162 lines
18 KiB
XML
162 lines
18 KiB
XML
<document id="1B5D0E20E9D1B015FC905BE0EC6D4575" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFCF1F5FFF50FC63FEBD8037" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax garciana Leon 1939" docType="treatment" docVersion="6" lastPageNumber="49" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="49" updateTime="1696268944350" updateUser="diego">
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<mods:titleInfo id="7E54634C38D70388224976CCE1CBCB9D">
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<mods:title id="98EBB6A42FB6B31DEB0AC3850D0B932E">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:name id="BF707C782CFC4B88F431E78F141F5F32" type="personal">
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<mods:namePart id="5C4DA8EE084C2E4B25D4BB98DF2AC469">Henderson, Andrew</mods:namePart>
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<mods:typeOfResource id="5B00AE0A81726D0C2B95D74725413A42">text</mods:typeOfResource>
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<mods:title id="D5306FD391C4FF0BD4BDC8D0BA34432C">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5A108EB4ED29E470B1EF316B6D314D33">
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<mods:date id="B720DB24C8B367B48C1F1CF3824233A1">2023</mods:date>
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<mods:number id="6AA05FE0AA01B8947B6B6063BDB261CD">2023-09-19</mods:number>
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<mods:number id="89758B95CB278928F13B387DDD697F45">614</mods:number>
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<mods:start id="76F2CA7EC807BA1EDD320F017594903E">1</mods:start>
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<mods:url id="E69F61F59CB37BF9A3F432DE94F5C72A">http://dx.doi.org/10.11646/phytotaxa.614.1.1</mods:url>
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<mods:classification id="BCE9AAE00D95E6C641790C4D5D26724C">journal article</mods:classification>
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<mods:identifier id="4135699A30476D8F4A84D88B294535AB" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
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<mods:identifier id="46513F55EA87A742626AB6AD573BBB62" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="50ABD0F4A92A9F89AEF679EDC28461A0" type="Zenodo-Dep">8389307</mods:identifier>
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</mods:mods>
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<treatment id="03B387DAFFCF1F5FFF50FC63FEBD8037" LSID="urn:lsid:plazi:treatment:03B387DAFFCF1F5FFF50FC63FEBD8037" httpUri="http://treatment.plazi.org/id/03B387DAFFCF1F5FFF50FC63FEBD8037" lastPageNumber="49" pageId="48" pageNumber="49">
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<subSubSection id="C3006547FFCF1F5FFF50FC63FD558BEF" box="[136,684,985,1013]" pageId="48" pageNumber="49" type="nomenclature">
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<paragraph id="8BA536CCFFCF1F5FFF50FC63FD558BEF" blockId="48.[136,1451,985,1085]" box="[136,684,985,1013]" pageId="48" pageNumber="49">
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<emphasis id="B96EEADEFFCF1F5FFF50FC63FD558BEF" bold="true" box="[136,684,985,1013]" pageId="48" pageNumber="49">
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1.15.
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<taxonomicName id="4C1A4D4FFFCF1F5FFF1FFC63FD5F8BEF" ID-CoL="WLYG" authority="Leon (1939: 143)" authorityName="Leon" authorityPageNumber="143" authorityYear="1939" box="[199,678,985,1013]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="garciana">
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<emphasis id="B96EEADEFFCF1F5FFF1FFC63FE288BEF" bold="true" box="[199,465,986,1013]" italics="true" pageId="48" pageNumber="49">Coccothrinax garciana</emphasis>
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<bibRefCitation id="EF8B4B3DFFCF1F5FFE0FFC60FD658BEF" author="Leon" box="[471,668,985,1012]" pageId="48" pageNumber="49" pagination="107 - 156" refId="ref62742" refString="Leon (1939) Contribucion al estudio de las palmas de Cuba. III. Memorias de la Sociedad Cubana de Historia Natural " Felipe Poey " 13: 107 - 156." type="journal article" year="1939">León (1939: 143</bibRefCitation>
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)
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</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFCF1F5FFD6BFC62FE138C27" pageId="48" pageNumber="49" type="materials_examined">
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<paragraph id="8BA536CCFFCF1F5FFD6BFC62FE138C27" blockId="48.[136,1451,985,1085]" pageId="48" pageNumber="49">
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<materialsCitation id="3B723C91FFCF1F5FFD6BFC62FE138C27" collectingDate="1932-03-17" collectionCode="HAC, A, BH, GH, MICH, MT" collectorName="Fr. Leon" country="Cuba" location="Cerrito del Fraile" pageId="48" pageNumber="49" specimenCount="2" stateProvince="Holguin" typeStatus="lectotype">
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<typeStatus id="54A1886EFFCF1F5FFD6BFC62FCDA8BEE" box="[691,803,987,1013]" pageId="48" pageNumber="49">Lectotype</typeStatus>
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(designated by
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<bibRefCitation id="EF8B4B3DFFCF1F5FFC0BFC62FBAF8BEF" author="Moya, C." box="[979,1110,986,1013]" pageId="48" pageNumber="49" pagination="1 - 63" refId="ref63249" refString="Moya, C. (2020) An annotated checklist of Cuban palms 2. Coccothrinax, Pt. 1: 1816 - 1939. Nomenclature, typification, and distribution. PalmArbor 4: 1 - 63." type="journal article" year="2020">Moya 2020</bibRefCitation>
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):—
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<collectingCountry id="F30D765CFFCF1F5FFB58FC63FB2B8BEF" box="[1152,1234,986,1012]" name="Cuba" pageId="48" pageNumber="49">CUBA</collectingCountry>
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.
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<collectingRegion id="49DEF82EFFCF1F5FFB03FC63FACE8BEF" box="[1243,1335,986,1012]" country="Cuba" name="Holguin" pageId="48" pageNumber="49">Holguín</collectingRegion>
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(Oriente),
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<location id="8EC56017FFCF1F5FFF50FC47FEB78C03" LSID="urn:lsid:plazi:treatment:03B387DAFFCF1F5FFF50FC63FEBD8037:8EC56017FFCF1F5FFF50FC47FEB78C03" box="[136,334,1022,1048]" country="Cuba" name="Cerrito del Fraile" pageId="48" pageNumber="49" stateProvince="Holguin">Cerrito del Fraile</location>
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,
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<date id="FFA4100CFFCF1F5FFE82FC47FDF38C03" box="[346,522,1022,1048]" pageId="48" pageNumber="49" value="1932-03-17">
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<collectingDate id="EFE0E9E4FFCF1F5FFE82FC47FDF38C03" box="[346,522,1022,1048]" pageId="48" pageNumber="49" value="1932-03-17">17 March 1932</collectingDate>
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</date>
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,
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<emphasis id="B96EEADEFFCF1F5FFDCEFC46FD3D8C03" box="[534,708,1022,1049]" italics="true" pageId="48" pageNumber="49">
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<collectorName id="26EF531AFFCF1F5FFDCEFC46FD8F8C02" box="[534,630,1023,1049]" pageId="48" pageNumber="49">Fr. León</collectorName>
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15476
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</emphasis>
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(
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<typeStatus id="54A1886EFFCF1F5FFD0BFC47FCC58C03" box="[723,828,1022,1048]" pageId="48" pageNumber="49">lectotype</typeStatus>
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<collectionCode id="ED0BAE09FFCF1F5FFC9BFC47FC7F8C03" box="[835,902,1022,1048]" country="Cuba" httpUri="http://grbio.org/cool/cenz-te88" name="Instituto de Ecología y Sistemática" pageId="48" pageNumber="49" type="Herbarium">HAC</collectionCode>
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!, isolectotypes
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<collectionCode id="ED0BAE09FFCF1F5FFBF7FC47FBB38C03" box="[1071,1098,1022,1048]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="48" pageNumber="49" type="Herbarium">A</collectionCode>
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!,
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<collectionCode id="ED0BAE09FFCF1F5FFB84FC47FB728C03" box="[1116,1163,1022,1048]" country="USA" lsid="urn:lsid:biocol.org:col:15518" name="L. H. Bailey Hortorium, Cornell University" pageId="48" pageNumber="49" type="Herbarium">BH</collectionCode>
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!,
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<collectionCode id="ED0BAE09FFCF1F5FFB43FC47FB328C03" box="[1179,1227,1022,1048]" country="USA" lsid="urn:lsid:biocol.org:col:15631" name="Harvard University - Gray Herbarium" pageId="48" pageNumber="49" type="Herbarium">GH</collectionCode>
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!,
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<collectionCode id="ED0BAE09FFCF1F5FFB04FC47FADC8C03" box="[1244,1317,1022,1048]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
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<emphasis id="B96EEADEFFCF1F5FFAF5FC46FAAA8C03" box="[1325,1363,1023,1048]" italics="true" pageId="48" pageNumber="49">
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<taxonomicNameLabel id="A25D57A5FFCF1F5FFAF5FC46FAAA8C03" box="[1325,1363,1023,1048]" pageId="48" pageNumber="49" rank="variety">n.v.</taxonomicNameLabel>
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</emphasis>
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,
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<collectionCode id="ED0BAE09FFCF1F5FFABAFC47FA528C03" box="[1378,1451,1022,1048]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
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image!,
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<collectionCode id="ED0BAE09FFCF1F5FFF3CFB9AFEF48C26" box="[228,269,1059,1085]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="48" pageNumber="49">MT</collectionCode>
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<emphasis id="B96EEADEFFCF1F5FFECCFB9AFECF8C27" box="[276,310,1059,1084]" italics="true" pageId="48" pageNumber="49">n.v</emphasis>
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., US!).
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<figureCitation id="13212A49FFCF1F5FFE57FB9BFE138C27" box="[399,490,1058,1084]" captionStart="PLATE 12" captionStartId="49.[136,214,1930,1951]" captionTargetBox="[151,1433,360,1904]" captionTargetId="figure-16@49.[150,1435,193,1906]" captionTargetPageId="49" captionText="PLATE 12. Coccothrinax garciana, Cuba, with stems covered with persistent dead leaves, giving a ‘skirt’ of dead leaves. (Image by Mayté Pernús Alvarez)." figureDoi="http://doi.org/10.5281/zenodo.8389381" httpUri="https://zenodo.org/record/8389381/files/figure.png" pageId="48" pageNumber="49">Plate 12</figureCitation>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFCF1F5FFF50FBD3FE718EBB" pageId="48" pageNumber="49" type="description">
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<paragraph id="8BA536CCFFCF1F5FFF50FBD3FE718EBB" blockId="48.[136,1452,1130,2093]" pageId="48" pageNumber="49">
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<emphasis id="B96EEADEFFCF1F5FFF50FBD3FF298C9F" bold="true" box="[136,208,1130,1156]" pageId="48" pageNumber="49">Stems</emphasis>
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length and diameter not recorded, solitary.
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<emphasis id="B96EEADEFFCF1F5FFD6FFBD3FCF08C9F" bold="true" box="[695,777,1130,1156]" pageId="48" pageNumber="49">Leaves</emphasis>
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persistent, giving a ‘skirt’ of dead leaves covering the stem; leaf sheath fibers 2.2(1.8–2.7) mm diameter, stout, woody, loosely woven, the inner and outer layer combining at the apices to
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<taxonomicName id="4C1A4D4FFFCF1F5FFF2BFB0BFE938CD6" box="[243,362,1202,1229]" form="erect" pageId="48" pageNumber="49" rank="form">form erect</taxonomicName>
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, spine-like fibers; petioles 8.6(6.6–11.2) mm diameter just below the apex; palmans 4.0(2.0–7.2) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 24(21–27) per leaf, the middle ones 29.1(23.0–34.0) cm long and 3.1(2.0–4.0) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, reddish-brown center, without transverse veinlets.
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<emphasis id="B96EEADEFFCF1F5FFADAFA17FA528DD3" bold="true" box="[1282,1451,1454,1480]" pageId="48" pageNumber="49">Inflorescences</emphasis>
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erect amongst or above the leaves, with few partial inflorescences at apex of inflorescence; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 3(2–4); proximalmost rachillae straight, 6.6(4.4–9.0) cm long and 1.2(0.9–1.4) mm diameter in fruit; rachillae glabrous at or near anthesis; stamens 6; fruit pedicels 1.4(1.0–1.9) mm long;
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<emphasis id="B96EEADEFFCF1F5FFD74F987FD148E43" bold="true" box="[684,749,1598,1624]" pageId="48" pageNumber="49">fruits</emphasis>
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7.0(6.1–9.3) mm long and 7.4(6.1–8.7) mm diameter, purple; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFCF1F5FFF65F913FDF78EF3" pageId="48" pageNumber="49" type="distribution">
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<paragraph id="8BA536CCFFCF1F5FFF65F913FDF78EF3" blockId="48.[136,1452,1130,2093]" pageId="48" pageNumber="49">
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<emphasis id="B96EEADEFFCF1F5FFF65F913FDF88EDF" bold="true" box="[189,513,1706,1732]" pageId="48" pageNumber="49">Distribution and habitat:—</emphasis>
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Eastern
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<collectingCountry id="F30D765CFFCF1F5FFD83F913FD6F8EDF" box="[603,662,1706,1732]" name="Cuba" pageId="48" pageNumber="49">Cuba</collectingCountry>
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(
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<collectingRegion id="49DEF82EFFCF1F5FFD7AF913FCF88EDF" box="[674,769,1706,1732]" country="Cuba" name="Holguin" pageId="48" pageNumber="49">Holguín</collectingRegion>
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) (
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<figureCitation id="13212A49FFCF1F5FFCCEF913FC9F8EDF" box="[790,870,1706,1732]" captionStart="FIGURE 12" captionStartId="47.[136,229,1777,1799]" captionTargetBox="[155,1429,469,1747]" captionTargetId="figure-147@47.[150,1434,464,1752]" captionTargetPageId="47" captionText="FIGURE 12. Distribution of Coccothrinax fagildei, C. fragrans, C. garciana, and C. gonaivensis." figureDoi="http://doi.org/10.5281/zenodo.8389379" httpUri="https://zenodo.org/record/8389379/files/figure.png" pageId="48" pageNumber="49">Fig. 12</figureCitation>
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) in dry shrublands, dry hills, or barrens on serpentine soils at 140(120–160) m elevation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFCF1F5FFF65F94BFEBD8037" pageId="48" pageNumber="49" type="discussion">
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<paragraph id="8BA536CCFFCF1F5FFF65F94BFEBD8037" blockId="48.[136,1452,1130,2093]" pageId="48" pageNumber="49">
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<emphasis id="B96EEADEFFCF1F5FFF65F94BFE578F17" bold="true" box="[189,430,1778,1804]" pageId="48" pageNumber="49">Taxonomic notes:—</emphasis>
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As a preliminary species,
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<taxonomicName id="4C1A4D4FFFCF1F5FFD00F94BFC248F17" box="[728,989,1778,1804]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="garciana">
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<emphasis id="B96EEADEFFCF1F5FFD00F94BFC248F17" box="[728,989,1778,1804]" italics="true" pageId="48" pageNumber="49">Coccothrinax garciana</emphasis>
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</taxonomicName>
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has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is notable for its abaxial leaf segment surfaces with the hairs with reddish-brown centers. The presence of a ‘skirt’ of dead leaves persisting on the stem is scored from an image of plants from the
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<typeStatus id="54A1886EFFCF1F5FFE96F8E6FE878F62" box="[334,382,1887,1913]" pageId="48" pageNumber="49">type</typeStatus>
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locality in
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<bibRefCitation id="EF8B4B3DFFCF1F5FFE2FF8E6FD728F63" author="Moya, C." box="[503,651,1886,1913]" pageId="48" pageNumber="49" pagination="1 - 63" refId="ref63249" refString="Moya, C. (2020) An annotated checklist of Cuban palms 2. Coccothrinax, Pt. 1: 1816 - 1939. Nomenclature, typification, and distribution. PalmArbor 4: 1 - 63." type="journal article" year="2020">Moya (2020)</bibRefCitation>
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||
.
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||
<taxonomicName id="4C1A4D4FFFCF1F5FFD4BF8E7FC6C8F63" box="[659,917,1886,1912]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="garciana">
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<emphasis id="B96EEADEFFCF1F5FFD4BF8E7FC6C8F63" box="[659,917,1886,1912]" italics="true" pageId="48" pageNumber="49">Coccothrinax garciana</emphasis>
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</taxonomicName>
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appears similar to
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<taxonomicName id="4C1A4D4FFFCF1F5FFBB9F8E6FAED8F63" box="[1121,1300,1886,1913]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="pseudorigida">
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<emphasis id="B96EEADEFFCF1F5FFBB9F8E6FAED8F63" box="[1121,1300,1886,1913]" italics="true" pageId="48" pageNumber="49">C. pseudorigida</emphasis>
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</taxonomicName>
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. Both species also have exceptionally large spine-like leaf sheath fibers, relatively small leaves, erect, projecting inflorescences with few, distal partial inflorescences, and relatively short rachillae and large fruits.
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<taxonomicName id="4C1A4D4FFFCF1F5FFBCCF81FFAE38FDB" box="[1044,1306,1958,1984]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="garciana">
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<emphasis id="B96EEADEFFCF1F5FFBCCF81FFAE38FDB" box="[1044,1306,1958,1984]" italics="true" pageId="48" pageNumber="49">Coccothrinax garciana</emphasis>
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</taxonomicName>
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differs from
|
||
<taxonomicName id="4C1A4D4FFFCF1F5FFF50F872FEC78FFF" box="[136,318,1994,2021]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="pseudorigida">
|
||
<emphasis id="B96EEADEFFCF1F5FFF50F872FEC78FFF" box="[136,318,1994,2021]" italics="true" pageId="48" pageNumber="49">C. pseudorigida</emphasis>
|
||
</taxonomicName>
|
||
in its solitary stems and reddish-brown centers to the hairs on the abaxial surface of the leaves. Its sheath spines tend to dry almost black and its leaves tend to dry flat, as opposed to brown and curled, respectively, in
|
||
<emphasis id="B96EEADEFFCF1F5FFF50F7AAFEBD8037" box="[136,324,2066,2093]" italics="true" pageId="48" pageNumber="49">
|
||
<taxonomicName id="4C1A4D4FFFCF1F5FFF50F7AAFEB98037" box="[136,320,2066,2093]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="pseudorigida">C. pseudorigida</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |