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<document id="9F0D6E90FBE0B1B833122A130CA7240D" ID-CLB-Dataset="276843" ID-DOI="10.5852/ejt.2023.901.2317" ID-GBIF-Dataset="f82db014-3609-4e5e-a3d9-c90c9de565e3" ID-ISSN="2118-9773" ID-Zenodo-Dep="10088606" ID-ZooBank="FB29CFDF-0064-4C6B-9C8A-3673B9437837" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1699006741891" checkinUser="plazi" docAuthor="Dhadwal, Tarun &amp; Bharti, Himender" docDate="2023" docId="60418785FFAFA33BFDCAF9276E55FB01" docLanguage="en" docName="EJT.2023.901.1-51.pdf" docOrigin="European Journal of Taxonomy 901 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2317/10057" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Camponotus simoni Emery 1893" docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="9C78FFFDFFB5A324FF98FFBE6C3DFFB8" masterDocTitle="Two new species of the genus Camponotus Mayr, 1861 (Hymenoptera: Formicidae) with five new records from India" masterLastPageNumber="51" masterPageNumber="1" pageNumber="27" updateTime="1699700754509" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="52A6FF6D1B2161EEA513153170F0F4E0">Two new species of the genus Camponotus Mayr, 1861 (Hymenoptera: Formicidae) with five new records from India</mods:title>
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<mods:namePart id="A4F05FE532038676C03E89D7A4F04349">Dhadwal, Tarun</mods:namePart>
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<mods:affiliation id="89428FBA4E5D6EF1341CEF186533A633">Department of Zoology and Environmental Sciences, Punjabi University, Patiala, Punjab, India.</mods:affiliation>
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<mods:namePart id="64EDB60E872F0987E4C96D0DA8BD5BBC">Bharti, Himender</mods:namePart>
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<mods:affiliation id="70222D4CCF704874E519BCFA77C12213">Department of Zoology and Environmental Sciences, Punjabi University, Patiala, Punjab, India.</mods:affiliation>
<mods:nameIdentifier id="1FF032E37EEDE70F9191E348875760BF" type="email">himenderbharti@gmail.com</mods:nameIdentifier>
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<mods:title id="BE5F0CD1AAFD1A66143CA19C7DE8C9A2">European Journal of Taxonomy</mods:title>
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<mods:date id="F34F0D772465DC324F6AAFD2739DC007">2023</mods:date>
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<treatment id="60418785FFAFA33BFDCAF9276E55FB01" ID-GBIF-Taxon="217349317" LSID="urn:lsid:plazi:treatment:60418785FFAFA33BFDCAF9276E55FB01" httpUri="http://treatment.plazi.org/id/60418785FFAFA33BFDCAF9276E55FB01" lastPageId="31" lastPageNumber="32" pageId="26" pageNumber="27">
<subSubSection id="A0F26518FFAFA33EFDCAF9276FDFF90C" box="[594,994,1689,1716]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="E8573693FFAFA33EFDCAF9276FDFF90C" blockId="26.[594,994,1689,1751]" box="[594,994,1689,1716]" pageId="26" pageNumber="27">
<heading id="B31F81FFFFAFA33EFDCAF9276FDFF90C" box="[594,994,1689,1716]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="2FE84D10FFAFA33EFDCAF9276FDFF90C" ID-CoL="5X8L4" authority="Emery, 1893" authorityName="Emery" authorityYear="1893" box="[594,994,1689,1716]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="DA9CEA81FFAFA33EFDCAF9276F78F90B" bold="true" box="[594,837,1689,1715]" italics="true" pageId="26" pageNumber="27">Camponotus simoni</emphasis>
<bibRefCitation id="8C794B62FFAFA33EFCD4F9246FDFF90C" author="Emery C." box="[844,994,1690,1716]" pageId="26" pageNumber="27" pagination="239 - 258" refId="ref14541" refString="Emery C. 1893. Voyage de M. E. Simon a l'ile de Ceylan (janvier - fevrier 1892). Formicides. Annales de la Societe entomologique de France 62: 239 - 258." type="journal article" year="1893">Emery, 1893</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="A0F26518FFAFA33EFD40F9036F66F96F" box="[728,859,1725,1751]" pageId="26" pageNumber="27" type="description">
<paragraph id="E8573693FFAFA33EFD40F9036F66F96F" blockId="26.[594,994,1689,1751]" box="[728,859,1725,1751]" pageId="26" pageNumber="27">
<figureCitation id="70D32A16FFAFA33EFD40F9036F66F96F" box="[728,859,1725,1751]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="28.[189,232,1963,1989]" captionStartId-1="29.[189,232,1963,1989]" captionStartId-2="30.[189,232,1963,1989]" captionTargetBox-0="[426,1161,265,1922]" captionTargetBox-1="[416,1171,265,1915]" captionTargetBox-2="[316,1270,265,1915]" captionTargetId-0="figure-21@28.[426,1161,265,1922]" captionTargetId-1="figure-11@29.[416,1171,265,1915]" captionTargetId-2="figure-21@30.[316,1270,265,1915]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="Fig. 16. Camponotus simoni Emery, 1893, major worker (PUAC T123). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." captionText-1="Fig. 17. Camponotus simoni Emery, 1893, minor worker (PUAC T127). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." captionText-2="Fig. 18. Camponotus simoni Emery, 1893, gyne (PUAC T133). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.10088638" figureDoi-1="http://doi.org/10.5281/zenodo.10088640" figureDoi-2="http://doi.org/10.5281/zenodo.10088642" httpUri-0="https://zenodo.org/record/10088638/files/figure.png" httpUri-1="https://zenodo.org/record/10088640/files/figure.png" httpUri-2="https://zenodo.org/record/10088642/files/figure.png" pageId="26" pageNumber="27">Figs 16–18</figureCitation>
</paragraph>
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<subSubSection id="A0F26518FFAFA33EFF25F9416F7AF8C5" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="E8573693FFAFA33EFF25F9416EBAF8A1" blockId="26.[189,647,1790,1817]" box="[189,647,1790,1817]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="C8F811BDFFAFA33EFF25F9416EBAF8A1" box="[189,647,1790,1817]" pageId="26" pageNumber="27">
<treatmentCitation id="69491082FFAFA33EFF25F9416EBEF8A1" ID-DOI="http://doi.org/10.5281/zenodo.6293175" ID-Zenodo-Dep="6293175" author="Emery C." box="[189,643,1790,1817]" httpUri="http://treatment.plazi.org/id/A35B580488883275FF91DCA05053F9C8" page="250" pageId="26" pageNumber="27" year="1893">
<taxonomicName id="2FE84D10FFAFA33EFF25F9416EBEF8A1" ID-CoL="5X8L4" authority="Emery, 1893: 250" authorityName="Emery" authorityPageNumber="250" authorityYear="1893" box="[189,643,1790,1817]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="DA9CEA81FFAFA33EFF25F9416D9AF8A0" box="[189,423,1791,1817]" italics="true" pageId="26" pageNumber="27">Camponotus simoni</emphasis>
<bibRefCitation id="8C794B62FFAFA33EFE36F9416EBEF8A1" author="Emery C." box="[430,643,1790,1817]" pageId="26" pageNumber="27" pagination="239 - 258" refId="ref14541" refString="Emery C. 1893. Voyage de M. E. Simon a l'ile de Ceylan (janvier - fevrier 1892). Formicides. Annales de la Societe entomologique de France 62: 239 - 258." type="journal article" year="1893">Emery, 1893: 250</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E8573693FFAFA33EFF25F8FE6F6EF8E1" blockId="26.[189,851,1855,1918]" box="[189,851,1855,1882]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="C8F811BDFFAFA33EFF25F8FE6F6EF8E1" box="[189,851,1855,1882]" pageId="26" pageNumber="27">
<taxonomicName id="2FE84D10FFAFA33EFF25F8FE6E36F8E1" ID-CoL="5X8L4" baseAuthorityName="Emery" baseAuthorityYear="1893" box="[189,523,1855,1882]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="dorycus" subSpecies="simoni">
<emphasis id="DA9CEA81FFAFA33EFF25F8FE6E36F8E1" box="[189,523,1855,1882]" italics="true" pageId="26" pageNumber="27">Camponotus dorycus simoni</emphasis>
</taxonomicName>
–
<treatmentCitation id="69491082FFAFA33EFDB0F8816EDAF8E2" author="Forel A." box="[552,743,1855,1882]" page="288" pageId="26" pageNumber="27" year="1902">
<bibRefCitation id="8C794B62FFAFA33EFDB0F8816EDAF8E2" author="Forel A." box="[552,743,1855,1882]" pageId="26" pageNumber="27" pagination="284 - 296" refId="ref14849" refString="Forel A. 1902. Varietes myrmecologiques. Annales de la Societe entomologique de Belgique 46: 284 - 296." type="journal article" year="1902">Forel 1902: 288</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="69491082FFAFA33EFD6BF8FE6F72F8E1" author="Forel A." box="[755,847,1855,1882]" page="6" pageId="26" pageNumber="27" year="1908">
<bibRefCitation id="8C794B62FFAFA33EFD6BF8FE6F72F8E1" author="Forel A." box="[755,847,1855,1882]" pageId="26" pageNumber="27" pagination="1 - 22" refId="ref14870" refString="Forel A. 1908. Fourmis de Ceylan et d'Egypte recoltees par le Prof. E. Bugnion. Lasius carniolicus. Fourmis de Kerguelen. Pseudandrie? Strongylognathus testaceus. Bulletin de la Societe vaudoise des Sciences naturelles 44: 1 - 22." type="journal article" year="1908">1908: 6</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E8573693FFAFA33EFF25F8DD6F7AF8C5" blockId="26.[189,851,1855,1918]" box="[189,839,1891,1918]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="C8F811BDFFAFA33EFF25F8DD6F7AF8C5" box="[189,839,1891,1918]" pageId="26" pageNumber="27">
<taxonomicName id="2FE84D10FFAFA33EFF25F8DD6E58F8C5" ID-CoL="5X8L4" authorityName="Emery" authorityYear="1893" box="[189,613,1891,1917]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="simoni" subGenus="Tanaemyrmex">
<emphasis id="DA9CEA81FFAFA33EFF25F8DD6D6CF8C5" box="[189,337,1891,1917]" italics="true" pageId="26" pageNumber="27">Camponotus</emphasis>
(
<emphasis id="DA9CEA81FFAFA33EFEFAF8DA6E38F8C5" box="[354,517,1892,1917]" italics="true" pageId="26" pageNumber="27">Tanaemyrmex</emphasis>
)
<emphasis id="DA9CEA81FFAFA33EFD8FF8DA6E58F8C5" box="[535,613,1892,1917]" italics="true" pageId="26" pageNumber="27">simoni</emphasis>
</taxonomicName>
–
<treatmentCitation id="69491082FFAFA33EFD1BF8DD6F7EF8C5" author="Emery C." box="[643,835,1891,1918]" page="90" pageId="26" pageNumber="27" year="1925">
<bibRefCitation id="8C794B62FFAFA33EFD1BF8DD6F7EF8C5" author="Emery C." box="[643,835,1891,1918]" pageId="26" pageNumber="27" pagination="1 - 302" refId="ref14704" refString="Emery C. 1925. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183: 1 - 302. Available from https: // www. biodiversitylibrary. org / page / 53206942 [accessed 31 Jul. 2022]." type="journal article" year="1925">Emery 1925: 90</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAEA33FFF25FEB66DF1FDFF" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="E8573693FFAEA33FFF25FEB66D0BFE9A" blockId="27.[189,310,264,290]" box="[189,310,264,290]" pageId="27" pageNumber="28">
<heading id="B31F81FFFFAEA33FFF25FEB66D0BFE9A" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="DA9CEA81FFAEA33FFF25FEB66D0BFE9A" bold="true" box="[189,310,264,290]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FE896DF1FDFF" blockId="27.[187,1400,311,584]" pageId="27" pageNumber="28">
This species resembles
<taxonomicName id="2FE84D10FFAEA33FFE4AFE896EEAFEEA" authority="(Smith, 1858)" baseAuthorityName="F. Smith" baseAuthorityYear="1858" box="[466,727,311,338]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="DA9CEA81FFAEA33FFE4AFE896E10FEE9" box="[466,557,311,337]" italics="true" pageId="27" pageNumber="28">C. mitis</emphasis>
(Smith, 1858)
</taxonomicName>
(major worker) but can be distinguished by its elongated head; the clypeal margin is shaped as a short lobe anteriorly truncate; the masticatory margin of mandibles with 6 teeth; the head and gaster are dark reddish-brown, with yellowish markings on the dorsal surface of the gaster; mesosoma, antennal flagella and legs are yellowish brown. While in
<taxonomicName id="2FE84D10FFAEA33FFBC5FE1F688AFE02" baseAuthorityName="F. Smith" baseAuthorityYear="1858" box="[1117,1207,417,443]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitis">
<emphasis id="DA9CEA81FFAEA33FFBC5FE1F688AFE02" box="[1117,1207,417,443]" italics="true" pageId="27" pageNumber="28">C. mitis</emphasis>
</taxonomicName>
(major worker), the head is subtriangular in shape; the clypeal margin is shaped as a short lobe anteriorly transverse; the masticatory margin of mandibles bears 7 teeth; the head and scape are dark brown to black; the mesosoma, petiolar and gastral colouration is variable from yellow-brown to dark brown; the tibiae and tarsi are usually darker.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAEA33FFF25FDD3680DFD60" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="E8573693FFAEA33FFF25FDD36D97FD3F" blockId="27.[189,426,621,647]" box="[189,426,621,647]" pageId="27" pageNumber="28">
<heading id="B31F81FFFFAEA33FFF25FDD36D97FD3F" bold="true" box="[189,426,621,647]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="DA9CEA81FFAEA33FFF25FDD36D97FD3F" bold="true" box="[189,426,621,647]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FD25680DFD60" blockId="27.[189,1399,667,728]" pageId="27" pageNumber="28">
<materialsCitation id="58803CCEFFAEA33FFF25FD256810FD60" ID-GBIF-Occurrence="4436650303" collectingDate="2017-01-30" collectionCode="T, PUAC" collectorName="Dhadwal" country="India" elevation="600" latitude="10.3834" location="Parambikulum National Park" longLatPrecision="7" longitude="77.0831" pageId="27" pageNumber="28" specimenCode="PUAC T121, PUAC T122, PUAC T123, PUAC T124, PUAC T125, PUAC T126, PUAC T127, PUAC T128, PUAC T129, PUAC T130, PUAC T131, PUAC T132, PUAC T133, PUAC T134, PUAC T135" specimenCount="15" specimenCount-female="1" specimenCount-worker="14" stateProvince="Kerala">
<collectingCountry id="90FF7603FFAEA33FFF25FD256D2DFD0D" box="[189,272,667,693]" name="India" pageId="27" pageNumber="28">INDIA</collectingCountry>
•
<specimenCount id="FEEEFD1AFFAEA33FFEB0FD256D8DFD0D" box="[296,432,667,693]" count="14" pageId="27" pageNumber="28" type="worker">14 workers</specimenCount>
,
<specimenCount id="FEEEFD1AFFAEA33FFE23FD256DD4FD0E" box="[443,489,667,694]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
;
<collectingRegion id="2A2CF871FFAEA33FFE60FD256E74FD0D" box="[504,585,667,693]" country="India" name="Kerala" pageId="27" pageNumber="28">Kerala</collectingRegion>
,
<location id="ED376048FFAEA33FFDCDFD256F8CFD0D" LSID="urn:lsid:plazi:treatment:60418785FFAFA33BFDCAF9276E55FB01:ED376048FFAEA33FFDCDFD256F8CFD0D" box="[597,945,667,693]" country="India" latitude="10.3834" longLatPrecision="7" longitude="77.0831" name="Parambikulum National Park" pageId="27" pageNumber="28" stateProvince="Kerala">Parambikulum National Park</location>
;
<geoCoordinate id="8DDC5054FFAEA33FFC25FD25687AFD0E" box="[957,1095,667,694]" degrees="10.3834" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="5" value="10.3834">10.3834° N</geoCoordinate>
,
<geoCoordinate id="8DDC5054FFAEA33FFBCCFD2568E5FD0E" box="[1108,1240,667,694]" degrees="77.0831" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="5" value="77.0831">77.0831° E</geoCoordinate>
; elev.
<quantity id="2F109B76FFAEA33FFABCFD25694CFD0E" box="[1316,1393,667,694]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="m" value="600.0">
<elevation id="63C5D1A0FFAEA33FFABCFD25694CFD0E" box="[1316,1393,667,694]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="27" pageNumber="28" unit="m" value="600.0">600 m</elevation>
</quantity>
;
<date id="9C561053FFAEA33FFF26FD006D6BFD60" box="[190,342,702,728]" pageId="27" pageNumber="28" value="2017-01-30">
<collectingDate id="8C12E9BBFFAEA33FFF26FD006D6BFD60" box="[190,342,702,728]" pageId="27" pageNumber="28" value="2017-01-30">30 Jan. 2017</collectingDate>
</date>
;
<collectionCode id="8EF9AE56FFAEA33FFEFAFD006D48FD60" box="[354,373,702,728]" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="28">T</collectionCode>
.
<collectorName id="451D5345FFAEA33FFE18FD036DD7FD6F" box="[384,490,701,727]" pageId="27" pageNumber="28">Dhadwal</collectorName>
leg.; hand picking method;
<collectionCode id="8EF9AE56FFAEA33FFCABFD006FBCFD60" box="[819,897,702,728]" pageId="27" pageNumber="28">PUAC</collectionCode>
<specimenCode id="B84E9EE8FFAEA33FFC10FD006FFBFD60" box="[904,966,702,728]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="28">T121</specimenCode>
to
<specimenCode id="B84E9EE8FFAEA33FFC73FD036810FD60" box="[1003,1069,701,728]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="27" pageNumber="28">T135</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAEA33BFF25FD406D5AFD07" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28" type="description">
<paragraph id="E8573693FFAEA33FFF25FD406D48FCA0" blockId="27.[189,373,766,792]" box="[189,373,766,792]" pageId="27" pageNumber="28">
<heading id="B31F81FFFFAEA33FFF25FD406D48FCA0" bold="true" box="[189,373,766,792]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="DA9CEA81FFAEA33FFF25FD406D48FCA0" bold="true" box="[189,373,766,792]" pageId="27" pageNumber="28">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAEA33FFF72FC956DCEFCFD" blockId="27.[189,1398,811,940]" box="[234,499,811,838]" pageId="27" pageNumber="28">
<emphasis id="DA9CEA81FFAEA33FFF72FC956DA1FCFD" bold="true" box="[234,412,811,837]" pageId="27" pageNumber="28">Major worker</emphasis>
(n = 7)
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FCF06E36FC14" blockId="27.[189,1398,811,940]" pageId="27" pageNumber="28">
HL 2.52–2.70; HW 2.05–2.13; EL 0.45–0.49; SL 2.74–2.87; PW 1.43–1.47; WL 3.07–3.19; MTL 2.13– 2.46; HTL 3.09–3.19; PL 0.65–0.69;
<collectingCountry id="90FF7603FFAEA33FFDE2FCCE6EA2FC32" box="[634,671,880,906]" name="Philippines" pageId="27" pageNumber="28">PH</collectingCountry>
0.63–0.65; GL 2.46–2.58; TL 8.70–9.16; CI 78–81; SI 133– 134; REL 17–18; PrI 69–70.
</paragraph>
<paragraph id="E8573693FFAEA33FFF72FC6C6DCEFC54" blockId="27.[189,1399,978,1107]" box="[234,499,978,1005]" pageId="27" pageNumber="28">
<emphasis id="DA9CEA81FFAEA33FFF72FC6C6DA1FC54" bold="true" box="[234,412,978,1004]" pageId="27" pageNumber="28">Minor worker</emphasis>
(n = 7)
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FC4B6DC1FBEB" blockId="27.[189,1399,978,1107]" pageId="27" pageNumber="28">
HL 1.76–1.92; HW 0.77–0.94; EL 0.41–0.45; SL 3.07–3.15; PW 1.21–1.33; WL 2.95–3.11; MTL 2.21– 2.58; HTL 3.23–3.29; PL 0.57–0.61;
<collectingCountry id="90FF7603FFAEA33FFDE2FBA96EA2FB89" box="[634,671,1047,1073]" name="Philippines" pageId="27" pageNumber="28">PH</collectingCountry>
0.61–0.65; GL 2.13–2.27; TL 7.41–7.91; CI 43–48; SI 335– 398; REL 23; PrI 141–157.
</paragraph>
<paragraph id="E8573693FFAEA33FFF72FBC66DB8FB2B" blockId="27.[191,1399,1144,1240]" box="[234,389,1144,1171]" pageId="27" pageNumber="28">
<emphasis id="DA9CEA81FFAEA33FFF72FBC66D10FB2A" bold="true" box="[234,301,1144,1170]" pageId="27" pageNumber="28">Gyne</emphasis>
(n = 1)
</paragraph>
<paragraph id="E8573693FFAEA33FFF5FFB256E75FB6F" blockId="27.[191,1399,1144,1240]" pageId="27" pageNumber="28">
HL 2.79; HW 1.98; EL 0.68; SL 4.15; WL 4.46; MTL 2.41; HTL 3.03; PL 0.80;
<collectingCountry id="90FF7603FFAEA33FFB0FFB256881FB0D" box="[1175,1212,1179,1205]" name="Philippines" pageId="27" pageNumber="28">PH</collectingCountry>
1.05; GL 3.34; TL 11.39; CI 70; SI 209; REL 24.
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FB436D73FAAF" blockId="27.[189,334,1277,1303]" box="[189,334,1277,1303]" pageId="27" pageNumber="28">
<heading id="B31F81FFFFAEA33FFF25FB436D73FAAF" bold="true" box="[189,334,1277,1303]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="DA9CEA81FFAEA33FFF25FB436D73FAAF" bold="true" box="[189,334,1277,1303]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAEA33FFF72FA956E37FAFD" blockId="27.[188,1399,1323,1691]" box="[234,522,1323,1349]" pageId="27" pageNumber="28">
<emphasis id="DA9CEA81FFAEA33FFF72FA956DA1FAFD" bold="true" box="[234,412,1323,1349]" pageId="27" pageNumber="28">Major worker</emphasis>
(
<figureCitation id="70D32A16FFAEA33FFE34FA956E3CFAFD" box="[428,513,1323,1349]" captionStart="Fig" captionStartId="28.[189,232,1963,1989]" captionTargetBox="[426,1161,265,1922]" captionTargetId="figure-21@28.[426,1161,265,1922]" captionTargetPageId="28" captionText="Fig. 16. Camponotus simoni Emery, 1893, major worker (PUAC T123). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088638" httpUri="https://zenodo.org/record/10088638/files/figure.png" pageId="27" pageNumber="28">Fig. 16</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFAEA33FFF25FAF06ED8F922" blockId="27.[188,1399,1323,1691]" pageId="27" pageNumber="28">
<smallCapsWord id="EEB1A04FFFAEA33FFF25FAF06D20FAD0" baselines="1377,1377" box="[189,285,1358,1384]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="27" pageNumber="28">HABITUS</smallCapsWord>
. In full-face view, head elongated, longer than broad (CI 78–81), posterior margin emarginated in the middle, occipital corners round, lateral margins convex anteriorly; clypeus carinate in the middle, clypeal margin produced as a short lobe anteriorly truncate; mandibles moderately broad with 6 teeth; eyes moderate in size, placed in front over the mid-length of the head; antennae long, slender and 12-segmented, scape long (SI 133–134), surpassing the posterior margin of head by ¹/5 of its length. In dorsal view, mesosoma elongated, pronotum anteriorly narrow (PrI 69–70) pro-mesonotal suture and metanotal groove distinct; mesonotum and propodeum compressed laterally; mesosoma strongly convex in lateral view; propodeal declivity slightly concave, propodeal spiracle round placed below the margin of propodeal declivity; petiole node thick, convex anteriorly and flat posteriorly, tapering towards the tip; tibiae compressed; gaster oval.
</paragraph>
<paragraph id="E8573693FFAEA33FFF25F97F6D56F945" blockId="27.[189,1399,1728,1789]" pageId="27" pageNumber="28">
<smallCapsWord id="EEB1A04FFFAEA33FFF25F97F6D05F963" baselines="1748,1748" box="[189,312,1729,1755]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Sculpture" pageId="27" pageNumber="28">SCULPTURE</smallCapsWord>
. Head, mesosoma and gaster minutely reticulated and matte; mandibles smooth and shiny with scattered.
</paragraph>
<paragraph id="E8573693FFAEA33FFF25F89A68A6F83A" blockId="27.[188,1399,1827,1922]" pageId="27" pageNumber="28">
<smallCapsWord id="EEB1A04FFFAEA33FFF25F89A6D23F886" baselines="1847,1847" box="[189,286,1828,1854]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pilosity" pageId="27" pageNumber="28">PILOSITY</smallCapsWord>
<smallCapsWord id="EEB1A04FFFAEA33FFEB4F8976D64F884" baselines="1847" box="[300,345,1833,1852]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="27" pageNumber="28">AND</smallCapsWord>
<smallCapsWord id="EEB1A04FFFAEA33FFEFEF8976DD6F884" baselines="1847" box="[358,491,1833,1852]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pubescence" pageId="27" pageNumber="28">PUBESCENCE</smallCapsWord>
. Body pilose, long erect abundant yellowish hair on the vertex of head, mesosoma and gaster, a few short hairs present on anterior of head and clypeus; hind tibia without a row of spiny bristles on the ventral margin, but with 3–4 erect setae close to apical spurs.
</paragraph>
<paragraph id="E8573693FFAEA33FFF25F8166F41F85C" blockId="27.[189,1399,1960,2020]" pageId="27" pageNumber="28">
<smallCapsWord id="EEB1A04FFFAEA33FFF25F8166D6AF87A" baselines="1979,1979" box="[189,343,1960,1986]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colouration" pageId="27" pageNumber="28">COLOURATION</smallCapsWord>
. Head and gaster dark reddish brown, with yellowish stripes on the dorsal side of the gaster; mesosoma, antennomeres and legs yellowish brown.
</paragraph>
<caption id="BC97661BFFA9A338FF25F8156EC8F850" ID-DOI="http://doi.org/10.5281/zenodo.10088638" ID-Zenodo-Dep="10088638" httpUri="https://zenodo.org/record/10088638/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1963,1989]" targetBox="[426,1161,265,1922]" targetPageId="28">
<paragraph id="E8573693FFA9A338FF25F8156EC8F850" blockId="28.[189,1399,1963,2025]" pageId="28" pageNumber="29">
<emphasis id="DA9CEA81FFA9A338FF25F8156D23F87D" bold="true" box="[189,286,1963,1989]" pageId="28" pageNumber="29">Fig. 16.</emphasis>
<taxonomicName id="2FE84D10FFA9A338FEB1F8126E80F87D" authority="Emery, 1893" authorityName="Emery" authorityYear="1893" box="[297,701,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="DA9CEA81FFA9A338FEB1F8126E2BF87D" box="[297,534,1964,1990]" italics="true" pageId="28" pageNumber="29">Camponotus simoni</emphasis>
<bibRefCitation id="8C794B62FFA9A338FDB9F8126E80F87D" author="Emery C." box="[545,701,1963,1990]" pageId="28" pageNumber="29" pagination="239 - 258" refId="ref14541" refString="Emery C. 1893. Voyage de M. E. Simon a l'ile de Ceylan (janvier - fevrier 1892). Formicides. Annales de la Societe entomologique de France 62: 239 - 258." type="journal article" year="1893">Emery, 1893</bibRefCitation>
</taxonomicName>
, major worker (PUAC T123).
<emphasis id="DA9CEA81FFA9A338FBAEF8156876F87D" bold="true" box="[1078,1099,1963,1989]" pageId="28" pageNumber="29">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA9A338FF25F8706CECF850" bold="true" box="[189,209,1998,2024]" pageId="28" pageNumber="29">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA9A338FE46F8706DCEF850" bold="true" box="[478,499,1998,2024]" pageId="28" pageNumber="29">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<caption id="BC97661BFFA8A339FF25F8156EC8F850" ID-DOI="http://doi.org/10.5281/zenodo.10088640" ID-Zenodo-Dep="10088640" httpUri="https://zenodo.org/record/10088640/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1963,1989]" targetBox="[416,1171,265,1915]" targetPageId="29">
<paragraph id="E8573693FFA8A339FF25F8156EC8F850" blockId="29.[189,1399,1963,2025]" pageId="29" pageNumber="30">
<emphasis id="DA9CEA81FFA8A339FF25F8156D23F87D" bold="true" box="[189,286,1963,1989]" pageId="29" pageNumber="30">Fig. 17.</emphasis>
<taxonomicName id="2FE84D10FFA8A339FEB1F8126E81F87D" authority="Emery, 1893" authorityName="Emery" authorityYear="1893" box="[297,700,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="DA9CEA81FFA8A339FEB1F8126E2BF87D" box="[297,534,1964,1990]" italics="true" pageId="29" pageNumber="30">Camponotus simoni</emphasis>
<bibRefCitation id="8C794B62FFA8A339FDB9F8126E81F87D" author="Emery C." box="[545,700,1963,1990]" pageId="29" pageNumber="30" pagination="239 - 258" refId="ref14541" refString="Emery C. 1893. Voyage de M. E. Simon a l'ile de Ceylan (janvier - fevrier 1892). Formicides. Annales de la Societe entomologique de France 62: 239 - 258." type="journal article" year="1893">Emery, 1893</bibRefCitation>
</taxonomicName>
, minor worker (PUAC T127).
<emphasis id="DA9CEA81FFA8A339FBAFF8156871F87D" bold="true" box="[1079,1100,1963,1989]" pageId="29" pageNumber="30">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFA8A339FF25F8706CECF850" bold="true" box="[189,209,1998,2024]" pageId="29" pageNumber="30">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFA8A339FE46F8706DCEF850" bold="true" box="[478,499,1998,2024]" pageId="29" pageNumber="30">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<caption id="BC97661BFFABA33AFF25F8156E53F850" ID-DOI="http://doi.org/10.5281/zenodo.10088642" ID-Zenodo-Dep="10088642" httpUri="https://zenodo.org/record/10088642/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1963,1989]" targetBox="[316,1270,265,1915]" targetPageId="30">
<paragraph id="E8573693FFABA33AFF25F8156E53F850" blockId="30.[189,1399,1963,2025]" pageId="30" pageNumber="31">
<emphasis id="DA9CEA81FFABA33AFF25F8156D26F87D" bold="true" box="[189,283,1963,1989]" pageId="30" pageNumber="31">Fig. 18.</emphasis>
<taxonomicName id="2FE84D10FFABA33AFEBCF8126E8FF87D" authority="Emery, 1893" authorityName="Emery" authorityYear="1893" box="[292,690,1963,1990]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="DA9CEA81FFABA33AFEBCF8126E32F87D" box="[292,527,1964,1990]" italics="true" pageId="30" pageNumber="31">Camponotus simoni</emphasis>
<bibRefCitation id="8C794B62FFABA33AFD81F8126E8FF87D" author="Emery C." box="[537,690,1963,1990]" pageId="30" pageNumber="31" pagination="239 - 258" refId="ref14541" refString="Emery C. 1893. Voyage de M. E. Simon a l'ile de Ceylan (janvier - fevrier 1892). Formicides. Annales de la Societe entomologique de France 62: 239 - 258." type="journal article" year="1893">Emery, 1893</bibRefCitation>
</taxonomicName>
, gyne (PUAC T133).
<emphasis id="DA9CEA81FFABA33AFC21F8156FF0F87D" bold="true" box="[953,973,1963,1989]" pageId="30" pageNumber="31">A</emphasis>
. Head in full face view.
<emphasis id="DA9CEA81FFABA33AFB6DF815693AF87D" bold="true" box="[1269,1287,1963,1989]" pageId="30" pageNumber="31">B</emphasis>
. Body in profile view.
<emphasis id="DA9CEA81FFABA33AFECFF8706D51F850" bold="true" box="[343,364,1998,2024]" pageId="30" pageNumber="31">C</emphasis>
. Body in dorsal view.
</paragraph>
</caption>
<paragraph id="E8573693FFAAA33BFF72FEB76E37FE9B" blockId="31.[188,1399,265,465]" box="[234,522,265,291]" pageId="31" pageNumber="32">
<emphasis id="DA9CEA81FFAAA33BFF72FEB76DA1FE9B" bold="true" box="[234,412,265,291]" pageId="31" pageNumber="32">Minor worker</emphasis>
(
<figureCitation id="70D32A16FFAAA33BFE34FEB76E3CFE9B" box="[428,513,265,291]" captionStart="Fig" captionStartId="29.[189,232,1963,1989]" captionTargetBox="[416,1171,265,1915]" captionTargetId="figure-11@29.[416,1171,265,1915]" captionTargetPageId="29" captionText="Fig. 17. Camponotus simoni Emery, 1893, minor worker (PUAC T127). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088640" httpUri="https://zenodo.org/record/10088640/files/figure.png" pageId="31" pageNumber="32">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFAAA33BFF25FE92683DFE68" blockId="31.[188,1399,265,465]" pageId="31" pageNumber="32">All characteristics as of major worker except: in minor worker head is comparatively small (CI 43– 48) with round posterior margin and subparallel lateral margins converging anteriorly; clypeus margin anteriorly produced as a short round lobe; mandibles with 5 teeth; eyes moderate in size, placed laterally over the mid-length of the head; scape distinctly long (SI 335–398), surpassing posterior margin of head by half of its length; mesosoma short as compared to the major worker.</paragraph>
<paragraph id="E8573693FFAAA33BFF72FE496DA9FDA9" blockId="31.[189,1400,503,703]" box="[234,404,503,529]" pageId="31" pageNumber="32">
<emphasis id="DA9CEA81FFAAA33BFF72FE496D10FDA9" bold="true" box="[234,301,503,529]" pageId="31" pageNumber="32">Gyne</emphasis>
(
<figureCitation id="70D32A16FFAAA33BFEA5FE496DB6FDA9" box="[317,395,503,529]" captionStart="Fig" captionStartId="30.[189,232,1963,1989]" captionTargetBox="[316,1270,265,1915]" captionTargetId="figure-21@30.[316,1270,265,1915]" captionTargetPageId="30" captionText="Fig. 18. Camponotus simoni Emery, 1893, gyne (PUAC T133). A. Head in full face view. B. Body in profile view. C. Body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10088642" httpUri="https://zenodo.org/record/10088642/files/figure.png" pageId="31" pageNumber="32">Fig.18</figureCitation>
)
</paragraph>
<paragraph id="E8573693FFAAA33BFF25FDA46D5AFD07" blockId="31.[189,1400,503,703]" pageId="31" pageNumber="32">Similar to the major worker with few modifications indicating the caste and the following differences: head narrower (CI 70) with subparallel lateral margins, posterior margin straight; cephalic dorsum with 3 prominent ocelli; mandibles with 6 teeth; scapes surpassing the posterior margin of head by half of their length (SI 209); propodeal declivity smoothly convex; head, scutellum and gaster with erect or suberect hairs.</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAAA33BFF25FD586DD9FC96" pageId="31" pageNumber="32" type="distribution">
<paragraph id="E8573693FFAAA33BFF25FD586D90FCB8" blockId="31.[189,429,742,768]" box="[189,429,742,768]" pageId="31" pageNumber="32">
<heading id="B31F81FFFFAAA33BFF25FD586D90FCB8" bold="true" box="[189,429,742,768]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="DA9CEA81FFAAA33BFF25FD586D90FCB8" bold="true" box="[189,429,742,768]" pageId="31" pageNumber="32">Global distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAAA33BFF25FCAA6DD9FC96" blockId="31.[189,484,788,814]" box="[189,484,788,814]" pageId="31" pageNumber="32">
<collectingCountry id="90FF7603FFAAA33BFF25FCAA6D0DFC96" box="[189,304,788,814]" name="Sri Lanka" pageId="31" pageNumber="32">Sri Lanka</collectingCountry>
(
<typeStatus id="37538831FFAAA33BFEA7FCAA6D4EFC96" box="[319,371,788,814]" pageId="31" pageNumber="32">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAAA33BFF25FCEB6F24FB9F" pageId="31" pageNumber="32" type="biology_ecology">
<paragraph id="E8573693FFAAA33BFF25FCEB6D20FCD7" blockId="31.[189,285,853,879]" box="[189,285,853,879]" pageId="31" pageNumber="32">
<heading id="B31F81FFFFAAA33BFF25FCEB6D20FCD7" bold="true" box="[189,285,853,879]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="DA9CEA81FFAAA33BFF25FCEB6D20FCD7" bold="true" box="[189,285,853,879]" pageId="31" pageNumber="32">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAAA33BFF25FC3D6F24FB9F" blockId="31.[189,1399,899,1064]" pageId="31" pageNumber="32">
During the field survey, the species was collected from Karianchola (Parambikulam National Park),
<collectingRegion id="2A2CF871FFAAA33BFF25FC1B6D32FC07" box="[189,271,933,959]" country="India" name="Kerala" pageId="31" pageNumber="32">Kerala</collectingRegion>
. The nest was located in the deep forest. It was mound
<typeStatus id="37538831FFAAA33BFC2CFC186FDAFC78" box="[948,999,934,960]" pageId="31" pageNumber="32">type</typeStatus>
located above the ground shared by termites also. The association is not determined yet. On disturbing, the workers started coming out, otherwise, no foraging worker was observed. The region has an average daily temperature of 32°C and is comprised of intact tropical wet evergreen forest.
</paragraph>
</subSubSection>
<subSubSection id="A0F26518FFAAA33BFF25FBF16E55FB01" pageId="31" pageNumber="32" type="discussion">
<paragraph id="E8573693FFAAA33BFF25FBF16D01FBD1" blockId="31.[189,316,1103,1129]" box="[189,316,1103,1129]" pageId="31" pageNumber="32">
<heading id="B31F81FFFFAAA33BFF25FBF16D01FBD1" bold="true" box="[189,316,1103,1129]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="DA9CEA81FFAAA33BFF25FBF16D01FBD1" bold="true" box="[189,316,1103,1129]" pageId="31" pageNumber="32">Relevance</emphasis>
</heading>
</paragraph>
<paragraph id="E8573693FFAAA33BFF25FBC26E55FB01" blockId="31.[188,1399,1148,1209]" pageId="31" pageNumber="32">
This species represents a new record for
<collectingCountry id="90FF7603FFAAA33BFD0BFBC26EEEFB2E" box="[659,723,1148,1174]" name="India" pageId="31" pageNumber="32">India</collectingCountry>
. Formerly it was reported from
<collectingCountry id="90FF7603FFAAA33BFBDAFBC26885FB2E" box="[1090,1208,1148,1174]" name="Sri Lanka" pageId="31" pageNumber="32">Sri Lanka</collectingCountry>
. The gyne of the species is described for the first time
</paragraph>
</subSubSection>
</treatment>
</document>
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