167 lines
21 KiB
XML
167 lines
21 KiB
XML
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<mods:title id="90C3E8659FBF2EFE9EF0473F16712118">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<mods:namePart id="A3F827E983242A25F649009157F490A6">Ho, - Z.</mods:namePart>
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<mods:title id="602DCF5B09236032DEF3FDEB76D823C3">Zootaxa</mods:title>
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<mods:date id="F71147DFDAC17587B2F6F77E57B5A416">2019</mods:date>
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<treatment id="6C3DA91C514D1820FF0EFCF6EE3F1B5C" ID-DOI="http://doi.org/10.5281/zenodo.4688919" ID-GBIF-Taxon="159757410" ID-Zenodo-Dep="4688919" LSID="urn:lsid:plazi:treatment:6C3DA91C514D1820FF0EFCF6EE3F1B5C" httpUri="http://treatment.plazi.org/id/6C3DA91C514D1820FF0EFCF6EE3F1B5C" lastPageNumber="119" pageId="118" pageNumber="119">
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<subSubSection id="AC8E4B81514D1820FF0EFCF6ED201FC4" box="[151,536,873,900]" pageId="118" pageNumber="119" type="nomenclature">
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<paragraph id="E42B180A514D1820FF0EFCF6ED201FC4" blockId="118.[151,536,873,935]" box="[151,536,873,900]" pageId="118" pageNumber="119">
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<heading id="BF63AF66514D1820FF0EFCF6ED201FC4" bold="true" box="[151,536,873,900]" fontSize="11" level="1" pageId="118" pageNumber="119" reason="1">
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<taxonomicName id="23946389514D1820FF0EFCF6ED201FC4" ID-CoL="75SS" authority="(Ballantyne 1968)" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[151,536,873,900]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FF0EFCF6ED201FC4" bold="true" box="[151,536,873,900]" pageId="118" pageNumber="119">
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<emphasis id="D6E0C418514D1820FF0EFCF6EE0B1FC3" bold="true" box="[151,307,873,900]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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(
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<bibRefCitation id="800565FB514D1820FEDAFCF6ED371FC4" author="Ballantyne" box="[323,527,873,899]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="103 - 139" refId="ref113808" refString="Ballantyne, L. A. (1968) Revisional Studies of Australian and Indomalayan Luciolini (Coleoptera, Lampyridae, Luciolinae). University of Queensland Papers, Department of Entomology, University of Queensland Press, II (6), 103 - 139." type="journal article" year="1968">Ballantyne 1968</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B81514D1820FF0EFC12EE2E1FE0" box="[151,278,909,935]" pageId="118" pageNumber="119" type="description">
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<paragraph id="E42B180A514D1820FF0EFC12EE2E1FE0" blockId="118.[151,536,873,935]" box="[151,278,909,935]" pageId="118" pageNumber="119">
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<figureCitation id="7CAF048F514D1820FF0EFC12EE2E1FE0" box="[151,278,909,935]" captionStart="FIGURES 30−37" captionStartId="15.[151,264,1560,1585]" captionTargetBox="[173,1434,300,1530]" captionTargetId="figure@15.[151,1435,300,1540]" captionTargetPageId="15" captionText="FIGURES 30−37. Key to genera characters 5. 30, 34, 35 Pygatypella undulata; 31 Pygat. wisselmerenia; 32, 36 Pygat. obsoleta; 33 Pygat. peculiaris; 37 Pygat. kiunga paratype male. 30−34, 37 dorsal (34 diagrammatic representation of colour pattern); 35 aedeagal sheath, ventral; 36 aedeagus ventral, line outline." figureDoi="http://doi.org/10.5281/zenodo.3508112" httpUri="https://zenodo.org/record/3508112/files/figure.png" pageId="118" pageNumber="119">Figs 30−37</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B81514D1820FF0EFC4EEA21184F" pageId="118" pageNumber="119" type="reference_group">
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<paragraph id="E42B180A514D1820FF0EFC4EED511FAE" blockId="118.[151,1305,976,1063]" box="[151,617,976,1001]" pageId="118" pageNumber="119">
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<treatmentCitationGroup id="C4843F24514D1820FF0EFC4EED511FAE" box="[151,617,976,1001]" pageId="118" pageNumber="119">
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<taxonomicName id="23946389514D1820FF0EFC4EED5D1FAE" ID-CoL="8NYG9" authority="Ballantyne 1968: 109" authorityName="Ballantyne" authorityPageNumber="109" authorityYear="1968" box="[151,613,976,1001]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="subGenus" subGenus="Pygatyphella">
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<emphasis id="D6E0C418514D1820FF0EFC4EEE411FAE" box="[151,377,977,1001]" italics="true" pageId="118" pageNumber="119">Luciola (Pygatyphella</emphasis>
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)
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<treatmentCitation id="65353E1B514D1820FE11FC4FED5D1FAE" author="Ballantyne, L. A." box="[392,613,976,1001]" page="109" pageId="118" pageNumber="119" year="1968">
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<bibRefCitation id="800565FB514D1820FE11FC4FED5D1FAE" author="Ballantyne" box="[392,613,976,1001]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="103 - 139" refId="ref113808" refString="Ballantyne, L. A. (1968) Revisional Studies of Australian and Indomalayan Luciolini (Coleoptera, Lampyridae, Luciolinae). University of Queensland Papers, Department of Entomology, University of Queensland Press, II (6), 103 - 139." type="journal article" year="1968">Ballantyne 1968: 109</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="E42B180A514D1820FF0EFC6FEA21184F" blockId="118.[151,1305,976,1063]" box="[151,1305,1007,1032]" pageId="118" pageNumber="119">
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<treatmentCitationGroup id="C4843F24514D1820FF0EFC6FEA21184F" box="[151,1305,1007,1032]" pageId="118" pageNumber="119">
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<taxonomicName id="23946389514D1820FF0EFC6FEE9D184F" ID-CoL="75SS" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[151,421,1007,1032]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FF0EFC6FEE25184F" box="[151,285,1008,1032]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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(Ballantyne)
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</taxonomicName>
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.
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<treatmentCitation id="65353E1B514D1820FE37FC70EDC21840" author="Ballantyne, L. A. & Lambkin, C. L." box="[430,762,1007,1032]" httpUri="http://treatment.plazi.org/id/0394D665BE60FFF3FF3C531326D6EA7A" page="77" pageId="118" pageNumber="119" year="2009">
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<bibRefCitation id="800565FB514D1820FE37FC70EDC21840" author="Ballantyne & Lambkin" box="[430,762,1007,1032]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 77</bibRefCitation>
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</treatmentCitation>
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, figs 23–26, 61, 69–72, 255–328, 377–396, 441–487.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B81514D1820FF0EFB90EDBA1860" box="[151,642,1038,1063]" pageId="118" pageNumber="119" type="type_taxon">
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<paragraph id="E42B180A514D1820FF0EFB90EDBA1860" blockId="118.[151,1305,976,1063]" box="[151,642,1038,1063]" pageId="118" pageNumber="119">
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<emphasis id="D6E0C418514D1820FF0EFB90EE261860" bold="true" box="[151,286,1038,1063]" pageId="118" pageNumber="119">
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<typeStatus id="3B2FA6A8514D1820FF0EFB90EFF41860" box="[151,204,1039,1063]" pageId="118" pageNumber="119">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="23946389514D1820FEB3FB91ED461860" authority="Olivier 1911 b" authorityName="Olivier" authorityYear="1911" box="[298,638,1038,1063]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="D6E0C418514D1820FEB3FB91EED01861" box="[298,488,1038,1062]" italics="true" pageId="118" pageNumber="119">Atyphella obsoleta</emphasis>
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<bibRefCitation id="800565FB514D1820FE77FB91ED461860" author="Olivier" box="[494,638,1038,1063]" firstAuthor="Olivier" pageId="118" pageNumber="119" pagination="171 - 175" refId="ref120903" refString="Olivier, E. (1911 b) Le genre Atyphella Oll. et descriptions des especes nouvelle (Col. Lampyridae). Bulletin de la Societe Entomologique de France, 8, 171 - 175. https: // doi. org / 10.5962 / bhl. part. 2971" type="journal article" year="1911" yearSuffix="b">Olivier 1911b</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B81514D1820FF0EFBCBEE3F1B5C" pageId="118" pageNumber="119" type="diagnosis">
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<paragraph id="E42B180A514D1820FF0EFBCBEE3F1B5C" blockId="118.[151,1437,1108,1819]" pageId="118" pageNumber="119">
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<emphasis id="D6E0C418514D1820FF0EFBCBEE2A1828" bold="true" box="[151,274,1108,1135]" pageId="118" pageNumber="119">Diagnosis.</emphasis>
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Both
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<taxonomicName id="23946389514D1820FEC5FBCAEEC81828" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[348,496,1109,1135]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FEC5FBCAEEC81828" box="[348,496,1109,1135]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="23946389514D1820FDB5FBCAEDB11828" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[556,649,1109,1135]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FDB5FBCAEDB11828" box="[556,649,1109,1135]" italics="true" pageId="118" pageNumber="119">Pacifica</emphasis>
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</taxonomicName>
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occur in New
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<collectingCountry id="9C83589A514D1820FCA3FBCAECB31828" box="[826,907,1109,1135]" name="Guinea-Bissau" pageId="118" pageNumber="119">Guinea</collectingCountry>
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and Pacific islands, and both have LO in
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<date id="902A3ECA514D1820FAE3FBCAEAA41828" box="[1402,1436,1109,1135]" pageId="118" pageNumber="119">V7</date>
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retracted to the anterior half or less; the posterior clear portion of
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<date id="902A3ECA514D1820FC1FFBE6EC9018D4" box="[902,936,1145,1171]" pageId="118" pageNumber="119">V7</date>
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is frequently arched and underlying muscle impressions can be clearly seen. The two genera are most obviously distinguished by their dorsal colouration,
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<taxonomicName id="23946389514D1820FF0EFB5EEE13189C" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[151,299,1217,1243]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FF0EFB5EEE13189C" box="[151,299,1217,1243]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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</taxonomicName>
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having a pattern resembling bird droppings, where most
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<taxonomicName id="23946389514D1820FC28FB5EEB36189C" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[945,1038,1217,1243]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FC28FB5EEB36189C" box="[945,1038,1217,1243]" italics="true" pageId="118" pageNumber="119">Pacifica</emphasis>
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</taxonomicName>
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species have orange pronotum and dark brown elytra which may be pale margined.
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<bibRefCitation id="800565FB514D1820FD56FB7AEB1418B8" author="Ballantyne & Lambkin" box="[719,1068,1253,1279]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin (2009)</bibRefCitation>
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distinguished two forms within
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<taxonomicName id="23946389514D1820FF0EFA96EE131964" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[151,299,1289,1315]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FF0EFA96EE131964" box="[151,299,1289,1315]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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</taxonomicName>
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they named
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<taxonomicName id="23946389514D1820FE27FA96ED6A1964" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[446,594,1289,1315]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FE27FA96ED6A1964" box="[446,594,1289,1315]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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</taxonomicName>
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A and B. Subsequently
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<taxonomicName id="23946389514D1820FCFFFA96ECC21964" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[870,1018,1289,1315]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FCFFFA96ECC21964" box="[870,1018,1289,1315]" italics="true" pageId="118" pageNumber="119">Pygatyphella</emphasis>
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</taxonomicName>
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B was described from 5 species as a distinct genus
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<taxonomicName id="23946389514D1820FEA3FAB2EEAF1900" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2013" box="[314,407,1325,1351]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FEA3FAB2EEAF1900" box="[314,407,1325,1351]" italics="true" pageId="118" pageNumber="119">Pacifica</emphasis>
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</taxonomicName>
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. The following diagnosis of
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<taxonomicName id="23946389514D1820FD47FAB2EA2D1900" authority="sensu Ballantyne & Lambkin (2013)" authorityName="Ballantyne & Lambkin" authorityYear="2013" box="[734,1301,1325,1351]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C418514D1820FD47FAB2EC801900" box="[734,952,1325,1351]" italics="true" pageId="118" pageNumber="119">Pygatyphella sensu</emphasis>
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<bibRefCitation id="800565FB514D1820FC26FAB2EA2D1900" author="Ballantyne & Lambkin" box="[959,1301,1325,1351]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. & Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne & Lambkin (2013)</bibRefCitation>
|
||
</taxonomicName>
|
||
is modified from
|
||
<bibRefCitation id="800565FB514D1820FF4EFACEED0D192C" author="Ballantyne & Lambkin" box="[215,565,1361,1387]" firstAuthor="Ballantyne" pageId="118" pageNumber="119" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin (2009)</bibRefCitation>
|
||
: Dorsal colouration usually cryptic, very pale brown with dark markings on pronotum, MS, MN and elytral base and apex; pronotum often with angulate convergence along lateral margins, and small depression often present just anterior to posterolateral corner; lacking irregularities at the corners; posterolateral corners may project beyond median posterior margin; at least two interstitial lines usually well–defined; elytral margins convex or parallel-sided; posterior area of
|
||
<date id="902A3ECA514D1820FCBDFA7EEC7E19BC" box="[804,838,1505,1531]" pageId="118" pageNumber="119">V7</date>
|
||
may reach into LO (which is medially emarginated); apex of MPP rounded, pointed, or squarely or obliquely truncate; MPP lacking median dorsal ridge (except in
|
||
<emphasis id="D6E0C418514D1820FACDF999EAAD1A58" box="[1364,1429,1542,1567]" italics="true" pageId="118" pageNumber="119">Pygat</emphasis>
|
||
.
|
||
<emphasis id="D6E0C418514D1820FF0EF9B6EFD61A04" box="[151,238,1577,1603]" italics="true" pageId="118" pageNumber="119">karimui</emphasis>
|
||
), lacking curved, slightly off-centre posterior ridge on ventral surface of T8; dorsal surface of posterior area of MPP faced with cuticle that is not attached to the ventral surface of
|
||
<date id="902A3ECA514D1820FC2CF9D2ECEF1A20" box="[949,983,1613,1639]" pageId="118" pageNumber="119">V7</date>
|
||
and ends just behind the area of muscle attachment posterior to the LO; the anterior margins of this cuticle are reflexed in
|
||
<emphasis id="D6E0C418514D1820FBDBF9EDEA3B1ACC" box="[1090,1283,1649,1675]" italics="true" pageId="118" pageNumber="119">Pygat. peculiaris</emphasis>
|
||
; median area of this facing cuticle lacking longitudinal split; T8 anterolateral prolongations lacking ventral projections at their base; aedeagal sheath not subparallel-sided in basal third; aedeagus L/W usually>3; LL/ML not wide; ML sometimes asymmetrical in a horizontal plane only, never with lateral teeth; anterior margin of LL often asymmetrically produced.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |