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<document id="BC7A3DFD0638A93B743DCD60D92BDC31" ID-CLB-Dataset="35149" ID-DOI="http://dx.doi.org/10.3897/zookeys.536.5982" ID-GBIF-Dataset="f3546db1-d96a-40d8-ac8c-fe806547f201" ID-PMC="PMC4714036" ID-Pensoft-Pub="1313-2970-536-1" ID-PubMed="26798233" ID-ZooBank="A8B92CDD0B8C4384AAC559648BB45AA5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-536-1" ModsDocOrigin="ZooKeys 536" ModsDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" checkinTime="1451243741663" checkinUser="pensoft" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2015" docId="F09D5BDE68EDC9E0920F91C9952461C9" docLanguage="en" docName="ZooKeys 536: 1-92" docOrigin="ZooKeys 536" docSource="http://dx.doi.org/10.3897/zookeys.536.5982" docTitle="Pterostichus (Circinatus) cavazzutianus Shi &amp; Liang, 2015, s. str." docType="treatment" docUuid="89769B18-853F-4F85-B671-C1ADE830CF14" docUuidSource="ZooBank" docVersion="8" lastPageNumber="35" masterDocId="FFEAFF976442467ED413FFC17531FF81" masterDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="35" updateTime="1731888672137" updateUser="admin">
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<mods:title id="F983913F1A3A202896188D7A0D6DF2F3">The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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<mods:namePart id="2A10AFCF7DDF236270CA5874FC48ABA8">Shi, Hongliang</mods:namePart>
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<treatment id="F09D5BDE68EDC9E0920F91C9952461C9" ID-GBIF-Taxon="152063160" LSID="urn:lsid:zoobank.org:act:89769B18-853F-4F85-B671-C1ADE830CF14" httpUri="http://treatment.plazi.org/id/F09D5BDE68EDC9E0920F91C9952461C9" lastPageNumber="35" pageId="34" pageNumber="35">
<subSubSection id="03FECFE24D1A16D4C51F63752D82CEC6" pageId="34" pageNumber="35" type="multiple">
<paragraph id="8C1F25C841B56C3A3C1FB316C042D504" pageId="34" pageNumber="35">
Taxon classification Animalia
<taxonomicName id="24A05A0177C81104F736C2D29A8DB9C2" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="582823272CA39040B542300B599A8F34" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E667F2B284EEF4C3E093D1A03A121895" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="D5BF2250B66FE626408A8D46CFF97D9D" pageId="34" pageNumber="35">
<taxonomicName id="C2E410DCFD4EFED0B7757B558FA87909" ID-CoL="8MFK5" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" sensu="stricto" species="cavazzutianus" subGenus="Circinatus">
Pterostichus (Circinatus) cavazzutianus
<taxonomicNameLabel id="F637CF405606E06C82285B69E3B69CFC" sensu="stricto">s. str.</taxonomicNameLabel>
</taxonomicName>
<taxonomicNameLabel id="F816CDF351EA923DA5FCB131DEB8F893" pageId="34" pageNumber="35">replacement name</taxonomicNameLabel>
Figures 5, 35, 57, 72, 91, 114
</paragraph>
</subSubSection>
<subSubSection id="64FD3725269C43530ECBAC906ADA9FCE" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="BD11EC69469F24FEDFA9C78E84D3B7C5" pageId="34" pageNumber="35">
<taxonomicName id="A4E49081FB79F4C15976CB6097800CE1" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus) cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" sensu="stricto" species="cavazzutianus" subGenus="Circinatus">
Pterostichus (Circinatus) cavazzutianus
<taxonomicNameLabel id="032A1467C1223FF3960B4E343A23C9F5" sensu="stricto">s. str.</taxonomicNameLabel>
</taxonomicName>
Synonym:
<taxonomicName id="7ABFE7A9ECC522B8A82738FB0FE64C5F" authority="Allegro &amp; Sciaky, 2010: 8" authorityName="Allegro &amp; Sciaky" authorityPageNumber="8" authorityYear="2010" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutii" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzutii">cavazzutii Allegro &amp; Sciaky, 2010: 8</taxonomicName>
(Original:
<taxonomicName id="A11497AA750A43A2263FB0AC13001480" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Pterostichus (Circinatus)</taxonomicName>
; holotype deposited in collection of Allegro). Junior homonym of
<taxonomicName id="C1E7A075F40E1A917F097A1756968386" authority="Sciaky &amp; Facchini, 2003" authorityName="Sciaky &amp; Facchini" authorityYear="2003" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Sinosteropus) barbarae" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="barbarae" subGenus="Sinosteropus" subSpecies="cavazzutii">Pterostichus (Sinosteropus) barbarae cavazzutii Sciaky &amp; Facchini, 2003</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="AB69854DBCB8D8A4511416DF9C3A44EC" pageId="34" pageNumber="35" type="type locality">
<paragraph id="192D3941512CC27F08B1C47CEACCF10A" pageId="34" pageNumber="35">Type locality.</paragraph>
<paragraph id="6289905FB82A6B7473EF0210658FC426" pageId="34" pageNumber="35">Sichuan: Ebian, Yizhi Yakou (= Yizi pass), Xiaoliang Shan, Si He (= Xihe, or Heizhugou Town), 2800-3100 m.</paragraph>
</subSubSection>
<subSubSection id="DCFC9B69B296144FB9445DEC575EF3BA" pageId="34" pageNumber="35" type="type material">
<paragraph id="FB707488528223E5E62D205EA4C61653" pageId="34" pageNumber="35">Type material examined.</paragraph>
<paragraph id="CBFE564C729ABB5326A8D3F89EDDB3BC" pageId="34" pageNumber="35">
Paratype of
<taxonomicName id="CFD105AA16D5D32574CBFFCB35E60C15" authority="Allegro &amp; Sciaky" authorityName="Allegro &amp; Sciaky" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutii" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzutii">Pterostichus cavazzutii Allegro &amp; Sciaky</taxonomicName>
(CRS): male, board mounted, genitalia dissected and glued on board, pinned under specimen, &quot;CHINA S Sichuan / Pass btw Xihe/Hongxi / Ta
<taxonomicName id="365F1C92BC0AF0C2DD532BE6BA38E577" authority="Ping" authorityName="Ping" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="yan">Yan Ping</taxonomicName>
, 17.-25.6. / 1999 leg. V. Benes&quot;; &quot;PARATYPUS ♂/
<taxonomicName id="8381761661CDBCDE046B49E9687B1B43" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
/ (
<taxonomicName id="F7C5376F0582167D28523643916B18F0" class="Insecta" family="Carabidae" genus="Circinatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
)/
<taxonomicName id="102654A55E1A65E59FDE76428341CD9E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cavazzutii" status="sp. nov.">cavazzutii</taxonomicName>
<taxonomicNameLabel id="F10640D271520847455508A642158D3C" rank="species">sp. n.</taxonomicNameLabel>
/ Allegro &amp; Sciaky det. 2010&quot; [red label].
</paragraph>
</subSubSection>
<subSubSection id="05672AEEA9E01958AB70D13991498DE2" pageId="34" pageNumber="35" type="type material">
<paragraph id="9A3AC909410700FC103A0AD4DF01C740" pageId="34" pageNumber="35">Non-type material examined</paragraph>
<paragraph id="C114AC25AF458641BD6D149A4F46F9B6" pageId="34" pageNumber="35">
(19 specimens). 8 males, 3 females (IZAS), &quot;China, Sichuan Prov., Yizi pass btw. Meigu county and Ebian county, mixed forest;
<geoCoordinate id="912F630F645D6167DC6D23CAB5D49C1A" direction="north" orientation="latitude" precision="1" value="28.67477">N28.67477</geoCoordinate>
<geoCoordinate id="548933188BEBB579505A8C0DA09E4AB4" direction="east" orientation="longitude" precision="1" value="103.05248">E103.05248</geoCoordinate>
; 2923 m; 2012.VI.15; by pitfall trap; SHI Hongliang &amp; LIU Ye lgt.&quot;; 1 female (IZAS), &quot;China, Sichuan Prov., Yizi pass btw. Meigu county and Ebian county, mixed forest;
<geoCoordinate id="F4F659388509BD4DC447574CC8D700DC" direction="north" orientation="latitude" precision="1" value="28.67477">N28.67477</geoCoordinate>
<geoCoordinate id="9B2B4EF40F0774811970325124D6801B" direction="east" orientation="longitude" precision="1" value="103.05248">E103.05248</geoCoordinate>
; 2923 m; 2012.VI.12;in dead log; SHI Hongliang &amp; LIU Ye lgt.&quot;; 4 males, 1 female (IZAS), &quot;China, Sichuan Prov., Meigu county, Dafengding nat. cons., Hongxi station, mixed forest; 2541 m,
<geoCoordinate id="AD1ADB992307F27961E6C3CEBCECCAF3" direction="north" orientation="latitude" precision="1" value="28.6585">N28.65850</geoCoordinate>
<geoCoordinate id="24E8499223E32E3563B9497E5123C326" direction="east" orientation="longitude" precision="1" value="103.06123">E103.06123</geoCoordinate>
; 2012.VI.15; by pitfall trap; SHI Hongliang &amp; LIU Ye lgt.&quot;; 1 male, 1 female (IZAS), &quot;China, Sichuan Prov., Ebian county, Heizhugu forestry park, 2888 m,
<geoCoordinate id="CD97DA921B109A368460FA9636790E7E" direction="north" orientation="latitude" precision="1" value="29.02373">N29.02373</geoCoordinate>
<geoCoordinate id="3A61D78E276BC57591482C8D30870F33" direction="east" orientation="longitude" precision="1" value="102.98473">E102.98473</geoCoordinate>
; 2012.VI.17; by pitfall trap; rhododendra forest with bamboo, YANG Ganyan &amp; LIU Ye lgt.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="BCAFDAA0AAE1879CCC026AA4DFF5AC40" pageId="34" pageNumber="35" type="nomenclature notes">
<paragraph id="3F9F0F31C0878A851BEBDA6A42D92706" pageId="34" pageNumber="35">Nomenclature notes.</paragraph>
<paragraph id="65697BFDEE0906BF03189640330DA172" pageId="34" pageNumber="35">
We here propose
<taxonomicName id="3C1FF87A3BF62C0BB1F2AF14405D040F" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzuttianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzuttianus">Pterostichus cavazzuttianus</taxonomicName>
as a replacement name for
<taxonomicName id="84A84D749A189DC0530D9F170EBDC525" authority="Allegro &amp; Sciaky, 2010" authorityName="Allegro &amp; Sciaky" authorityYear="2010" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzuttii" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzuttii">Pterostichus cavazzuttii Allegro &amp; Sciaky, 2010</taxonomicName>
, preoccupied by
<taxonomicName id="41E968569B056B20CF4DCFAC32B0DF55" authority="Sciaky and Facchini, 2003" authorityName="Sciaky and Facchini" authorityYear="2003" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzuttii" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzuttii">Pterostichus cavazzuttii Sciaky and Facchini, 2003</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="192490B2C921E39F362EC14CCA97E3D6" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="DB8E5B5325494E380EC55C029DB8AEA7" pageId="34" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="D37D5CA39683CE92C89AD19464629834" pageId="34" pageNumber="35">Pronotum usually with three mid-lateral setae; hind angle completely rounded; basal fovea punctate; lateral margin of elytron narrow and deep; males with terminal sternum slightly depressed in middle, not rugose in depression; apical lamella of aedeagus located on right side of aedeagal apex, basal width approx same as length.</paragraph>
<paragraph id="92BB77B86EF5D4A06DE0A84E0AE8DFFA" pageId="34" pageNumber="35">
For comparisons between
<taxonomicName id="EF51AC295D0AA4A757FD70B12145CB6C" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" sensu="stricto" species="cavazzutianus">Pterostichus cavazzutianus</taxonomicName>
<taxonomicNameLabel id="A519E39AD0D93A1273D2C9E42920F81A" sensu="stricto">s. str.</taxonomicNameLabel>
and the subspecies
<taxonomicName id="E72AB742AD31CFAEB6DC4616CBD8551E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="cavazzutianus" status="subsp. nov." subSpecies="mianningensis">Pterostichus cavazzutianus mianningensis</taxonomicName>
<taxonomicNameLabel id="CF2ECBBB143F2411A379A86FC0463D05" rank="subSpecies">subsp. n.</taxonomicNameLabel>
, see the diagnosis of
<taxonomicName id="D58A5B8B727C353587A5FA55E528C765" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="cavazzutianus" status="subsp. nov." subSpecies="mianningensis">Pterostichus cavazzutianus mianningensis</taxonomicName>
<taxonomicNameLabel id="534A6FC00372969C95D080C409555779" rank="subSpecies">ssp. n.</taxonomicNameLabel>
This species is similar to and sympatric with
<taxonomicName id="5C3C69B3DEE97ACE3ED57FCA08565F15" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="adelphus" status="sp. nov.">Pterostichus adelphus</taxonomicName>
<taxonomicNameLabel id="2EDAE1F41D4340F6AACAF21801450C6D" rank="species">sp. n.</taxonomicNameLabel>
For their comparisons see the diagnosis of
<taxonomicName id="B948884FC9C13A934C498C2D898E525A" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="adelphus" status="sp. nov.">Pterostichus adelphus</taxonomicName>
<taxonomicNameLabel id="18B18FAC61DABB79C1E1517D425B6C7F" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="EB8859E9ED4813E819A980460EF46C5E" pageId="34" pageNumber="35" type="supplemental description">
<paragraph id="AE208832102B20669EF58107409C2205" pageId="34" pageNumber="35">Supplemental description.</paragraph>
<paragraph id="C1037AD7B57BF3E302045B802900571F" pageId="34" pageNumber="35">Male sternum. Penultimate sternum unmodified; terminal sternum slightly depressed in middle, apex of sternum weakly but distinctly bending downwards, almost smooth inside sternum depression (Fig. 114). Endophallus (Figs 35D, E, F) short, bent to ventral side across apical lamella, and then turned to aedeagal base; gonopore (gp) located at approx same level as apical lamella. Three distinct lobes recognized: ventral-basal lobe I (vb-I) small, close to base of apical lamella, compressed, upper surface heavily scaled, apex pointed; ventral-basal lobe II (vb-II) large and long, close to apical lamella, pointing to ventral face of median lobe, upper surface strongly chitinized, lower surface membranous, apex capitate and scaled; ventral-apical lobe (va) at right side of vb-II, its main body strongly chitinized, forming a large piece with notched apex; right lobe (rl) membranous, very small and indistinct. Female genitalia. Spermatheca with seminal canal approx three times as long as receptaculum; receptaculum capitate (Fig. 57), club approx half length of receptaculum; seminal canal inserted at base of common oviduct, base of seminal canal sclerotized. Stylomere II with two ensiform setae at outer margin, and one at basal third of inner margin; two very short nematiform setae located in a furrow near apex. Female sternum VIII (Fig. 72B) evenly chitinized; posterior margin with fine and sparse setae, almost straight, notched in middle; middle transparent region small, approx semicircular, without oblique extensions, adjacent to anterior and posterior notches in middle. Female tergum VIII (Fig. 72A) semi-chitinized, posterior region without denser pigmentation, anterior margin without notch in middle, posterior margin evenly arcuate.</paragraph>
</subSubSection>
<subSubSection id="5551E048FE6E2E2175C8C04E278D346C" pageId="34" pageNumber="35" type="distribution">
<paragraph id="DE4555639BB870A241828A3F2BF4F33E" pageId="34" pageNumber="35">Distribution.</paragraph>
<paragraph id="596C0E08C2B489B51D30F6AC892F8AF9" pageId="34" pageNumber="35">
The nominotypical subspecies is only recorded from localities near the border between Meigu and Ebian County in south Sichuan (Map 2). It is sympatric with three other
<taxonomicName id="4C8C0C83FF83C7A0C631F330CFF7F51F" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
species,
<taxonomicName id="2E434886FC521E497DC5539A4CBF8FB8" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus bullatus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="bullatus">Pterostichus bullatus</taxonomicName>
,
<taxonomicName id="75326FC7F5D5E54CCBF81F53626B949E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="adelphus" status="sp. nov.">Pterostichus adelphus</taxonomicName>
<taxonomicNameLabel id="74FE284793EFDFC4BFADF3507FC71DB1" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="DB535630671A72817B648C85EB5A7F9A" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="zhygealu" status="sp. nov.">Pterostichus zhygealu</taxonomicName>
<taxonomicNameLabel id="DE66D819F857294DA833AB9534712CC9" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F7DB1AFCD7DADC900D3FC0A7610E932E" pageId="34" pageNumber="35" type="etymology">
<paragraph id="5C51FED93E1143D462D2A9033FB5821B" pageId="34" pageNumber="35">Etymology.</paragraph>
<paragraph id="1F838927548CB7ECE6F90ED676F1687F" pageId="34" pageNumber="35">
The name
<taxonomicName id="1459CBD3E5F8A8C546AB254D5C9C87D0" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutii" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzutii">Pterostichus cavazzutii</taxonomicName>
is preoccupied by a subspecies
<taxonomicName id="D263F81C52229190DDF08589FE263178" authority="Sciaky &amp; Facchini, 2003" authorityName="Sciaky &amp; Facchini" authorityYear="2003" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus (Sinosteropus) barbarae" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="barbarae" subGenus="Sinosteropus" subSpecies="cavazzutii">Pterostichus (Sinosteropus) barbarae cavazzutii Sciaky &amp; Facchini, 2003</taxonomicName>
. We therefore propose a replacement name
<taxonomicName id="D3572DAD9699725EAC41D6F593B263FE" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus cavazzutianus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="cavazzutianus">Pterostichus cavazzutianus</taxonomicName>
for this species, respecting the
<normalizedToken id="3ADE1021183D332ABCA5D8F93C1064AF" originalValue="authors’">authors'</normalizedToken>
original notion by changing only the suffix.
</paragraph>
</subSubSection>
<subSubSection id="4DB10180FE9FD8B2A31A1812307F8ED9" pageId="34" pageNumber="35" type="affinities">
<paragraph id="1227A2AAD83A03ABE77EA124590B650B" pageId="34" pageNumber="35">Affinities.</paragraph>
<paragraph id="A92C0C72A3A6223CE0EB59DF0D9D46EE" pageId="34" pageNumber="35">
This species seems to be close to
<taxonomicName id="7CD744563FBBB19D3266F59659BCC2B7" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="adelphus" status="sp. nov.">Pterostichus adelphus</taxonomicName>
<taxonomicNameLabel id="E7E51ECB34CC1845284ECBC941C84789" rank="species">sp. n.</taxonomicNameLabel>
in their similarities of elytral microsculpture and short apical lamella of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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