446 lines
58 KiB
XML
446 lines
58 KiB
XML
<document id="81A471CC4FDA03C6D17E73B5032B7A2F" ID-CLB-Dataset="21955" ID-DOI="10.5852/ejt.2017.368" ID-GBIF-Dataset="7d707e31-1104-4ed3-8493-ef9302b98796" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838412" ID-ZooBank="E4AEAA16-A546-48B8-99BA-2E0152C8C6F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590087673171" checkinUser="carolina" docAuthor="Zlatkov, Boyan & Huemer, Peter" docDate="2017" docId="03E9996C14424E6CB68C68718652DE55" docLanguage="en" docName="ejt-2017-368.pdf.imf" docOrigin="European Journal of Taxonomy 368" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phtheochroa frigidana Zlatkov & Huemer 2017, stat. rev." docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFD0E114144F4E7DB7706C11845EDC35" masterDocTitle="Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="14" updateTime="1698840706466" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C4A3DE94B67E9D45A0845AAB24B4DB4E">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:namePart id="EE4E6894C6CB154738CDF9187CC1440E">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="D6DE135FF053DFFD3407D7F5A1B65FDB">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. & National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. & urn: lsid: zoobank. org: author: 3538 D 3 F 5 - 6822 - 403 F-B 9 E 6 - E 8 A 95 E 884404 & Corresponding author: bzlatkov @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="6F7DB6DB93956A4B053E7A11083F5DD7" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6CF7EA4FE9E700C926442F44F7D9A781">Huemer, Peter</mods:namePart>
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<mods:affiliation id="F351DCB2FD5B992A5F1EA9CC98FCEB96">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria. & Email: p. huemer @ tiroler-landesmuseen. at & urn: lsid: zoobank. org: author: 4 E 6 F 001 E-DB 9 D- 460 C-AB 39 - 0 CE 7 CC 773 B 59</mods:affiliation>
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<mods:title id="66FA53459203965B4EF7AFDE702D7E11">European Journal of Taxonomy</mods:title>
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<mods:date id="86C9D1808420E0B939DFD4151BC1A8FB">2017</mods:date>
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<mods:number id="28D8D487DD02D39908C76FB92BAE19A4">2017-11-21</mods:number>
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<treatment id="03E9996C14424E6CB68C68718652DE55" ID-DOI="http://doi.org/10.5281/zenodo.3852266" ID-GBIF-Taxon="164246402" ID-Zenodo-Dep="3852266" LSID="urn:lsid:plazi:treatment:03E9996C14424E6CB68C68718652DE55" httpUri="http://treatment.plazi.org/id/03E9996C14424E6CB68C68718652DE55" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
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<heading id="D0B79F1614424E70B68C68718066D84E" box="[508,1080,1120,1147]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
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<taxonomicName id="4C4053F914424E70B68C68718790D84E" ID-CoL="4GX97" authority="(Guenee, 1845)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[508,974,1120,1147]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frigidana" status="stat. nov.">
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<emphasis id="B934F46814424E70B68C6871874ED84E" bold="true" box="[508,784,1120,1147]" italics="true" pageId="13" pageNumber="14">Phtheochroa frigidana</emphasis>
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(Guenée, 1845)
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<taxonomicNameLabel id="A207491314424E70B4A568708066D84E" box="[981,1080,1121,1147]" pageId="13" pageNumber="14">stat. rev.</taxonomicNameLabel>
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<paragraph id="8BFF287A14424E70B5506892804DD8A8" blockId="13.[508,1080,1120,1182]" box="[544,1043,1155,1182]" pageId="13" pageNumber="14">
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<figureCitation id="137B34FF14424E70B550689286DED8A8" box="[544,640,1155,1181]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="13" pageNumber="14">Figs 1H</figureCitation>
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,
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<figureCitation id="137B34FF14424E70B5FD689286EED8A8" box="[653,688,1155,1181]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="13" pageNumber="14">2E</figureCitation>
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,
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<figureCitation id="137B34FF14424E70B5CC68928681D8A8" box="[700,735,1155,1181]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="13" pageNumber="14">3E</figureCitation>
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,
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<figureCitation id="137B34FF14424E70B59B6892874CD8A8" box="[747,786,1155,1181]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="13" pageNumber="14">4H</figureCitation>
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,
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<figureCitation id="137B34FF14424E70B46E68958772D8AB" box="[798,812,1156,1182]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="13" pageNumber="14">7</figureCitation>
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A–B, 9, 10;
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<tableCitation id="C6C21DC114424E70B4CC6892804DD8A8" box="[956,1043,1155,1181]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="13" pageNumber="14" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
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</paragraph>
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<treatmentCitation id="0AE10E6B14424E70B7CD68D486F9D8EA" author="Guenee A." box="[189,679,1221,1248]" page="298" pageId="13" pageNumber="14" year="1845">
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<taxonomicName id="4C4053F914424E70B7CD68D486F9D8EA" ID-CoL="B53V2" authority="Guenee, 1845 a: 298" authorityName="Guenee" authorityPageNumber="298" authorityYear="1845" box="[189,679,1221,1248]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frigidana">
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<emphasis id="B934F46814424E70B7CD68D485EED8EA" box="[189,432,1221,1247]" italics="true" pageId="13" pageNumber="14">Eupoecilia frigidana</emphasis>
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<bibRefCitation id="EFD1558B14424E70B6C868D486F9D8EA" author="Guenee A." box="[440,679,1221,1248]" pageId="13" pageNumber="14" pagination="297 - 344" refId="ref11382" refString="Guenee A. 1845 a. Essai sur une nouvelle classification des microlepidopteres et catalogue des especes europeennes connues jusqu'a jour. Annales de la Societe entomologique de France 2 (3): 297 - 344. Available from https: // biodiversitylibrary. org / page / 8310469 [accessed 16 Oct. 2017]." type="journal article" year="1845">Guenée, 1845a: 298</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BFF287A14424E70B7CD68F986DBD937" blockId="13.[189,871,1221,1353]" box="[189,645,1256,1283]" pageId="13" pageNumber="14">
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<treatmentCitation id="0AE10E6B14424E70B7CD68F986DCD937" author="Guenee A." box="[189,642,1256,1283]" page="66" pageId="13" pageNumber="14" year="1845">
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<taxonomicName id="4C4053F914424E70B7CD68F986DCD937" ID-CoL="96DGT" authority="Guenee, 1845 b: 66" authorityName="Guenee" authorityPageNumber="66" authorityYear="1845" box="[189,642,1256,1283]" class="Insecta" family="Tortricidae" genus="Cochylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="flavidana">
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<emphasis id="B934F46814424E70B7CD68F985C6D937" box="[189,408,1256,1282]" italics="true" pageId="13" pageNumber="14">Cochylis flavidana</emphasis>
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<bibRefCitation id="EFD1558B14424E70B6EF68F986DCD937" author="Guenee A." box="[415,642,1256,1283]" pageId="13" pageNumber="14" refId="ref11440" refString="Guenee A. 1845 b. Europaeorum Microlepidopterorum Index Methodicus, sive Pyrales, Tortrices, Tineae et Alucitae Linnaei, secundum novum naturalemque Ordinem dispositae, Nominibus Genuinis restitutis, Synomimia accurate elucidata, Locis indicatis, novisque Speciebus aut Larvis brevi descriptis. Pars prima: Sistens Tortrices, Phycidas, Crambidas, Tinearumque initiumaut Larvis brevi descriptis. Pars prima: Sistens Tortrices, Phycidas, Crambidas, Tinearumque initium. Roret, Paris, France." type="book" year="1845">Guenée, 1845b: 66</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BFF287A14424E70B7CD691A86CCD910" blockId="13.[189,871,1221,1353]" box="[189,658,1291,1318]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB500F5414424E70B7CD691A86CCD910" box="[189,658,1291,1318]" pageId="13" pageNumber="14">
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<treatmentCitation id="0AE10E6B14424E70B7CD691A86D1D910" author="Guenee A." box="[189,655,1291,1318]" page="67" pageId="13" pageNumber="14" year="1845">
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<taxonomicName id="4C4053F914424E70B7CD691A86D1D910" ID-CoL="95XDW" authority="Guenee, 1845 b: 67" authorityName="Guenee" authorityPageNumber="67" authorityYear="1845" box="[189,655,1291,1318]" class="Insecta" family="Tortricidae" genus="Aphelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sulphurana">
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<emphasis id="B934F46814424E70B7CD691A85F8D910" box="[189,422,1291,1317]" italics="true" pageId="13" pageNumber="14">Aphelia sulphurana</emphasis>
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<bibRefCitation id="EFD1558B14424E70B6DD691A86D1D910" author="Guenee A." box="[429,655,1291,1318]" pageId="13" pageNumber="14" refId="ref11440" refString="Guenee A. 1845 b. Europaeorum Microlepidopterorum Index Methodicus, sive Pyrales, Tortrices, Tineae et Alucitae Linnaei, secundum novum naturalemque Ordinem dispositae, Nominibus Genuinis restitutis, Synomimia accurate elucidata, Locis indicatis, novisque Speciebus aut Larvis brevi descriptis. Pars prima: Sistens Tortrices, Phycidas, Crambidas, Tinearumque initiumaut Larvis brevi descriptis. Pars prima: Sistens Tortrices, Phycidas, Crambidas, Tinearumque initium. Roret, Paris, France." type="book" year="1845">Guenée, 1845b: 67</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BFF287A14424E70B7CD693F8739D97C" blockId="13.[189,871,1221,1353]" box="[189,871,1326,1353]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB500F5414424E70B7CD693F8739D97C" box="[189,871,1326,1353]" pageId="13" pageNumber="14">
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<treatmentCitation id="0AE10E6B14424E70B7CD693F86FAD97D" box="[189,676,1326,1353]" pageId="13" pageNumber="14">
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||
<taxonomicName id="4C4053F914424E70B7CD693F86FAD97D" ID-CoL="96F5C" authority="Milliere, 1865: 167" authorityName="Milliere" authorityPageNumber="167" authorityYear="1865" box="[189,676,1326,1353]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="andorrana">
|
||
<emphasis id="B934F46814424E70B7CD693F85E9D97D" box="[189,439,1326,1352]" italics="true" pageId="13" pageNumber="14">Conchylis andorrana</emphasis>
|
||
Millière, 1865: 167
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, pl. 69, figs 4–5.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B7CD696087A3D9BF" blockId="13.[189,1021,1392,1454]" box="[189,1021,1392,1418]" pageId="13" pageNumber="14">
|
||
<treatmentCitationGroup id="AB500F5414424E70B7CD696087A3D9BF" box="[189,1021,1392,1418]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C4053F914424E70B7CD696085F1D9BF" box="[189,431,1392,1418]" class="Insecta" family="Tortricidae" genus="Hysterosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F46814424E70B7CD696085F1D9BF" box="[189,431,1392,1418]" italics="true" pageId="13" pageNumber="14">Hysterosia frigidana</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE10E6B14424E70B6BD696186ECD9BF" author="Razowski J." box="[461,690,1392,1418]" page="73" pageId="13" pageNumber="14" year="1970">
|
||
<bibRefCitation id="EFD1558B14424E70B6BD696186ECD9BF" author="Razowski J." box="[461,690,1392,1418]" pageId="13" pageNumber="14" pagination="3" refId="ref12243" refString="Razowski J. 1970. Cochylidae. In: Amsel H. G., Gregor F. & Reisser H. (eds) Microlepidoptera Palaearctica 3. Georg Fromme, Vienna, Austria." type="book chapter" year="1970">Razowski 1970: 73</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 29, fig. 3, pl. 112, fig. 3.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B7CD698286B5D998" blockId="13.[189,1021,1392,1454]" box="[189,747,1427,1454]" pageId="13" pageNumber="14">
|
||
<treatmentCitationGroup id="AB500F5414424E70B7CD698286B5D998" box="[189,747,1427,1454]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C4053F914424E70B7CD69828596D998" ID-CoL="4GX97" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[189,456,1427,1453]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F46814424E70B7CD69828596D998" box="[189,456,1427,1453]" italics="true" pageId="13" pageNumber="14">Phtheochroa frigidana</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE10E6B14424E70B695698286B8D998" author="Razowski J." box="[485,742,1427,1453]" page="103" pageId="13" pageNumber="14" year="1991">
|
||
<bibRefCitation id="EFD1558B14424E70B695698286B8D998" author="Razowski J." box="[485,742,1427,1453]" pageId="13" pageNumber="14" pagination="99 - 162" refId="ref12279" refString="Razowski J. 1991 a. The catalogue of the species of Tortricidae (Lepidoptera). Part I: Palaearctic Chlidanothinae and Tortricinae: Cochylini, Tortricini, Ceracini, Cnephasiini. Acta Zoologica Cracoviensia 34 (1): 99 - 162." type="journal article" year="1991">Razowski 1991a: 103</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B7CD69C486A7D9DA" blockId="13.[189,761,1492,1519]" box="[189,761,1492,1519]" pageId="13" pageNumber="14">
|
||
<treatmentCitationGroup id="AB500F5414424E70B7CD69C486A7D9DA" box="[189,761,1492,1519]" pageId="13" pageNumber="14">
|
||
non
|
||
<taxonomicName id="4C4053F914424E70B78069C585ACD9DB" ID-CoL="BDJ49" box="[240,498,1492,1518]" class="Insecta" family="Tortricidae" genus="Aphelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sulphurosana">
|
||
<emphasis id="B934F46814424E70B78069C585ACD9DB" box="[240,498,1492,1518]" italics="true" pageId="13" pageNumber="14">Aphelia sulphurosana</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AE10E6B14424E70B57F69C586ABD9DA" author="Razowski J." box="[527,757,1492,1519]" page="73" pageId="13" pageNumber="14" year="1970">
|
||
<bibRefCitation id="EFD1558B14424E70B57F69C586ABD9DA" author="Razowski J." box="[527,757,1492,1519]" pageId="13" pageNumber="14" pagination="3" refId="ref12243" refString="Razowski J. 1970. Cochylidae. In: Amsel H. G., Gregor F. & Reisser H. (eds) Microlepidoptera Palaearctica 3. Georg Fromme, Vienna, Austria." type="book chapter" year="1970">Razowski 1970: 73</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114424E70B7CD6A07863ADB3B" pageId="13" pageNumber="14" type="diagnosis">
|
||
<paragraph id="8BFF287A14424E70B7CD6A078568DA05" blockId="13.[189,310,1558,1584]" box="[189,310,1558,1584]" pageId="13" pageNumber="14">
|
||
<heading id="D0B79F1614424E70B7CD6A078568DA05" bold="true" box="[189,310,1558,1584]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B934F46814424E70B7CD6A078568DA05" bold="true" box="[189,310,1558,1584]" pageId="13" pageNumber="14">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B7CD6A55863ADB3B" blockId="13.[189,1399,1604,1807]" pageId="13" pageNumber="14">
|
||
Externally, this species is practically indistinguishable from
|
||
<taxonomicName id="4C4053F914424E70B4FC6A548067DA6B" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[908,1081,1604,1630]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F46814424E70B4FC6A548067DA6B" box="[908,1081,1604,1630]" italics="true" pageId="13" pageNumber="14">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314424E70B3336A5480C1DA6A" box="[1091,1183,1605,1631]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The characters of the phallus and vesica easily separate this species from all other members of the group: the phallic process is straight, in contrast to the other species with a curved process; the vesica has a single long dorsal diverticulum, the other species have two diverticula. The paler fore- and hindwing upperside colour distinguishes
|
||
<taxonomicName id="4C4053F914424E70B6DF6AC38666DADE" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[431,568,1745,1771]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F46814424E70B6DF6AC38666DADE" box="[431,568,1745,1771]" italics="true" pageId="13" pageNumber="14">P. frigidana</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C4053F914424E70B50F6AC38778DADE" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[639,806,1745,1771]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F46814424E70B50F6AC38778DADE" box="[639,806,1745,1771]" italics="true" pageId="13" pageNumber="14">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4053F914424E70B42E6AC3805BDADE" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[862,1029,1746,1771]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
|
||
<emphasis id="B934F46814424E70B42E6AC3805BDADE" box="[862,1029,1746,1771]" italics="true" pageId="13" pageNumber="14">P. apenninana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314424E70B37B6AC38036DAD9" box="[1035,1128,1746,1772]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, both with deep yellow forewings and dark grey hindwings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114424E72B7CD6B278137DDCA" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BFF287A14424E70B7CD6B2785F4DB65" blockId="13.[189,426,1846,1872]" box="[189,426,1846,1872]" pageId="13" pageNumber="14">
|
||
<heading id="D0B79F1614424E70B7CD6B2785F4DB65" bold="true" box="[189,426,1846,1872]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B934F46814424E70B7CD6B2785F4DB65" bold="true" box="[189,426,1846,1872]" pageId="13" pageNumber="14">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B79A6B74850FDB4A" blockId="13.[234,337,1893,1919]" box="[234,337,1893,1919]" pageId="13" pageNumber="14">
|
||
<emphasis id="B934F46814424E70B79A6B74850FDB4A" bold="true" box="[234,337,1893,1919]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54FB96D814424E70B79A6B74850FDB4A" box="[234,337,1893,1919]" pageId="13" pageNumber="14" type="neotype">Neotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14424E70B7CD6B828129DBE4" blockId="13.[189,1399,1939,2001]" pageId="13" pageNumber="14">
|
||
<collectingCountry id="F35768EA14424E70B7CD6B82850DDB98" box="[189,339,1939,1965]" name="Andorra" pageId="13" pageNumber="14">ANDORRA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D46E3F314424E70B62F6B828529DB98" box="[351,375,1939,1965]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
|
||
, pinned, with partially spread wings, slightly worn, four labels: //
|
||
<collectingCountry id="F35768EA14424E70B3F96B828146DB98" box="[1161,1304,1939,1965]" name="Andorra" pageId="13" pageNumber="14">ANDORRA</collectingCountry>
|
||
<location id="8E9F7EA114424E70B2516B828557DBE5" LSID="urn:lsid:plazi:treatment:03E9996C14424E6CB68C68718652DE55:8E9F7EA114424E70B2516B828557DBE5" country="Andorra" latitude="42.545834" longLatPrecision="19" longitude="1.4202777" name="Port de Cabus" pageId="13" pageNumber="14">
|
||
Port
|
||
<collectorName id="26B54DAC14424E70B22B6B828557DBE5" pageId="13" pageNumber="14">de Cabús</collectorName>
|
||
</location>
|
||
,
|
||
<quantity id="4CB8859F14424E70B6676BA7852CDBE4" box="[279,370,1974,2001]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="13" pageNumber="14" unit="m" value="2290.0">
|
||
<elevation id="006DCF4914424E70B6676BA7852CDBE4" box="[279,370,1974,2001]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="13" pageNumber="14" unit="m" value="2290.0">2290 m</elevation>
|
||
</quantity>
|
||
<geoCoordinate id="EE744EBD14424E70B60B6BA5865EDBE4" box="[379,512,1972,2001]" degrees="1" direction="east" minutes="25" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="13" value="1.4202777">1°25′13′′ E</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE744EBD14424E70B57F6BA586FADBE4" box="[527,676,1972,2001]" degrees="42" direction="north" minutes="32" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="45" value="42.545834">42°32′45′′ N</geoCoordinate>
|
||
<date id="FFFE0EBA14424E70B5DD6BA7876DDBE5" box="[685,819,1974,2000]" pageId="13" pageNumber="14" value="2012-07-16">
|
||
<collectingDate id="EFBAF75214424E70B5DD6BA7876DDBE5" box="[685,819,1974,2000]" pageId="13" pageNumber="14" value="2012-07-16">16.vii.2012</collectingDate>
|
||
</date>
|
||
leg.
|
||
<collectorName id="26B54DAC14424E70B4016BA68791DBE4" box="[881,975,1975,2001]" pageId="13" pageNumber="14">Huemer</collectorName>
|
||
TLMF 2012-011 // BC TLMF
|
||
<collectorName id="26B54DAC14424E70B2396BA68129DBE4" box="[1353,1399,1975,2001]" pageId="13" pageNumber="14">Lep</collectorName>
|
||
</paragraph>
|
||
<caption id="DF3F78F214414E73B7CD6B748541DBDF" ID-DOI="http://doi.org/10.5281/zenodo.3838426" ID-Zenodo-Dep="3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1893,1919]" targetBox="[250,1346,269,1849]" targetPageId="14">
|
||
<paragraph id="8BFF287A14414E73B7CD6B748541DBDF" blockId="14.[189,1399,1893,2026]" pageId="14" pageNumber="15">
|
||
<emphasis id="B934F46814414E73B7CD6B748550DBB5" bold="true" box="[189,270,1893,1920]" pageId="14" pageNumber="15">Fig. 7.</emphasis>
|
||
Phalli with vesica everted of
|
||
<taxonomicName id="4C4053F914414E73B5F36B748747DB4A" authorityName="Stephens" authorityYear="1829" box="[643,793,1893,1919]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934F46814414E73B5F36B748747DB4A" box="[643,793,1893,1919]" italics="true" pageId="14" pageNumber="15">Phtheochroa</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B934F46814414E73B4106B7487C9DB4A" bold="true" box="[864,919,1893,1919]" pageId="14" pageNumber="15">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F914414E73B4DA6B7780A3DBB5" authority="(Guenee, 1845)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[938,1277,1893,1920]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="frigidana" status="stat. nov.">
|
||
<emphasis id="B934F46814414E73B4DA6B778066DB4A" box="[938,1080,1893,1919]" italics="true" pageId="14" pageNumber="15">P. frigidana</emphasis>
|
||
(Guenée, 1845)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314414E73B2796B77812CDBB5" box="[1289,1394,1894,1920]" pageId="14" pageNumber="15">stat. rev.</taxonomicNameLabel>
|
||
, Andorra, Pyrenees, neotype of
|
||
<taxonomicName id="4C4053F914414E73B54D6B988768DB96" authorityName="Guenee" authorityYear="1845" box="[573,822,1929,1955]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F46814414E73B54D6B988768DB96" box="[573,822,1929,1955]" italics="true" pageId="14" pageNumber="15">Eupoecilia frigidana</emphasis>
|
||
</taxonomicName>
|
||
. –
|
||
<emphasis id="B934F46814414E73B4146B9987C3DB97" bold="true" box="[868,925,1928,1954]" pageId="14" pageNumber="15">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F914414E73B4C16B9B803FDB96" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[945,1121,1929,1955]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F46814414E73B4C16B9B803FDB96" box="[945,1121,1929,1955]" italics="true" pageId="14" pageNumber="15">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314414E73B31E6B98808EDB96" box="[1134,1232,1929,1955]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF287A14404E72B7CD6D1A801CDD7C" blockId="15.[189,1398,267,329]" pageId="15" pageNumber="16">
|
||
08349” [green label] // Gen. prep.
|
||
<specimenCount id="9D46E3F314404E72B5186D1D8620DD13" box="[616,638,268,294]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
|
||
1/
|
||
<date id="FFFE0EBA14404E72B5EB6D1A874BDD10" box="[667,789,267,293]" pageId="15" pageNumber="16" value="2012-07-16">
|
||
<collectingDate id="EFBAF75214404E72B5EB6D1A874BDD10" box="[667,789,267,293]" pageId="15" pageNumber="16" value="2012-07-16">16.7.2012</collectingDate>
|
||
</date>
|
||
[genitalia slide number] //
|
||
<typeStatus id="54FB96D814404E72B3176D1A80B3DD10" box="[1127,1261,267,293]" pageId="15" pageNumber="16" type="neotype">NEOTYPE</typeStatus>
|
||
<taxonomicName id="4C4053F914404E72B3876D1A875DDD7C" authority="Guenee, 1845 Zlatkov & Huemer, 2017" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F46814404E72B3876D1A8574DD7C" italics="true" pageId="15" pageNumber="16">Eupoecilia frigidana</emphasis>
|
||
Guenée, 1845 Zlatkov & Huemer, 2017
|
||
</taxonomicName>
|
||
des. [red label] // (
|
||
<collectionCode id="ED51B0BF14404E72B4916D3E8068DD7C" box="[993,1078,303,329]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="15" pageNumber="16">TLMF</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B79A6D6185F8DDBF" blockId="15.[234,422,368,394]" box="[234,422,368,394]" pageId="15" pageNumber="16">
|
||
<emphasis id="B934F46814404E72B79A6D6185F8DDBF" bold="true" box="[234,422,368,394]" pageId="15" pageNumber="16">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6D8E8137DDCA" blockId="15.[189,1398,414,512]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="F35768EA14404E72B7CD6D8E856BDD8C" box="[189,309,415,441]" name="France" pageId="15" pageNumber="16">FRANCE</collectingCountry>
|
||
:
|
||
<specimenCount id="9D46E3F314404E72B6306D8E8533DD8C" box="[320,365,415,441]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
|
||
, Ariège, Teilhet, D119 riv. Hers,
|
||
<date id="FFFE0EBA14404E72B59E6D8F87DFDD8C" box="[750,897,414,441]" pageId="15" pageNumber="16" value="2002-07-26">26 Jul. 2002</date>
|
||
, J. Nel leg. (
|
||
<collectionCode id="ED51B0BF14404E72B3616D8E803DDD8C" box="[1041,1123,415,441]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="15" pageNumber="16">TLMF</collectionCode>
|
||
) (slide 14258 JN);
|
||
<specimenCount id="9D46E3F314404E72B2346D8E812FDD8C" box="[1348,1393,415,441]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
|
||
, Pyrénées Orientales, Formiguères, le Galbe,
|
||
<quantity id="4CB8859F14404E72B5B46DD38742DDE9" box="[708,796,450,476]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="16" unit="m" value="1700.0">1700 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA14404E72B4176DD387A9DDE9" box="[871,1015,450,476]" pageId="15" pageNumber="16" value="1993-08-03">3 Aug. 1993</date>
|
||
, J. Nel leg. (
|
||
<collectionCode id="ED51B0BF14404E72B3F66DD38086DDE9" box="[1158,1240,450,476]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="15" pageNumber="16">TLMF</collectionCode>
|
||
) (slide 01646 JN);
|
||
<specimenCount id="9D46E3F314404E72B7876DF4857BDDCA" box="[247,293,485,511]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
|
||
, Pyrénées Orientales, Col de Puymorens,
|
||
<quantity id="4CB8859F14404E72B4616DF48734DDCA" box="[785,874,485,512]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="15" pageNumber="16" unit="m" value="1950.0">1950 m</quantity>
|
||
a.s.l.,
|
||
<date id="FFFE0EBA14404E72B4C76DF4800EDDCA" box="[951,1104,485,511]" pageId="15" pageNumber="16" value="2003-06-30">30 Jun. 2003</date>
|
||
, J. Junnilainen leg. (
|
||
<collectionCode id="ED51B0BF14404E72B2316DF48102DDCA" box="[1345,1372,485,511]" pageId="15" pageNumber="16">JJ</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114404E72B7CD6E378017DA8E" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BFF287A14404E72B7CD6E378535DE75" blockId="15.[189,363,550,576]" box="[189,363,550,576]" pageId="15" pageNumber="16">
|
||
<heading id="D0B79F1614404E72B7CD6E378535DE75" bold="true" box="[189,363,550,576]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B934F46814404E72B7CD6E378535DE75" bold="true" box="[189,363,550,576]" pageId="15" pageNumber="16">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B79A6E4485C1DE5A" blockId="15.[234,415,597,623]" box="[234,415,597,623]" pageId="15" pageNumber="16">
|
||
<emphasis id="B934F46814404E72B79A6E448574DE5A" bold="true" box="[234,298,597,623]" pageId="15" pageNumber="16">Male</emphasis>
|
||
(
|
||
<figureCitation id="137B34FF14404E72B64A6E4485CADE5A" box="[314,404,597,623]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="15" pageNumber="16">Fig. 1H</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6E958097DEF4" blockId="15.[189,1398,643,705]" pageId="15" pageNumber="16">HEAD. Most scales from frons and vertex missing, remaining white. Labial palps long, pointed anterad, with white scales dorsally and ochreous laterally. Antennae filiform, with creamy scales.</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6EF885C5DFE0" blockId="15.[189,1399,744,981]" pageId="15" pageNumber="16">
|
||
THORAX. Dorsally covered with white scales, ventrally with whitish scales. Tegula whitish, laterally with ochreous scales. Forewing length
|
||
<quantity id="4CB8859F14404E72B5296F1A86A6DF13" box="[601,760,779,806]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.2" metricValueMin="8.8" pageId="15" pageNumber="16" unit="mm" value="9.5" valueMax="10.2" valueMin="8.8">8.8–10.2 mm</quantity>
|
||
(x =9.6, n=4). Forewing long, narrow, with pointed apex, upperside ground colour creamy, more intense yellowish at base and along proximal half of costa; no pattern recognizable in studied specimens, but a specimen figured in Millière (1865) has rust dorsal blotch and subterminal fascia. Underside dark grey, with scattered white scales, especially in apical area. Cilia whitish, with darker basal line. Hindwing upperside pale grey, underside white, with scattered grey scales. Cilia white.
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6FEC8524D822" blockId="15.[189,378,1021,1047]" box="[189,378,1021,1047]" pageId="15" pageNumber="16">ABDOMEN. Grey.</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD682F8546D9A0" blockId="15.[189,1399,1086,1429]" pageId="15" pageNumber="16">
|
||
GENITALIA (
|
||
<figureCitation id="137B34FF14404E72B635682F85FBD86D" box="[325,421,1086,1112]" captionStart="Fig" captionStartId="5.[189,232,1813,1839]" captionTargetBox="[230,1355,468,1774]" captionTargetId="figure@5.[189,1398,417,1778]" captionTargetPageId="5" captionText="Fig. 2. Male genitalia (without phalli) of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov. A. Bulgaria, Rila Mts. B. Republic of Macedonia, Korab Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838416" httpUri="https://zenodo.org/record/3838416/files/figure.png" pageId="15" pageNumber="16">Fig. 2E</figureCitation>
|
||
). Curvature of valva prominent. Rest of valva, uncus and socii resembles those of
|
||
<taxonomicName id="4C4053F914404E72B7AF687385D4D84E" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[223,394,1121,1147]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="schawerdae">
|
||
<emphasis id="B934F46814404E72B7AF687385D4D84E" box="[223,394,1121,1147]" italics="true" pageId="15" pageNumber="16">P. schawerdae</emphasis>
|
||
</taxonomicName>
|
||
. Transtilla of
|
||
<typeStatus id="54FB96D814404E72B546687386CDD849" box="[566,659,1122,1148]" pageId="15" pageNumber="16" type="neotype">neotype</typeStatus>
|
||
(
|
||
<figureCitation id="137B34FF14404E72B5D66870875FD84E" box="[678,769,1121,1147]" captionStart="Fig" captionStartId="6.[188,231,1807,1833]" captionTargetBox="[191,1389,1348,1764]" captionTargetId="figure@6.[188,1397,1344,1765]" captionTargetPageId="6" captionText="Fig. 3. Transtilla of Phtheochroa spp. A–B. P. schawerdae (Rebel, 1908) comb. nov., Bulgaria, Rila Mts. – C. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype. – E. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – F. P. cantabriana sp. nov., Spain, Picos de Europa National Park, paratype. Scale bar = 250 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838418" httpUri="https://zenodo.org/record/3838418/files/figure.png" pageId="15" pageNumber="16">Fig. 3E</figureCitation>
|
||
) short and wide at base, narrow and round apically, but in other specimens with angular apical part. Phallus (
|
||
<figureCitation id="137B34FF14404E72B419689587F1D8AA" box="[873,943,1156,1183]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
|
||
A–B) almost straight, with very large, straight, slender, blunt medioventral process (
|
||
<figureCitation id="137B34FF14404E72B59268B9871AD8F7" box="[738,836,1192,1218]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="15" pageNumber="16">Fig. 4H</figureCitation>
|
||
). Vesica with ovoid part protruded to the right bearing anteroventrally directed cornutus. Long, slender, bent at basal third, anterodorsally directed diverticulum with cornutus emerges from the left dorsal part of vesica. Gonopore located dorsally, at base of diverticulum, with sclerotized plate and collar-like fold expanding laterally. Cornuti unequal, ventral one shorter, thicker and curved, dorsal one long, slender, sharp-tipped and only slightly curved. Acanthae present in small area on dorsal part, near gonopore, and scarcely around attachment areas of cornuti.
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B79A69AC851BD9E2" blockId="15.[234,325,1469,1495]" box="[234,325,1469,1495]" pageId="15" pageNumber="16">
|
||
<emphasis id="B934F46814404E72B79A69AC851BD9E2" bold="true" box="[234,325,1469,1495]" pageId="15" pageNumber="16">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD69FA85A8DA1C" blockId="15.[189,1398,1515,1577]" pageId="15" pageNumber="16">
|
||
A drawing by
|
||
<bibRefCitation id="EFD1558B14404E72B61569FA8677DA30" author="Razowski J." box="[357,553,1515,1541]" pageId="15" pageNumber="16" pagination="3" refId="ref12243" refString="Razowski J. 1970. Cochylidae. In: Amsel H. G., Gregor F. & Reisser H. (eds) Microlepidoptera Palaearctica 3. Georg Fromme, Vienna, Austria." type="book chapter" year="1970">Razowski (1970</bibRefCitation>
|
||
: pl. 112, fig. 3) may represent the female genitalia of a specimen of this species from the Pyrenees.
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6A41861FDA5F" blockId="15.[189,577,1615,1642]" box="[189,577,1615,1642]" pageId="15" pageNumber="16">
|
||
<heading id="D0B79F1614404E72B7CD6A418525DA5F" box="[189,379,1616,1642]" pageId="15" pageNumber="16">
|
||
<emphasis id="B934F46814404E72B7CD6A418525DA5F" bold="true" box="[189,379,1616,1642]" pageId="15" pageNumber="16">Molecular data</emphasis>
|
||
</heading>
|
||
(
|
||
<tableCitation id="C6C21DC114404E72B6FB6A5E85BADA5F" box="[395,484,1615,1642]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionTargetPageId="7" captionText="Table 1. Minimal uncorrected COI p-distances between different taxa of the P. frigidana s. lat. species complex." httpUri="http://table.plazi.org/id/DF3F78F214484E7AB7CD6D1D8574DD7D" pageId="15" pageNumber="16" tableUuid="DF3F78F214484E7AB7CD6D1D8574DD7D">Table 1</tableCitation>
|
||
,
|
||
<figureCitation id="137B34FF14404E72B6826A5E8669DA5F" box="[498,567,1615,1642]" captionStart="Fig" captionStartId="18.[189,232,1930,1956]" captionTargetBox="[190,1397,1281,1884]" captionTargetId="figure@18.[172,1412,1251,1902]" captionTargetPageId="18" captionText="Fig. 9. Maximum likelihood tree (built with MEGA6) of cytochrome c oxidase subunit I (COI) barcode fragments. Values at the nodes are bootstrap support values based on 500 replicates." figureDoi="http://doi.org/10.5281/zenodo.3838430" httpUri="https://zenodo.org/record/3838430/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6A6E8017DA8E" blockId="15.[189,1399,1662,1724]" pageId="15" pageNumber="16">
|
||
BIN URI: BOLD:ACA9598. The intraspecific divergence of the barcode region is unknown (n =1). The minimum distance to the nearest neighbour,
|
||
<taxonomicName id="4C4053F914404E72B5B66AB3872CDA8E" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[710,882,1697,1723]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
|
||
<emphasis id="B934F46814404E72B5B66AB3872CDA8E" box="[710,882,1697,1723]" italics="true" pageId="15" pageNumber="16">P. cantabriana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207491314404E72B40A6AB3878ADA89" box="[890,980,1698,1724]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is 3.69%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF114404E72B7CD6AF285BDDBA0" pageId="15" pageNumber="16" type="biology_ecology">
|
||
<paragraph id="8BFF287A14404E72B7CD6AF2857EDAC8" blockId="15.[189,288,1763,1789]" box="[189,288,1763,1789]" pageId="15" pageNumber="16">
|
||
<heading id="D0B79F1614404E72B7CD6AF2857EDAC8" bold="true" box="[189,288,1763,1789]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B934F46814404E72B7CD6AF2857EDAC8" bold="true" box="[189,288,1763,1789]" pageId="15" pageNumber="16">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A14404E72B7CD6B0085BDDBA0" blockId="15.[189,1399,1809,1941]" pageId="15" pageNumber="16">
|
||
Preimaginal stages and larval host plant are unknown. According to the label data, the moths fly from the middle of July to the beginning of August. Guenée (1846) reported May and June. The habitat of the
|
||
<typeStatus id="54FB96D814404E72B7CD6B498544DB47" box="[189,282,1880,1906]" pageId="15" pageNumber="16" type="neotype">neotype</typeStatus>
|
||
was alpine meadows above the tree-line, but based on supplementary material this species also occurs at lower altitudes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3F78F2145F4E6DB7CD6B9B8658DBDE" ID-DOI="http://doi.org/10.5281/zenodo.3838428" ID-Zenodo-Dep="3838428" httpUri="https://zenodo.org/record/3838428/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1930,1956]" targetBox="[188,1400,281,1861]" targetPageId="16">
|
||
<paragraph id="8BFF287A145F4E6DB7CD6B9B8658DBDE" blockId="16.[189,1399,1930,2027]" pageId="16" pageNumber="17">
|
||
<emphasis id="B934F468145F4E6DB7CD6B9B8554DB91" bold="true" box="[189,266,1930,1956]" pageId="16" pageNumber="17">Fig. 8.</emphasis>
|
||
Female genitalia of
|
||
<taxonomicName id="4C4053F9145F4E6DB6886B9B86D0DB91" authorityName="Stephens" authorityYear="1829" box="[504,654,1930,1956]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934F468145F4E6DB6886B9B86D0DB91" box="[504,654,1930,1956]" italics="true" pageId="16" pageNumber="17">Phtheochroa</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B934F468145F4E6DB5BC6B9B86BFDB91" bold="true" box="[716,737,1930,1956]" pageId="16" pageNumber="17">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9145F4E6DB59F6B9A8063DB91" authority="(Rebel, 1908)" authorityName="Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[751,1085,1930,1956]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="schawerdae" status="comb. nov.">
|
||
<emphasis id="B934F468145F4E6DB59F6B9A87C8DB91" box="[751,918,1930,1956]" italics="true" pageId="16" pageNumber="17">P. schawerdae</emphasis>
|
||
(Rebel, 1908)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913145F4E6DB3346B9B8099DB91" box="[1092,1223,1930,1956]" pageId="16" pageNumber="17" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, Bulgaria, Rila Mts. –
|
||
<emphasis id="B934F468145F4E6DB67D6BBC857FDBF2" bold="true" box="[269,289,1965,1991]" pageId="16" pageNumber="17">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C4053F9145F4E6DB65E6BBF85ECDBF2" authority="Zlatkov & Huemer, 2017" authorityName="Zlatkov & Huemer" authorityYear="2017" box="[302,434,1965,1991]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
|
||
<emphasis id="B934F468145F4E6DB65E6BBF85ECDBF2" box="[302,434,1965,1991]" italics="true" pageId="16" pageNumber="17">P. alpinana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2074913145F4E6DB6C86BBC864ADBF2" box="[440,532,1965,1991]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, France, Alpes Maritimes, paratype. Arrow: ventral diverticulum of ductus bursae. Scale bar = 250 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35A7BF1145E4E6CB7CD6D1A878BDD6D" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="8BFF287A145E4E6CB7CD6D1A859ADD10" blockId="17.[189,452,267,293]" box="[189,452,267,293]" pageId="17" pageNumber="18">
|
||
<heading id="D0B79F16145E4E6CB7CD6D1A8508DD10" box="[189,342,267,293]" pageId="17" pageNumber="18">
|
||
<emphasis id="B934F468145E4E6CB7CD6D1A8508DD10" bold="true" box="[189,342,267,293]" pageId="17" pageNumber="18">Distribution</emphasis>
|
||
</heading>
|
||
(
|
||
<figureCitation id="137B34FF145E4E6CB6166D1A85E5DD10" box="[358,443,267,293]" captionStart="Fig" captionStartId="20.[189,232,1990,2016]" captionTargetBox="[189,1398,1296,1948]" captionTargetId="figure@20.[183,1404,1290,1954]" captionTargetPageId="20" captionText="Fig. 10. Distribution of the Phtheochroa frigidana s. lat. species complex based on examined material." figureDoi="http://doi.org/10.5281/zenodo.3838432" httpUri="https://zenodo.org/record/3838432/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFF287A145E4E6CB7CD6D2F878BDD6D" blockId="17.[189,981,318,344]" box="[189,981,318,344]" pageId="17" pageNumber="18">Pyrenees. All previous records from other mountains need revision.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A7BF1145E4E6CB7CD6D968652DE55" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="8BFF287A145E4E6CB7CD6D968571DD94" blockId="17.[189,303,391,417]" box="[189,303,391,417]" pageId="17" pageNumber="18">
|
||
<heading id="D0B79F16145E4E6CB7CD6D968571DD94" bold="true" box="[189,303,391,417]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B934F468145E4E6CB7CD6D968571DD94" bold="true" box="[189,303,391,417]" pageId="17" pageNumber="18">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BFF287A145E4E6CB7CD6DA88652DE55" blockId="17.[189,1398,441,608]" pageId="17" pageNumber="18">
|
||
The erroneous type locality of “
|
||
<collectingRegion id="4984E698145E4E6CB54C6DA886E4DDE6" box="[572,698,441,467]" country="Sweden" name="Dalarna" pageId="17" pageNumber="18">Dalecarlia</collectingRegion>
|
||
” in the original description of
|
||
<taxonomicName id="4C4053F9145E4E6CB35F6DAB80E5DDE6" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[1071,1211,441,467]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F468145E4E6CB35F6DAB80E5DDE6" box="[1071,1211,441,467]" italics="true" pageId="17" pageNumber="18">P. frigidana</emphasis>
|
||
</taxonomicName>
|
||
is a major hindrance in the interpretation of this taxon. Therefore, in order to maintain the stability of the nomenclature, we have designated a
|
||
<typeStatus id="54FB96D8145E4E6CB6B26E118641DE2F" box="[450,543,512,538]" pageId="17" pageNumber="18" type="neotype">neotype</typeStatus>
|
||
for
|
||
<taxonomicName id="4C4053F9145E4E6CB5226DEE8718DE2C" authorityName="Guenee, 1845 Zlatkov & Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[594,838,511,537]" class="Insecta" family="Tortricidae" genus="Eupoecilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="frigidana">
|
||
<emphasis id="B934F468145E4E6CB5226DEE8718DE2C" box="[594,838,511,537]" italics="true" pageId="17" pageNumber="18">Eupoecilia frigidana</emphasis>
|
||
</taxonomicName>
|
||
in accordance with past usage of this name for populations from the Pyrenees (
|
||
<bibRefCitation id="EFD1558B145E4E6CB5496E3386A9DE08" author="Razowski J." box="[569,759,546,573]" pageId="17" pageNumber="18" pagination="3" refId="ref12243" refString="Razowski J. 1970. Cochylidae. In: Amsel H. G., Gregor F. & Reisser H. (eds) Microlepidoptera Palaearctica 3. Georg Fromme, Vienna, Austria." type="book chapter" year="1970">Razowski 1970</bibRefCitation>
|
||
), with
|
||
<taxonomicName id="4C4053F9145E4E6CB43A6E328079DE08" authorityName="Guenee" authorityYear="1845" box="[842,1063,547,573]" class="Insecta" family="Tortricidae" genus="Cochylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavidana">
|
||
<emphasis id="B934F468145E4E6CB43A6E328079DE08" box="[842,1063,547,573]" italics="true" pageId="17" pageNumber="18">Cochylis flavidana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4053F9145E4E6CB3146E32815BDE08" authorityName="Milliere" authorityYear="1865" box="[1124,1285,547,573]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andorrana">
|
||
<emphasis id="B934F468145E4E6CB3146E32815BDE08" box="[1124,1285,547,573]" italics="true" pageId="17" pageNumber="18">C. andorrana</emphasis>
|
||
</taxonomicName>
|
||
as junior synonyms (
|
||
<bibRefCitation id="EFD1558B145E4E6CB6336E57865EDE55" author="Razowski J." box="[323,512,582,608]" pageId="17" pageNumber="18" refId="ref12358" refString="Razowski J. 2002. Tortricidae of Europe. Volume 1. Tortricinae and Chlidanothinae. Frantisek Slamka, Bratislava, Slovakia." type="book" year="2002">Razowski 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |