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<document id="4CED6198812245BFDCE5D2AF639BD8CE" ID-CLB-Dataset="35780" ID-DOI="http://dx.doi.org/10.3897/zookeys.466.8047" ID-GBIF-Dataset="470bc8d8-7432-47cb-bae3-88617b4e91a9" ID-PMC="PMC4296484" ID-Pensoft-Pub="1313-2970-466-77" ID-PubMed="25610337" ID-ZooBank="19C20324B7A548EBBBA45D849697EA23" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-466-77" ModsDocOrigin="ZooKeys 466" ModsDocTitle="Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China" checkinTime="1451244919976" checkinUser="pensoft" docAuthor="Liu, Sulin, Shih, Chungkun &amp; Ren, Dong" docDate="2014" docId="F22B24A45E0C292FA7FB43781B3BB741" docLanguage="en" docName="ZooKeys 466: 77-94" docOrigin="ZooKeys 466" docSource="http://dx.doi.org/10.3897/zookeys.466.8047" docTitle="Exilibittacus foliaceus Liu, Shih &amp; Ren, 2014, sp. n." docType="treatment" docUuid="98FF3AC2-E493-45B7-B351-27B42685200A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="84" masterDocId="FF84FF8DFFDAFFD6F05B8E6A350B1F27" masterDocTitle="Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China" masterLastPageNumber="94" masterPageNumber="77" pageNumber="84" updateTime="1732699651032" updateUser="ExternalLinkService">
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<mods:title id="21C6DEDFC469474E0D6FAB7F2F944A8B">Four new species of hangingflies (Insecta, Mecoptera, Bittacidae) from the Middle Jurassic of northeastern China</mods:title>
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<mods:namePart id="01E03F64B8B69028D59D886FE0945741">Liu, Sulin</mods:namePart>
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<mods:namePart id="A791B1C641E3CABB74440964FB6055DD">Shih, Chungkun</mods:namePart>
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<mods:namePart id="6914E98BC1B4BD893B4A30064A897A30">Ren, Dong</mods:namePart>
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<mods:date id="50C47505D6B25538B81693026605CC67">2014</mods:date>
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<treatment id="F22B24A45E0C292FA7FB43781B3BB741" ID-GBIF-Taxon="152057235" LSID="urn:lsid:zoobank.org:act:98FF3AC2-E493-45B7-B351-27B42685200A" httpUri="http://treatment.plazi.org/id/F22B24A45E0C292FA7FB43781B3BB741" lastPageNumber="84" pageId="7" pageNumber="84" scope_class="Insecta" scope_family="Bittacidae" scope_order="Mecoptera">
<subSubSection id="680D1EA619E2C517CDA15549F603E487" pageId="7" pageNumber="84" type="multiple">
<paragraph id="3E6757945D910E8820C95FB0901FAA21" pageId="7" pageNumber="84">Taxon classification Animalia Mecoptera Bittacidae</paragraph>
</subSubSection>
<subSubSection id="A525413239FB872E9DC5B8E8D92CFA68" pageId="7" pageNumber="84" type="nomenclature">
<paragraph id="D0A9882C7DEBBD71C558300B839812DE" pageId="7" pageNumber="84">
<taxonomicName id="505291DBD242F37304EF141F5433B714" LSID="http://zoobank.org/98FF3AC2-E493-45B7-B351-27B42685200A" genus="Exilibittacus" lsidName="Exilibittacus foliaceus" pageId="7" pageNumber="84" rank="species" species="foliaceus">Exilibittacus foliaceus</taxonomicName>
<taxonomicNameLabel id="4A79F27A6932821946AC47F1A23DF208" pageId="7" pageNumber="84">sp. n.</taxonomicNameLabel>
Fig. 7
</paragraph>
</subSubSection>
<subSubSection id="1695AD62C6C003F0696CE9142120743A" pageId="7" pageNumber="84" type="etymology">
<paragraph id="34ED20FC88B2AC34DD9AF665D630331A" pageId="7" pageNumber="84">Etymology.</paragraph>
<paragraph id="54942BCBA04860905AD452A862A091FC" pageId="7" pageNumber="84">
The Latin word of
<normalizedToken id="D3ABB38AFEA0AA984D7168120226977F" originalValue="“foliaceus”">&quot;foliaceus&quot;</normalizedToken>
means folliform, referring to the shape of the wings like leaves.
</paragraph>
</subSubSection>
<subSubSection id="88C920735503198AD0AC1306839E7AC9" pageId="7" pageNumber="84" type="holotype">
<paragraph id="F112FE1A2C764E1FB481F4B60183518B" pageId="7" pageNumber="84">Holotype.</paragraph>
<paragraph id="EA17C299B216C277BA8FD30409781985" pageId="7" pageNumber="84">Female, CNU-MEC-NN2013010, in dorsal view. Body length 12.9 mm, forewing 11.7 mm long and 2.9 mm wide; hind wing 9.7 mm long and 2.6 mm wide.</paragraph>
</subSubSection>
<subSubSection id="5C26F38703EFA6A82F32DA98768858E4" pageId="7" pageNumber="84" type="horizon and locality">
<paragraph id="FC71BB18495D3292655A7726CF338801" pageId="7" pageNumber="84">Horizon and locality.</paragraph>
<paragraph id="95FD35CB78AD2BCBEF363866FC67193E" pageId="7" pageNumber="84">Jiulongshan Formation, late Middle Jurassic, Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.</paragraph>
</subSubSection>
<subSubSection id="AF48CDEF10B802A596152B61FEF5666B" pageId="7" pageNumber="84" type="diagnosis">
<paragraph id="219FC1FCAB6D733E46746517287A10CB" pageId="7" pageNumber="84">Diagnosis.</paragraph>
<paragraph id="772F32C28E5AD7AD6548BB729CB9A0A5" pageId="7" pageNumber="84">Forewing: pterostigmal crossveins (Pcv) absent, but 2A present. Hind wing: Rs with four branches and the bifurcation of Rs at the same level of the bifurcation of M.</paragraph>
</subSubSection>
<subSubSection id="24BF56669E17983F4374E994206A716E" pageId="7" pageNumber="84" type="description">
<paragraph id="F428AD3B59B766CBA47E9C767A7E3782" pageId="7" pageNumber="84">Description.</paragraph>
<paragraph id="347E5567C32432148D6B24D8ECD5AC77" pageId="7" pageNumber="84">A female holotype preserved in dorsal view. Antenna filiform, scape, pedicel and part of other antennomeres preserved. The vertex of the head raised. Legs not well-preserved, covered with short setae; the fifth tarsomere folded against the fourth, a claw present (Fig. 7A, B).</paragraph>
<caption id="84EADA62596EFE46D0D7CB89896E2EFE" pageId="7" pageNumber="84">
<paragraph id="4F3B1D478A110F810D42050368FD635C" pageId="7" pageNumber="84">
Figure 7.
<taxonomicName id="472B32F63E2F1380128A015A080A2177" genus="Exilibittacus" lsidName="Exilibittacus foliaceus" pageId="7" pageNumber="84" rank="species" species="foliaceus">Exilibittacus foliaceus</taxonomicName>
sp. n., holotype, CNU-MEC-NN2013010. A photo B line drawing C line drawing of left forewing D line drawing of left hind wing E photo of genitalia in dorsal view, under alcohol F line drawing of genitalia in dorsal view. Scale bars represent 3 mm in A, B, 2 mm in C, D, 0.5 mm in E, F. Abbreviations: T9, the ninth tergum; T10, the tenth tergum; c, cercus; spa, supraanale.
</paragraph>
</caption>
<paragraph id="C309AE70E1E4AD962C042206E3A622AF" pageId="7" pageNumber="84">
Forewing. The base of wings narrow, pterostigma slightly dark. Sc terminating at the anterior margin proximad of the R4+5 forking; one subcostal crossvein (Scv) between Sc and R1; R1 running straight through pterostigma, without sagging; one crossvein between R1 and R2+3; Rs with four branches, R4 slightly curved at beginning and then parallel with R5; one crossvein between R2+3 and R4 and one crossvein between R4 and R5; M with four branches, M3+4 forking far proximad of the bifurcation of M1+2; the 'bittacid
<normalizedToken id="EBFD9134987FE639B957607460CD4664" originalValue="cross">cross'</normalizedToken>
aligned and gently curved, posterior part of the 'bittacid
<normalizedToken id="E147AC3EC9ADA7254E3E5B5E858E4F77" originalValue="cross">cross'</normalizedToken>
reaching M3 distad of the M3+4 forking point; one crossvein between R5 and M1, one between M1 and M2 and one between M2 and M3; Cu1 and Cu2 almost parallel with two crossveins between them, one crossvein between M4 and Cu1; 1A and 2A simple and one crossvein between them; 1A reaching the posterior margin at the same level of the origination of Rs from R1; 2A reaching the posterior margin at the same level of the origination point of M (Fig. 7C).
</paragraph>
<paragraph id="8EC8950EF874C72926AA9688F0F5472C" pageId="7" pageNumber="84">
Hind wing. With the same shape as the forewing. R1 running smoothly through pterostigma; pterostigmal crossveins (Pcv) absent; Rs with four branches; one crossvein between R2+3 and R4 and one between R4 and R5; the 'bittacid
<normalizedToken id="031338EF7E9C77B33B673CF6000E05A4" originalValue="cross">cross'</normalizedToken>
aligned; M divided into three branches; two crossveins between R5 and M1, one between M1 and M2, one between M2 and M3 and one between M3 and Cu1; Cu1 and Cu2 parallel and with one crossvein between them. Vein 1A reaching the posterior margin at the level slightly proximad of the Rs originating from R1, one crossvein between Cu2 and 1A (Fig. 7D).
</paragraph>
<paragraph id="65199C8808737E900CADAEDAB8C85B53" pageId="7" pageNumber="84">Abdomen. Abdomen 9.1 mm long, with ten visible segments. Female genital structure well-preserved from the dorsal view. Supraanale and cercus covered with small and short setae (Fig. 7A, B, E, F).</paragraph>
</subSubSection>
<subSubSection id="032FD5D0C9F857B852E9879E106FD220" pageId="7" pageNumber="84" type="remarks">
<paragraph id="D2B97FD000058DBE93C285C3153C14B2" pageId="7" pageNumber="84">Remarks.</paragraph>
<paragraph id="82C0A82E19598939A23D5D1A713EB0F7" pageId="7" pageNumber="84">
<taxonomicName id="E245C6E9FA43AA8AE15318367360CAE9" genus="Exilibittacus" lsidName="Exilibittacus foliaceus" pageId="7" pageNumber="84" rank="species" species="foliaceus">Exilibittacus foliaceus</taxonomicName>
sp. n. (Fig. 7) is assigned to the genus
<taxonomicName id="65595AF89568997162D1BE0E2CC7E3E5" genus="Exilibittacus" lsidName="Exilibittacus" pageId="7" pageNumber="84" rank="genus">Exilibittacus</taxonomicName>
Yang, Ren &amp; Shih, 2012 based on the following generic diagnostic characters: in forewing, Sc reaching the anterior margin proximad of the forking of R4+5 and the 'bittacid
<normalizedToken id="46451C8CA3892CF776429CFBA3C004A0" originalValue="cross">cross'</normalizedToken>
aligned; and in hind wing, Rs with four branches while M with three branches.
<taxonomicName id="6100C46F4943483DDA4C9DFFA3ED79DF" genus="Exilibittacus" lsidName="Exilibittacus foliaceus" pageId="7" pageNumber="84" rank="species" species="foliaceus">Exilibittacus foliaceus</taxonomicName>
sp. n. is distinguished from the other two species as shown by the key below.
</paragraph>
</subSubSection>
</treatment>
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