="[928,1171,1631,1652]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="324" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFA9FFC4ABE2F9C8ED97F9E3" box="[928,1171,1631,1652]" italics="true" pageId="6" pageNumber="324">Scaptolenus acrognathus</emphasis>
</taxonomicName>
shows well-marked sexual dimorphism, with the main differences in the total lengths and relative sizes of the appendages, the female being smaller and with shorter appendages. These differences in sizes are more evident in certain appendages such as the tarsomeres (
<figureCitation id="DCFF2A4AFFA8FFC5A9D9FBBCE8F8FBD7" box="[411,508,1067,1089]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="7" pageNumber="325">Figs. 1–3</figureCitation>
, 
<figureCitation id="DCFF2A4AFFA8FFC5AA45FBBCEB43FBD7" box="[519,583,1067,1088]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="7" pageNumber="325">11–13</figureCitation>
), tibiae (
<figureCitation id="DCFF2A4AFFA8FFC5A8F7FBD2E834FBCC" box="[181,304,1093,1115]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="7" pageNumber="325">Figs. 21, 22</figureCitation>
) and in the antennae (
<figureCitation id="DCFF2A4AFFA8FFC5AA4CFBD2EB8CFBCC" box="[526,648,1093,1115]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="7" pageNumber="325">Figs. 19, 20</figureCitation>
), which only reach the apical part of the pronotum. The pronotum is wider than long in the male (
<figureCitation id="DCFF2A4AFFA8FFC5AA0DFBECEB81FB07" box="[591,645,1147,1169]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="7" pageNumber="325">Fig. 1</figureCitation>
). The elytra of the female only cover the first four abdominal segments (
<figureCitation id="DCFF2A4AFFA8FFC5A939FB27E8BBFB52" box="[379,447,1200,1222]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="7" pageNumber="325">Fig. 13</figureCitation>
), while in the male the elytra cover the entire abdomen (
<figureCitation id="DCFF2A4AFFA8FFC5A9AEFB5CEB4DFB77" box="[492,585,1227,1249]" captionStart="Figs" captionStartId="4.[176,215,1672,1691]" captionTargetBox="[156,1169,193,1652]" captionTargetId="figure-7@4.[151,1171,193,1655]" captionTargetPageId="4" captionText="Figs. 1–10. Scaptolenus acrognathus (male). 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Hind wing;" figureDoi="http://doi.org/10.5281/zenodo.13243122" httpUri="https://zenodo.org/record/13243122/files/figure.png" pageId="7" pageNumber="325">Figs. 1–3</figureCitation>
). There are also differences in the shape of the metacoxal plates (
<figureCitation id="DCFF2A4AFFA8FFC5A8B3FA97E867FA82" box="[241,355,1280,1302]" captionStart="Figs" captionStartId="6.[176,215,1250,1269]" captionTargetBox="[177,1152,223,1182]" captionTargetId="figure-245@6.[151,1171,193,1233]" captionTargetPageId="6" captionText="Figs. 19–26. Scaptolenus acrognathus. Antenna: 19) Male; 20) Female. Protibia: 21) Male; 22) Female. Metacoxal plate: 23) Male; 24) Female. Maxillary palp: 25) Male; 26) Female." figureDoi="http://doi.org/10.5281/zenodo.13243126" httpUri="https://zenodo.org/record/13243126/files/figure.png" pageId="7" pageNumber="325">Figs. 23, 24</figureCitation>
) and in the complexity (number of veins present) of the wing venation, with the female wing venation being the simplest (
<figureCitation id="DCFF2A4AFFA8FFC5AA02FAA2EB8CFADC" box="[576,648,1333,1355]" captionStart="Figs" captionStartId="5.[197,238,1685,1704]" captionTargetBox="[173,1185,203,1656]" captionTargetId="figure-6@5.[173,1193,193,1667]" captionTargetPageId="5" captionText="Figs. 11–18. Scaptolenus acrognathus (female). 11) Dorsal habitus; 12) Ventral habitus; 13) Lateral habitus;" figureDoi="http://doi.org/10.5281/zenodo.13243124" httpUri="https://zenodo.org/record/13243124/files/figure.png" pageId="7" pageNumber="325">Fig. 14</figureCitation>
).
</paragraph>
<caption id="10BB6647FFA8FFC5A887FC33EB78FC20" ID-DOI="http://doi.org/10.5281/zenodo.13243130" ID-Zenodo-Dep="13243130" box="[197,636,932,951]" httpUri="https://zenodo.org/record/13243130/files/figure.png" pageId="7" pageNumber="325" startId="7.[197,228,932,951]" targetBox="[200,1168,232,879]" targetPageId="7" targetType="figure">
<paragraph id="447B36CFFFA8FFC5A887FC33EB78FC20" blockId="7.[197,636,932,951]" box="[197,636,932,951]" pageId="7" pageNumber="325">
<emphasis id="76B0EADDFFA8FFC5A887FC33E80CFC20" bold="true" box="[197,264,932,951]" pageId="7" pageNumber="325">Fig. 27.</emphasis>
Distribution of 
<taxonomicName id="83C44D4CFFA8FFC5A9E3FC33EB7CFC20" authorityName="Champion" authorityYear="1896" box="[417,632,932,951]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="325" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFA8FFC5A9E3FC33EB7CFC20" box="[417,632,932,951]" italics="true" pageId="7" pageNumber="325">Scaptolenus acrognathus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="447B36CFFFA8FFC5A887FAC7EA1CFBD6" blockId="7.[173,660,1013,1729]" lastBlockId="7.[705,1192,1013,1729]" pageId="7" pageNumber="325">
<emphasis id="76B0EADDFFA8FFC5A887FAC7E877FAF2" bold="true" box="[197,371,1360,1382]" pageId="7" pageNumber="325">Ecological Notes.</emphasis>
Few prior works provide biological notes for any species in the genus other than their edaphic habits and occurrence during intense summer rains (
<bibRefCitation id="20554B3EFFA8FFC5A97AFA37E890FA22" author="Fall, H. C." box="[312,404,1440,1462]" pageId="7" pageNumber="325" pagination="56 - 62" refId="ref5303" refString="Fall, H. C. 1932. New Coleoptera XV. The Canadian Entomologist 64: 56 - 62." type="journal article" year="1932">Fall 1932</bibRefCitation>
; 
<bibRefCitation id="20554B3EFFA8FFC5A9DFFA36EB1CFA21" author="Werner, F. G." box="[413,536,1440,1462]" pageId="7" pageNumber="325" pagination="26 - 27" refId="ref6163" refString="Werner, F. G. 1969. Two flights of Scaptolenus (Coleoptera: Cebrionidae). The Coleopterists Bulletin 23 (1): 26 - 27." type="journal article" year="1969">Werner 1969</bibRefCitation>
), and we feel that habitat information is essential for 
<taxonomicName id="83C44D4CFFA8FFC5AA5DFA2CE9FDFA7C" class="Insecta" family="Elateridae" genus="Scaptolenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="325" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="76B0EADDFFA8FFC5AA5DFA2CEB97FA47" box="[543,659,1467,1488]" italics="true" pageId="7" pageNumber="325">Scaptolenus</emphasis>
species.
</taxonomicName>
Recently, 
<bibRefCitation id="20554B3EFFA8FFC5A919FA41EB74FA7C" author="Martinez-Luque, E. O." box="[347,624,1493,1517]" pageId="7" pageNumber="325" pagination="267 - 278" refId="ref5880" refString="Martinez-Luque, E. O., R. W. Jones, and S. Nino- Maldonado. 2022. New species, notes on the biology of Scaptolenus LeConte (Coleoptera: Elateridae), and a key to the &quot; vagans-group &quot;. Zootaxa 5169 (3): 267 - 278." type="journal article" year="2022">
MartÍnez-Luque 
<emphasis id="76B0EADDFFA8FFC5A9B9FA41EB28FA7D" box="[507,556,1493,1515]" italics="true" pageId="7" pageNumber="325">et al.</emphasis>
(2022)
</bibRefCitation>
described an ephemeral lifespan for males of an undescribed 
<taxonomicName id="83C44D4CFFA8FFC5A953F99CE8D5F9B6" box="[273,465,1547,1569]" pageId="7" pageNumber="325">
<emphasis id="76B0EADDFFA8FFC5A953F99CE881F9B7" box="[273,389,1547,1568]" italics="true" pageId="7" pageNumber="325">Scaptolenus</emphasis>
species
</taxonomicName>
(less than 24 hours after their emergence from the ground). Immature stages develop underground, but nothing is known of their feeding habits. Copulation takes place at the ground surface with the female within her burrow and abdomen available to the male. For these edaphic animals, it may be important to study soil characteristics and their ecological associations to better define the habitats where each species is found.
</paragraph>
<paragraph id="447B36CFFFA8FFC5AA99FBD2EA9AF9B6" blockId="7.[705,1192,1013,1729]" pageId="7" pageNumber="325">
The specimens from 
<collectingRegion id="8600F82DFFA8FFC5ABE3FBD1ED05FBCC" box="[929,1025,1094,1115]" country="Mexico" name="Queretaro de Arteaga" pageId="7" pageNumber="325">Querétaro</collectingRegion>
were collected in the Sierra de Amealco Natural Region with an altitude of 
<quantity id="833C9B2AFFA8FFC5AB4BFBECEA9DFB07" box="[777,921,1147,1168]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.8" metricValueMin="1.2" pageId="7" pageNumber="325" unit="m" value="2000.0" valueMax="2800.0" valueMin="1200.0">
<geoCoordinate id="21F05008FFA8FFC5AB4BFBECEA3AFB07" box="[777,830,1147,1168]" degrees="1,200" direction="north" orientation="latitude" pageId="7" pageNumber="325" precision="55" value="1.2">1,200</geoCoordinate>
<geoCoordinate id="21F05008FFA8FFC5AB7CFBECEA85FB07" box="[830,897,1147,1168]" degrees="2,800" direction="west" orientation="longitude" pageId="7" pageNumber="325" precision="55" value="-2.8">–2,800</geoCoordinate>
m
</quantity>
in oak forests surrounded by agricultural areas (
<bibRefCitation id="20554B3EFFA8FFC5ABC0FB02ED54FB3C" author="Bayona-Celis, A." box="[898,1104,1173,1195]" pageId="7" pageNumber="325" pagination="15 - 22" refId="ref4767" refString="Bayona-Celis, A. 2016 a. Capitulo 1. El Estado de Queretaro [pp. 15 - 22]. In: Historia Natural de Queretaro (W. R. Jones and V. Serrano-Cardenas, editors). Universidad Autonoma de Queretaro, Editorial Universitaria, Coleccion Academia, Serie Nodos, Mexico." type="book chapter" year="2016">Bayona-Celis 2016a</bibRefCitation>
; GarcÍa-Sánchez and Meraz 2016). In this region there is a sub-humid temperate climate with summer rains providing an average annual precipitation of 
<quantity id="833C9B2AFFA8FFC5AA9CFA97EA41FA81" box="[734,837,1280,1302]" metricMagnitude="-1" metricUnit="m" metricValue="8.615" pageId="7" pageNumber="325" unit="mm" value="861.5">861.5 mm</quantity>
, May to October, with periodic well-marked periods of drought (
<bibRefCitation id="20554B3EFFA8FFC5AB98FA8CEDA1FAA6" author="Bayona-Celis, A." box="[986,1189,1307,1329]" pageId="7" pageNumber="325" pagination="15 - 22" refId="ref4767" refString="Bayona-Celis, A. 2016 a. Capitulo 1. El Estado de Queretaro [pp. 15 - 22]. In: Historia Natural de Queretaro (W. R. Jones and V. Serrano-Cardenas, editors). Universidad Autonoma de Queretaro, Editorial Universitaria, Coleccion Academia, Serie Nodos, Mexico." type="book chapter" year="2016">Bayona-Celis 2016a</bibRefCitation>
; Luna-Soria and Suzán-Azpiri 2016). The average annual temperature recorded is 14.4 °C with a minimum temperature between 3.8 and 9.8 °C and maximum temperatures from 19.2 °C to 24.8 °C (Luna-Soria and Suzán-Azpiri 2016). The soil at the sampling locality belongs to the Feozems group, which is characterized by being dark-colored with porous properties and is rich in organic matter (
<bibRefCitation id="20554B3EFFA8FFC5AA8BF99CEA97F9B6" author="Bayona-Celis, A." box="[713,915,1547,1569]" pageId="7" pageNumber="325" pagination="67 - 87" refId="ref4825" refString="Bayona-Celis, A. 2016 b. CapItulo 4. Los suelos del Estado de Queretaro [pp. 67 - 87]. In: Historia Natural de Queretaro (W. R. Jones and V. Serrano-Cardenas, editors). Universidad Autonoma de Queretaro, Editorial Universitaria, Coleccion Academia, Serie Nodos, Mexico." type="book chapter" year="2016">Bayona-Celis 2016b</bibRefCitation>
).
</paragraph>
<paragraph id="447B36CFFFA8FFCAAA99F9B2E822FEE0" blockId="7.[705,1192,1013,1729]" lastBlockId="8.[151,639,193,1202]" lastPageId="8" lastPageNumber="326" pageId="7" pageNumber="325">
The locality in the 
<collectingRegion id="8600F82DFFA8FFC5ABD9F9B2ED49F9AC" box="[923,1101,1573,1595]" country="Mexico" name="Mexico" pageId="7" pageNumber="325">Estado de México</collectingRegion>
is in the Valle de Bravo basin, in the physiographic province Eje Neovolcánico Transversal. This region has a temperate sub-humid climate with summer rains, with well-marked dry and rainy seasons (
<bibRefCitation id="20554B3EFFA8FFC5AC27F907EBF3F957" author="INEGI &amp; Instituto Nacional de Estadistica" pageId="7" pageNumber="325" refId="ref5443" refString="INEGI (Instituto Nacional de Estadistica y Geografia). 1977. Carta topografica Valle de Bravo E 14 - A 46, Escala 1: 50000. Mexico." type="journal volume" year="1977">INEGI 1977</bibRefCitation>
). The average annual rainfall ranges from 
<quantity id="833C9B2AFFA7FFCAA8DAFF55E854FF40" box="[152,336,193,215]" metricMagnitude="0" metricUnit="m" metricValue="1.405" metricValueMax="1.5" metricValueMin="1.31" pageId="8" pageNumber="326" unit="mm" value="1405.0" valueMax="1500.0" valueMin="1310.0">1,310 to 1,500 mm</quantity>
. The main 
<typeStatus id="9B7F886DFFA7FFCAA9F8FF55E8E7FF40" box="[442,483,194,215]" pageId="8" pageNumber="326">type</typeStatus>
of vegetation is pine-oak forest (
<bibRefCitation id="20554B3EFFA7FFCAA972FF4BE8D4FF66" author="Rzedowski, J." box="[304,464,220,242]" pageId="8" pageNumber="326" refId="ref6128" refString="Rzedowski, J. 1981. Vegetacion de Mexico. Cuarta reimpresion, Editorial Limusa, Mexico, 432 pp." type="book" year="1981">Rzedowski 1981</bibRefCitation>
). This 
<typeStatus id="9B7F886DFFA7FFCAAA4CFF4AEB33FF65" box="[526,567,221,242]" pageId="8" pageNumber="326">type</typeStatus>
of vegetation is associated in this region with an altitude of 
<quantity id="833C9B2AFFA7FFCAA8F0FE85E842FEB0" box="[178,326,274,295]" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="8" pageNumber="326" unit="m" value="2600.0" valueMax="3000.0" valueMin="2200.0">
<geoCoordinate id="21F05008FFA7FFCAA8F0FE85E9ECFEB0" box="[178,232,274,295]" degrees="2,200" direction="north" orientation="latitude" pageId="8" pageNumber="326" precision="55" value="2.2">2,200</geoCoordinate>
<geoCoordinate id="21F05008FFA7FFCAA8AAFE85E82EFEB0" box="[232,298,274,295]" degrees="3,000" direction="west" orientation="longitude" pageId="8" pageNumber="326" precision="55" value="-3.0">–3,000</geoCoordinate>
m
</quantity>
. Predominant soil in the basin is Andosol, which is formed from volcanic ash and glass, and is dark in color, porous, and very fertile (
<bibRefCitation id="20554B3EFFA7FFCAA8DCFEF5E81FFEE0" author="INEGI &amp; Instituto Nacional de Estadistica" box="[158,283,353,375]" pageId="8" pageNumber="326" refId="ref5501" refString="INEGI (Instituto Nacional de Estadistica y Geografia). 2015. GuIa para la interpretacion de cartografIa: EdafologIa: Escala 1: 250 000. Serie III / Instituto Nacional de EstadIstica y GeografIa, Mexico, 32 pp." type="book" year="2015">INEGI 2015</bibRefCitation>
).
</paragraph>
<paragraph id="447B36CFFFA7FFCAA8F2FEEBEB7BFCAB" blockId="8.[151,639,193,1202]" pageId="8" pageNumber="326">
The record of 
<taxonomicName id="83C44D4CFFA7FFCAA97BFEEAE8CCFE06" authorityName="Champion" authorityYear="1896" box="[313,456,380,402]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="326" phylum="Arthropoda" rank="species" species="acrognathus">
<emphasis id="76B0EADDFFA7FFCAA97BFEEAE8CCFE06" box="[313,456,380,402]" italics="true" pageId="8" pageNumber="326">S. acrognathus</emphasis>
</taxonomicName>
in 
<collectingRegion id="8600F82DFFA7FFCAA9A5FEEBEB79FE03" box="[487,637,380,404]" country="Mexico" name="San Luis Potosi" pageId="8" pageNumber="326">San Luis PotosÍ</collectingRegion>
lacks specific geographic location data or other habitat information for its collecting site. The municipality of Matehuala is located in the northern part of the state, in the area of the Altiplano Mexicano (
<bibRefCitation id="20554B3EFFA7FFCAA8DCFD95E870FD80" author="Pedraza-Montes, J. F." box="[158,372,514,536]" pageId="8" pageNumber="326" refId="ref6022" refString="Pedraza-Montes, J. F. 1994. Sinopsis Historica de los Municipios del Estado de San Luis PotosI. Centro de Desarrollo Municipal, Gobierno del Estado de San Luis PotosI, Mexico, 61 pp." type="book" year="1994">Pedraza-Montes 1994</bibRefCitation>
). This municipality presents a topography mainly of undulating plains, with scattered hills and ravines. These conditions correspond to sub-humid temperate climates, located at altitudes of 
<quantity id="833C9B2AFFA7FFCAA8AEFDFBE87BFD15" box="[236,383,620,642]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="2.1" metricValueMin="1.2" pageId="8" pageNumber="326" unit="m" value="1650.0" valueMax="2100.0" valueMin="1200.0">
<geoCoordinate id="21F05008FFA7FFCAA8AEFDFBE825FD16" box="[236,289,620,641]" degrees="1,200" direction="north" orientation="latitude" pageId="8" pageNumber="326" precision="55" value="1.2">1,200</geoCoordinate>
<geoCoordinate id="21F05008FFA7FFCAA963FDFBE860FD16" box="[289,356,620,641]" degrees="2,100" direction="west" orientation="longitude" pageId="8" pageNumber="326" precision="55" value="-2.1">–2,100</geoCoordinate>
m
</quantity>
, where we find associated pine-oak vegetation (
<bibRefCitation id="20554B3EFFA7FFCAA938FD10EB2EFD0A" author="Rzedowski, J." box="[378,554,647,669]" pageId="8" pageNumber="326" pagination="5 - 112" refId="ref6060" refString="Rzedowski, J. 1960. Notas sobre la flora y la vegetacion del estado de san Luis PotosI. VII. Vegetacion del Valle de San Luis PotosI. Acta CientIfica Potosina 4 (1): 5 - 112." type="journal article" year="1960">Rzedowski 1960</bibRefCitation>
, 
<bibRefCitation id="20554B3EFFA7FFCAAA7DFD10EB77FD0A" author="Rzedowski, J." box="[575,627,647,669]" pageId="8" pageNumber="326" pagination="5 - 291" refId="ref6102" refString="Rzedowski, J. 1965. Vegetacion del estado de San Luis PotosI. Acta CientIfica Potosina 5 (2): 5 - 291." type="journal article" year="1965">1965</bibRefCitation>
). Unfortunately, this vegetation is currently considered an isolated relict within the municipality of Matehuala. In 
<collectingRegion id="8600F82DFFA7FFCAA966FD40E8BBFD79" box="[292,447,727,750]" country="Mexico" name="San Luis Potosi" pageId="8" pageNumber="326">San Luis PotosÍ</collectingRegion>
, these forests are in the northern part, on the eastern slope of Sierra de Catorce, and extend to the southeast of the state, in a portion of the Sierra Madre Oriental (
<bibRefCitation id="20554B3EFFA7FFCAA9BEFCB0EB70FCAB" author="INEGI &amp; Instituto Nacional de Estadistica" box="[508,628,807,828]" pageId="8" pageNumber="326" refId="ref5473" refString="INEGI (Instituto Nacional de Estadistica y Geografia). 2002. SIntesis de informacion geografica del estado de San Luis PotosI. Mexico, 83 pp." type="book" year="2002">INEGI 2002</bibRefCitation>
).
</paragraph>
<paragraph id="447B36CFFFA7FFCAA8F2FCD5E8F7FB26" blockId="8.[151,639,193,1202]" pageId="8" pageNumber="326">
In the case of the 
<typeStatus id="9B7F886DFFA7FFCAA91BFCD5E886FCC0" box="[345,386,834,855]" pageId="8" pageNumber="326">type</typeStatus>
locality, this was recorded by 
<bibRefCitation id="20554B3EFFA7FFCAA8FAFCCBE865FCE5" author="Champion, G. C." box="[184,353,860,882]" pageId="8" pageNumber="326" pagination="557 - 573" refId="ref4964" refString="Champion, G. C. 1896. Fam. Cebrionidae [pp. 557 - 573, pl. 25]. In: Biologia Centrali-Americana, Insecta, Coleoptera, Vol. 3, part 1, Serricornia (F. D. Godman and O. Salvin, editors). R. H. Porter, London. xv + [1] + 690 pp., 27 pl." type="book chapter" year="1896">Champion (1896)</bibRefCitation>
as Omilteme, 
<collectingRegion id="8600F82DFFA7FFCAA9B3FCCBEB4DFCE6" box="[497,585,860,881]" country="Mexico" name="Guerrero" pageId="8" pageNumber="326">Guerrero</collectingRegion>
. This locality corresponds to the community of Omiltemi located to the west of the municipality of Chilpancingo de los Bravo in the center of the state of 
<collectingRegion id="8600F82DFFA7FFCAAA52FC3BEB61FC56" box="[528,613,940,961]" country="Mexico" name="Guerrero" pageId="8" pageNumber="326">Guerrero</collectingRegion>
in southeastern 
<collectingCountry id="3CD3765FFFA7FFCAA953FC50E858FC4A" box="[273,348,967,989]" name="Mexico" pageId="8" pageNumber="326">Mexico</collectingCountry>
. This locality has a very rugged topography with elevations of 
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<geoCoordinate id="21F05008FFA7FFCAA9A6FC75EB1CFC60" box="[484,536,994,1015]" degrees="1,900" direction="north" orientation="latitude" pageId="8" pageNumber="326" precision="55" value="1.9">1,900</geoCoordinate>
<geoCoordinate id="21F05008FFA7FFCAAA5AFC75EB5EFC60" box="[536,602,994,1015]" degrees="2,800" direction="west" orientation="longitude" pageId="8" pageNumber="326" precision="55" value="-2.8">–2,800</geoCoordinate>
m
</quantity>
(
<bibRefCitation id="20554B3EFFA7FFCAA8DCFC6AE86DFB86" author="Garcia-Rendon, M. M." box="[158,361,1020,1044]" pageId="8" pageNumber="326" refId="ref5324" refString="Garcia-Rendon, M. M. 1988. Distribucion y estructura de las comunidades arboreas del Parque Ecologico Estatal Omiltemi, Guerrero; una contribucion a su planificacion. Tesis Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, Mexico, CDMX, 107 pp." type="book" year="1988">GarcÍa-Rendón 1988</bibRefCitation>
). At the highest points, there is a sub-humid temperate climate, with average annual temperatures of 15 °C to 24 °C. The average annual rainfall is 
<quantity id="833C9B2AFFA7FFCAA97CFBDBE8A6FBF5" box="[318,418,1100,1122]" metricMagnitude="0" metricUnit="m" metricValue="1.65" pageId="8" pageNumber="326" unit="mm" value="1650.0">1,650 mm</quantity>
, with well-marked wet and dry seasons (Miranda and Sharp 1950). The 
<typeStatus id="9B7F886DFFA7FFCAA8DAFB15E9C5FB00" box="[152,193,1154,1175]" pageId="8" pageNumber="326">type</typeStatus>
of vegetation is pine-oak forest, with soil of volcanic origin (
<bibRefCitation id="20554B3EFFA7FFCAA977FB0BE8ECFB26" author="Meave, J. &amp; Soto, L. M. &amp; Calvo-Irabien, H." box="[309,488,1180,1202]" pageId="8" pageNumber="326" pagination="31 - 77" refId="ref5936" refString="Meave, J., M. A. Soto, L. M. Calvo-Irabien, H. Paz-Hernandez, and S. Valencia-Avalos. 1992. Analisis sicecologico del bosque mesofilo de montana de Omiltemi, Guerrero. Boletin de la Sociedad Botanica de Mexico 52: 31 - 77." type="journal article" year="1992">
Meave 
<emphasis id="76B0EADDFFA7FFCAA93CFB0AE8ABFB26" box="[382,431,1180,1202]" italics="true" pageId="8" pageNumber="326">et al.</emphasis>
1992
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>