196 lines
17 KiB
XML
196 lines
17 KiB
XML
<document id="2FA6F51D74FBFEA4DB444689EB55D97C" ID-DOI="10.3897/zookeys.1203.123757" ID-ZooBank="6FE93D4B-8104-45F4-850A-C1B78CA10A9D" ID-publisher-id="123757" URI-arpha="E0E2E4CE-0D0E-533C-9BD4-A1327BB57504" URI-zoobank="http://zoobank.org/6FE93D4B-8104-45F4-850A-C1B78CA10A9D" article-type="research-article" checkinTime="1716923959232" checkinUser="pensoft" docAuthor="Jordal, Bjarte H." docDate="2024" docId="E94738BF8DCA5D94B01E4A87085E3EF4" docLanguage="en" docName="ZooKeys 1203: 95-130" docOrigin="ZooKeys 1203" docSource="https://zookeys.pensoft.net/article/123757/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Ctonoxylon tuberculatum Jordal 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="130" masterDocId="E0E2E4CE0D0E533C9BD4A1327BB57504" masterDocTitle="An integrated taxonomic revision of Ctonoxylon (Coleoptera, Curculionidae, Scolytinae) reveals new Malagasy species originating from multiple recent colonisations of the island" masterLastPageNumber="130" masterPageNumber="95" pageNumber="95" updateTime="1716924051971" updateUser="pensoft">
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<mods:titleInfo id="F80ACFD1C80CFFCABA5573699D363F7E">
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<mods:title id="63BA89DEC6FC4398E4B0F826091B76E2">An integrated taxonomic revision of Ctonoxylon (Coleoptera, Curculionidae, Scolytinae) reveals new Malagasy species originating from multiple recent colonisations of the island</mods:title>
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<mods:name id="067828944AFF5B3FA9FCB21699D59695" type="personal">
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<mods:roleTerm id="DEBAD0B0A5E67391ADAA2574DD3174C2">Author</mods:roleTerm>
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<mods:namePart id="E663C5B2B0C2015E8C69DE2D361F2D68">Jordal, Bjarte H.</mods:namePart>
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<mods:nameIdentifier id="4770BEE78AC59B1141264F5DD9074FB4" type="ORCID">0000-0001-6082-443X</mods:nameIdentifier>
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<mods:affiliation id="DC587533D448F60404EB4E66847AC6BD">Department of Natural History, University Museum of Bergen, University of Bergen, P. O. 7800, NO- 5020 Bergen, Norway</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="47368482D55BAD9744732D92356CBBDD">text</mods:typeOfResource>
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<mods:titleInfo id="ECB264487101F8DCEDEBB2CA24B6DE48">
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<mods:title id="4E1E6DED717E46BDE9E0C768413B2B79">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="86A24C4EB1981C4CB78FECF1C7BA4A3F">
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<mods:date id="F618AFBF48813519C7E01DDA6DBE4B26">2024</mods:date>
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<mods:detail id="30DB33B90C94C9607C87697E5F63C3A8" type="pubDate">
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<mods:number id="9780B83792D314E03259393D93B7B5F7">2024-05-28</mods:number>
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<mods:detail id="DB349D5B5F128D7A3BDAA8F648939F2F" type="volume">
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<mods:number id="01C6AB04AAD4D10D63A3748E31A478AA">1203</mods:number>
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<mods:extent id="2F7240322E080E483556B291FF62B92C" unit="page">
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<mods:start id="57E4404EC22BB0579B37D22DE4276AFD">95</mods:start>
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<mods:end id="FB1C16D48D8461AB55B11901A6AC3C56">130</mods:end>
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<mods:classification id="3E1013E8A1710E3F71492B93A299079A">journal article</mods:classification>
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<mods:identifier id="024C56BB9C066A1F26D9015BD0F2B137" type="DOI">10.3897/zookeys.1203.123757</mods:identifier>
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<mods:identifier id="1AE474F4EA72A512811003B3937E614E" type="ZooBank">6FE93D4B-8104-45F4-850A-C1B78CA10A9D</mods:identifier>
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</mods:mods>
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<treatment id="E94738BF8DCA5D94B01E4A87085E3EF4" ID-arpha="E94738BF-8DCA-5D94-B01E-4A87085E3EF4" ID-zoobank="https://zoobank.org/0921A825-BD54-4309-90AD-2345527F0034" LSID="urn:lsid:plazi:treatment:E94738BF8DCA5D94B01E4A87085E3EF4" LSID-ZBK="urn:lsid:zoobank.org:act:0921A825-BD54-4309-90AD-2345527F0034" httpUri="http://treatment.plazi.org/id/E94738BF8DCA5D94B01E4A87085E3EF4">
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<subSubSection id="C578A3D0DBDC0483D4DDFC5E27D16909" type="nomenclature">
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<paragraph id="2ECFAF54440DD42B87AEB6C202E3B8CE">
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<taxonomicName id="991B49E19E4568CA497177D4A68367C6" authority="Jordal, 2024" authorityName="Jordal" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Ctonoxylon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tuberculatum" status="sp. nov.">
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<emphasis id="407A21692B98BB88FB81811D532E03F7" italics="true">Ctonoxylon tuberculatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5B974DA7F2B3024E6A5260BB9DD4011A" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="E96FB03FB56FB5D8898FFB536E6377B3" type="description">
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<paragraph id="AF97E070AA1405A3061D072EC3ADC169">
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<figureCitation id="F0ED8E2CDD81B21D3D63E22275AF22D5" captionStart="Figures 30–38" captionStartId="F5" captionText="Figures 30–38. Dorsal, lateral, and front views of 30, 33, 36 Ctonoxylon tuberculatum sp. nov. male holotype 31, 34, 37 Ctonoxylon longipilum female (= C. montanum), and 32, 35, 38 Ctonoxylon nodosum male holotype (= C. montanum)." figureDoi="10.3897/zookeys.1203.123757.figures30-38" httpUri="https://binary.pensoft.net/fig/1057223">Figs 30</figureCitation>
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,
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<figureCitation id="033AC9172F82ED8477F6962750B899FF" captionStart="Figures 30–38" captionStartId="F5" captionText="Figures 30–38. Dorsal, lateral, and front views of 30, 33, 36 Ctonoxylon tuberculatum sp. nov. male holotype 31, 34, 37 Ctonoxylon longipilum female (= C. montanum), and 32, 35, 38 Ctonoxylon nodosum male holotype (= C. montanum)." figureDoi="10.3897/zookeys.1203.123757.figures30-38" httpUri="https://binary.pensoft.net/fig/1057223">33</figureCitation>
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,
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<figureCitation id="8011E5EB5556702DAF5F1FAD6C0B184C" captionStart="Figures 30–38" captionStartId="F5" captionText="Figures 30–38. Dorsal, lateral, and front views of 30, 33, 36 Ctonoxylon tuberculatum sp. nov. male holotype 31, 34, 37 Ctonoxylon longipilum female (= C. montanum), and 32, 35, 38 Ctonoxylon nodosum male holotype (= C. montanum)." figureDoi="10.3897/zookeys.1203.123757.figures30-38" httpUri="https://binary.pensoft.net/fig/1057223">36</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0ERMAI" type="type material">
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<paragraph id="AAF21F9B1786144D5B0A4595A3FC25DB">
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<heading id="36549DC822A6ED56655BF14D0308A3EE" reason="title">Type material.</heading>
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</paragraph>
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<paragraph id="C268D7ECD0222BDF3214075C106CCAE6">
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<materialsCitation id="9E02071285A3D280B749B30832BC00FD" collectingDate="2019-11-03" collectionCode="ZMUB" collectorName="B. Jordal" country="Madagascar" elevation="1000" location="Diana prov." municipality="Montagne d'Ambre" specimenCount="1" typeStatus="holotype">
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<emphasis id="4D0A5B574DF8348CC049166BD3B6113E" bold="true" italics="true">
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<typeStatus id="6F93E60CF620865490E1DE23BFD0336B" type="holotype">Holotype</typeStatus>
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</emphasis>
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, male:
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<collectingCountry id="29550D759E80DAD7BFD007411BB052B3" name="Madagascar">Madagascar</collectingCountry>
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,
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<location id="01BB458F85CCA9B9A0BC32FC1683CA7B" LSID="urn:lsid:plazi:treatment:E94738BF8DCA5D94B01E4A87085E3EF4:01BB458F85CCA9B9A0BC32FC1683CA7B" country="Madagascar" municipality="Montagne d'Ambre" name="Diana prov.">Diana prov.</location>
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,
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<collectingMunicipality id="78968CD2363AE525EDDDB623E9B3E1EA">Montagne d’Ambre</collectingMunicipality>
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[GIS:
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6MAI" specific-use="{"type":"Point","coordinates":[49.170000,-12.540000]}">- 12.54, 49.17</named-content>
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],
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<quantity id="39AEFA2CD13F9CD054699F9B469E40BE" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
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<elevation id="E0C87384A07006B556ED61B016589A19" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
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</quantity>
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alt., ex
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<taxonomicName id="802D2D667B56B8586CE2D2788638C779" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="genus">
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<emphasis id="F0CB9124AFF406467BE139FE51707E05" italics="true">Ficus</emphasis>
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</taxonomicName>
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branch,
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<date id="10A5A8C27146E3E1E2F02A81EDC6CB51" value="2019-11-03">
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<collectingDate id="7DD9466F7FFA6F220F6295BBE00610F6" value="2019-11-03">03. 11. 2019</collectingDate>
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</date>
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,
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<collectorName id="52BF462EE809D1BBE09FDBBA42862227">B. Jordal</collectorName>
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, leg. [
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<named-content id="E6B2DFC53C293BE9F0FA6CCC736D7C98" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-zoology-university-bergen-vertebrate-collections" xlink_title="University Museum of Bergen, Norway">
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<collectionCode id="A59CB0FE05F7679A57B89C9047F262F2" country="Norway" httpUri="http://grbio.org/cool/5b4j-820w" name="Museum of Zoology at the University of Bergen, Vertebrate collections" type="">ZMUB</collectionCode>
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</named-content>
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]
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</materialsCitation>
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.
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<materialsCitation id="485D5937B67D6FE362477F512BF58C47" collectingDate="2019-11-03" collectionCode="ZMUB, NHMW" collectorName="B. Jordal" country="Madagascar" elevation="1000" location="Diana prov." municipality="Montagne d'Ambre" specimenCount="2" typeStatus="allotype">
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<typeStatus id="36CAB0D2AB66E5407972F588980D5107" type="allotype">Allotype</typeStatus>
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[
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<named-content id="DC99C140CC9E28BD60A990D3371A36B0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-zoology-university-bergen-vertebrate-collections" xlink_title="University Museum of Bergen, Norway">
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<collectionCode id="6911511B0F22A6AF5F9DD23DB51AB33C" country="Norway" httpUri="http://grbio.org/cool/5b4j-820w" name="Museum of Zoology at the University of Bergen, Vertebrate collections" type="">ZMUB</collectionCode>
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</named-content>
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] and
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<typeStatus id="A2E88B4DE95603F1B6C29F46F25102CB" type="paratype">paratype</typeStatus>
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[
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<named-content id="F7FF9538260B34CD4853FCD6F639724F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum, Vienna, Austria">
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<collectionCode id="E84B5F09BB3904525D06999F93E4F030" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
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</named-content>
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]: same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E1NAI" type="diagnosis">
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<paragraph id="84F94050CB65B87189EABE5E9E5374AC">
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<heading id="E833724D957403BEA8D29EBA399ABFEA" reason="title">Diagnosis.</heading>
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</paragraph>
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<paragraph id="129BDE3463B5D9CACBACA3E57C655279">Two teeth along the anterior margin of pronotum separated by more than width of a tooth; scattered interstrial setae separated within uniseriate rows by more than their length; posterior margin of elytra and declivital interstriae with sharp tubercles, largest tubercles on interstriae 1 and 3.</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E6NAI" type="description">
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<paragraph id="92FE7A89347F8F3B9AC4A8F4FAECE920">
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<heading id="ED65123A75DE8EB138D5F922F44989AE" reason="title">Description.</heading>
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</paragraph>
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<paragraph id="9E419183B617C78978FA578325636D29">
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<emphasis id="327476B4A93E89D35F5ACC47C98EC20E" bold="true">Male.</emphasis>
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Body length 3.0–
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<quantity id="035CDFCEC8C08BB7E1058C537CDCFD98" metricMagnitude="-3" metricUnit="m" metricValue="3.1" unit="mm" value="3.1">3.1 mm</quantity>
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, 2.1–2.2 × as long as broad; colour brown.
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<emphasis id="C5DC9CF3211F98128D2F7DA5E294485C" bold="true" italics="true">Frons</emphasis>
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flattened from upper level of eyes to epistoma, surface shrivelled; vestiture of fine short setae. Eyes divided, each part separated by a little more than size of upper half. Antennal funiculus 7 - segmented; club setose, sutures and septum barely indicated.
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<emphasis id="069C172F9A6D569EA28B453221B15301" bold="true" italics="true">Pronotum</emphasis>
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coarsely asperate on anterior three quarters, two front teeth separated by little more than width of a tooth, located behind the margin. Anterior lateral margin of prothorax at middle with prominent eye scraper; a sharp carina running from eye scraper to near procoxa; propleural pit absent.
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<emphasis id="C84F1174F84B1745532E438C8ABDCE01" bold="true" italics="true">Scutellar shield</emphasis>
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wider than long, oval.
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<emphasis id="CC5F31C225C395CEEC7AF8FBAF41EE39" bold="true" italics="true">Elytral</emphasis>
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striae impressed, punctures irregular and small; interstriae rounded, densely micropunctate, vestiture consisting of uniseriate rows of slightly curved bristle-like setae and fine dense ground vestiture along the suture; elytral suture straight; elytral apex slightly emarginated, along the posterior margin and at each declivital interstriae with small sharp tubercles, at interstriae 1 and 3 tubercles 2–3 × larger.
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<emphasis id="8DD57A99384707B56037229B42ED2C0B" bold="true" italics="true">Metanepisternum</emphasis>
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with mixed short bifid and longer bristle-like setae; metaventrite with simple hair-like setae; these sclerites on its anterior third partly glabrous and with small bifid setae, and with a swollen trace of mesotibia’s resting position.
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<emphasis id="D507A8BE1A194C859DF55FF5311B9389" bold="true">Female</emphasis>
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as in male, except anterior pair of teeth more closely placed and located along the anterior margin of the pronotum, and surface of the frons smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E4OAI" type="etymology">
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<paragraph id="EFAD7D33C56EB80B14583D6D0FA0D767">
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<heading id="2BD3B8405E509D2E7FAC5433DDFB01A2" reason="title">Etymology.</heading>
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</paragraph>
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<paragraph id="2467B1B6957AE2AEB2E0C7FA02B424B3">
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The Latin adjective
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<emphasis id="18E0B668C00C50CC6E08BC46406AEF76" italics="true">tuberculatus</emphasis>
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in its neuter form, reflecting the tubercles on declivital interstriae which are more prominent than in related species.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0ENPAI" type="distribution">
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<paragraph id="0AEC3210899A9C4AE2D51339746F6F0B">
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<heading id="AB2C4EEC430F795EA36BCC1E7ACEAA80" reason="title">Distribution and biology.</heading>
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</paragraph>
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<paragraph id="4CE532AA066D147F547B257CD0EFF523">
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Only known from the
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<typeStatus id="C78D44AE32E291FA9EB5069BD4C3F7F1" type="holotype">holotype</typeStatus>
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locality in
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<collectingCountry id="3D4794E41158C48995D27A4595271F11" name="Madagascar">Madagascar</collectingCountry>
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where it was collected from very thick bark of a fallen
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<taxonomicName id="7EB047054459E9875CB8FE2477071414" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="genus">
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<emphasis id="CF2B161643B35AF26987E58F268DD785" italics="true">Ficus</emphasis>
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</taxonomicName>
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branch (
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<taxonomicName id="89D0FC359128B20AC0CD7ADC58A2A8F8" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
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),
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<quantity id="141D651569B6EFFE79780CBBBA8FFD6C" metricMagnitude="-2" metricUnit="m" metricValue="7.0" unit="cm" value="7.0">7 cm</quantity>
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in diameter. Two pairs were collected at the early stage of tunnel construction, including a mating niche with a short egg tunnel.
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E6PAI" type="comments">
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<paragraph id="C9897E1AC01BD0EFB2A8304409F6DD91">
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<heading id="E2E25FEAB13087DE010BCF2195F98806" reason="title">Comments.</heading>
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</paragraph>
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<paragraph id="41698B37E90356D1417A6FAC050F03DF">
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Previous records of
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<taxonomicName id="120CFA353AF953D65302423540C4A804" authorityName="Hagedorn" authorityYear="1910" class="Insecta" family="Curculionidae" genus="Ctonoxylon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="flavescens">
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<emphasis id="7C48A94F0B685A23CCA776F41CB84395" italics="true">C. flavescens</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="BA2DFD217AD30C587EBAF659F62D69CE" name="Madagascar">Madagascar</collectingCountry>
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are most likely
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<taxonomicName id="546241BF1EACAAA292460FE76E4DD480" authorityName="Jordal" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Ctonoxylon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="tuberculatum">
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<emphasis id="754962506B6B1B50CDCEF4D8C905D28E" italics="true">C. tuberculatum</emphasis>
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</taxonomicName>
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as these two species are very similar. This could also be the case for records of
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<taxonomicName id="A1845FCC0F03EF3188A34E29D9572523" authorityName="Eggers" authorityYear="1922" class="Insecta" family="Curculionidae" genus="Ctonoxylon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="montanum">
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<emphasis id="AFAA051E596C77E74B5C65854B4F4B59" italics="true">C. montanum</emphasis>
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</taxonomicName>
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(recorded as
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<taxonomicName id="A706FE8B9051E27F5D9CB05512C3F53F" authorityName="Eggers" authorityYear="1935" class="Insecta" family="Curculionidae" genus="Ctonoxylon" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="longipilum">
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<emphasis id="D4CDA977C3B2E8FDEAA010C8CDDEEBC9" italics="true">C. longipilum</emphasis>
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</taxonomicName>
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), in which the female mainly differs by the more closely set anterior teeth on the pronotum, and the stouter body. These two species are therefore removed from the list of Malagasy species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |