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<document id="5E1D2D238ED0C90B37788DD10B10EC24" ID-DOI="10.12782/specdiv.23.209" ID-ISSN="2189-7301" ID-Zenodo-Dep="5738171" ID-ZooBank="CCA2E8E2-81F1-4559-995E-412EC65752F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638228499649" checkinUser="felipe" docAuthor="Fujimoto, Shinta" docDate="2018" docId="637687BAFFE08337468BB1710263F9E4" docLanguage="en" docName="SpeciesDiversity.23.2.209-213.pdf" docOrigin="Species Diversity 23" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Tanarctus ittanmomen Fujimoto 2018, sp. nov." docType="treatment" docUuid="E6F48ECA-1DE3-42CE-8B8D-C707DE85DE27" docUuidSource="ZooBank" docVersion="3" lastPageNumber="212" masterDocId="9F4FFFC2FFE08334454CB448070AFFB1" masterDocTitle="A New Species of Tanarctus (Heterotardigrada: Arthrotardigrada: Tanarctidae) from Oku-Matsushima, Japan" masterLastPageNumber="213" masterPageNumber="209" pageNumber="209" updateTime="1699433806081" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="13819294FB8C0C9B64369CDE86449394" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C8EC2CBF58750302CA4D520D55678F2C">
<mods:title id="9F5125F8B44D16C3205413146365DFAE">A New Species of Tanarctus (Heterotardigrada: Arthrotardigrada: Tanarctidae) from Oku-Matsushima, Japan</mods:title>
</mods:titleInfo>
<mods:name id="4940A86ECF8399E74BF0DCE2277BFCDD" type="personal">
<mods:role id="5F3ECB840A7CEA3877832DC23C1DD9A9">
<mods:roleTerm id="9913DD7B70519FEBCE89FD9EC12C8E9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="322BA43FA8EBCE3C44E1F2CEEF8A7EA8">Fujimoto, Shinta</mods:namePart>
</mods:name>
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<mods:title id="ED55E7714BBC78712D83D3F7BFB438A6">Species Diversity</mods:title>
</mods:titleInfo>
<mods:part id="4FD738315EE0F78638C9E7616F7DF82B">
<mods:date id="7550C814B989A2172C74DA9DFE669B7D">2018</mods:date>
<mods:detail id="FFF94C5F61CC79D33739E86452F88B83" type="pubDate">
<mods:number id="44EB1D5F3784DE429832024B7CB6A21C">2018-11-25</mods:number>
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<mods:number id="5E70F5DC9577444489226E7532869E76">23</mods:number>
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<mods:classification id="E1CB0D00E6065C060BC9328D1A5F4A5E">journal article</mods:classification>
<mods:identifier id="4948E71F3E73EC4E702D83C7396AB2E1" type="DOI">10.12782/specdiv.23.209</mods:identifier>
<mods:identifier id="B0FD3D24E933E09D685924F95E8EC3C1" type="ISSN">2189-7301</mods:identifier>
<mods:identifier id="8214A4720775EE7B473E1A2D2C7BBF50" type="Zenodo-Dep">5738171</mods:identifier>
<mods:identifier id="B45CA6DD57340B4C77F74D4036B8EFA2" type="ZooBank">CCA2E8E2-81F1-4559-995E-412EC65752F1</mods:identifier>
</mods:mods>
<treatment id="637687BAFFE08337468BB1710263F9E4" LSID="urn:lsid:zoobank.org:act:E6F48ECA-1DE3-42CE-8B8D-C707DE85DE27" httpUri="http://treatment.plazi.org/id/637687BAFFE08337468BB1710263F9E4" lastPageId="3" lastPageNumber="212" pageId="0" pageNumber="209">
<subSubSection id="A3C56527FFE08334468BB17103C8FAC3" pageId="0" pageNumber="209" type="nomenclature">
<paragraph id="EB6036ACFFE08334468BB17103C8FAC3" blockId="0.[967,1286,1337,1394]" pageId="0" pageNumber="209">
<heading id="B02881C0FFE08334468BB17103C8FAC3" centered="true" fontSize="10" level="2" pageId="0" pageNumber="209" reason="2">
<taxonomicName id="2CDF4D2FFFE08334468BB17103BEFAE0" authority="Fujimoto, 2018" authorityName="Fujimoto" authorityYear="2018" box="[967,1204,1337,1361]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="0" pageNumber="209" phylum="Tardigrada" rank="species" species="ittanmomen" status="sp. nov.">
<emphasis id="D9ABEABEFFE08334468BB17103BEFAE0" bold="true" box="[967,1204,1337,1361]" italics="true" pageId="0" pageNumber="209">Tanarctus ittanmomen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C29857C5FFE0833441F6B172020CFAE0" box="[1210,1286,1338,1361]" pageId="0" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="73E42A29FFE08334415DB1120344FAC3" box="[1041,1102,1370,1394]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="0" pageNumber="209">Figs 1</figureCitation>
,
<figureCitation id="73E42A29FFE083344116B1120362FAC3" box="[1114,1128,1370,1394]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="0" pageNumber="209">2</figureCitation>
;
<tableCitation id="A65D0317FFE08334413DB11203B2FAC3" box="[1137,1208,1370,1394]" captionStart="Table 1" captionStartId="3.[156,204,190,212]" captionText="Table 1. Measurements (µm) of Tanarctus ittanmomen sp.nov." pageId="0" pageNumber="209">Table 1</tableCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A3C56527FFE083344602B1D203D8F906" pageId="0" pageNumber="209" type="diagnosis">
<paragraph id="EB6036ACFFE083344602B1D203D8F906" blockId="0.[821,1432,1434,2077]" pageId="0" pageNumber="209">
<emphasis id="D9ABEABEFFE083344602B1D204B4FA03" bold="true" box="[846,958,1434,1458]" pageId="0" pageNumber="209">Diagnosis.</emphasis>
<taxonomicName id="2CDF4D2FFFE08334469BB1D30333FA03" authorityName="Renaud-Debyser" authorityYear="1959" box="[983,1081,1435,1458]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="0" pageNumber="209" phylum="Tardigrada" rank="genus">
<emphasis id="D9ABEABEFFE08334469BB1D30333FA03" box="[983,1081,1435,1458]" italics="true" pageId="0" pageNumber="209">Tanarctus</emphasis>
</taxonomicName>
with the scapi of the internal cirri embedded in the cuticle; smooth primary clavae; flat secondary clavae each with three internally directed swellings; leg IV appendages each consisting of a main branch exceeding double the body length with variable numbers of flexible, mostly long secondary branches; all branches with unmodified tips; seminal receptacles open overlapping gonopore and each terminates in a laterally situated round vesicle with a duct of approximately even width.
</paragraph>
</subSubSection>
<subSubSection id="A3C56527FFE083344602B2F7023EF84D" pageId="0" pageNumber="209" type="materials_examined">
<paragraph id="EB6036ACFFE083344602B2F7030DF80A" blockId="0.[821,1432,1434,2077]" pageId="0" pageNumber="209">
<emphasis id="D9ABEABEFFE083344602B2F704E5F966" bold="true" box="[846,1007,1727,1751]" pageId="0" pageNumber="209">Type material.</emphasis>
<materialsCitation id="5BB73CF1FFE083344144B2F704B6F888" collectingDate="2017-11-20" collectionCode="KUZ" country="Japan" county="Oku-Matsushima" location="Ohama Beach" municipality="Miyatojima" pageId="0" pageNumber="209" specimenCode="Z1954" specimenCount="1" typeStatus="holotype">
<typeStatus id="3464880EFFE083344144B2F70363F966" box="[1032,1129,1727,1751]" pageId="0" pageNumber="209" type="holotype">Holotype</typeStatus>
:
<collectionCode id="8DCEAE69FFE083344137B28803A4F966" box="[1147,1198,1728,1751]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE0833441F1B2F703F5F966" box="[1213,1279,1727,1751]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">Z1954</specimenCode>
, adult female mounted in glycerol collected at
<location id="EE006077FFE0833441D3B2A8023DF949" LSID="urn:lsid:plazi:treatment:637687BAFFE08337468BB1710263F9E4:EE006077FFE0833441D3B2A8023DF949" box="[1183,1335,1760,1784]" country="Japan" county="Oku-Matsushima" municipality="Miyatojima" name="Ohama Beach" pageId="0" pageNumber="209">Ohama Beach</location>
,
<collectingMunicipality id="0B04ACD6FFE08334400BB2A8046EF8A9" pageId="0" pageNumber="209">Miyatojima</collectingMunicipality>
,
<collectingCounty id="02014E20FFE083344638B3480322F8A9" box="[884,1064,1792,1816]" pageId="0" pageNumber="209">Oku-Matsushima</collectingCounty>
,
<collectingCountry id="93C8763CFFE083344174B3490379F8A9" box="[1080,1139,1793,1816]" name="Japan" pageId="0" pageNumber="209">Japan</collectingCountry>
, by the author on
<date id="9F61106CFFE083344006B34904B6F888" pageId="0" pageNumber="209" value="2017-11-20">
<collectingDate id="8F25E984FFE083344006B34904B6F888" pageId="0" pageNumber="209" value="2017-11-20">20 November 2017</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="5BB73CF1FFE08334468BB369031BF82B" collectionCode="KUZ" country="Japan" pageId="0" pageNumber="209" specimenCode="Z1955, Z1957, Z1958, Z1959" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
<typeStatus id="3464880EFFE08334468BB3690321F888" box="[967,1067,1825,1849]" pageId="0" pageNumber="209" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="8DCEAE69FFE08334417BB3690360F889" box="[1079,1130,1825,1848]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE08334413EB36903BBF888" box="[1138,1201,1825,1849]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">Z1955</specimenCode>
and Z1956,
<specimenCount id="FDD9FD25FFE08334407AB3690292F888" box="[1334,1432,1825,1849]" pageId="0" pageNumber="209" type="adult">two adult</specimenCount>
females;
<collectionCode id="8DCEAE69FFE0833446DDB30A04CEF8E8" box="[913,964,1858,1881]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE083344682B309031AF8E8" box="[974,1040,1857,1881]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">Z1957</specimenCode>
, juvenile moulting into adult female;
<collectionCode id="8DCEAE69FFE083344679B32B0462F8CB" box="[821,872,1891,1914]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE083344621B32A04A5F8CB" box="[877,943,1890,1914]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">Z1958</specimenCode>
, adult male; and
<collectionCode id="8DCEAE69FFE083344110B32B0385F8CB" box="[1116,1167,1891,1914]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE0833441D9B32A03DCF8CB" box="[1173,1238,1890,1914]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="0" pageNumber="209">Z1959</specimenCode>
, four-digit juvenile of undetermined sex
</materialsCitation>
. All collected with the
<typeStatus id="3464880EFFE083344042B3CA026DF82B" box="[1294,1383,1922,1946]" pageId="0" pageNumber="209" type="holotype">holotype</typeStatus>
and mounted in glycerol.
</paragraph>
<paragraph id="EB6036ACFFE083344602B38C023EF84D" blockId="0.[821,1432,1434,2077]" pageId="0" pageNumber="209">
<emphasis id="D9ABEABEFFE083344602B38C04E6F86A" bold="true" box="[846,1004,1987,2012]" pageId="0" pageNumber="209">
<typeStatus id="3464880EFFE083344602B38C0488F86D" box="[846,898,1988,2012]" pageId="0" pageNumber="209">Type</typeStatus>
locality.
</emphasis>
<materialsCitation id="5BB73CF1FFE08334414AB38C023BF84D" country="Japan" county="Matsushima" latitude="38.32789" location="Ohama Beach" longLatPrecision="1" longitude="141.16277" municipality="Miyato-jima" pageId="0" pageNumber="209" specimenCount="1" typeStatus="holotype">
<location id="EE006077FFE08334414AB38C03A3F86D" LSID="urn:lsid:plazi:treatment:637687BAFFE08337468BB1710263F9E4:EE006077FFE08334414AB38C03A3F86D" box="[1030,1193,1988,2012]" country="Japan" county="Matsushima" latitude="38.32789" longLatPrecision="1" longitude="141.16277" municipality="Miyato-jima" name="Ohama Beach" pageId="0" pageNumber="209">Ohama Beach</location>
,
<collectingMunicipality id="0B04ACD6FFE083344189B38C024FF86D" box="[1221,1349,1988,2012]" pageId="0" pageNumber="209">Miyato-jima</collectingMunicipality>
, Oku-
<collectingCounty id="02014E20FFE083344679B3AC04B9F84D" box="[821,947,2020,2044]" pageId="0" pageNumber="209">Matsushima</collectingCounty>
,
<collectingCountry id="93C8763CFFE0833446F0B3AD04FEF84D" box="[956,1012,2021,2044]" name="Japan" pageId="0" pageNumber="209">Japan</collectingCountry>
(
<geoCoordinate id="8EEB506BFFE08334414EB3AC0386F84D" box="[1026,1164,2019,2044]" degrees="38" direction="north" minutes="19" orientation="latitude" pageId="0" pageNumber="209" precision="1" seconds="40.4" value="38.32789">38°19′40.4″N</geoCoordinate>
,
<geoCoordinate id="8EEB506BFFE0833441DAB3AC0223F84D" box="[1174,1321,2019,2044]" degrees="141" direction="east" minutes="09" orientation="longitude" pageId="0" pageNumber="209" precision="1" seconds="46.0" value="141.16277">141°09′46.0″E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3C56527FFE083364602BC4D05AEFEAE" lastPageId="2" lastPageNumber="211" pageId="0" pageNumber="209" type="etymology">
<paragraph id="EB6036ACFFE083364602BC4D05AEFEAE" blockId="0.[821,1432,1434,2077]" lastBlockId="2.[156,768,198,2077]" lastPageId="2" lastPageNumber="211" pageId="0" pageNumber="209">
<emphasis id="D9ABEABEFFE083344602BC4D04CFF7AC" bold="true" box="[846,965,2053,2077]" pageId="0" pageNumber="209">Etymology.</emphasis>
The specific epithet refers to ‘Ittan-momen’ (
<emphasis id="D9ABEABEFFE2833645E9B48F07CCFF6F" box="[165,198,199,222]" italics="true" pageId="2" pageNumber="211">e.g.</emphasis>
,
<bibRefCitation id="8F4E4B5DFFE28336459EB48E065CFF6F" author="Mizuki, S." box="[210,342,198,222]" pageId="2" pageNumber="211" refId="ref3005" refString="Mizuki, S. 1994. Zusetsu Nihon Yokai Taizen [Illustrated Encyclopedia of Japanese Yokai]. Kodansha, Tokyo, 500 pp. [In Japanese]" type="book" year="1994">Mizuki 1994</bibRefCitation>
). The appearance of the new species gives the impression that it may drift and twine like this yokai (a term for mysterious beings from Japanese folklore).
</paragraph>
</subSubSection>
<caption id="BFA06624FFE1833545D0B3DB0338F84C" ID-DOI="http://doi.org/10.5281/zenodo.5738173" ID-Zenodo-Dep="5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="1" pageNumber="210" startId="1.[156,187,1939,1961]" targetBox="[201,1387,208,1898]" targetPageId="1">
<paragraph id="EB6036ACFFE1833545D0B3DB0338F84C" blockId="1.[156,1432,1939,2046]" pageId="1" pageNumber="210">
Fig. 1. Drawings of
<taxonomicName id="2CDF4D2FFFE18335442AB3DC053BF818" authority="Fujimoto, 2018" authorityName="Fujimoto" authorityYear="2018" box="[358,561,1940,1961]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="1" pageNumber="210" phylum="Tardigrada" rank="species" species="ittanmomen" status="sp. nov.">
<emphasis id="D9ABEABEFFE18335442AB3DC053BF818" box="[358,561,1940,1961]" italics="true" pageId="1" pageNumber="210">Tanarctus ittanmomen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C29857C5FFE183354774B3DC0577F818" box="[568,637,1940,1961]" pageId="1" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l
<subScript id="775B34E9FFE183354585B39C07DAF853" attach="both" box="[201,208,2004,2018]" fontSize="6" pageId="1" pageNumber="210">4</subScript>
a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so
<subScript id="775B34E9FFE183354061B39C0249F853" attach="left" box="[1325,1347,2004,2018]" fontSize="6" pageId="1" pageNumber="210">1–3</subScript>
, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw
<subScript id="775B34E9FFE183354648B3B80410F84F" attach="left" box="[772,794,2032,2046]" fontSize="6" pageId="1" pageNumber="210">1–3</subScript>
, swellings of secondary clavae.
</paragraph>
</caption>
<subSubSection id="A3C56527FFE2833745F9B560059AF8EB" lastPageId="3" lastPageNumber="212" pageId="2" pageNumber="211" type="description">
<paragraph id="EB6036ACFFE2833645F9B5600521FE10" blockId="2.[156,768,198,2077]" pageId="2" pageNumber="211">
<emphasis id="D9ABEABEFFE2833645F9B56006B3FEF1" bold="true" box="[181,441,296,320]" pageId="2" pageNumber="211">
Description of
<typeStatus id="3464880EFFE28336441AB56006BFFEF1" box="[342,437,296,320]" pageId="2" pageNumber="211" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female (
<figureCitation id="73E42A29FFE28336472FB56005BFFEF1" box="[611,693,296,320]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="2" pageNumber="211">Figs 1A</figureCitation>
,
<figureCitation id="73E42A29FFE28336478CB56005F1FE8E" box="[704,763,296,320]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="2" pageNumber="211">2A, B</figureCitation>
;
<tableCitation id="A65D0317FFE2833645D0B50007ECFED0" box="[156,230,328,353]" captionStart="Table 1" captionStartId="3.[156,204,190,212]" captionText="Table 1. Measurements (µm) of Tanarctus ittanmomen sp.nov." pageId="2" pageNumber="211">Table 1</tableCitation>
). Dorsoventrally flattened body 145 µm long, 71 µm wide between legs II and III. Dorsal and ventral cuticles smooth with visible epicuticular pillars.
</paragraph>
<paragraph id="EB6036ACFFE2833645F9B5E205F2FAE0" blockId="2.[156,768,198,2077]" pageId="2" pageNumber="211">
Cephalic region (
<figureCitation id="73E42A29FFE28336442AB5E206BDFE73" box="[358,439,426,450]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="2" pageNumber="211">Figs 1A</figureCitation>
,
<figureCitation id="73E42A29FFE28336448FB5E206E8FE73" box="[451,482,426,450]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="2" pageNumber="211">2A</figureCitation>
) with unpaired median cirrus, paired internal cirri, paired external cirri, paired lateral cirri, paired primary clavae, and paired secondary clavae. Median cirrus (13 µm) inserted dorsally, 17 µm from anterior margin of cephalic region, without clear subdivision, but constricted tip. Internal cirrus (37 µm) inserted dorsally on anterior margin of cephalic region, consisting of scapus (6 µm), long tubular portion (22 µm), and flagellum (10 µm). Cuticle present between two scapi, more than half the length of the scapi embedded. External cirrus (22 µm) inserted externally to internal cirri on ventral side of cephalic region, consisting of scapus (11 µm), tubular portion (7 µm), and flagellum (3 µm). Lateral cirrus (16 µm) and primary clava (195 µm) inserted on short, common cirrophore at lateral margin of cephalic region. Lateral cirrus positioned dorsoposterior to primary clava, without clear subdivision, basally swollen and distally constricted. Primary clava long, smooth, with van der Land’s body at base. Dense, rod-like particles present near tips of median and lateral cirri and near tips of tubular portions of internal and external cirri. Mouth cone protrudes anteroventrally. Buccal apparatus partially recognised; pharyngeal bulb 9 µm long, 14 µm wide. Narrow buccal tube, with stylets and stylet supports. No furcae or placoids recognised. Secondary clava present as area without epicuticular pillars surrounding mouth. Thorough observation of this structure obstructed by protruding mouth cone. Seven amoebocytes distributed posteriorly in cephalic region. Possible disturbed amoebocytes present anteriorly in cephalic region (not drawn in
<figureCitation id="73E42A29FFE2833647ECB17105E6FAE0" box="[672,748,1337,1361]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="2" pageNumber="211">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="EB6036ACFFE2833645F9B1110526FA20" blockId="2.[156,768,198,2077]" pageId="2" pageNumber="211">
Paired cirri E (30 µm) (
<figureCitation id="73E42A29FFE2833644D1B11106E3FAC3" box="[413,489,1369,1394]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="2" pageNumber="211">Fig. 1A</figureCitation>
), with proximal annulation arising dorsally between legs III and IV.
</paragraph>
<paragraph id="EB6036ACFFE2833745F9B1D2064CFC3B" blockId="2.[156,768,198,2077]" lastBlockId="3.[156,768,880,2077]" lastPageId="3" lastPageNumber="212" pageId="2" pageNumber="211">
Each leg consists of indistinct coxa, femur, lance-like tibia, conical tarsus, and four digits, each with claw. Leg I femur has tapering sensory organ (18 µm) consisting of proximal (12 µm) and distal (6 µm) portions.
<materialsCitation id="5BB73CF1FFE2833647DDB1B50695F9E4" collectionCode="II, III" collectorName="Femurs" pageId="2" pageNumber="211" specimenCount="1">
<collectorName id="462A537AFFE2833647DDB1B505D7F9A5" box="[657,733,1533,1556]" pageId="2" pageNumber="211">Femurs</collectorName>
of legs
<collectionCode id="8DCEAE69FFE283364580B25507D7F985" box="[204,221,1565,1588]" pageId="2" pageNumber="211">II</collectionCode>
and
<collectionCode id="8DCEAE69FFE28336445BB255063BF985" box="[279,305,1565,1588]" pageId="2" pageNumber="211">III</collectionCode>
have sensory organs as tapering spines (21 and 24 µm, respectively)
</materialsCitation>
.
<materialsCitation id="5BB73CF1FFE2833644E1B27605F6F927" collectionCode="IV" collectorName="Leg" pageId="2" pageNumber="211" specimenCount="1">
<collectorName id="462A537AFFE2833644E1B27606DBF9E7" box="[429,465,1598,1622]" pageId="2" pageNumber="211">Leg</collectorName>
<collectionCode id="8DCEAE69FFE283364491B27606FDF9E4" box="[477,503,1598,1621]" pageId="2" pageNumber="211">IV</collectionCode>
sensory organ modified into long, branching appendage arising from short process (5 µm) positioned dorsally at base of leg
<collectionCode id="8DCEAE69FFE283364704B2370568F927" box="[584,610,1663,1686]" pageId="2" pageNumber="211">IV</collectionCode>
(
<figureCitation id="73E42A29FFE28336473EB23605CFF927" box="[626,709,1662,1686]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="2" pageNumber="211">Figs 1A</figureCitation>
,
<figureCitation id="73E42A29FFE28336479FB23605FEF927" box="[723,756,1662,1686]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="2" pageNumber="211">2A</figureCitation>
)
</materialsCitation>
. Secondary branches (9 and
<quantity id="2C279B49FFE28336448FB2D706F0F906" box="[451,506,1695,1719]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="211" unit="in" value="12.0">12 in</quantity>
number, respectively) of variable lengths (44–194 and 20–92 µm, respectively) grow on the main branches (170 and 301 µm, respectively; shorter appendage deformed [see
<emphasis id="D9ABEABEFFE2833644F0B3480609F888" bold="true" pageId="2" pageNumber="211">
Additional information from
<typeStatus id="3464880EFFE2833645D0B3690609F888" box="[156,259,1825,1849]" pageId="2" pageNumber="211" type="paratype">paratypes</typeStatus>
</emphasis>
]). Secondary branches of longer appendage grow exteriorly on proximal half of main branch. Paired internal digits arise from common pretarsus; each terminates in crescent-shaped claws with strong calcar and minute, dorsal spur. Paired external digits each with basal cuticle fold, terminating in crescent-shaped claws with strong calcar.
<materialsCitation id="5BB73CF1FFE283374585B3AC0648FC3B" collectionCode="II, III, IV" collectorName="Internal" lastPageId="3" lastPageNumber="212" pageId="2" pageNumber="211" specimenCount="1">
<collectorName id="462A537AFFE283364585B3AC0610F84D" box="[201,282,2020,2044]" pageId="2" pageNumber="211">Internal</collectorName>
digits always longer than external digits (legs I: 12 µm&gt; 8 µm;
<collectionCode id="8DCEAE69FFE283364419BC4D0663F7AD" box="[341,361,2053,2076]" pageId="2" pageNumber="211">II</collectionCode>
: 14 µm&gt; 9 µm;
<collectionCode id="8DCEAE69FFE28336475ABC4D0538F7AD" box="[534,562,2053,2076]" pageId="2" pageNumber="211">III</collectionCode>
: 16 µm&gt; 9 µm;
<collectionCode id="8DCEAE69FFE2833647ACBC4D05F1F7AD" box="[736,763,2053,2076]" pageId="2" pageNumber="211">IV</collectionCode>
: 17 µm&gt; 10 µm)
</materialsCitation>
.
</paragraph>
<caption id="BFA06624FFE283364679B298024BF7AB" ID-DOI="http://doi.org/10.5281/zenodo.5738175" ID-Zenodo-Dep="5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="2" pageNumber="211" startId="2.[821,852,1744,1766]" targetBox="[861,1391,206,1705]" targetPageId="2">
<paragraph id="EB6036ACFFE283364679B298024BF7AB" blockId="2.[821,1432,1744,2074]" pageId="2" pageNumber="211">
Fig. 2. DIC micrographs of
<taxonomicName id="2CDF4D2FFFE283364119B299022FF957" authority="Fujimoto, 2018" authorityName="Fujimoto" authorityYear="2018" box="[1109,1317,1745,1766]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="211" phylum="Tardigrada" rank="species" species="ittanmomen" status="sp. nov.">
<emphasis id="D9ABEABEFFE283364119B299022FF957" box="[1109,1317,1745,1766]" italics="true" pageId="2" pageNumber="211">Tanarctus ittanmomen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C29857C5FFE283364063B2990273F957" box="[1327,1401,1745,1766]" pageId="2" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so
<subScript id="775B34E9FFE2833646B0B39D0309F852" attach="left" box="[1020,1027,2005,2019]" fontSize="6" pageId="2" pageNumber="211">2</subScript>
, leg II sensory organ;
<subScript id="775B34E9FFE283364181B38503ECF852" attach="right" box="[1229,1254,1997,2019]" fontSize="6" pageId="2" pageNumber="211">srd</subScript>
, seminal receptacle duct;
<subScript id="775B34E9FFE283364625B3A1048AF84E" attach="left" box="[873,896,2025,2047]" fontSize="6" pageId="2" pageNumber="211">sro</subScript>
, seminal receptacle opening; sw
<subScript id="775B34E9FFE2833641E4B3B903B4F84E" attach="left" box="[1192,1214,2033,2047]" fontSize="6" pageId="2" pageNumber="211">1–3</subScript>
, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle.
</paragraph>
</caption>
<caption id="BFA06624FFE3833745D0B4F605ECFF65" box="[156,742,190,212]" pageId="3" pageNumber="212" startId="3.[156,204,190,212]">
<paragraph id="EB6036ACFFE3833745D0B4F605ECFF65" blockId="3.[156,742,190,212]" box="[156,742,190,212]" pageId="3" pageNumber="212">
Table 1. Measurements (µm) of
<taxonomicName id="2CDF4D2FFFE38337449EB4F70596FF65" authority="Fujimoto, 2018" authorityName="Fujimoto" authorityYear="2018" box="[466,668,191,212]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="ittanmomen" status="sp. nov.">
<emphasis id="D9ABEABEFFE38337449EB4F70596FF65" box="[466,668,191,212]" italics="true" pageId="3" pageNumber="212">Tanarctus ittanmomen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C29857C5FFE3833747EDB4F705ECFF65" box="[673,742,191,212]" pageId="3" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="EB6036ACFFE383374499B4BA025BFCF2" pageId="3" pageNumber="212">
<table id="99DFC40CFFE37CCB45FDB4BA027FFCF2" box="[177,1397,242,835]" gridcols="7" gridrows="18" pageId="3" pageNumber="212">
<tr id="55EF34EEFFE37CCB45FDB4BA027FFEB4" box="[177,1397,242,261]" gridrow="0" pageId="3" pageNumber="212" rowspan-0="1">
<th id="163E5D92FFE37CCB448AB4BA053EFEB4" box="[454,564,242,261]" gridcol="1" gridrow="0" pageId="3" pageNumber="212">Holotype</th>
<th id="163E5D92FFE37CCB4721B4BA05D6FEB4" box="[621,732,242,261]" gridcol="2" gridrow="0" pageId="3" pageNumber="212">Paratype</th>
<th id="163E5D92FFE37CCB4659B4BA0489FEB4" box="[789,899,242,261]" gridcol="3" gridrow="0" pageId="3" pageNumber="212">Paratype</th>
<th id="163E5D92FFE37CCB46EBB4BA0335FEB4" box="[935,1087,242,261]" gridcol="4" gridrow="0" pageId="3" pageNumber="212">Paratype</th>
<th id="163E5D92FFE37CCB4124B4BA03C4FEB4" box="[1128,1230,242,261]" gridcol="5" gridrow="0" pageId="3" pageNumber="212">Paratype</th>
<th id="163E5D92FFE37CCB4043B4BA027FFEB4" box="[1295,1397,242,261]" gridcol="6" gridrow="0" pageId="3" pageNumber="212">Paratype</th>
</tr>
<tr id="55EF34EEFFE37CCB45FDB55E027FFE98" box="[177,1397,278,297]" gridrow="1" pageId="3" pageNumber="212" rowspan-0="1">
<td id="163E5D92FFE37CCB448AB55E053EFE98" box="[454,564,278,297]" gridcol="1" gridrow="1" pageId="3" pageNumber="212">KUZ Z1954</td>
<td id="163E5D92FFE37CCB4721B55E05D6FE98" box="[621,732,278,297]" gridcol="2" gridrow="1" pageId="3" pageNumber="212">KUZ Z1955</td>
<td id="163E5D92FFE37CCB4659B55E0489FE98" box="[789,899,278,297]" gridcol="3" gridrow="1" pageId="3" pageNumber="212">KUZ Z1956</td>
<td id="163E5D92FFE37CCB46EBB55E0335FE98" box="[935,1087,278,297]" gridcol="4" gridrow="1" pageId="3" pageNumber="212">KUZ Z1957</td>
<td id="163E5D92FFE37CCB4124B55E03C4FE98" box="[1128,1230,278,297]" gridcol="5" gridrow="1" pageId="3" pageNumber="212">KUZ Z1958</td>
<td id="163E5D92FFE37CCB4043B55E027FFE98" box="[1295,1397,278,297]" gridcol="6" gridrow="1" pageId="3" pageNumber="212">KUZ Z1959</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB572027FFED9" box="[177,1397,314,360]" gridrow="2" pageId="3" pageNumber="212" rowspan-0="1">
<td id="163E5D92FFE37CCB448AB572053EFED9" box="[454,564,314,360]" gridcol="1" gridrow="2" pageId="3" pageNumber="212">Adult female</td>
<td id="163E5D92FFE37CCB4721B57205D6FED9" box="[621,732,314,360]" gridcol="2" gridrow="2" pageId="3" pageNumber="212">Adult female</td>
<td id="163E5D92FFE37CCB4659B5720489FED9" box="[789,899,314,360]" gridcol="3" gridrow="2" pageId="3" pageNumber="212">Adult female</td>
<td id="163E5D92FFE37CCB46EBB5720335FED9" box="[935,1087,314,360]" gridcol="4" gridrow="2" pageId="3" pageNumber="212">Juvenile moulting into adult female</td>
<td id="163E5D92FFE37CCB4124B57203C4FED9" box="[1128,1230,314,360]" gridcol="5" gridrow="2" pageId="3" pageNumber="212">Adult male</td>
<td id="163E5D92FFE37CCB4043B572027FFED9" box="[1295,1397,314,360]" gridcol="6" gridrow="2" pageId="3" pageNumber="212">Four-digit juvenile</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB536027FFE23" box="[177,1397,382,402]" gridrow="3" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB5360698FE23" box="[177,402,382,402]" gridcol="0" gridrow="3" pageId="3" pageNumber="212">Body length</th>
<td id="163E5D92FFE37CCB448AB536053EFE23" box="[454,564,382,402]" gridcol="1" gridrow="3" pageId="3" pageNumber="212">145</td>
<td id="163E5D92FFE37CCB4721B53605D6FE23" box="[621,732,382,402]" gridcol="2" gridrow="3" pageId="3" pageNumber="212">132</td>
<td id="163E5D92FFE37CCB4659B5360489FE23" box="[789,899,382,402]" gridcol="3" gridrow="3" pageId="3" pageNumber="212">131</td>
<td id="163E5D92FFE37CCB46EBB5360335FE23" box="[935,1087,382,402]" gridcol="4" gridrow="3" pageId="3" pageNumber="212">74</td>
<td id="163E5D92FFE37CCB4124B53603C4FE23" box="[1128,1230,382,402]" gridcol="5" gridrow="3" pageId="3" pageNumber="212">109</td>
<td id="163E5D92FFE37CCB4043B536027FFE23" box="[1295,1397,382,402]" gridcol="6" gridrow="3" pageId="3" pageNumber="212">113</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB5D5027FFE00" box="[177,1397,413,433]" gridrow="4" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB5D50698FE00" box="[177,402,413,433]" gridcol="0" gridrow="4" pageId="3" pageNumber="212">Body width</th>
<td id="163E5D92FFE37CCB448AB5D5053EFE00" box="[454,564,413,433]" gridcol="1" gridrow="4" pageId="3" pageNumber="212">71</td>
<td id="163E5D92FFE37CCB4721B5D505D6FE00" box="[621,732,413,433]" gridcol="2" gridrow="4" pageId="3" pageNumber="212">65</td>
<td id="163E5D92FFE37CCB4659B5D50489FE00" box="[789,899,413,433]" gridcol="3" gridrow="4" pageId="3" pageNumber="212">65</td>
<td id="163E5D92FFE37CCB46EBB5D50335FE00" box="[935,1087,413,433]" gridcol="4" gridrow="4" pageId="3" pageNumber="212">61</td>
<td id="163E5D92FFE37CCB4124B5D503C4FE00" box="[1128,1230,413,433]" gridcol="5" gridrow="4" pageId="3" pageNumber="212">55</td>
<td id="163E5D92FFE37CCB4043B5D5027FFE00" box="[1295,1397,413,433]" gridcol="6" gridrow="4" pageId="3" pageNumber="212">56</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB5F4027FFE61" box="[177,1397,444,464]" gridrow="5" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB5F40698FE61" box="[177,402,444,464]" gridcol="0" gridrow="5" pageId="3" pageNumber="212">Median cirrus</th>
<td id="163E5D92FFE37CCB448AB5F4053EFE61" box="[454,564,444,464]" gridcol="1" gridrow="5" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4721B5F405D6FE61" box="[621,732,444,464]" gridcol="2" gridrow="5" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4659B5F40489FE61" box="[789,899,444,464]" gridcol="3" gridrow="5" pageId="3" pageNumber="212">10</td>
<td id="163E5D92FFE37CCB46EBB5F40335FE61" box="[935,1087,444,464]" gridcol="4" gridrow="5" pageId="3" pageNumber="212">8</td>
<td id="163E5D92FFE37CCB4124B5F403C4FE61" box="[1128,1230,444,464]" gridcol="5" gridrow="5" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4043B5F4027FFE61" box="[1295,1397,444,464]" gridcol="6" gridrow="5" pageId="3" pageNumber="212">9</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB593027FFE5E" box="[177,1397,475,495]" gridrow="6" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB5930698FE5E" box="[177,402,475,495]" gridcol="0" gridrow="6" pageId="3" pageNumber="212">Internal cirrus</th>
<td id="163E5D92FFE37CCB448AB593053EFE5E" box="[454,564,475,495]" gridcol="1" gridrow="6" pageId="3" pageNumber="212">37</td>
<td id="163E5D92FFE37CCB4721B59305D6FE5E" box="[621,732,475,495]" gridcol="2" gridrow="6" pageId="3" pageNumber="212">37</td>
<td id="163E5D92FFE37CCB4659B5930489FE5E" box="[789,899,475,495]" gridcol="3" gridrow="6" pageId="3" pageNumber="212">34</td>
<td id="163E5D92FFE37CCB46EBB5930335FE5E" box="[935,1087,475,495]" gridcol="4" gridrow="6" pageId="3" pageNumber="212">33</td>
<td id="163E5D92FFE37CCB4124B59303C4FE5E" box="[1128,1230,475,495]" gridcol="5" gridrow="6" pageId="3" pageNumber="212">—</td>
<td id="163E5D92FFE37CCB4043B593027FFE5E" box="[1295,1397,475,495]" gridcol="6" gridrow="6" pageId="3" pageNumber="212">31</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB5B2027FFDBC" box="[177,1397,506,525]" gridrow="7" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB5B20698FDBC" box="[177,402,506,525]" gridcol="0" gridrow="7" pageId="3" pageNumber="212">External cirrus</th>
<td id="163E5D92FFE37CCB448AB5B2053EFDBC" box="[454,564,506,525]" gridcol="1" gridrow="7" pageId="3" pageNumber="212">22</td>
<td id="163E5D92FFE37CCB4721B5B205D6FDBC" box="[621,732,506,525]" gridcol="2" gridrow="7" pageId="3" pageNumber="212">24</td>
<td id="163E5D92FFE37CCB4659B5B20489FDBC" box="[789,899,506,525]" gridcol="3" gridrow="7" pageId="3" pageNumber="212">21</td>
<td id="163E5D92FFE37CCB46EBB5B20335FDBC" box="[935,1087,506,525]" gridcol="4" gridrow="7" pageId="3" pageNumber="212">19</td>
<td id="163E5D92FFE37CCB4124B5B203C4FDBC" box="[1128,1230,506,525]" gridcol="5" gridrow="7" pageId="3" pageNumber="212">19</td>
<td id="163E5D92FFE37CCB4043B5B2027FFDBC" box="[1295,1397,506,525]" gridcol="6" gridrow="7" pageId="3" pageNumber="212">18</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB651027FFD9D" box="[177,1397,537,556]" gridrow="8" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB6510698FD9D" box="[177,402,537,556]" gridcol="0" gridrow="8" pageId="3" pageNumber="212">Lateral cirrus</th>
<td id="163E5D92FFE37CCB448AB651053EFD9D" box="[454,564,537,556]" gridcol="1" gridrow="8" pageId="3" pageNumber="212">16</td>
<td id="163E5D92FFE37CCB4721B65105D6FD9D" box="[621,732,537,556]" gridcol="2" gridrow="8" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4659B6510489FD9D" box="[789,899,537,556]" gridcol="3" gridrow="8" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB46EBB6510335FD9D" box="[935,1087,537,556]" gridcol="4" gridrow="8" pageId="3" pageNumber="212">12</td>
<td id="163E5D92FFE37CCB4124B65103C4FD9D" box="[1128,1230,537,556]" gridcol="5" gridrow="8" pageId="3" pageNumber="212">12</td>
<td id="163E5D92FFE37CCB4043B651027FFD9D" box="[1295,1397,537,556]" gridcol="6" gridrow="8" pageId="3" pageNumber="212">13</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB670027FFDFA" box="[177,1397,568,587]" gridrow="9" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB6700698FDFA" box="[177,402,568,587]" gridcol="0" gridrow="9" pageId="3" pageNumber="212">Primary clava</th>
<td id="163E5D92FFE37CCB448AB670053EFDFA" box="[454,564,568,587]" gridcol="1" gridrow="9" pageId="3" pageNumber="212">195</td>
<td id="163E5D92FFE37CCB4721B67005D6FDFA" box="[621,732,568,587]" gridcol="2" gridrow="9" pageId="3" pageNumber="212">179</td>
<td id="163E5D92FFE37CCB4659B6700489FDFA" box="[789,899,568,587]" gridcol="3" gridrow="9" pageId="3" pageNumber="212">162</td>
<td id="163E5D92FFE37CCB46EBB6700335FDFA" box="[935,1087,568,587]" gridcol="4" gridrow="9" pageId="3" pageNumber="212">153</td>
<td id="163E5D92FFE37CCB4124B67003C4FDFA" box="[1128,1230,568,587]" gridcol="5" gridrow="9" pageId="3" pageNumber="212">187</td>
<td id="163E5D92FFE37CCB4043B670027FFDFA" box="[1295,1397,568,587]" gridcol="6" gridrow="9" pageId="3" pageNumber="212">158</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB61F027FFDDA" box="[177,1397,599,619]" gridrow="10" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB61F0698FDDA" box="[177,402,599,619]" gridcol="0" gridrow="10" pageId="3" pageNumber="212">Cirrus E</th>
<td id="163E5D92FFE37CCB448AB61F053EFDDA" box="[454,564,599,619]" gridcol="1" gridrow="10" pageId="3" pageNumber="212">30</td>
<td id="163E5D92FFE37CCB4721B61F05D6FDDA" box="[621,732,599,619]" gridcol="2" gridrow="10" pageId="3" pageNumber="212">—</td>
<td id="163E5D92FFE37CCB4659B61F0489FDDA" box="[789,899,599,619]" gridcol="3" gridrow="10" pageId="3" pageNumber="212">29</td>
<td id="163E5D92FFE37CCB46EBB61F0335FDDA" box="[935,1087,599,619]" gridcol="4" gridrow="10" pageId="3" pageNumber="212">26</td>
<td id="163E5D92FFE37CCB4124B61F03C4FDDA" box="[1128,1230,599,619]" gridcol="5" gridrow="10" pageId="3" pageNumber="212">28</td>
<td id="163E5D92FFE37CCB4043B61F027FFDDA" box="[1295,1397,599,619]" gridcol="6" gridrow="10" pageId="3" pageNumber="212">26</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB63E027FFD38" box="[177,1397,630,649]" gridrow="11" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB63E0698FD38" box="[177,402,630,649]" gridcol="0" gridrow="11" pageId="3" pageNumber="212">Leg I sensory organ</th>
<td id="163E5D92FFE37CCB448AB63E053EFD38" box="[454,564,630,649]" gridcol="1" gridrow="11" pageId="3" pageNumber="212">18</td>
<td id="163E5D92FFE37CCB4721B63E05D6FD38" box="[621,732,630,649]" gridcol="2" gridrow="11" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4659B63E0489FD38" box="[789,899,630,649]" gridcol="3" gridrow="11" pageId="3" pageNumber="212">14</td>
<td id="163E5D92FFE37CCB46EBB63E0335FD38" box="[935,1087,630,649]" gridcol="4" gridrow="11" pageId="3" pageNumber="212">14</td>
<td id="163E5D92FFE37CCB4124B63E03C4FD38" box="[1128,1230,630,649]" gridcol="5" gridrow="11" pageId="3" pageNumber="212">10</td>
<td id="163E5D92FFE37CCB4043B63E027FFD38" box="[1295,1397,630,649]" gridcol="6" gridrow="11" pageId="3" pageNumber="212">12</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB6DD027FFD19" box="[177,1397,661,680]" gridrow="12" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB6DD0698FD19" box="[177,402,661,680]" gridcol="0" gridrow="12" pageId="3" pageNumber="212">Leg II sensory organ</th>
<td id="163E5D92FFE37CCB448AB6DD053EFD19" box="[454,564,661,680]" gridcol="1" gridrow="12" pageId="3" pageNumber="212">21</td>
<td id="163E5D92FFE37CCB4721B6DD05D6FD19" box="[621,732,661,680]" gridcol="2" gridrow="12" pageId="3" pageNumber="212">15</td>
<td id="163E5D92FFE37CCB4659B6DD0489FD19" box="[789,899,661,680]" gridcol="3" gridrow="12" pageId="3" pageNumber="212">18</td>
<td id="163E5D92FFE37CCB46EBB6DD0335FD19" box="[935,1087,661,680]" gridcol="4" gridrow="12" pageId="3" pageNumber="212">10</td>
<td id="163E5D92FFE37CCB4124B6DD03C4FD19" box="[1128,1230,661,680]" gridcol="5" gridrow="12" pageId="3" pageNumber="212">13</td>
<td id="163E5D92FFE37CCB4043B6DD027FFD19" box="[1295,1397,661,680]" gridcol="6" gridrow="12" pageId="3" pageNumber="212">15</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB6FC027FFD76" box="[177,1397,692,711]" gridrow="13" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB6FC0698FD76" box="[177,402,692,711]" gridcol="0" gridrow="13" pageId="3" pageNumber="212">Leg III sensory organ</th>
<td id="163E5D92FFE37CCB448AB6FC053EFD76" box="[454,564,692,711]" gridcol="1" gridrow="13" pageId="3" pageNumber="212">24</td>
<td id="163E5D92FFE37CCB4721B6FC05D6FD76" box="[621,732,692,711]" gridcol="2" gridrow="13" pageId="3" pageNumber="212">—</td>
<td id="163E5D92FFE37CCB4659B6FC0489FD76" box="[789,899,692,711]" gridcol="3" gridrow="13" pageId="3" pageNumber="212">23</td>
<td id="163E5D92FFE37CCB46EBB6FC0335FD76" box="[935,1087,692,711]" gridcol="4" gridrow="13" pageId="3" pageNumber="212">12</td>
<td id="163E5D92FFE37CCB4124B6FC03C4FD76" box="[1128,1230,692,711]" gridcol="5" gridrow="13" pageId="3" pageNumber="212">16</td>
<td id="163E5D92FFE37CCB4043B6FC027FFD76" box="[1295,1397,692,711]" gridcol="6" gridrow="13" pageId="3" pageNumber="212">19</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB69B027FFD57" box="[177,1397,723,742]" gridrow="14" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB69B0698FD57" box="[177,402,723,742]" gridcol="0" gridrow="14" pageId="3" pageNumber="212">Leg IV appendage</th>
<td id="163E5D92FFE37CCB448AB69B053EFD57" box="[454,564,723,742]" gridcol="1" gridrow="14" pageId="3" pageNumber="212">301</td>
<td id="163E5D92FFE37CCB4721B69B05D6FD57" box="[621,732,723,742]" gridcol="2" gridrow="14" pageId="3" pageNumber="212">284</td>
<td id="163E5D92FFE37CCB4659B69B0489FD57" box="[789,899,723,742]" gridcol="3" gridrow="14" pageId="3" pageNumber="212">307</td>
<td id="163E5D92FFE37CCB46EBB69B0335FD57" box="[935,1087,723,742]" gridcol="4" gridrow="14" pageId="3" pageNumber="212">259</td>
<td id="163E5D92FFE37CCB4124B69B03C4FD57" box="[1128,1230,723,742]" gridcol="5" gridrow="14" pageId="3" pageNumber="212">253</td>
<td id="163E5D92FFE37CCB4043B69B027FFD57" box="[1295,1397,723,742]" gridcol="6" gridrow="14" pageId="3" pageNumber="212">—</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB6BA027FFCB4" box="[177,1397,754,773]" gridrow="15" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB6BA0698FCB4" box="[177,402,754,773]" gridcol="0" gridrow="15" pageId="3" pageNumber="212">No. of secondary branches</th>
<td id="163E5D92FFE37CCB448AB6BA053EFCB4" box="[454,564,754,773]" gridcol="1" gridrow="15" pageId="3" pageNumber="212">9, 12</td>
<td id="163E5D92FFE37CCB4721B6BA05D6FCB4" box="[621,732,754,773]" gridcol="2" gridrow="15" pageId="3" pageNumber="212">16, 14</td>
<td id="163E5D92FFE37CCB4659B6BA0489FCB4" box="[789,899,754,773]" gridcol="3" gridrow="15" pageId="3" pageNumber="212">12, 10</td>
<td id="163E5D92FFE37CCB46EBB6BA0335FCB4" box="[935,1087,754,773]" gridcol="4" gridrow="15" pageId="3" pageNumber="212">9, —</td>
<td id="163E5D92FFE37CCB4124B6BA03C4FCB4" box="[1128,1230,754,773]" gridcol="5" gridrow="15" pageId="3" pageNumber="212">8, 6</td>
<td id="163E5D92FFE37CCB4043B6BA027FFCB4" box="[1295,1397,754,773]" gridcol="6" gridrow="15" pageId="3" pageNumber="212">—</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB759027FFC95" box="[177,1397,785,804]" gridrow="16" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB7590698FC95" box="[177,402,785,804]" gridcol="0" gridrow="16" pageId="3" pageNumber="212">Gonopore-anus distance</th>
<td id="163E5D92FFE37CCB448AB759053EFC95" box="[454,564,785,804]" gridcol="1" gridrow="16" pageId="3" pageNumber="212">8</td>
<td id="163E5D92FFE37CCB4721B75905D6FC95" box="[621,732,785,804]" gridcol="2" gridrow="16" pageId="3" pageNumber="212">8</td>
<td id="163E5D92FFE37CCB4659B7590489FC95" box="[789,899,785,804]" gridcol="3" gridrow="16" pageId="3" pageNumber="212">8</td>
<td id="163E5D92FFE37CCB46EBB7590335FC95" box="[935,1087,785,804]" gridcol="4" gridrow="16" pageId="3" pageNumber="212">—</td>
<td id="163E5D92FFE37CCB4124B75903C4FC95" box="[1128,1230,785,804]" gridcol="5" gridrow="16" pageId="3" pageNumber="212">0</td>
<td id="163E5D92FFE37CCB4043B759027FFC95" box="[1295,1397,785,804]" gridcol="6" gridrow="16" pageId="3" pageNumber="212">—</td>
</tr>
<tr id="55EF34EEFFE37CCB45FDB778027FFCF2" box="[177,1397,816,835]" gridrow="17" pageId="3" pageNumber="212">
<th id="163E5D92FFE37CCB45FDB7780698FCF2" box="[177,402,816,835]" gridcol="0" gridrow="17" pageId="3" pageNumber="212">No. of amoebocytes</th>
<td id="163E5D92FFE37CCB448AB778053EFCF2" box="[454,564,816,835]" gridcol="1" gridrow="17" pageId="3" pageNumber="212">7</td>
<td id="163E5D92FFE37CCB4721B77805D6FCF2" box="[621,732,816,835]" gridcol="2" gridrow="17" pageId="3" pageNumber="212">2</td>
<td id="163E5D92FFE37CCB4659B7780489FCF2" box="[789,899,816,835]" gridcol="3" gridrow="17" pageId="3" pageNumber="212">0</td>
<td id="163E5D92FFE37CCB46EBB7780335FCF2" box="[935,1087,816,835]" gridcol="4" gridrow="17" pageId="3" pageNumber="212">7</td>
<td id="163E5D92FFE37CCB4124B77803C4FCF2" box="[1128,1230,816,835]" gridcol="5" gridrow="17" pageId="3" pageNumber="212">4</td>
<td id="163E5D92FFE37CCB4043B778027FFCF2" box="[1295,1397,816,835]" gridcol="6" gridrow="17" pageId="3" pageNumber="212">8</td>
</tr>
</table>
</paragraph>
<paragraph id="EB6036ACFFE3833745F9B7DA05BFFB9D" blockId="3.[156,768,880,2077]" pageId="3" pageNumber="212">
Female genital structure (
<figureCitation id="73E42A29FFE3833744F6B7DA0501FC1B" box="[442,523,914,938]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="3" pageNumber="212">Figs 1A</figureCitation>
,
<figureCitation id="73E42A29FFE38337475AB7DA0539FC1B" box="[534,563,914,938]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="3" pageNumber="212">2B</figureCitation>
) consists of rosettelike gonopore (7 µm in diameter), paired seminal receptacles with anus 8 µm posterior to gonopore. Seminal receptacle opens ventrally, overlapping gonopore, with duct of even width running laterally, terminating in round vesicle.
</paragraph>
<paragraph id="EB6036ACFFE3833745F9B07C059AF8EB" blockId="3.[156,768,880,2077]" pageId="3" pageNumber="212">
<emphasis id="D9ABEABEFFE3833745F9B07C0563FBFC" bold="true" box="[181,617,1076,1101]" pageId="3" pageNumber="212">
Additional information from
<typeStatus id="3464880EFFE3833744B0B07D056FFBFC" box="[508,613,1077,1101]" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
.
</emphasis>
Dense, rodlike particles embedded in the cephalic cirri (
<figureCitation id="73E42A29FFE383374736B01D05C3FBDC" box="[634,713,1109,1133]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="3" pageNumber="212">Fig. 2C</figureCitation>
) and in the leg I sensory organ were confirmed in all of the
<typeStatus id="3464880EFFE383374785B03E07DEFB1E" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
.
<materialsCitation id="5BB73CF1FFE3833745ADB0DE07CCFB41" collectionCode="KUZ" collectorName="The" pageId="3" pageNumber="212" specimenCode="Z1957" specimenCount="1">
<collectorName id="462A537AFFE3833745ADB0DE060DFB1F" box="[225,263,1174,1198]" pageId="3" pageNumber="212">The</collectorName>
complete contours of the flat secondary clavae were recognised in the old cuticle of
<collectionCode id="8DCEAE69FFE38337476CB0FF0559FB7F" box="[544,595,1207,1230]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE383374710B0FF0591FB7E" box="[604,667,1207,1231]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">Z1957</specimenCode>
(
<figureCitation id="73E42A29FFE3833747E7B0FF05F1FB7E" box="[683,763,1207,1231]" captionStart="Fig" captionStartId="1.[156,187,1939,1961]" captionTargetBox="[201,1387,208,1898]" captionTargetId="figure-3@1.[198,1390,205,1901]" captionTargetPageId="1" captionText="Fig. 1. Drawings of Tanarctus ittanmomen sp. nov. A, habitus of holotype, KUZ Z1954; B, secondary clavae of paratype, KUZ Z1957. Abbreviations: am, amoebocyte; an, anus; bt, buccal tube; cE, cirrus E; ec, external cirrus; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; l4a, leg IV appendage; mc, median cirrus; mo, mouth opening; pb, pharyngeal bulb; pc, primary clava; sc, secondary clava; so1–3, leg I–III sensory organs; sr, seminal receptacle; ss, stylet support; st, stylet; sw1–3, swellings of secondary clavae." figureDoi="http://doi.org/10.5281/zenodo.5738173" httpUri="https://zenodo.org/record/5738173/files/figure.png" pageId="3" pageNumber="212">Figs 1B</figureCitation>
,
<figureCitation id="73E42A29FFE3833745D0B09007B4FB41" box="[156,190,1240,1264]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="3" pageNumber="212">2D</figureCitation>
)
</materialsCitation>
. The paired clavae each had two swellings anterior to the mouth and one posterior swelling. All swellings were directed internally, but never fused with their counterparts. Amoebocytes were distributed both anteriorly and posteriorly in the cephalic region in the
<typeStatus id="3464880EFFE3833744BBB1120552FAC3" box="[503,600,1370,1394]" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
(see
<tableCitation id="A65D0317FFE3833747DCB11105D0FAC3" box="[656,730,1369,1394]" captionStart="Table 1" captionStartId="3.[156,204,190,212]" captionText="Table 1. Measurements (µm) of Tanarctus ittanmomen sp.nov." pageId="3" pageNumber="212">Table 1</tableCitation>
for number of amoebocytes).
<materialsCitation id="5BB73CF1FFE3833744FDB13207F3FA45" collectionCode="KUZ" collectorName="The" pageId="3" pageNumber="212" specimenCode="Z1955" specimenCount="1">
<collectorName id="462A537AFFE3833744FDB13206DDFA23" box="[433,471,1402,1426]" pageId="3" pageNumber="212">The</collectorName>
dorsal spur on the internal claws and the well-developed calcar on all claws were best observed when
<collectionCode id="8DCEAE69FFE383374473B1F40678FA62" box="[319,370,1468,1491]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE383374435B1F306B3FA62" box="[377,441,1467,1491]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">Z1955</specimenCode>
was squeezed in distilled water (
<figureCitation id="73E42A29FFE3833745EFB19407FBFA45" box="[163,241,1500,1524]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="3" pageNumber="212">Fig. 2E</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="5BB73CF1FFE383374448B19407F6F9A5" collectionCode="KUZ" collectorName="The" pageId="3" pageNumber="212" specimenCode="Z1958" specimenCount="1">
<collectorName id="462A537AFFE383374448B1940620FA45" box="[260,298,1500,1524]" pageId="3" pageNumber="212">The</collectorName>
male gonopore was observed in
<collectionCode id="8DCEAE69FFE3833747C8B19405BDFA42" box="[644,695,1500,1523]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">KUZ</collectionCode>
<specimenCode id="BB799ED7FFE3833747F3B19305F5FA42" box="[703,767,1499,1523]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="212">Z1958</specimenCode>
(
<figureCitation id="73E42A29FFE3833745EFB1B407FEF9A5" box="[163,244,1532,1556]" captionStart="Fig" captionStartId="2.[821,852,1744,1766]" captionTargetBox="[861,1391,206,1705]" captionTargetId="figure-547@2.[860,1392,205,1706]" captionTargetPageId="2" captionText="Fig. 2. DIC micrographs of Tanarctus ittanmomen sp. nov. A, habitus of holotypic female, KUZ Z1954; B, genital structure of holotypic female, KUZ Z1954; C, cephalic sensory organs of paratypic female squeezed in distilled water, KUZ Z1955; D, leg II of paratypic female squeezed in distilled water, KUZ Z1955; E, secondary clavae of paratypic female, KUZ Z1957; F, genital structure of paratypic male, KUZ Z1958. Abbreviations: an, anus; ca, calcar; ds, dorsal spur; ec, external cirrus; fe, femur; fg, female gonopore; ic, internal cirrus; lc, lateral cirrus; mg, male gonopore; pc, primary clava; pt, pretarsus; so2, leg II sensory organ; srd, seminal receptacle duct; sro, seminal receptacle opening; sw1–3, swellings of secondary clavae; ta, tarsus; ti, tibia; white arrowhead, dense particle." figureDoi="http://doi.org/10.5281/zenodo.5738175" httpUri="https://zenodo.org/record/5738175/files/figure.png" pageId="3" pageNumber="212">Fig. 2F</figureCitation>
)
</materialsCitation>
. The longitudinally long, oval gonopore (length 6 µm, width 5 µm) opens immediately anterior to the anus. As only
<specimenCount id="FDD9FD25FFE3833745B8B2760658F9E4" box="[244,338,1597,1621]" pageId="3" pageNumber="212" type="male">one male</specimenCount>
was observed, sexual dimorphism in the quantitative traits was not investigated.
<materialsCitation id="5BB73CF1FFE38337477AB2160510F966" collectionCode="IV" collectorName="The" pageId="3" pageNumber="212" specimenCount="2" typeStatus="paratype">
<collectorName id="462A537AFFE38337477AB2160556F9C7" box="[566,604,1630,1654]" pageId="3" pageNumber="212">The</collectorName>
leg
<collectionCode id="8DCEAE69FFE3833747C4B21605A8F9C4" box="[648,674,1630,1653]" pageId="3" pageNumber="212">IV</collectionCode>
appendages of the
<typeStatus id="3464880EFFE38337445FB237067EF926" box="[275,372,1663,1687]" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
resemble the longer appendage of the
<typeStatus id="3464880EFFE3833745D0B2D707FDF906" box="[156,247,1695,1719]" pageId="3" pageNumber="212" type="holotype">holotype</typeStatus>
, but exhibited variation in the number and length of the secondary branches (
<tableCitation id="A65D0317FFE38337448DB2F70505F966" box="[449,527,1727,1751]" captionStart="Table 1" captionStartId="3.[156,204,190,212]" captionText="Table 1. Measurements (µm) of Tanarctus ittanmomen sp.nov." pageId="3" pageNumber="212">Table 1</tableCitation>
)
</materialsCitation>
. Unlike the
<typeStatus id="3464880EFFE3833747ECB2F705F1F966" box="[672,763,1727,1751]" pageId="3" pageNumber="212" type="holotype">holotype</typeStatus>
, no apparent difference was observed between the lengths of the two main branches in the
<typeStatus id="3464880EFFE38337449FB349053DF8A8" box="[467,567,1793,1817]" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
.
<materialsCitation id="5BB73CF1FFE38337470EB3480586F8EB" collectionCode="IV" collectorName="The" pageId="3" pageNumber="212" specimenCount="2" typeStatus="holotype">
<collectorName id="462A537AFFE38337470EB3480562F8A9" box="[578,616,1792,1816]" pageId="3" pageNumber="212">The</collectorName>
shorter leg
<collectionCode id="8DCEAE69FFE3833747A8B34905F4F8A9" box="[740,766,1793,1816]" pageId="3" pageNumber="212">IV</collectionCode>
appendage of the
<typeStatus id="3464880EFFE38337441FB36906A6F888" box="[339,428,1825,1849]" pageId="3" pageNumber="212" type="holotype">holotype</typeStatus>
is deemed to be deformed as no similar appendage was observed in the
<typeStatus id="3464880EFFE383374764B30A0586F8EB" box="[552,652,1858,1882]" pageId="3" pageNumber="212" type="paratype">paratypes</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3C56527FFE3833745F9B32A0263F9E4" pageId="3" pageNumber="212" type="discussion">
<paragraph id="EB6036ACFFE3833745F9B32A0263F9E4" blockId="3.[156,768,880,2077]" lastBlockId="3.[820,1432,881,1621]" pageId="3" pageNumber="212">
<emphasis id="D9ABEABEFFE3833745F9B32A0613F8CB" bold="true" box="[181,281,1890,1914]" pageId="3" pageNumber="212">Remarks.</emphasis>
The new species resembles
<taxonomicName id="2CDF4D2FFFE383374718B32B0696F82A" authority="Lindgren, 1971" authorityName="Lindgren" authorityYear="1971" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="arborspinosus">
<emphasis id="D9ABEABEFFE383374718B32B07E4F82B" italics="true" pageId="3" pageNumber="212">Tanarctus arborspinosus</emphasis>
<bibRefCitation id="8F4E4B5DFFE3833745B7B3CA0696F82A" author="Lindgren, E. W." box="[251,412,1922,1947]" pageId="3" pageNumber="212" pagination="481 - 496" refId="ref2936" refString="Lindgren, E. W. 1971. Psammolittoral marine tardigrades from North Carolina and their conformity to worldwide zonation patterns. Cahiers de Biologie Marine 12: 481 - 496." type="journal article" year="1971">Lindgren, 1971</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2CDF4D2FFFE3833744E0B3CB07C5F80A" authority="Renaud-Mornant, 1980" authorityName="Renaud-Mornant" authorityYear="1980" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="dendriticus">
<emphasis id="D9ABEABEFFE3833744E0B3CB06B1F82B" box="[428,443,1923,1946]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE383374481B3CA0536F82B" box="[461,572,1922,1946]" italics="true" pageId="3" pageNumber="212">dendriticus</emphasis>
<bibRefCitation id="8F4E4B5DFFE383374704B3CA07C5F80A" author="Renaud-Mornant, J." pageId="3" pageNumber="212" pagination="129 - 141" refId="ref3128" refString="Renaud-Mornant, J. 1980. Description de trois especies nouvelles du genre Tanarctus Renaud-Debyser, 1959, et creation de la sousfamille des Tanarctinae subfam. nov. (Tardigrada, Heterotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 2: 129 - 141." type="book chapter" year="1980">Renaud-Mornant, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2CDF4D2FFFE3833745AFB3EC0695F80A" authorityName="Jorgensen, Boesgaard, Mobjerg &amp; Kristensen" authorityYear="2014" box="[227,415,1955,1979]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="hirsutospinosus">
<emphasis id="D9ABEABEFFE3833745AFB3EC07F8F80A" box="[227,242,1956,1979]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE383374444B3EB0695F80A" box="[264,415,1955,1979]" italics="true" pageId="3" pageNumber="212">hirsutospinosus</emphasis>
</taxonomicName>
<bibRefCitation id="8F4E4B5DFFE3833744FCB3EA0673F86D" author="JOrgensen, A. &amp; Boesgaard, T. M. &amp; MObjerg, N. &amp; Kristensen, R. M." pageId="3" pageNumber="212" pagination="401 - 443" refId="ref2786" refString="JOrgensen, A., Boesgaard, T. M., MObjerg, N., and Kristensen, R. M. 2014. The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada. Zootaxa 3802: 401 - 443." type="journal article" year="2014">JØrgensen, Boesgaard, MØbjerg and Kristensen, 2014</bibRefCitation>
, and
<taxonomicName id="2CDF4D2FFFE3833744F8B38C07F8F7AC" authority="Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi, 1982" authorityName="Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi" authorityYear="1982" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="longisetosus">
<emphasis id="D9ABEABEFFE3833744F8B38C06C9F86A" box="[436,451,1988,2011]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337449EB38B054DF86A" box="[466,583,1987,2011]" italics="true" pageId="3" pageNumber="212">longisetosus</emphasis>
<bibRefCitation id="8F4E4B5DFFE38337471CB38B07F8F7AC" author="Grimaldi de Zio, S. &amp; D'Addabbo Gallo, M. &amp; Morone de Lucia, M. R. &amp; Vaccarella, R. &amp; Grimaldi, P." pageId="3" pageNumber="212" pagination="415 - 426" refId="ref2723" refString="Grimaldi de Zio, S., D'Addabbo Gallo, M., Morone de Lucia, M. R., Vaccarella, R., and Grimaldi, P. 1982. Quattro nuove specie di Halechiniscidae rinvenute in due grotte sottomarine dell'Italia meridionale. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine 23: 415 - 426." type="journal article" year="1982">Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi, 1982</bibRefCitation>
</taxonomicName>
, due to its branching leg IV appendages with unmodified tips (
<bibRefCitation id="8F4E4B5DFFE3833746AEB739038CFC38" author="Lindgren, E. W." box="[994,1158,881,905]" pageId="3" pageNumber="212" pagination="481 - 496" refId="ref2936" refString="Lindgren, E. W. 1971. Psammolittoral marine tardigrades from North Carolina and their conformity to worldwide zonation patterns. Cahiers de Biologie Marine 12: 481 - 496." type="journal article" year="1971">Lindgren 1971</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE3833741D1B739029EFC38" author="Renaud-Mornant, J." box="[1181,1428,881,905]" pageId="3" pageNumber="212" pagination="129 - 141" refId="ref3128" refString="Renaud-Mornant, J. 1980. Description de trois especies nouvelles du genre Tanarctus Renaud-Debyser, 1959, et creation de la sousfamille des Tanarctinae subfam. nov. (Tardigrada, Heterotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 2: 129 - 141." type="book chapter" year="1980">Renaud-Mornant 1980</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374678B7DA035AFC1B" author="Grimaldi de Zio, S. &amp; D'Addabbo Gallo, M. &amp; Morone de Lucia, M. R. &amp; Vaccarella, R. &amp; Grimaldi, P." box="[820,1104,914,938]" pageId="3" pageNumber="212" pagination="415 - 426" refId="ref2723" refString="Grimaldi de Zio, S., D'Addabbo Gallo, M., Morone de Lucia, M. R., Vaccarella, R., and Grimaldi, P. 1982. Quattro nuove specie di Halechiniscidae rinvenute in due grotte sottomarine dell'Italia meridionale. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine 23: 415 - 426." type="journal article" year="1982">
Grimaldi de Zio
<emphasis id="D9ABEABEFFE3833746A8B7DB0305FC1B" box="[996,1039,914,938]" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 1982
</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374117B7D80239FC1B" author="JOrgensen, A. &amp; Boesgaard, T. M. &amp; MObjerg, N. &amp; Kristensen, R. M." box="[1115,1331,912,938]" pageId="3" pageNumber="212" pagination="401 - 443" refId="ref2786" refString="JOrgensen, A., Boesgaard, T. M., MObjerg, N., and Kristensen, R. M. 2014. The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada. Zootaxa 3802: 401 - 443." type="journal article" year="2014">
JØrgensen
<emphasis id="D9ABEABEFFE38337418BB7DB03F8FC1B" box="[1223,1266,914,938]" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 2014
</bibRefCitation>
) [see
<bibRefCitation id="8F4E4B5DFFE38337403DB7D80365FC7A" author="JOrgensen, A. &amp; Kristensen, R. M." pageId="3" pageNumber="212" pagination="425 - 439" refId="ref2836" refString="JOrgensen, A. and Kristensen, R. M. 2001. A new tanarctid arthrotardigrade with buoyant bodies. Zoologischer Anzeiger 240: 425 - 439." type="journal article" year="2001">JØrgensen and Kristensen (2001)</bibRefCitation>
for a discussion on the leg IV appendages]. However, the new species differs from
<taxonomicName id="2CDF4D2FFFE383374678B7BC04D7FBBA" authorityName="Lindgren" authorityYear="1971" box="[820,989,1011,1035]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="arborspinosus">
<emphasis id="D9ABEABEFFE383374678B7BC0449FBBA" box="[820,835,1012,1035]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE383374618B7BB04D7FBBA" box="[852,989,1011,1035]" italics="true" pageId="3" pageNumber="212">arborspinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="2CDF4D2FFFE383374154B7BC03ACFBBA" authorityName="Renaud-Mornant" authorityYear="1980" box="[1048,1190,1011,1035]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="dendriticus">
<emphasis id="D9ABEABEFFE383374154B7BC032DFBBA" box="[1048,1063,1012,1035]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337417BB7BB03ACFBBA" box="[1079,1190,1011,1035]" italics="true" pageId="3" pageNumber="212">dendriticus</emphasis>
</taxonomicName>
in the lack of tertiary branches (
<bibRefCitation id="8F4E4B5DFFE3833746EDB05C0334FB9D" author="Lindgren, E. W." box="[929,1086,1044,1068]" pageId="3" pageNumber="212" pagination="481 - 496" refId="ref2936" refString="Lindgren, E. W. 1971. Psammolittoral marine tardigrades from North Carolina and their conformity to worldwide zonation patterns. Cahiers de Biologie Marine 12: 481 - 496." type="journal article" year="1971">Lindgren 1971</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374101B05C0234FB9D" author="Renaud-Mornant, J." box="[1101,1342,1044,1068]" pageId="3" pageNumber="212" pagination="129 - 141" refId="ref3128" refString="Renaud-Mornant, J. 1980. Description de trois especies nouvelles du genre Tanarctus Renaud-Debyser, 1959, et creation de la sousfamille des Tanarctinae subfam. nov. (Tardigrada, Heterotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 2: 129 - 141." type="book chapter" year="1980">Renaud-Mornant 1980</bibRefCitation>
). Unlike the new species, the other two species have only short secondary branches (
<bibRefCitation id="8F4E4B5DFFE3833746A3B01D0204FBDC" author="Grimaldi de Zio, S. &amp; D'Addabbo Gallo, M. &amp; Morone de Lucia, M. R. &amp; Vaccarella, R. &amp; Grimaldi, P." box="[1007,1294,1109,1133]" pageId="3" pageNumber="212" pagination="415 - 426" refId="ref2723" refString="Grimaldi de Zio, S., D'Addabbo Gallo, M., Morone de Lucia, M. R., Vaccarella, R., and Grimaldi, P. 1982. Quattro nuove specie di Halechiniscidae rinvenute in due grotte sottomarine dell'Italia meridionale. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine 23: 415 - 426." type="journal article" year="1982">
Grimaldi de Zio
<emphasis id="D9ABEABEFFE3833741EDB01E03C7FBDC" box="[1185,1229,1109,1133]" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 1982
</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374056B01C0483FB3F" author="JOrgensen, A. &amp; Boesgaard, T. M. &amp; MObjerg, N. &amp; Kristensen, R. M." pageId="3" pageNumber="212" pagination="401 - 443" refId="ref2786" refString="JOrgensen, A., Boesgaard, T. M., MObjerg, N., and Kristensen, R. M. 2014. The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada. Zootaxa 3802: 401 - 443." type="journal article" year="2014">
JØrgensen
<emphasis id="D9ABEABEFFE3833740CAB01E044CFB3F" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 2014
</bibRefCitation>
). The new species could also be distinguished from
<taxonomicName id="2CDF4D2FFFE383374678B0DF04D6FB1F" authorityName="Lindgren" authorityYear="1971" box="[820,988,1174,1198]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="arborspinosus">
<emphasis id="D9ABEABEFFE383374678B0DF0449FB1F" box="[820,835,1175,1198]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337461FB0DE04D6FB1F" box="[851,988,1174,1198]" italics="true" pageId="3" pageNumber="212">arborspinosus</emphasis>
</taxonomicName>
,
<taxonomicName id="2CDF4D2FFFE3833746A0B0DF03ABFB1F" authorityName="Jorgensen, Boesgaard, Mobjerg &amp; Kristensen" authorityYear="2014" box="[1004,1185,1174,1198]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="hirsutospinosus">
<emphasis id="D9ABEABEFFE3833746A0B0DF04F1FB1F" box="[1004,1019,1175,1198]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE383374146B0DE03ABFB1F" box="[1034,1185,1174,1198]" italics="true" pageId="3" pageNumber="212">hirsutospinosus</emphasis>
</taxonomicName>
, and
<taxonomicName id="2CDF4D2FFFE3833741ADB0DF027FFB1F" authorityName="Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi" authorityYear="1982" box="[1249,1397,1174,1198]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="longisetosus">
<emphasis id="D9ABEABEFFE3833741ADB0DF03FAFB1F" box="[1249,1264,1175,1198]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337404CB0DE027FFB1F" box="[1280,1397,1174,1198]" italics="true" pageId="3" pageNumber="212">longisetosus</emphasis>
</taxonomicName>
by its smooth primary clavae (unknown in
<taxonomicName id="2CDF4D2FFFE3833741B0B0FF0285FB7E" authorityName="Renaud-Mornant" authorityYear="1980" box="[1276,1423,1207,1231]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="dendriticus">
<emphasis id="D9ABEABEFFE3833741B0B0FF0201FB7F" box="[1276,1291,1207,1230]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337406CB0FF0285FB7E" box="[1312,1423,1207,1231]" italics="true" pageId="3" pageNumber="212">dendriticus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="8F4E4B5DFFE383374677B09F04DEFB5E" author="Lindgren, E. W." box="[827,980,1239,1263]" pageId="3" pageNumber="212" pagination="481 - 496" refId="ref2936" refString="Lindgren, E. W. 1971. Psammolittoral marine tardigrades from North Carolina and their conformity to worldwide zonation patterns. Cahiers de Biologie Marine 12: 481 - 496." type="journal article" year="1971">Lindgren 1971</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE3833746ACB09F03C6FB5E" author="Renaud-Mornant, J." box="[992,1228,1239,1263]" pageId="3" pageNumber="212" pagination="129 - 141" refId="ref3128" refString="Renaud-Mornant, J. 1980. Description de trois especies nouvelles du genre Tanarctus Renaud-Debyser, 1959, et creation de la sousfamille des Tanarctinae subfam. nov. (Tardigrada, Heterotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 2: 129 - 141." type="book chapter" year="1980">Renaud-Mornant 1980</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE38337419BB09F0483FAA1" author="Grimaldi de Zio, S. &amp; D'Addabbo Gallo, M. &amp; Morone de Lucia, M. R. &amp; Vaccarella, R. &amp; Grimaldi, P." pageId="3" pageNumber="212" pagination="415 - 426" refId="ref2723" refString="Grimaldi de Zio, S., D'Addabbo Gallo, M., Morone de Lucia, M. R., Vaccarella, R., and Grimaldi, P. 1982. Quattro nuove specie di Halechiniscidae rinvenute in due grotte sottomarine dell'Italia meridionale. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine 23: 415 - 426." type="journal article" year="1982">
Grimaldi de Zio
<emphasis id="D9ABEABEFFE3833740CAB090044CFAA1" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 1982
</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE3833746D9B0BE037EFAA1" author="JOrgensen, A. &amp; Boesgaard, T. M. &amp; MObjerg, N. &amp; Kristensen, R. M." box="[917,1140,1270,1296]" pageId="3" pageNumber="212" pagination="401 - 443" refId="ref2786" refString="JOrgensen, A., Boesgaard, T. M., MObjerg, N., and Kristensen, R. M. 2014. The tardigrade fauna of Australian marine caves: With descriptions of nine new species of Arthrotardigrada. Zootaxa 3802: 401 - 443." type="journal article" year="2014">
JØrgensen
<emphasis id="D9ABEABEFFE38337414EB0B00325FAA1" box="[1026,1071,1272,1296]" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 2014
</bibRefCitation>
). Besides the prominent leg IV appendage and the primary clava used above for species identification, it is noteworthy that the large, flat secondary clavae similar to that of the new species have been report- ed in
<taxonomicName id="2CDF4D2FFFE383374638B13303F1FA23" authority="Renaud-Mornant, 1984" authorityName="Renaud-Mornant" authorityYear="1984" box="[884,1275,1402,1426]" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="helleouetae">
<emphasis id="D9ABEABEFFE383374638B1330489FA23" box="[884,899,1403,1426]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE3833746DFB132030BFA23" box="[915,1025,1402,1426]" italics="true" pageId="3" pageNumber="212">helleouetae</emphasis>
<bibRefCitation id="8F4E4B5DFFE383374146B13203F1FA23" author="Renaud-Mornant, J." box="[1034,1275,1402,1426]" pageId="3" pageNumber="212" pagination="975 - 988" refId="ref3181" refString="Renaud-Mornant, J. 1984. Nouveaux Arthrotardigrades des Antilles. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 6: 975 - 988." type="book chapter" year="1984">Renaud-Mornant, 1984</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="2CDF4D2FFFE383374078B13303F7FA03" authority="McKirdy, Schmidt and McGinty-Bayly, 1976" authorityName="McKirdy, Schmidt and McGinty-Bayly" authorityYear="1976" class="Heterotardigrada" family="Halechiniscidae" genus="Tanarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="velatus">
<emphasis id="D9ABEABEFFE383374078B1330249FA23" box="[1332,1347,1403,1426]" italics="true" pageId="3" pageNumber="212">T</emphasis>
.
<emphasis id="D9ABEABEFFE38337401FB1320292FA23" box="[1363,1432,1402,1426]" italics="true" pageId="3" pageNumber="212">velatus</emphasis>
<bibRefCitation id="8F4E4B5DFFE383374678B1D203F7FA03" author="McKirdy, D. J. &amp; Schmidt, P. &amp; McGinty-Bayly, M." box="[820,1277,1434,1458]" pageId="3" pageNumber="212" pagination="409 - 449" refId="ref2968" refString="McKirdy, D. J., Schmidt, P., and McGinty-Bayly, M. 1976. Interstitielle Fauna von Galapagos XVI. Tardigrada. Mikrofauna des Meeres- bodens 58: 409 - 449." type="journal article" year="1976">McKirdy, Schmidt and McGinty-Bayly, 1976</bibRefCitation>
</taxonomicName>
, and also from confamilial
<taxonomicName id="2CDF4D2FFFE3833746FDB1F3029EFA62" authority="Renaud-Mornant, 1979" authorityName="Renaud-Mornant" authorityYear="1979" box="[945,1428,1467,1491]" class="Heterotardigrada" family="Halechiniscidae" genus="Actinarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="lyrophorus">
<emphasis id="D9ABEABEFFE3833746FDB1F3039CFA62" box="[945,1174,1467,1491]" italics="true" pageId="3" pageNumber="212">Actinarctus lyrophorus</emphasis>
<bibRefCitation id="8F4E4B5DFFE3833741ECB1F3029EFA62" author="Renaud-Mornant, J." box="[1184,1428,1467,1491]" pageId="3" pageNumber="212" pagination="257 - 277" refId="ref3093" refString="Renaud-Mornant, J. 1979. Tardigrades marins de Madagascar. I. Halechiniscidae et Batillipedidae. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 1: 257 - 277." type="book chapter" year="1979">Renaud-Mornant, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2CDF4D2FFFE383374678B19403BFF9A5" authority="Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi, 1982" authorityName="Grimaldi de Zio, D'Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi" authorityYear="1982" class="Heterotardigrada" family="Halechiniscidae" genus="Actinarctus" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="neretinus">
<emphasis id="D9ABEABEFFE383374678B194044FFA42" box="[820,837,1500,1523]" italics="true" pageId="3" pageNumber="212">A</emphasis>
.
<emphasis id="D9ABEABEFFE38337461EB19404A4FA42" box="[850,942,1500,1523]" italics="true" pageId="3" pageNumber="212">neretinus</emphasis>
<bibRefCitation id="8F4E4B5DFFE3833746F9B19303BFF9A5" author="Grimaldi de Zio, S. &amp; D'Addabbo Gallo, M. &amp; Morone de Lucia, M. R. &amp; Vaccarella, R. &amp; Grimaldi, P." pageId="3" pageNumber="212" pagination="415 - 426" refId="ref2723" refString="Grimaldi de Zio, S., D'Addabbo Gallo, M., Morone de Lucia, M. R., Vaccarella, R., and Grimaldi, P. 1982. Quattro nuove specie di Halechiniscidae rinvenute in due grotte sottomarine dell'Italia meridionale. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine 23: 415 - 426." type="journal article" year="1982">Grimaldi de Zio, D’Addabbo Gallo, Morone De Lucia, Vaccarella and Grimaldi, 1982</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="2CDF4D2FFFE3833741A3B1B40329F984" authority="Renaud-Mornant, 1987" authorityName="Renaud-Mornant" authorityYear="1987" class="Heterotardigrada" family="Halechiniscidae" genus="Zioella" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="212" phylum="Tardigrada" rank="species" species="pavonina">
<emphasis id="D9ABEABEFFE3833741A3B1B4029DF9A5" box="[1263,1431,1532,1556]" italics="true" pageId="3" pageNumber="212">Zioella pavonina</emphasis>
<bibRefCitation id="8F4E4B5DFFE383374678B2540329F984" author="Renaud-Mornant, J." box="[820,1059,1564,1589]" pageId="3" pageNumber="212" pagination="353 - 373" refId="ref3210" refString="Renaud-Mornant, J. 1987. Halechiniscidae nouveaux de sables coralliens tropicaux (Tardigrada, Arthrotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 9: 353 - 373." type="book chapter" year="1987">Renaud-Mornant, 1987</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="8F4E4B5DFFE38337417DB254020BF985" author="McKirdy, D. J. &amp; Schmidt, P. &amp; McGinty-Bayly, M." box="[1073,1281,1564,1588]" pageId="3" pageNumber="212" pagination="409 - 449" refId="ref2968" refString="McKirdy, D. J., Schmidt, P., and McGinty-Bayly, M. 1976. Interstitielle Fauna von Galapagos XVI. Tardigrada. Mikrofauna des Meeres- bodens 58: 409 - 449." type="journal article" year="1976">
McKirdy
<emphasis id="D9ABEABEFFE3833741D9B25503CAF985" box="[1173,1216,1564,1588]" italics="true" pageId="3" pageNumber="212">et al</emphasis>
. 1976
</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374040B2540491F9E4" author="Renaud-Mornant, J." pageId="3" pageNumber="212" pagination="257 - 277" refId="ref3093" refString="Renaud-Mornant, J. 1979. Tardigrades marins de Madagascar. I. Halechiniscidae et Batillipedidae. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 1: 257 - 277." type="book chapter" year="1979">Renaud-Mornant 1979</bibRefCitation>
,
<bibRefCitation id="8F4E4B5DFFE3833746E8B27504DDF9E4" author="Renaud-Mornant, J." box="[932,983,1597,1621]" pageId="3" pageNumber="212" pagination="975 - 988" refId="ref3181" refString="Renaud-Mornant, J. 1984. Nouveaux Arthrotardigrades des Antilles. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 6: 975 - 988." type="book chapter" year="1984">1984</bibRefCitation>
,
<bibRefCitation id="8F4E4B5DFFE3833746ACB2750319F9E4" author="Renaud-Mornant, J." box="[992,1043,1597,1621]" pageId="3" pageNumber="212" pagination="353 - 373" refId="ref3210" refString="Renaud-Mornant, J. 1987. Halechiniscidae nouveaux de sables coralliens tropicaux (Tardigrada, Arthrotardigrada). Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e 9: 353 - 373." type="book chapter" year="1987">1987</bibRefCitation>
;
<bibRefCitation id="8F4E4B5DFFE383374150B2750254F9E4" author="Boesgaard, T. M. &amp; Kristensen, R. M." box="[1052,1374,1597,1621]" pageId="3" pageNumber="212" pagination="253 - 264" refId="ref2634" refString="Boesgaard, T. M. and Kristensen, R. M. 2001. Tardigrades from Australian marine caves. With a redescription of Actinarctus neretinus (Arthrotardigrada). Zoologischer Anzeiger 240: 253 - 264." type="journal article" year="2001">Boesgaard and Kristensen 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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