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<document id="B80383AD5E91FA3A133461FC3DA3255F" ID-CLB-Dataset="8966" ID-DOI="10.11646/zootaxa.4820.3.8" ID-GBIF-Dataset="f702f2bf-e817-4863-96f7-f20015f61de3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4398204" ID-ZooBank="B8AAC629-F431-49E2-B15C-80CCC0CE070C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609213172128" checkinUser="plazi" docAuthor="Alekseev, Vitalii Igorevich, Bukejs, Andris &amp; Pollock, Darren Andrew" docDate="2020" docId="17568786FFE2735A23A8050CFD8FC6BD" docLanguage="en" docName="zootaxa.4820.3.8.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Europoeurypus Alekseev &amp; Bukejs &amp; Pollock 2020, gen. nov." docType="treatment" docUuid="7E559053-7D87-4EF1-A2BD-D9BD160E0A51" docUuidSource="ZooBank" docVersion="8" lastPageNumber="552" masterDocId="EB6FFFFEFFE3735B233F0116FFEBC10E" masterDocTitle="A new fossil Eurypinae genus and sexual dimorphism in the representatives of Omineus (Coleoptera: Mycteridae) from Tertiary amber of Europe" masterLastPageNumber="562" masterPageNumber="551" pageNumber="552" updateTime="1698848390187" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4DB308C1038BD3B9B40C23458B464AF3">A new fossil Eurypinae genus and sexual dimorphism in the representatives of Omineus (Coleoptera: Mycteridae) from Tertiary amber of Europe</mods:title>
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<mods:namePart id="8860A84379127106D32448F394261605">Alekseev, Vitalii Igorevich</mods:namePart>
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<mods:affiliation id="F0F5E797ECC0F5FF4FAC561F1EE125C7">Shirshov Institute of Oceanology, Russian Academy of Sciences, Nahimovskiy prospekt 36, 117997 Moscow, Russia. alekseew 0802 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4390 - 5443 &amp; Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia.</mods:affiliation>
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<mods:namePart id="F97FA6D0353D8F09BE551F048A41521D">Bukejs, Andris</mods:namePart>
<mods:affiliation id="54DFF8588BEEA2C5FF117A39DD23E2DF">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<mods:namePart id="329724C4FE265C051EFF0B4581A67131">Pollock, Darren Andrew</mods:namePart>
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<subSubSection id="D7E5651BFFE2735A23A8050CFDF4C53B" box="[151,543,1050,1077]" pageId="1" pageNumber="552" type="nomenclature">
<paragraph id="9F403690FFE2735A23A8050CFDF4C53B" blockId="1.[151,543,1050,1077]" box="[151,543,1050,1077]" pageId="1" pageNumber="552">
<heading id="C40881FCFFE2735A23A8050CFDF4C53B" bold="true" box="[151,543,1050,1077]" fontSize="11" level="1" pageId="1" pageNumber="552" reason="1">
<emphasis id="AD8BEA82FFE2735A23A8050CFDF4C53B" bold="true" box="[151,543,1050,1077]" pageId="1" pageNumber="552">
Genus
<taxonomicName id="58FF4D13FFE2735A23D1050DFE42C53B" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[238,425,1051,1077]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD8BEA82FFE2735A23D1050DFE42C53B" bold="true" box="[238,425,1051,1077]" italics="true" pageId="1" pageNumber="552">Europoeurypus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6B857F9FFE2735A228E050DFDF4C53B" box="[433,543,1051,1077]" pageId="1" pageNumber="552" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D7E5651BFFE2735A23A80576FC8DC579" box="[151,870,1119,1144]" pageId="1" pageNumber="552" type="materials_examined">
<paragraph id="9F403690FFE2735A23A80576FC8DC579" blockId="1.[151,870,1119,1144]" box="[151,870,1119,1144]" pageId="1" pageNumber="552">
<typeStatus id="40448832FFE2735A23A80576FF22C576" box="[151,201,1120,1144]" pageId="1" pageNumber="552">Type</typeStatus>
species:
<taxonomicName id="58FF4D13FFE2735A22190576FDF5C579" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[294,542,1119,1143]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="inglaeso" status="sp. nov.">
<emphasis id="AD8BEA82FFE2735A22190576FDF5C579" box="[294,542,1119,1143]" italics="true" pageId="1" pageNumber="552">Europoeurypus inglaeso</emphasis>
</taxonomicName>
<emphasis id="AD8BEA82FFE2735A211B0549FD9EC579" bold="true" box="[548,629,1119,1143]" pageId="1" pageNumber="552">
<taxonomicNameLabel id="B6B857F9FFE2735A211B0549FD9EC579" box="[548,629,1119,1143]" pageId="1" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation
</paragraph>
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<subSubSection id="D7E5651BFFE2735A23A805B3FEBCC60E" pageId="1" pageNumber="552" type="diagnosis">
<paragraph id="9F403690FFE2735A23A805B3FBE1C47A" blockId="1.[151,1437,1189,1971]" pageId="1" pageNumber="552">
<emphasis id="AD8BEA82FFE2735A23A805B3FEF9C5B1" bold="true" box="[151,274,1189,1215]" pageId="1" pageNumber="552">Diagnosis.</emphasis>
<taxonomicName id="58FF4D13FFE2735A222505B0FE2DC5B1" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[282,454,1190,1215]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD8BEA82FFE2735A222505B0FE2DC5B1" box="[282,454,1190,1215]" italics="true" pageId="1" pageNumber="552">Europoeurypus</emphasis>
</taxonomicName>
<emphasis id="AD8BEA82FFE2735A22F005B3FDD1C5B1" bold="true" box="[463,570,1189,1215]" pageId="1" pageNumber="552">
<taxonomicNameLabel id="B6B857F9FFE2735A22F005B3FDD1C5B1" box="[463,570,1189,1215]" pageId="1" pageNumber="552" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="07C42A15FFE2735A217405B3FD5DC5CE" box="[587,694,1189,1216]" captionStart-0="FIGURES 12" captionStart-1="FIGURE 34" captionStart-2="FIGURE 5" captionStartId-0="2.[151,264,1899,1923]" captionStartId-1="3.[151,250,1050,1074]" captionStartId-2="4.[151,250,1955,1979]" captionTargetBox-0="[151,1435,179,1872]" captionTargetBox-1="[151,1436,181,1026]" captionTargetBox-2="[252,1339,315,1931]" captionTargetId-0="figure-23@2.[151,1435,179,1872]" captionTargetId-1="figure-18@3.[151,1436,181,1026]" captionTargetId-2="figure-78@4.[245,1342,309,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 12. Europoeurypus inglaeso gen. et sp. nov., holotype, No MAIG 6081: 1—habitus, dorsal view; 2—habitus, ven- tral view. Scale bars represent 4.0 mm." captionText-1="FIGURE 34. Europoeurypus inglaeso gen. et sp. nov., holotype, No MAIG 6081: 3—abdomen, showing ventrites without discernible sex patches; 4—details of forebody, dorsal view. Abbreviations: v1v5—abdominal ventrites 15 respectively; icp—intercoxal process of ventrite 1. Scale bars represent 1.0 mm." captionText-2="FIGURE 5. Reconstruction of Europoeurypus inglaeso gen. et sp. nov. from Baltic amber: habitus, dorsal view. Body length = 10.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4398206" figureDoi-1="http://doi.org/10.5281/zenodo.4398208" figureDoi-2="http://doi.org/10.5281/zenodo.4398210" httpUri-0="https://zenodo.org/record/4398206/files/figure.png" httpUri-1="https://zenodo.org/record/4398208/files/figure.png" httpUri-2="https://zenodo.org/record/4398210/files/figure.png" pageId="1" pageNumber="552">Figs. 15</figureCitation>
) can be distinguished from all other described
<taxonomicName id="58FF4D13FFE2735A27E905B3FAA2C5B1" authorityName="Thomson" authorityYear="1860" box="[1238,1353,1189,1215]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
genera by the following characters: (1) ocular (suborbital) groove not distinct; (2) head with straight, subparallel frontal furrows; (3) eyes protuberant, very slightly emarginated, without intrafacetal setae; (4) lateral carina of pronotum present in basal two-thirds; (5) frontoclypeal suture discernible; (6) dorsal body surface evenly punctate and setose; (7) pronotal disc with pair of small basal triangular impressions; (8) mandibles bidentate apically; (9) tarsal claws thickened basally, without tooth; and (10) total body length larger than
<quantity id="58079B75FFE2735A208B044FFBEDC47A" box="[948,1030,1369,1396]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="552" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="9F403690FFE2735A23F80468FEBCC60E" blockId="1.[151,1437,1189,1971]" pageId="1" pageNumber="552">
Among extant
<taxonomicName id="58FF4D13FFE2735A224E046BFE0CC499" authorityName="Thomson" authorityYear="1860" box="[369,487,1405,1431]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
, the new genus has a similar form to species of both
<taxonomicName id="58FF4D13FFE2735A2771046BFA80C499" authority="Pic, 1931" authorityName="Pic" authorityYear="1931" box="[1102,1387,1405,1431]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A2771046BFB1DC499" box="[1102,1270,1405,1431]" italics="true" pageId="1" pageNumber="552">Thisiomorphus</emphasis>
Pic, 1931
</taxonomicName>
and
<taxonomicName id="58FF4D13FFE2735A23A804B4FE6EC4B5" authority="Kirby, 1819" authorityName="Kirby" authorityYear="1819" box="[151,389,1441,1467]" class="Insecta" family="Mycteridae" genus="Eurypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A23A804B4FF1FC4B5" box="[151,244,1442,1467]" italics="true" pageId="1" pageNumber="552">Eurypus</emphasis>
Kirby, 1819
</taxonomicName>
. Species of both genera have a relatively wide pronotum and an elongate-oval body (
<bibRefCitation id="FB6E4B61FFE2735A267A04B7FF3FC4EE" author="Pollock, D. A." pageId="1" pageNumber="552" pagination="301 - 322" refId="ref5270" refString="Pollock, D. A. (2016) Revision of the genus Thisiomorphus Pic (Coleoptera: Mycteridae: Eurypinae) with descriptions of eleven new species from Central and South America and a key to genera of Neotropical Eurypinae. Zootaxa, 4093 (3), 301 - 322. https: // doi. org / 10.11646 / zootaxa. 4093.3.1" type="journal article" year="2016">Pollock 2016</bibRefCitation>
).
<taxonomicName id="58FF4D13FFE2735A23D604D0FE7EC4D1" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[233,405,1478,1503]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD8BEA82FFE2735A23D604D0FE7EC4D1" box="[233,405,1478,1503]" italics="true" pageId="1" pageNumber="552">Europoeurypus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6B857F9FFE2735A22A104D3FDE0C4EE" box="[414,523,1477,1504]" pageId="1" pageNumber="552" rank="genus">
<emphasis id="AD8BEA82FFE2735A22A104D3FE23C4D1" bold="true" box="[414,456,1477,1503]" pageId="1" pageNumber="552">gen</emphasis>
.
<emphasis id="AD8BEA82FFE2735A22E704D3FDEFC4D1" bold="true" box="[472,516,1477,1503]" pageId="1" pageNumber="552">nov</emphasis>
.
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="58FF4D13FFE2735A207904D0FC48C4D1" authorityName="Kirby" authorityYear="1819" box="[838,931,1478,1503]" class="Insecta" family="Mycteridae" genus="Eurypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A207904D0FC48C4D1" box="[838,931,1478,1503]" italics="true" pageId="1" pageNumber="552">Eurypus</emphasis>
</taxonomicName>
by the eyes without setation, not especially widened penultimate tarsomere and shorter antennae, and from
<taxonomicName id="58FF4D13FFE2735A206704FFFBEBC70D" authorityName="Pic" authorityYear="1931" box="[856,1024,1513,1539]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A206704FFFBEBC70D" box="[856,1024,1513,1539]" italics="true" pageId="1" pageNumber="552">Thisiomorphus</emphasis>
</taxonomicName>
by the considerably larger body size, the shorter antennae, the smaller eyes, tarsal claws without sharp tooth, and distinct frontoclypeal suture. This new genus from Baltic amber differs from known Eocene
<taxonomicName id="58FF4D13FFE2735A21C70727FC80C745" authorityName="Thomson" authorityYear="1860" box="[760,875,1585,1611]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
(see also key below) as follows: from
<taxonomicName id="58FF4D13FFE2735A26170724FE16C761" authority="Kirejtshuk &amp; Nel, 2009" authorityName="Kirejtshuk &amp; Nel" authorityYear="2009" class="Insecta" family="Mycteridae" genus="Bertinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="gallicus">
<emphasis id="AD8BEA82FFE2735A26170724FF1BC761" italics="true" pageId="1" pageNumber="552">Bertinotus gallicus</emphasis>
<bibRefCitation id="FB6E4B61FFE2735A23C90743FE16C761" author="Kirejtshuk, A. G. &amp; Nel, A." box="[246,509,1621,1647]" pageId="1" pageNumber="552" pagination="103 - 118" refId="ref5051" refString="Kirejtshuk, A. G. &amp; Nel, A. (2009) New genera and species of Cucujiformia (Coleoptera, Polyphaga) from lowermost Eocene French amber. Denisia, 26, 103 - 118." type="journal article" year="2009">Kirejtshuk &amp; Nel, 2009</bibRefCitation>
</taxonomicName>
(Lowermost Eocene amber of Oise,
<collectingCountry id="E7E87600FFE2735A20A90740FC0FC77E" box="[918,996,1622,1648]" name="France" pageId="1" pageNumber="552">France</collectingCountry>
) in punctate head, bidentate mandibles, non-toothed claws, markedly larger body size; from
<taxonomicName id="58FF4D13FFE2735A21D1076FFF24C7B9" authority="Alekseev, Pollock &amp; Bukejs, 2019" authorityName="Alekseev, Pollock &amp; Bukejs" authorityYear="2019" class="Insecta" family="Mycteridae" genus="Glesoconomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="nachzehrer">
<emphasis id="AD8BEA82FFE2735A21D1076FFBA5C79D" box="[750,1102,1657,1683]" italics="true" pageId="1" pageNumber="552">Glesoconomorphus nachzehrer</emphasis>
<bibRefCitation id="FB6E4B61FFE2735A276A076FFF24C7B9" author="Alekseev, V. I. &amp; Pollock, D. A. &amp; Bukejs, A." pageId="1" pageNumber="552" pagination="67 - 78" refId="ref4733" refString="Alekseev, V. I., Pollock, D. A. &amp; Bukejs, A. (2019) Two new fossil representatives of Eurypinae (Coleoptera: Tenebrionoidea: Mycteridae) from Eocene Baltic amber and placement of Neopolypria nigra Abdullah, 1964. Zootaxa, 4551 (1), 67 - 78. https: // doi. org / 10.11646 / zootaxa. 4551.1.4" type="journal article" year="2019">Alekseev, Pollock &amp; Bukejs, 2019</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="58FF4D13FFE2735A223C0788FCB8C7B9" authority="Alekseev, Pollock &amp; Bukejs, 2019" authorityName="Alekseev, Pollock &amp; Bukejs" authorityYear="2019" box="[259,851,1693,1720]" class="Insecta" family="Mycteridae" genus="Omineus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="febribilis">
<emphasis id="AD8BEA82FFE2735A223C0788FE39C7B9" box="[259,466,1693,1720]" italics="true" pageId="1" pageNumber="552">Omineus febribilis</emphasis>
<bibRefCitation id="FB6E4B61FFE2735A22E9078BFCB8C7B9" author="Alekseev, V. I. &amp; Pollock, D. A. &amp; Bukejs, A." box="[470,851,1693,1719]" pageId="1" pageNumber="552" pagination="67 - 78" refId="ref4733" refString="Alekseev, V. I., Pollock, D. A. &amp; Bukejs, A. (2019) Two new fossil representatives of Eurypinae (Coleoptera: Tenebrionoidea: Mycteridae) from Eocene Baltic amber and placement of Neopolypria nigra Abdullah, 1964. Zootaxa, 4551 (1), 67 - 78. https: // doi. org / 10.11646 / zootaxa. 4551.1.4" type="journal article" year="2019">Alekseev, Pollock &amp; Bukejs, 2019</bibRefCitation>
</taxonomicName>
(both Eocene Baltic amber,
<collectingCountry id="E7E87600FFE2735A27B2078BFB32C7B6" box="[1165,1241,1693,1720]" name="Russia" pageId="1" pageNumber="552">Russia</collectingCountry>
) in ocular groove absent and frontal furrows present, lateral carina of pronotum present, distinctly larger body size, and comparatively shorter antennae.
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE2735A23F8061FFE74C665" pageId="1" pageNumber="552" type="etymology">
<paragraph id="9F403690FFE2735A23F8061FFE74C665" blockId="1.[151,1437,1189,1971]" pageId="1" pageNumber="552">
<emphasis id="AD8BEA82FFE2735A23F8061FFEA5C62D" bold="true" box="[199,334,1801,1827]" pageId="1" pageNumber="552">Etymology.</emphasis>
The name of the new genus is a compound word and combines “
<taxonomicName id="58FF4D13FFE2735A2719061CFB92C62D" authorityName="Thomson" authorityYear="1860" box="[1062,1145,1802,1827]" class="Insecta" family="Cerambycidae" genus="Europa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A2719061CFB92C62D" box="[1062,1145,1802,1827]" italics="true" pageId="1" pageNumber="552">Europa</emphasis>
</taxonomicName>
” (Europe, western part of the continent Eurasia from where the Baltic amber originates) and “
<taxonomicName id="58FF4D13FFE2735A20BE0638FC35C649" authorityName="Kirby" authorityYear="1819" box="[897,990,1838,1863]" class="Insecta" family="Mycteridae" genus="Eurypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="AD8BEA82FFE2735A20BE0638FC35C649" box="[897,990,1838,1863]" italics="true" pageId="1" pageNumber="552">Eurypus</emphasis>
</taxonomicName>
” (name of the
<typeStatus id="40448832FFE2735A27BF0638FB5BC646" box="[1152,1200,1838,1864]" pageId="1" pageNumber="552">type</typeStatus>
genus in the subfamily). Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE2735A23F80663FD8FC6BD" pageId="1" pageNumber="552" type="discussion">
<paragraph id="9F403690FFE2735A23F80663FD8FC6BD" blockId="1.[151,1437,1189,1971]" pageId="1" pageNumber="552">
<emphasis id="AD8BEA82FFE2735A23F80663FED0C681" bold="true" box="[199,315,1909,1935]" pageId="1" pageNumber="552">Remarks.</emphasis>
The new genus is monotypic, represented by the
<typeStatus id="40448832FFE2735A20600660FC64C69E" box="[863,911,1910,1936]" pageId="1" pageNumber="552">type</typeStatus>
species only. Therefore the generic description considerably overlaps that of the species.
</paragraph>
</subSubSection>
</treatment>
</document>