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<document id="C3C374DAF48EB57F12E88F6B6F77C451" ID-DOI="10.11646/zootaxa.4028.4.1" ID-GBIF-Dataset="6fe19834-981f-4094-aa8f-3784b3743522" ID-ISSN="1175-5326" ID-Zenodo-Dep="254170" ID-ZooBank="DE76F18B-422D-4D97-93FD-F211F691F591" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461287286300" checkinUser="plazi" docAuthor="Dupérré, Nadine" docDate="2015" docId="BC6587896A7EFFAC7397FB2BFCC6355A" docLanguage="en" docName="zt04028p484.pdf" docOrigin="Zootaxa 4028 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chococtenus kashakara Dupérré, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="472" masterDocId="405CFFF16A6DFFB97300FFACFF9A3742" masterDocTitle="Description of a new genus and thirteen new species of Ctenidae (Araneae, Ctenidae) from the Chocó region of Ecuador" masterLastPageNumber="484" masterPageNumber="451" pageNumber="470" updateTime="1698615451779" updateUser="plazi">
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<mods:title id="753F1C4360A350919F3214D1B03B9AA8">Description of a new genus and thirteen new species of Ctenidae (Araneae, Ctenidae) from the Chocó region of Ecuador</mods:title>
</mods:titleInfo>
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<mods:namePart id="CBF7E8281031A59D61BDEA5AE8F8A82A">Dupérré, Nadine</mods:namePart>
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<mods:title id="BE5F8FA2EB0B626F1BB92BD6764034F9">Zootaxa</mods:title>
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<mods:date id="A6001FD55ECE38F9DF1AFA51FFEA0A2B">2015</mods:date>
<mods:detail id="C0A3935A8E1A4CD431B86FEE2EC086CD" type="volume">
<mods:number id="E7970109A2E3E4BE849ACD39DE0D3616">4028</mods:number>
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<mods:identifier id="3DF9F6A9CEF3CD20904A6183A9B6B219" type="DOI">10.11646/zootaxa.4028.4.1</mods:identifier>
<mods:identifier id="D1D34F10D38242F55B6B28D757B2EAED" type="GBIF-Dataset">6fe19834-981f-4094-aa8f-3784b3743522</mods:identifier>
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<treatment id="BC6587896A7EFFAC7397FB2BFCC6355A" ID-DOI="http://doi.org/10.5281/zenodo.6097697" ID-GBIF-Taxon="119641647" ID-Zenodo-Dep="6097697" LSID="urn:lsid:plazi:treatment:BC6587896A7EFFAC7397FB2BFCC6355A" httpUri="http://treatment.plazi.org/id/BC6587896A7EFFAC7397FB2BFCC6355A" lastPageId="21" lastPageNumber="472" pageId="19" pageNumber="470">
<subSubSection id="7CD665146A7EFFAA7397FB2BFEDF3381" pageId="19" pageNumber="470" type="nomenclature">
<paragraph id="3473369F6A7EFFAA7397FB2BFDCB33E0" blockId="19.[151,593,1159,1219]" box="[151,593,1159,1186]" pageId="19" pageNumber="470">
<heading id="6F3B81F36A7EFFAA7397FB2BFDCB33E0" bold="true" box="[151,593,1159,1186]" fontSize="11" level="1" pageId="19" pageNumber="470" reason="1">
<emphasis id="06B8EA8D6A7EFFAA7397FB2BFDCB33E0" bold="true" box="[151,593,1159,1186]" pageId="19" pageNumber="470">
<taxonomicName id="F3CC4D1C6A7EFFAA7397FB2BFE2333E3" ID-CoL="69YMQ" box="[151,441,1159,1185]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="kashakara" status="sp. nov.">
<emphasis id="06B8EA8D6A7EFFAA7397FB2BFE2333E3" bold="true" box="[151,441,1159,1185]" italics="true" pageId="19" pageNumber="470">Chococtenus kashakara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D8B57F66A7EFFAA72C1FB24FDCB33E0" box="[449,593,1160,1186]" pageId="19" pageNumber="470" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3473369F6A7EFFAA7397FB06FEDF3381" blockId="19.[151,593,1159,1219]" box="[151,325,1194,1219]" pageId="19" pageNumber="470">
<figureCitation id="ACF72A1A6A7EFFAA7397FB06FE8D3381" box="[151,279,1194,1219]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Figs 3841</figureCitation>
,
<figureCitation id="ACF72A1A6A7EFFAA7222FB06FEDB3381" box="[290,321,1194,1219]" captionStart="FIGURE 57" captionStartId="30.[151,250,1231,1253]" captionTargetBox="[159,1427,201,1197]" captionTargetId="figure@30.[151,1436,193,1210]" captionTargetPageId="30" captionText="FIGURE 57. Map of Ecuador showing distribution records of the thirteen new species of the genus Chococtenus. The main collecting site, Otonga Biological Reserve (white box with x inside), holds 10 species (not found: C. suffuscus, C. otongachi and C. piemontana). Other known localities for: C. otonga (white lozenge), C. cappuccino (white x), C. lasdamas (white circle), C. suffuscus (white plus sign), C. otongachi (white rectangle) and C. piemontana (upside down white triangle)." httpUri="https://zenodo.org/record/254189/files/figure.png" pageId="19" pageNumber="470">57</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7CD665146A7EFFAA7397FB5EFC7C3145" pageId="19" pageNumber="470" type="materials_examined">
<paragraph id="3473369F6A7EFFAA7397FB5EFE543235" blockId="19.[151,1437,1266,1975]" pageId="19" pageNumber="470">
<emphasis id="06B8EA8D6A7EFFAA7397FB5EFED03249" bold="true" box="[151,330,1266,1291]" pageId="19" pageNumber="470">
<typeStatus id="EB77883D6A7EFFAA7397FB5EFF483249" box="[151,210,1266,1291]" pageId="19" pageNumber="470">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="EB77883D6A7EFFAA729DFB5EFE653249" box="[413,511,1266,1291]" pageId="19" pageNumber="470" type="holotype">holotype</typeStatus>
from
<collectingCountry id="4CDB760F6A7EFFAA714EFB5EFD373249" box="[590,685,1266,1291]" name="Ecuador" pageId="19" pageNumber="470">Ecuador</collectingCountry>
, Cotopaxi Province,
<taxonomicName id="F3CC4D1C6A7EFFAA70A9FB5FFC613249" box="[937,1019,1267,1291]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="483" phylum="Arthropoda" rank="species" species="otonga">Otonga</taxonomicName>
Biological Reserve (-
<geoCoordinate id="51F850586A7EFFAA77FFFB5EFA1C3249" box="[1279,1414,1266,1291]" direction="south" orientation="latitude" pageId="19" pageNumber="470" precision="1" value="-0.42261">00.42261°S</geoCoordinate>
-
<geoCoordinate id="51F850586A7EFFAA7397FABAFE86326D" box="[151,284,1302,1327]" direction="west" orientation="longitude" pageId="19" pageNumber="470" precision="5" value="-79.5107">79.5107°W</geoCoordinate>
),
<date id="4072105F6A7EFFAA7231FABAFD93326D" box="[305,521,1302,1327]" pageId="19" pageNumber="470" value="2014-08-16" valueMax="2014-09-05" valueMin="2014-08-16">16.viii.05.ix.2014</date>
, pitfall,
<quantity id="F3349B7A6A7EFFAA716BFABAFD5E326C" box="[619,708,1302,1327]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="19" pageNumber="470" unit="m" value="2225.0">2225 m</quantity>
, E. Tapia, C. Tapia, N. Dupérré (
<collectionCode id="52DDAE5A6A7EFFAA7746FABBFB03326D" box="[1094,1177,1303,1327]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="470">QCAZ</collectionCode>
). One female
<typeStatus id="EB77883D6A7EFFAA763CFABBFA06326D" box="[1340,1436,1303,1327]" pageId="19" pageNumber="470" type="paratype">paratype</typeStatus>
from
<collectingCountry id="4CDB760F6A7EFFAA73D9FA96FEA23211" box="[217,312,1338,1363]" name="Ecuador" pageId="19" pageNumber="470">Ecuador</collectingCountry>
, Cotopaxi Province,
<taxonomicName id="F3CC4D1C6A7EFFAA7130FA97FD193211" box="[560,643,1339,1363]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="483" phylum="Arthropoda" rank="species" species="otonga">Otonga</taxonomicName>
Biological Reserve,
<date id="4072105F6A7EFFAA707CFA96FBA03211" box="[892,1082,1338,1363]" pageId="19" pageNumber="470" value="2014-05-24" valueMax="2014-06-08" valueMin="2014-05-24">24.v.08.vi.2014</date>
, hand collecting, E. Tapia, C. Tapia, N. Dupérré (
<collectionCode id="52DDAE5A6A7EFFAA726FFAF3FE583235" box="[367,450,1375,1399]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="470">QCAZ</collectionCode>
).
</paragraph>
<paragraph id="3473369F6A7EFFAA73C7FA2EFC7C3145" blockId="19.[151,1437,1266,1975]" pageId="19" pageNumber="470">
<emphasis id="06B8EA8D6A7EFFAA73C7FA2EFDA532D9" bold="true" box="[199,575,1410,1435]" pageId="19" pageNumber="470">Additional material examined.</emphasis>
<collectingCountry id="4CDB760F6A7EFFAA714BFA2EFD3732D9" box="[587,685,1410,1435]" name="Ecuador" pageId="19" pageNumber="470">Ecuador</collectingCountry>
:
<emphasis id="06B8EA8D6A7EFFAA71BDFA2FFC0E32D9" box="[701,916,1411,1435]" italics="true" pageId="19" pageNumber="470">Cotopaxi Province</emphasis>
:
<taxonomicName id="F3CC4D1C6A7EFFAA70A9FA2FFC6132D9" box="[937,1019,1411,1435]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="483" phylum="Arthropoda" rank="species" species="otonga">Otonga</taxonomicName>
Biological Reserve (-
<geoCoordinate id="51F850586A7EFFAA77FFFA2EFA1C32D9" box="[1279,1414,1410,1435]" direction="south" orientation="latitude" pageId="19" pageNumber="470" precision="1" value="-0.42261">00.42261°S</geoCoordinate>
-
<geoCoordinate id="51F850586A7EFFAA7397FA0AFE8632FD" box="[151,284,1446,1471]" direction="west" orientation="longitude" pageId="19" pageNumber="470" precision="5" value="-79.5107">79.5107°W</geoCoordinate>
), 24.v.08.
<date id="4072105F6A7EFFAA7297FA0AFD9632FD" box="[407,524,1446,1471]" pageId="19" pageNumber="470" value="2014-06-01">vi.2014, 1</date>
♀, pitfall,
<quantity id="F3349B7A6A7EFFAA7185FA0AFD4C32FD" box="[645,726,1446,1471]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="19" pageNumber="470" unit="m" value="2225.0">2225m</quantity>
, E. Tapia, C. Tapia, N. Dupérré (DTC); 19.ix.02.
<date id="4072105F6A7EFFAA7619FA0AFA1D32FD" box="[1305,1415,1446,1471]" pageId="19" pageNumber="470" value="2014-10-01">
x.2014,
<specimenCount id="22CAFD166A7EFFAA7679FA0BFA1D32FD" box="[1401,1415,1447,1471]" pageId="19" pageNumber="470" type="generic">1</specimenCount>
</date>
♂ (DTC).
<taxonomicName id="F3CC4D1C6A7EFFAA73F0FA67FED932A1" box="[240,323,1483,1507]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="483" phylum="Arthropoda" rank="species" species="otonga">Otonga</taxonomicName>
Biological Reserve, 21.vi.02.
<date id="4072105F6A7EFFAA719DFA66FC8D32A1" box="[669,791,1482,1507]" pageId="19" pageNumber="470" value="2014-07-01">
vii.2014,
<specimenCount id="22CAFD166A7EFFAA700CFA67FC8D32A1" box="[780,791,1483,1507]" pageId="19" pageNumber="470" type="generic">1</specimenCount>
</date>
♂, pitfall, E. Tapia, C. Tapia, N. Dupérré (DTC);
<taxonomicName id="F3CC4D1C6A7EFFAA764AFA67FA0632A1" box="[1354,1436,1483,1507]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="483" phylum="Arthropoda" rank="species" species="otonga">Otonga</taxonomicName>
(-
<geoCoordinate id="51F850586A7EFFAA73A8FA42FE9F3145" box="[168,261,1518,1543]" direction="south" orientation="latitude" pageId="19" pageNumber="470" precision="925" value="-0.41666666">00°25S</geoCoordinate>
-
<geoCoordinate id="51F850586A7EFFAA7215FA43FE1A3145" box="[277,384,1519,1543]" direction="west" orientation="longitude" pageId="19" pageNumber="470" precision="925" value="-79.0">79°00W</geoCoordinate>
), 09.
<date id="4072105F6A7EFFAA72B6FA42FDAE3145" box="[438,564,1518,1543]" pageId="19" pageNumber="470" value="1997-08-01">
viii.1997,
<specimenCount id="22CAFD166A7EFFAA7129FA43FDAE3145" box="[553,564,1519,1543]" pageId="19" pageNumber="470" type="generic">1</specimenCount>
</date>
♂,
<quantity id="F3349B7A6A7EFFAA715BFA43FD313145" box="[603,683,1519,1543]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="470" unit="m" value="2000.0">2000m</quantity>
, I. Tapia, P. Ponce (
<collectionCode id="52DDAE5A6A7EFFAA7087FA43FC403145" box="[903,986,1519,1543]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="470">QCAZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="7CD665146A7EFFAA73C7F9BEFC8C310D" pageId="19" pageNumber="470" type="etymology">
<paragraph id="3473369F6A7EFFAA73C7F9BEFC8C310D" blockId="19.[151,1437,1266,1975]" pageId="19" pageNumber="470">
<emphasis id="06B8EA8D6A7EFFAA73C7F9BEFED43169" bold="true" box="[199,334,1554,1579]" pageId="19" pageNumber="470">Etymology.</emphasis>
The specific name is a noun in apposition taken from the Kichwa language referring to the typical tree found in the cloud forest (
<taxonomicName id="F3CC4D1C6A7EFFAA72EAF99BFDF8310D" box="[490,610,1591,1615]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
, genus
<taxonomicName id="F3CC4D1C6A7EFFAA71B4F99BFC9F310C" box="[692,773,1591,1614]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="genus">
<emphasis id="06B8EA8D6A7EFFAA71B4F99BFC9F310C" box="[692,773,1591,1614]" italics="true" pageId="19" pageNumber="470">Ocotea</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="7CD665146A7EFFAA73C7F9F6FD5E319D" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="3473369F6A7EFFAA73C7F9F6FD5E319D" blockId="19.[151,1437,1266,1975]" pageId="19" pageNumber="470">
<emphasis id="06B8EA8D6A7EFFAA73C7F9F6FEDB3131" bold="true" box="[199,321,1626,1651]" pageId="19" pageNumber="470">Diagnosis.</emphasis>
Males are diagnosed by their unique median apophysis with strong projection basally (
<figureCitation id="ACF72A1A6A7EFFAA7638F9F6FA153131" box="[1336,1423,1626,1651]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Fig. 38</figureCitation>
); from
<taxonomicName id="F3CC4D1C6A7EFFAA73DBF9D3FEF131D5" box="[219,363,1662,1687]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="lasdamas">
<emphasis id="06B8EA8D6A7EFFAA73DBF9D3FEF131D5" box="[219,363,1662,1687]" italics="true" pageId="19" pageNumber="470">C. lasdamas</emphasis>
</taxonomicName>
by their wide embolus base (
<figureCitation id="ACF72A1A6A7EFFAA71D9F9D2FCAB31D5" box="[729,817,1662,1687]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Fig. 38</figureCitation>
) narrow in the later species (
<figureCitation id="ACF72A1A6A7EFFAA779BF9D2FB6C31D5" box="[1179,1270,1662,1687]" captionStart="FIGURES 25 28" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[256,1332,203,1902]" captionTargetId="figure@14.[242,1346,193,1907]" captionTargetPageId="14" captionText="FIGURES 25 28. Chococtenus lasdamas new species. Male. 25, 26. Female. 27, 28. 25. Palp, ventral view. 26. Palp, retrolateral view. 27. Epigynum, ventral view. 28. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254177/files/figure.png" pageId="19" pageNumber="470">Fig. 25</figureCitation>
). Females are distinguished from all species of the genus by the medially positioned lateral processes and from
<taxonomicName id="F3CC4D1C6A7EFFAA77E6F90FFAE231F9" box="[1254,1400,1698,1723]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="otongachi">
<emphasis id="06B8EA8D6A7EFFAA77E6F90FFAE231F9" box="[1254,1400,1698,1723]" italics="true" pageId="19" pageNumber="470">C. otongachi</emphasis>
</taxonomicName>
by their medium sized unsclerotized sector (
<figureCitation id="ACF72A1A6A7EFFAA7163F96AFD2D319D" box="[611,695,1734,1759]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7CD665146A7EFFAC73C7F946FB3736B7" lastPageId="21" lastPageNumber="472" pageId="19" pageNumber="470" type="description">
<paragraph id="3473369F6A7EFFAA73C7F946FD9730F5" blockId="19.[151,1437,1266,1975]" pageId="19" pageNumber="470">
<emphasis id="06B8EA8D6A7EFFAA73C7F946FDBB3041" bold="true" box="[199,545,1770,1795]" pageId="19" pageNumber="470">
Description.
<emphasis id="06B8EA8D6A7EFFAA7261F946FD833041" bold="true" box="[353,537,1770,1795]" italics="true" pageId="19" pageNumber="470">
Male (
<typeStatus id="EB77883D6A7EFFAA72AEF946FD8A3041" box="[430,528,1770,1795]" pageId="19" pageNumber="470" type="holotype">holotype</typeStatus>
)
</emphasis>
.
</emphasis>
Carapace length: 4.1; carapace width: 3.7. COLORATION: Unknown, in bad shape. LEGS: All legs missing. GENITALIA: Palpal cymbial keel strong (
<figureCitation id="ACF72A1A6A7EFFAA7715F8A2FBF43065" box="[1045,1134,1806,1831]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Fig. 38</figureCitation>
). Ventral tibial apophysis elongated, thumb-shaped, apically transparent, retrolateral tibial apophysis wide basally, spine-like (
<figureCitation id="ACF72A1A6A7EFFAA760AF89EFA0A3009" box="[1290,1424,1842,1867]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Figs 38, 39</figureCitation>
). Tegulum oval; median apophysis with large projection basally; conductor lamelliform, short, hyaline; membranous tegular process hyaline, short, hidden; embolus base wide, not twisted; embolus wide basally, originating prolaterally, tip pointed (
<figureCitation id="ACF72A1A6A7EFFAA72ACF832FE6530F5" box="[428,511,1950,1975]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="19" pageNumber="470">Fig. 38</figureCitation>
).
</paragraph>
<caption id="60B366176A79FFAD7397F800FC5D30A3" httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="20" pageNumber="471" targetBox="[227,1334,198,1894]" targetPageId="20">
<paragraph id="3473369F6A79FFAD7397F800FC5D30A3" blockId="20.[151,1436,1964,2017]" pageId="20" pageNumber="471">
<emphasis id="06B8EA8D6A79FFAD7397F800FEC23080" bold="true" box="[151,344,1964,1987]" pageId="20" pageNumber="471">FIGURES 3841.</emphasis>
<taxonomicName id="F3CC4D1C6A79FFAD7263F800FDC13080" box="[355,603,1964,1986]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="kashakara" status="sp. nov.">
<emphasis id="06B8EA8D6A79FFAD7263F800FDC13080" box="[355,603,1964,1986]" italics="true" pageId="20" pageNumber="471">Chococtenus kashakara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D8B57F66A79FFAD7166F801FD7D3080" box="[614,743,1964,1986]" pageId="20" pageNumber="471" rank="species">new species</taxonomicNameLabel>
. Male. 38, 39. Female. 40, 41.
<emphasis id="06B8EA8D6A79FFAD7742F801FBF83080" bold="true" box="[1090,1122,1965,1986]" pageId="20" pageNumber="471">39.</emphasis>
Palp, ventral view.
<emphasis id="06B8EA8D6A79FFAD763EF801FAC43080" bold="true" box="[1342,1374,1965,1986]" pageId="20" pageNumber="471">39.</emphasis>
Palp, retrolateral view.
<emphasis id="06B8EA8D6A79FFAD7248F867FEF230A2" bold="true" box="[328,360,1995,2016]" pageId="20" pageNumber="471">40.</emphasis>
Epigynum, ventral view.
<emphasis id="06B8EA8D6A79FFAD716DF867FD1730A2" bold="true" box="[621,653,1995,2016]" pageId="20" pageNumber="471">41.</emphasis>
Internal genitalia, dorsal view.
</paragraph>
</caption>
<paragraph id="3473369F6A78FFAC73C7FF3BFA0636EE" blockId="21.[151,1437,151,536]" pageId="21" pageNumber="472">
<emphasis id="06B8EA8D6A78FFAC73C7FF3BFE3037F2" bold="true" box="[199,426,151,176]" pageId="21" pageNumber="472">
<emphasis id="06B8EA8D6A78FFAC73C7FF3BFE3937F2" bold="true" box="[199,419,151,176]" italics="true" pageId="21" pageNumber="472">
Female (
<typeStatus id="EB77883D6A78FFAC7236FF34FE0037F2" box="[310,410,152,176]" pageId="21" pageNumber="472" type="paratype">paratype</typeStatus>
)
</emphasis>
.
</emphasis>
Total length: 9.3; carapace length: 4.7; carapace width: 3.3; abdomen length: 4.6. COLORATION: Carapace as in
<taxonomicName id="F3CC4D1C6A78FFAC7114FF11FD203796" box="[532,698,189,212]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="cappuccino">
<emphasis id="06B8EA8D6A78FFAC7114FF11FD203796" box="[532,698,189,212]" italics="true" pageId="21" pageNumber="472">C. cappuccino</emphasis>
</taxonomicName>
. Legs with numerous black marks. Abdomen as in
<emphasis id="06B8EA8D6A78FFAC7622FF11FA063796" box="[1314,1436,189,212]" italics="true" pageId="21" pageNumber="472">
<taxonomicName id="F3CC4D1C6A78FFAC7622FF11FA023796" box="[1314,1432,189,212]" class="Arachnida" family="Ctenidae" genus="Chococtenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="otonga">C. otonga</taxonomicName>
.
</emphasis>
LEGS: Total length: I: 13.1; II: 12.5; III: 11.9; IV: 15.1. LEGS SPINATION: Femur I p1, r1-1; tibia I p0, r1, d0; metatarsus I p0, r0. Femur II p1-1-1, r1-1; tibia II p0, r1, d0; metatarsus II p0, r0. Femur III pl-1-1-1, r1; tibia III p1-1, r1-1, d1-1,
<date id="4072105F6A78FFAC7256FE84FE163602" box="[342,396,296,320]" pageId="21" pageNumber="472">v2-2</date>
-2; metatarsus III p1-1-1, r1-1-1,
<date id="4072105F6A78FFAC71FEFE84FCAD3602" box="[766,823,296,320]" pageId="21" pageNumber="472">v2-2</date>
-1. Femur IV pl-1-1, r1-1; tibia IV p1-1, r1-1, d1-1,
<date id="4072105F6A78FFAC7676FE84FF3D3626" pageId="21" pageNumber="472">v2- 2</date>
-2; metatarsus IV p1-1-1, r1-1-1,
<date id="4072105F6A78FFAC7123FEE0FDC63626" box="[547,604,332,356]" pageId="21" pageNumber="472">v2-2</date>
-2-1. GENITALIA: Epigynum with elliptical median sector, apically with medium sized unsclerotized sector, flat; lateral processes short, positioned medially (
<figureCitation id="ACF72A1A6A78FFAC776CFEC3FB5E36CA" box="[1132,1220,367,392]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="21" pageNumber="472">Fig. 40</figureCitation>
). Internal genitalia with elongated and curved copulatory ducts; spermathecae oval, with apical pore; fertilization ducts short (
<figureCitation id="ACF72A1A6A78FFAC763CFE38FA1536EE" box="[1340,1423,404,429]" captionStart="FIGURES 38 41" captionStartId="20.[151,264,1964,1987]" captionTargetBox="[227,1334,198,1894]" captionTargetId="figure@20.[223,1335,193,1907]" captionTargetPageId="20" captionText="FIGURES 38 41. Chococtenus kashakara new species. Male. 38, 39. Female. 40, 41. 39. Palp, ventral view. 39. Palp, retrolateral view. 40. Epigynum, ventral view. 41. Internal genitalia, dorsal view." httpUri="https://zenodo.org/record/254180/files/figure.png" pageId="21" pageNumber="472">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="3473369F6A78FFAC73C7FE1BFB3736B7" blockId="21.[151,1437,151,536]" pageId="21" pageNumber="472">
<emphasis id="06B8EA8D6A78FFAC73C7FE1BFE1E3692" bold="true" box="[199,388,439,464]" pageId="21" pageNumber="472">Natural history.</emphasis>
Specimens were collected in pitfalls placed at
<quantity id="F3349B7A6A78FFAC708EFE1BFC7F368D" box="[910,997,439,464]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="21" pageNumber="472" unit="m" value="2225.0">2225 m</quantity>
, at the top of the cloud forest, and hand collected at an elevation of
<quantity id="F3349B7A6A78FFAC72CCFE70FDB936B6" box="[460,547,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="472" unit="m" value="2000.0">2000 m</quantity>
, therefore they can be identified as cloud forest specialist.
</paragraph>
</subSubSection>
<subSubSection id="7CD665146A78FFAC73C7FE53FCC6355A" box="[199,860,511,536]" pageId="21" pageNumber="472" type="distribution">
<paragraph id="3473369F6A78FFAC73C7FE53FCC6355A" blockId="21.[151,1437,151,536]" box="[199,860,511,536]" pageId="21" pageNumber="472">
<emphasis id="06B8EA8D6A78FFAC73C7FE53FEFA355A" bold="true" box="[199,352,511,536]" pageId="21" pageNumber="472">Distribution.</emphasis>
Only known from the
<typeStatus id="EB77883D6A78FFAC7160FDACFD0A355A" box="[608,656,512,536]" pageId="21" pageNumber="472">type</typeStatus>
locality (
<figureCitation id="ACF72A1A6A78FFAC71FAFE53FCD5355A" box="[762,847,511,536]" captionStart="FIGURE 57" captionStartId="30.[151,250,1231,1253]" captionTargetBox="[159,1427,201,1197]" captionTargetId="figure@30.[151,1436,193,1210]" captionTargetPageId="30" captionText="FIGURE 57. Map of Ecuador showing distribution records of the thirteen new species of the genus Chococtenus. The main collecting site, Otonga Biological Reserve (white box with x inside), holds 10 species (not found: C. suffuscus, C. otongachi and C. piemontana). Other known localities for: C. otonga (white lozenge), C. cappuccino (white x), C. lasdamas (white circle), C. suffuscus (white plus sign), C. otongachi (white rectangle) and C. piemontana (upside down white triangle)." httpUri="https://zenodo.org/record/254189/files/figure.png" pageId="21" pageNumber="472">Fig. 57</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>