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<document id="721B683C657B3767B877AC29EF7FFF19" ID-CLB-Dataset="31691" ID-DOI="10.11646/zootaxa.4341.2.1" ID-GBIF-Dataset="e1cd4661-3d65-4dce-bdbc-757b73e88fde" ID-ISSN="1175-5326" ID-Zenodo-Dep="1039411" ID-ZooBank="4070D499-15BE-4ED0-8FC6-7A52E070D053" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509434476181" checkinUser="plazi" docAuthor="Boyd, David A., Nithirojpakdee, Patchara, Deein, Gridsada, Vidthayanon, Chavalit, Grudpan, Chaiwut, Tangjitjaroen, Weerapongse, Pfeiffer, John M., Randall, Zachary S., Srisombat, Tippamas &amp; Page, Lawrence M." docDate="2017" docId="537A87CE835BFFA5FF04F9E6FC1FFF7F" docLanguage="en" docName="zootaxa.4341.2.1.pdf" docOrigin="Zootaxa 4341 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantopsis rungthipae Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page 2017, new species" docType="treatment" docVersion="7" lastPageNumber="176" masterDocId="AF43FFB6834DFFBCFF93FFF2FFE1FF87" masterDocTitle="Revision of the horseface loaches (Cobitidae, Acantopsis), with descriptions of three new species from Southeast Asia" masterLastPageNumber="192" masterPageNumber="151" pageNumber="173" updateTime="1698470517910" updateUser="ExternalLinkService">
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<mods:title id="CBA5BD4E9FAF2F39797CFEDEE98EC1A8">Revision of the horseface loaches (Cobitidae, Acantopsis), with descriptions of three new species from Southeast Asia</mods:title>
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<mods:namePart id="A7424B14FCE5C5BFE0A3918CF2375887">Boyd, David A.</mods:namePart>
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<mods:namePart id="D463DF39CF5FB5E59D20B96633AA28E6">Nithirojpakdee, Patchara</mods:namePart>
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</mods:role>
<mods:namePart id="25D82496B5A3FB8FFA64BB85D864FDAD">Deein, Gridsada</mods:namePart>
</mods:name>
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<mods:namePart id="271CA94C5C5839E5E2AFB589BEBE2EDB">Vidthayanon, Chavalit</mods:namePart>
</mods:name>
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<mods:namePart id="04CD435449085BF09CA280C2EDCF71BA">Grudpan, Chaiwut</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="961982022DE48A35B9CF725C234D4E1D">Tangjitjaroen, Weerapongse</mods:namePart>
</mods:name>
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<mods:namePart id="0733AC974685F549B64D7884FA72B896">Pfeiffer, John M.</mods:namePart>
</mods:name>
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<mods:namePart id="A4C6CDC53DAEF574C94C14992BDA51B5">Randall, Zachary S.</mods:namePart>
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</mods:role>
<mods:namePart id="48614C3B1C0DFF4166A1B24F666ACA3D">Srisombat, Tippamas</mods:namePart>
</mods:name>
<mods:name id="FDED4C0A7FE1445EC3D6CE120A26BBB1" type="personal">
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<mods:namePart id="E5F2902FBFFFED21F15DDBDE412DEC62">Page, Lawrence M.</mods:namePart>
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<mods:typeOfResource id="F20F5646C456CB06AE643DF059B2E5A2">text</mods:typeOfResource>
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<mods:title id="BA3135E85E501DFF1F5ECB2AC639CA01">Zootaxa</mods:title>
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<mods:part id="258FFCC1730DFAE49B9F8480FC7AFFB3">
<mods:date id="5203E18FCE46DD957FD0E49A82377454">2017</mods:date>
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<mods:number id="9A144D488BFEE8DC8DE0D7EC168F8FD9">2017-10-31</mods:number>
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<mods:number id="30E066CB17D4FC822F6648328093A235">4341</mods:number>
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<mods:identifier id="F56D559DD7CEBAD5F18C9A05A08DE500" type="DOI">10.11646/zootaxa.4341.2.1</mods:identifier>
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<treatment id="537A87CE835BFFA5FF04F9E6FC1FFF7F" ID-DOI="http://doi.org/10.5281/zenodo.6010738" ID-GBIF-Taxon="135489601" ID-Zenodo-Dep="6010738" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F" httpUri="http://treatment.plazi.org/id/537A87CE835BFFA5FF04F9E6FC1FFF7F" lastPageId="25" lastPageNumber="176" pageId="22" pageNumber="173">
<subSubSection id="93C96553835BFFAAFF04F9E6FE51F9C8" pageId="22" pageNumber="173" type="nomenclature">
<paragraph id="DB6C36D8835BFFAAFF04F9E6FC3DF9A9" blockId="22.[151,988,1556,1651]" box="[151,988,1556,1582]" pageId="22" pageNumber="173">
<heading id="802481B4835BFFAAFF04F9E6FC3DF9A9" bold="true" box="[151,988,1556,1582]" fontSize="11" level="1" pageId="22" pageNumber="173" reason="1">
<emphasis id="E9A7EACA835BFFAAFF04F9E6FC3DF9A9" bold="true" box="[151,988,1556,1582]" pageId="22" pageNumber="173">
<taxonomicName id="1CD34D5B835BFFAAFF04F9E6FE46F9A9" ID-CoL="64FRJ" authority="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page, 2017" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[151,423,1556,1582]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="173" phylum="Chordata" rank="species" species="rungthipae" status="sp. nov.">
<emphasis id="E9A7EACA835BFFAAFF04F9E6FE46F9A9" bold="true" box="[151,423,1556,1582]" italics="true" pageId="22" pageNumber="173">Acantopsis rungthipae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F29457B1835BFFAAFE3CF9E6FDA0F9A9" box="[431,577,1556,1582]" pageId="22" pageNumber="173" rank="species">new species</taxonomicNameLabel>
, Boyd, Nithirojpakdee, and Page
</emphasis>
</heading>
</paragraph>
<paragraph id="DB6C36D8835BFFAAFF04F9C4FE51F9C8" blockId="22.[151,988,1556,1651]" box="[151,432,1590,1615]" pageId="22" pageNumber="173">Piebald Horseface Loach</paragraph>
</subSubSection>
<subSubSection id="93C96553835BFFAAFF04F9A8FF18F9F4" box="[151,249,1626,1651]" pageId="22" pageNumber="173" type="description">
<paragraph id="DB6C36D8835BFFAAFF04F9A8FF18F9F4" blockId="22.[151,988,1556,1651]" box="[151,249,1626,1651]" pageId="22" pageNumber="173">
(
<figureCitation id="43E82A5D835BFFAAFF0CF9A8FF10F9F4" box="[159,241,1626,1651]" captionStart="FIGURE 14" captionStartId="23.[151,250,1702,1724]" captionTargetBox="[151,1436,179,1691]" captionTargetId="figure@23.[151,1436,179,1691]" captionTargetPageId="23" captionText="FIGURE 14. Acantopsis rungthipae. (A) Lateral and dorsal VieWs, THNHM F 0013610, holotype, Welu RiVer, 107.4 mm SL. LiVe VieWs of (B) UF 235976, paratype, Welu RiVer, 91.7 mm SL, (C) UF 188117, Wang RiVer, 56.4 mm SL, (D) UF 188114, Songkhram RiVer, 90.6 mm SL, and (E) UF 236085, Tapi RiVer, 77.3 mm SL." httpUri="https://zenodo.org/record/1039439/files/figure.png" pageId="22" pageNumber="173">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="93C96553835BFFAAFF04F950FE74F830" pageId="22" pageNumber="173" type="materials_examined">
<paragraph id="DB6C36D8835BFFAAFF04F950FD3EF885" blockId="22.[151,1437,1698,1975]" pageId="22" pageNumber="173">
<materialsCitation id="6BBB3C85835BFFAAFF04F950FD3EF885" ID-GBIF-Occurrence="1676480044" collectingDate="2014-01-30" collectionCode="THNHM" collectorName="Nithirojpakdee, J." country="Thailand" county="Welu River" latitude="12.537" location="Khen Creek" longLatPrecision="77" longitude="102.244" municipality="Khlung District" pageId="22" pageNumber="173" specimenCount="1" stateProvince="Chanthaburi Province" typeStatus="holotype">
<typeStatus id="0468887A835BFFAAFF04F950FEE5F93C" box="[151,260,1698,1723]" pageId="22" pageNumber="173" type="holotype">
<emphasis id="E9A7EACA835BFFAAFF04F950FEE5F93C" bold="true" box="[151,260,1698,1723]" pageId="22" pageNumber="173">Holotype</emphasis>
</typeStatus>
.
<collectionCode id="BDC2AE1D835BFFAAFE8FF951FE63F93D" box="[284,386,1699,1722]" pageId="22" pageNumber="173">THNHM</collectionCode>
F0013610,
<quantity id="1C2B9B3D835BFFAAFD8EF951FD78F93D" box="[541,665,1699,1723]" metricMagnitude="-1" metricUnit="m" metricValue="1.074" pageId="22" pageNumber="173" unit="mm" value="107.4">107.4 mm</quantity>
SL female (ex. UF 235976),
<collectingCountry id="A3C47648835BFFAAFB8CF950FB64F93C" box="[1055,1157,1698,1723]" name="Thailand" pageId="22" pageNumber="173">Thailand</collectingCountry>
,
<collectingCounty id="320D4E54835BFFAAFB09F950FAC2F93C" box="[1178,1315,1698,1723]" pageId="22" pageNumber="173">Welu River</collectingCounty>
drainage,
<collectingRegion id="1917F83A835BFFAAFF04F934FE77F958" box="[151,406,1734,1759]" country="Thailand" name="Chanthaburi" pageId="22" pageNumber="173">Chanthaburi Province</collectingRegion>
,
<collectingMunicipality id="3B08ACA2835BFFAAFE34F934FD81F958" box="[423,608,1734,1759]" pageId="22" pageNumber="173">Khlung District</collectingMunicipality>
,
<location id="DE0C6003835BFFAAFDE1F934FCE2F958" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFDE1F934FCE2F958" box="[626,771,1734,1759]" country="Thailand" county="Welu River" latitude="12.537" longLatPrecision="77" longitude="102.244" municipality="Khlung District" name="Khen Creek" pageId="22" pageNumber="173" stateProvince="Chanthaburi Province">Khen Creek</location>
, tributary of
<location id="DE0C6003835BFFAAFC3BF934FB99F958" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFC3BF934FB99F958" box="[936,1144,1734,1759]" country="Thailand" county="Welu River" latitude="12.537" longLatPrecision="77" longitude="102.244" municipality="Khlung District" name="Trok Nong River" pageId="22" pageNumber="173" stateProvince="Chanthaburi Province">Trok Nong River</location>
, off Rt. 3277,
<geoCoordinate id="BEE7501F835BFFAAFAA7F934FA79F958" box="[1332,1432,1734,1759]" direction="north" orientation="latitude" pageId="22" pageNumber="173" precision="55" value="12.537">12.537N</geoCoordinate>
,
<geoCoordinate id="BEE7501F835BFFAAFF04F919FEE7F884" box="[151,262,1771,1795]" direction="east" orientation="longitude" pageId="22" pageNumber="173" precision="55" value="102.244">102.244E</geoCoordinate>
,
<collectingDate id="BF29E9F0835BFFAAFE82F919FE2CF884" box="[273,461,1771,1795]" pageId="22" pageNumber="173" value="2014-01-30">
<date id="AF6D1018835BFFAAFE82F919FE2CF884" box="[273,461,1771,1795]" pageId="22" pageNumber="173" value="2014-01-30">30 January 2014</date>
</collectingDate>
,
<collectorName id="7626530E835BFFAAFE4BF918FD41F885" box="[472,672,1770,1795]" pageId="22" pageNumber="173">Nithirojpakdee, J.</collectorName>
<emphasis id="E9A7EACA835BFFAAFD34F919FD39F884" box="[679,728,1771,1795]" italics="true" pageId="22" pageNumber="173">et al</emphasis>
.
</materialsCitation>
</paragraph>
<paragraph id="DB6C36D8835BFFAAFF54F8FCFE74F830" blockId="22.[151,1437,1698,1975]" pageId="22" pageNumber="173">
<materialsCitation id="6BBB3C85835BFFAAFF54F8FCFF25F8CC" ID-GBIF-Occurrence="1676480049" country="Thailand" location="Welu River" pageId="22" pageNumber="173" specimenCount="2" typeStatus="paratype">
<typeStatus id="0468887A835BFFAAFF54F8FCFEDEF8A0" box="[199,319,1806,1831]" pageId="22" pageNumber="173" type="paratype">
<emphasis id="E9A7EACA835BFFAAFF54F8FCFEDEF8A0" bold="true" box="[199,319,1806,1831]" pageId="22" pageNumber="173">Paratypes</emphasis>
</typeStatus>
.
<collectingCountry id="A3C47648835BFFAAFEDFF8FCFE55F8A0" box="[332,436,1806,1831]" name="Thailand" pageId="22" pageNumber="173">Thailand</collectingCountry>
:
<location id="DE0C6003835BFFAAFE2DF8FCFDDFF8A0" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFE2DF8FCFDDFF8A0" box="[446,574,1806,1831]" country="Thailand" name="Welu River" pageId="22" pageNumber="173">Welu River</location>
drainage: same locality and date as
<typeStatus id="0468887A835BFFAAFC41F8FCFBD9F8A0" box="[978,1080,1806,1831]" pageId="22" pageNumber="173" type="holotype">holotype</typeStatus>
: UF 235976 (6,
<quantity id="1C2B9B3D835BFFAAFB7EF8FDFA7CF8A1" box="[1261,1437,1807,1831]" metricMagnitude="-1" metricUnit="m" metricValue="1.048" metricValueMax="1.197" metricValueMin="0.899" pageId="22" pageNumber="173" unit="mm" value="104.8" valueMax="119.7" valueMin="89.9">89.9119.7 mm</quantity>
SL)
</materialsCitation>
; ZRC 55596 (2,
<quantity id="1C2B9B3D835BFFAAFE10F8C0FDD2F8CD" box="[387,563,1842,1867]" metricMagnitude="-1" metricUnit="m" metricValue="1.024" metricValueMax="1.116" metricValueMin="0.932" pageId="22" pageNumber="173" unit="mm" value="102.4" valueMax="111.6" valueMin="93.2">93.2111.6 mm</quantity>
SL); NIFI 5102 (2, 94.2118.0 mm SL).
<materialsCitation id="6BBB3C85835BFFAAFB9AF8C0FC7BF814" ID-GBIF-Occurrence="1676480041" collectingDate="2001-02-22" collectionCode="ANSP" collectorName="M. H. Sabaj &amp; M. Hardman" country="Thailand" county="Ao Luek" latitude="8.623" location="Tapi River" longLatPrecision="77" longitude="98.98" municipality="Rt." pageId="22" pageNumber="173" specimenCode="ANSP 179976" specimenCount="1" stateProvince="Krabi" typeStatus="paratype">
<location id="DE0C6003835BFFAAFB9AF8C0FB61F8CC" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFB9AF8C0FB61F8CC" box="[1033,1152,1842,1867]" country="Thailand" county="Ao Luek" latitude="8.623" longLatPrecision="77" longitude="98.98" municipality="Rt." name="Tapi River" pageId="22" pageNumber="173" stateProvince="Krabi">Tapi River</location>
drainage:
<specimenCode id="8B759EA3835BFFAAFB6BF8C1FA7DF8CC" box="[1272,1436,1842,1867]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="173">ANSP 179976</specimenCode>
(1, 119.0 mm SL),
<collectingRegion id="1917F83A835BFFAAFEFAF8A4FE48F8E8" box="[361,425,1878,1903]" country="Thailand" name="Krabi" pageId="22" pageNumber="173">Krabi</collectingRegion>
/
<collectingRegion id="1917F83A835BFFAAFE22F8A4FDD2F8E8" box="[433,563,1878,1903]" country="Thailand" name="Surat Thani" pageId="22" pageNumber="173">Surat Thani</collectingRegion>
Provinces,
<location id="DE0C6003835BFFAAFD25F8A5FCCFF8E8" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFD25F8A5FCCFF8E8" box="[694,814,1878,1903]" country="Thailand" county="Ao Luek" latitude="8.623" longLatPrecision="77" longitude="98.98" municipality="Rt." name="Ipan River" pageId="22" pageNumber="173" stateProvince="Krabi">Ipan River</location>
, trib. of
<location id="DE0C6003835BFFAAFC18F8A4FBE2F8E8" LSID="urn:lsid:plazi:treatment:537A87CE835BFFA5FF04F9E6FC1FFF7F:DE0C6003835BFFAAFC18F8A4FBE2F8E8" box="[907,1027,1878,1903]" country="Thailand" county="Ao Luek" latitude="8.623" longLatPrecision="77" longitude="98.98" municipality="Rt." name="Tapi River" pageId="22" pageNumber="173" stateProvince="Krabi">Tapi River</location>
, on
<collectingMunicipality id="3B08ACA2835BFFAAFBA3F8A5FBB0F8E9" box="[1072,1105,1879,1902]" pageId="22" pageNumber="173">Rt.</collectingMunicipality>
4035, 25 km NE of
<collectingCounty id="320D4E54835BFFAAFAA0F8A5FA79F8E8" box="[1331,1432,1878,1903]" pageId="22" pageNumber="173">Ao Luek</collectingCounty>
,
<geoCoordinate id="BEE7501F835BFFAAFF04F888FF1AF814" box="[151,251,1914,1939]" direction="north" orientation="latitude" pageId="22" pageNumber="173" precision="55" value="8.623">08.623N</geoCoordinate>
,
<geoCoordinate id="BEE7501F835BFFAAFE95F889FE86F814" box="[262,359,1915,1939]" direction="east" orientation="longitude" pageId="22" pageNumber="173" precision="55" value="98.98">98.980E</geoCoordinate>
,
<collectingDate id="BF29E9F0835BFFAAFEE0F889FDDDF814" box="[371,572,1914,1939]" pageId="22" pageNumber="173" value="2001-02-22">
<date id="AF6D1018835BFFAAFEE0F889FDDDF814" box="[371,572,1914,1939]" pageId="22" pageNumber="173" value="2001-02-22">22 February 2001</date>
</collectingDate>
,
<collectorName id="7626530E835BFFAAFDDBF889FD31F814" box="[584,720,1914,1939]" pageId="22" pageNumber="173">M. H. Sabaj</collectorName>
and
<collectorName id="7626530E835BFFAAFC94F889FC7BF814" box="[775,922,1914,1939]" pageId="22" pageNumber="173">M. Hardman</collectorName>
</materialsCitation>
.
<materialsCitation id="6BBB3C85835BFFAAFC36F889FE74F830" ID-GBIF-Occurrence="1676480054" collectingDate="2001-02-22" collectionCode="INHS, ANSP" collectorName="M. H. Sabaj &amp; M. Hardman" country="Thailand" county="Ao Luek" latitude="8.623" location="Tapi River" longLatPrecision="77" longitude="98.98" municipality="Rt." pageId="22" pageNumber="173" specimenCode="INHS 93550, ANSP 179976" specimenCount="1" stateProvince="Krabi" typeStatus="paratype">
<specimenCode id="8B759EA3835BFFAAFC36F889FBD2F814" box="[933,1075,1914,1939]" collectionCode="INHS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="22" pageNumber="173">INHS 93550</specimenCode>
(1, 109.0 SL) same locality and date as
<specimenCode id="8B759EA3835BFFAAFF78F86DFE71F830" box="[235,400,1950,1975]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="173">ANSP 179976</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="8FAC6650835AFFABFF04F954FC4CF97E" httpUri="https://zenodo.org/record/1039439/files/figure.png" pageId="23" pageNumber="174" targetBox="[151,1436,179,1691]" targetPageId="23">
<paragraph id="DB6C36D8835AFFABFF04F954FC4CF97E" blockId="23.[151,1436,1702,1785]" pageId="23" pageNumber="174">
<emphasis id="E9A7EACA835AFFABFF04F954FEC1F93B" bold="true" box="[151,288,1702,1724]" pageId="23" pageNumber="174">FIGURE 14.</emphasis>
<emphasis id="E9A7EACA835AFFABFEB4F955FDF5F93B" box="[295,532,1702,1724]" italics="true" pageId="23" pageNumber="174">
<taxonomicName id="1CD34D5B835AFFABFEB4F955FDEEF93B" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[295,527,1702,1724]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="rungthipae">Acantopsis rungthipae</taxonomicName>
.
</emphasis>
(A) Lateral and dorsal VieWs, THNHM F0013610, holotype, Welu RiVer, 107.4 mm SL. LiVe VieWs of (B) UF 235976, paratype, Welu RiVer, 91.7 mm SL, (C) UF 188117, Wang RiVer, 56.4 mm SL, (D) UF 188114, Songkhram RiVer, 90.6 mm SL, and (E) UF 236085, Tapi RiVer, 77.3 mm SL.
</paragraph>
</caption>
<subSubSection id="93C96553835AFFA4FF54F8D3FC1FFDBA" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="diagnosis">
<paragraph id="DB6C36D8835AFFABFF54F8D3FDB9F86A" blockId="23.[151,1437,1825,2030]" pageId="23" pageNumber="174">
<emphasis id="E9A7EACA835AFFABFF54F8D3FEA0F8BD" bold="true" box="[199,321,1825,1850]" pageId="23" pageNumber="174">Diagnosis.</emphasis>
<taxonomicName id="1CD34D5B835AFFABFEDAF8D3FDA7F8BE" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[329,582,1825,1849]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA835AFFABFEDAF8D3FDA7F8BE" box="[329,582,1825,1849]" italics="true" pageId="23" pageNumber="174">Acantopsis rungthipae</emphasis>
</taxonomicName>
differs from all other species of
<taxonomicName id="1CD34D5B835AFFABFC2AF8D3FBD5F8BF" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[953,1076,1825,1848]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="23" pageNumber="174" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA835AFFABFC2AF8D3FBD5F8BF" box="[953,1076,1825,1848]" italics="true" pageId="23" pageNumber="174">Acantopsis</emphasis>
</taxonomicName>
(
<tableCitation id="96510363835AFFABFBD0F8D3FB78F8BE" box="[1091,1177,1825,1850]" captionStart="TABLE 4" captionStartId="15.[120,195,534,553]" captionText="TABLE 4. Diagnostic features of species of Acantopsis. * Data from Siebert (1991 b), Sontirat (1999), or Vishwanath and Laisram (2004). Methods from various sources may not be consistent." pageId="23" pageNumber="174">Table 4</tableCitation>
) by combination of no exposed labial barbels, usually distinct small black spot on upper margin of caudal-fin base, 01 row of large spots on side of head, no large black spots on dorsal or caudal fins, no black specks beneath midlateral row of black spots (rarely present), usually 9½ branched dorsal rays and 10 pectoral rays, 1020 dark saddles along dorsal midline, 9 17 black spots along side of body, 4143 total vertebrae, body depth 9.014.2% SL, body width 5.310.3% SL, caudal-peduncle length 11.516.1% SL.
</paragraph>
<paragraph id="DB6C36D88355FFA4FF54FF65FC1FFDBA" blockId="24.[151,1437,151,573]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF54FF65FE8CFF37" bold="true" box="[199,365,151,176]" pageId="24" pageNumber="175">Comparisons.</emphasis>
<taxonomicName id="1CD34D5B8355FFA4FEE7FF6AFD93FF37" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[372,626,152,176]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA8355FFA4FEE7FF6AFD93FF37" box="[372,626,152,176]" italics="true" pageId="24" pageNumber="175">Acantopsis rungthipae</emphasis>
</taxonomicName>
is distinguished from all species of
<taxonomicName id="1CD34D5B8355FFA4FB9FFF6AFB66FF28" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[1036,1159,152,175]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA8355FFA4FB9FFF6AFB66FF28" box="[1036,1159,152,175]" italics="true" pageId="24" pageNumber="175">Acantopsis</emphasis>
</taxonomicName>
except
<taxonomicName id="1CD34D5B8355FFA4FB73FF6AFAC5FF28" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[1248,1316,152,175]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="ioa">
<emphasis id="E9A7EACA8355FFA4FB73FF6AFAC5FF28" box="[1248,1316,152,175]" italics="true" pageId="24" pageNumber="175">A. ioa</emphasis>
</taxonomicName>
by having no labial barbels. It is distinguished from
<taxonomicName id="1CD34D5B8355FFA4FDE1FF4EFD56FF53" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[626,695,188,212]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="ioa">
<emphasis id="E9A7EACA8355FFA4FDE1FF4EFD56FF53" box="[626,695,188,212]" italics="true" pageId="24" pageNumber="175">A. ioa</emphasis>
</taxonomicName>
by usually having a distinct small black spot on upper margin of caudal-fin base, 917 (vs. 1321) black spots along side of body, and 4143 (vs. 4648) total vertebrae (
<figureCitation id="43E82A5D8355FFA4FAB7FF2DFA96FF7F" box="[1316,1399,223,248]" captionStart="FIGURE 15" captionStartId="24.[151,250,1376,1398]" captionTargetBox="[151,1436,603,1365]" captionTargetId="figure@24.[151,1436,603,1365]" captionTargetPageId="24" captionText="FIGURE 15. CT-scans of (A) Acantopsis rungthipae, THNHM F 0013610, holotype, 107.4 mm SL. (B) A. dinema, THNHM F 0013611, holotype, 100.7 mm SL. (C) A. ioa, UF 188136, 95.5 mm SL. CT-scans proVided by E. L. Stanley (UF)." httpUri="https://zenodo.org/record/1039441/files/figure.png" pageId="24" pageNumber="175">Fig. 15</figureCitation>
). It further differs from
<taxonomicName id="1CD34D5B8355FFA4FEE6FEF6FE06FE9B" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[373,487,260,284]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="dinema">
<emphasis id="E9A7EACA8355FFA4FEE6FEF6FE06FE9B" box="[373,487,260,284]" italics="true" pageId="24" pageNumber="175">A. dinema</emphasis>
</taxonomicName>
by having large dark spots on side of head, usually lacking conspicuous row of black specks beneath midlateral row of black spots, having usually 9½ (vs. 10½) branched dorsal rays, a distinct small black spot (vs. no spot) on upper margin of caudal-fin base, and usually 46 faint, narrow bands (vs. no bands) on caudal fin. It further differs from
<taxonomicName id="1CD34D5B8355FFA4FD81FE82FD28FE00" authorityName="Siebert" authorityYear="1991" box="[530,713,368,391]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="octoactinotos">
<emphasis id="E9A7EACA8355FFA4FD81FE82FD28FE00" box="[530,713,368,391]" italics="true" pageId="24" pageNumber="175">A. octoactinotos</emphasis>
</taxonomicName>
by having usually 9½ (vs. 8½) branched dorsal rays, large dark spots on side of head, usually 10 pectoral rays. It differs from
<taxonomicName id="1CD34D5B8355FFA4FC07FE66FBD0FE2B" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[916,1073,404,428]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA8355FFA4FC07FE66FBD0FE2B" box="[916,1073,404,428]" italics="true" pageId="24" pageNumber="175">A. spectabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="1CD34D5B8355FFA4FBE0FE66FAFBFE2B" authorityName="Sontirat" authorityYear="1999" box="[1139,1306,404,428]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="thiemmedhi">
<emphasis id="E9A7EACA8355FFA4FBE0FE66FAFBFE2B" box="[1139,1306,404,428]" italics="true" pageId="24" pageNumber="175">A. thiemmedhi</emphasis>
</taxonomicName>
in lacking conspicuous black bands on the anal and paired fins and spots or blotches on the caudal fin. It further differs from
<taxonomicName id="1CD34D5B8355FFA4FF04FE2EFED8FE73" authorityName="Sontirat" authorityYear="1999" box="[151,313,476,500]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="thiemmedhi">
<emphasis id="E9A7EACA8355FFA4FF04FE2EFED8FE73" box="[151,313,476,500]" italics="true" pageId="24" pageNumber="175">A. thiemmedhi</emphasis>
</taxonomicName>
in lacking a large black blotch on the tip of the anterior rays of the dorsal fin and in having usually 9½ (vs. 10½) branched dorsal rays, and from
<taxonomicName id="1CD34D5B8355FFA4FD30FDF2FCD3FD9F" authorityName="van Hasselt" authorityYear="1823" box="[675,818,512,536]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="dialuzona">
<emphasis id="E9A7EACA8355FFA4FD30FDF2FCD3FD9F" box="[675,818,512,536]" italics="true" pageId="24" pageNumber="175">A. dialuzona</emphasis>
</taxonomicName>
by lacking a well developed ocellus on the margin of caudal-fin base and having usually 9½ (vs. usually10½) branched dorsal rays.
</paragraph>
</subSubSection>
<caption id="8FAC66508355FFA4FF04FA92FAC7FA13" httpUri="https://zenodo.org/record/1039441/files/figure.png" pageId="24" pageNumber="175" targetBox="[151,1436,603,1365]" targetPageId="24">
<paragraph id="DB6C36D88355FFA4FF04FA92FAC7FA13" blockId="24.[151,1436,1376,1429]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF04FA92FEC1FAF1" bold="true" box="[151,288,1376,1398]" pageId="24" pageNumber="175">FIGURE 15.</emphasis>
CT-scans of (A)
<taxonomicName id="1CD34D5B8355FFA4FE47FA93FD5BFAF1" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[468,698,1376,1398]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA8355FFA4FE47FA93FD5BFAF1" box="[468,698,1376,1398]" italics="true" pageId="24" pageNumber="175">Acantopsis rungthipae</emphasis>
</taxonomicName>
, THNHM F0013610, holotype, 107.4 mm SL. (B)
<taxonomicName id="1CD34D5B8355FFA4FB58FA93FAD3FAF1" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[1227,1330,1376,1398]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="dinema">
<emphasis id="E9A7EACA8355FFA4FB58FA93FAD3FAF1" box="[1227,1330,1376,1398]" italics="true" pageId="24" pageNumber="175">A. dinema</emphasis>
</taxonomicName>
, THNHM F0013611, holotype, 100.7 mm SL. (C)
<taxonomicName id="1CD34D5B8355FFA4FDBEFA8DFD8BFA12" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[557,618,1407,1429]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="ioa">
<emphasis id="E9A7EACA8355FFA4FDBEFA8DFD8BFA12" box="[557,618,1407,1429]" italics="true" pageId="24" pageNumber="175">A. ioa</emphasis>
</taxonomicName>
, UF 188136, 95.5 mm SL. CT-scans proVided by E. L. Stanley (UF).
</paragraph>
</caption>
<subSubSection id="93C965538355FFA4FF54FA4EFB86F8E7" pageId="24" pageNumber="175" type="description">
<paragraph id="DB6C36D88355FFA4FF54FA4EFA99F99B" blockId="24.[151,1437,1468,2032]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF54FA4EFEB8FA52" bold="true" box="[199,345,1468,1493]" pageId="24" pageNumber="175">Description.</emphasis>
As in description of genus;
<tableCitation id="965103638355FFA4FD0BFA4EFD16FA53" box="[664,759,1468,1493]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Frequency distributions of labial barbels and meristic characteristics in species of Acantopsis." pageId="24" pageNumber="175">Tables 1</tableCitation>
,
<tableCitation id="965103638355FFA4FC95FA4EFCF7FA53" box="[774,790,1468,1492]" captionStart="TABLE 2" captionStartId="6.[120,202,556,577]" captionText="TABLE 2. Morphometric data for recognized species of Acantopsis." pageId="24" pageNumber="175">2</tableCitation>
,
<tableCitation id="965103638355FFA4FCB1FA4EFCD2FA53" box="[802,819,1468,1492]" captionStart="TABLE 4" captionStartId="15.[120,195,534,553]" captionText="TABLE 4. Diagnostic features of species of Acantopsis. * Data from Siebert (1991 b), Sontirat (1999), or Vishwanath and Laisram (2004). Methods from various sources may not be consistent." pageId="24" pageNumber="175">4</tableCitation>
. Head long (21.625.6% SL); snout long (54.768.7% HL); no exposed labial barbels; 9½, rarely 10½, branched dorsal rays; 10, rarely 9 or 11, pectoral rays; 2931 abdominal + 1112 caudal = 4143 total vertebrae. Maximum SL =
<quantity id="1C2B9B3D8355FFA4FC07F9F6FBE9F99B" box="[916,1032,1540,1564]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="24" pageNumber="175" unit="mm" value="203.2">203.2 mm</quantity>
, TL =
<quantity id="1C2B9B3D8355FFA4FBC0F9F6FB27F99B" box="[1107,1222,1540,1564]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" pageId="24" pageNumber="175" unit="mm" value="241.3">241.3 mm</quantity>
(CAS 238293).
</paragraph>
<paragraph id="DB6C36D88355FFA4FF54F9D5FF00F972" blockId="24.[151,1437,1468,2032]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF54F9D5FEF1F9C7" bold="true" box="[199,272,1575,1600]" pageId="24" pageNumber="175">Color.</emphasis>
Dark spots in 12 rows on top of head, usually 1 row anteriorly, 2 posteriorly; dark spots on side of head usually in 1 row. Black line from eye to tip of snout. Ten20 dark saddles along dorsal midline; 917 large dark spots or blotches along lateral line from head to caudal fin; small dark spots on upper side of body; no or few dark spots on lower side of body below midlateral row of dark spots. Dorsal fin with 23 faint bands, caudal with 45 faint bands; no dark upper and lower margins on caudal fin. Small bold black spot near upper margin of caudal-fin origin.
</paragraph>
<paragraph id="DB6C36D88355FFA4FF54F90DFAE6F8BA" blockId="24.[151,1437,1468,2032]" pageId="24" pageNumber="175">Live individuals yellowish brown dorsally, white below, with large black blotches dorsally and laterally on body; reddish brown streak on lower caudal margin; iridescent yellow on side of head and lower fins.</paragraph>
<paragraph id="DB6C36D88355FFA4FF54F8B5FB86F8E7" blockId="24.[151,1437,1468,2032]" box="[199,1127,1863,1888]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF54F8B5FE56F8E7" bold="true" box="[199,439,1863,1888]" pageId="24" pageNumber="175">Sexual dimorphism.</emphasis>
Males have a longer, ramified first branched pectoral-fin ray.
</paragraph>
</subSubSection>
<subSubSection id="93C965538355FFA4FF54F89EFF0DF877" pageId="24" pageNumber="175" type="etymology">
<paragraph id="DB6C36D88355FFA4FF54F89EFF0DF877" blockId="24.[151,1437,1468,2032]" pageId="24" pageNumber="175">
<emphasis id="E9A7EACA8355FFA4FF54F89EFEAFF802" bold="true" box="[199,334,1900,1925]" pageId="24" pageNumber="175">Etymology.</emphasis>
This species is named in honor of Rungthip Plongsesthee, who was born on
<date id="AF6D10188355FFA4FB31F89EFAD6F803" box="[1186,1335,1900,1925]" pageId="24" pageNumber="175" value="1978-05-27">27 May 1978</date>
and died on
<date id="AF6D10188355FFA4FF2FF87DFEB4F82F" box="[188,341,1935,1960]" pageId="24" pageNumber="175" value="2014-07-16">16 July 2014</date>
. “Kae” was a dear friend, close colleague, a Ph.D. student of Dr. F. W. H. Beamish at Burapha University, Bangsaen,
<collectingCountry id="A3C476488355FFA4FE0AF846FE1EF84A" box="[409,511,1972,1997]" name="Thailand" pageId="24" pageNumber="175">Thailand</collectingCountry>
, and an extremely enthusiastic ichthyologist who is greatly missed by her many friends.
</paragraph>
</subSubSection>
<subSubSection id="93C965538354FFA5FF54FF65FC1FFF7F" pageId="25" pageNumber="176" type="distribution">
<paragraph id="DB6C36D88354FFA5FF54FF65FC1FFF7F" blockId="25.[151,1436,151,248]" pageId="25" pageNumber="176">
<emphasis id="E9A7EACA8354FFA5FF54FF65FE81FF37" bold="true" box="[199,352,151,176]" pageId="25" pageNumber="176">Distribution.</emphasis>
<taxonomicName id="1CD34D5B8354FFA5FEFAFF6AFD86FF37" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[361,615,152,176]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="25" pageNumber="176" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA8354FFA5FEFAFF6AFD86FF37" box="[361,615,152,176]" italics="true" pageId="25" pageNumber="176">Acantopsis rungthipae</emphasis>
</taxonomicName>
is the most widespread and common species of
<taxonomicName id="1CD34D5B8354FFA5FB01FF6AFAECFF28" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[1170,1293,152,175]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="25" pageNumber="176" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA8354FFA5FB01FF6AFAECFF28" box="[1170,1293,152,175]" italics="true" pageId="25" pageNumber="176">Acantopsis</emphasis>
</taxonomicName>
in mainland Asia. It is known from the Tapi, Mae Khlong, Chao Phraya, Bang Pakong, and Welu River basins in
<collectingCountry id="A3C476488354FFA5FA91FF4EFA89FF52" box="[1282,1384,188,213]" name="Thailand" pageId="25" pageNumber="176">Thailand</collectingCountry>
, and the Mekong River basin in
<collectingCountry id="A3C476488354FFA5FE5BFF2DFDCFFF7F" box="[456,558,223,248]" name="Thailand" pageId="25" pageNumber="176">Thailand</collectingCountry>
,
<collectingCountry id="A3C476488354FFA5FDA9FF2DFD51FF7F" box="[570,688,223,248]" name="Cambodia" pageId="25" pageNumber="176">Cambodia</collectingCountry>
,
<collectingCountry id="A3C476488354FFA5FD28FF12FD15FF70" box="[699,756,224,247]" name="Laos" pageId="25" pageNumber="176">Laos</collectingCountry>
, and
<collectingCountry id="A3C476488354FFA5FCBCFF2DFC6FFF7F" box="[815,910,223,248]" name="Vietnam" pageId="25" pageNumber="176">Vietnam</collectingCountry>
(
<figureCitation id="43E82A5D8354FFA5FC0EFF2DFC10FF7F" box="[925,1009,223,248]" captionStart="FIGURE 16" captionStartId="25.[151,250,1901,1923]" captionTargetBox="[186,1385,316,1868]" captionTargetId="figure@25.[175,1407,301,1895]" captionTargetPageId="25" captionText="FIGURE 16. Localities of specimens examined of Acantopsis rungthipae." httpUri="https://zenodo.org/record/1039443/files/figure.png" pageId="25" pageNumber="176">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>