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<document id="ACF5B5FC93C1765337C3DB98DCE290D0" ID-DOI="10.26107/RBZ-2022-0008" ID-ISSN="2345-7600" ID-Zenodo-Dep="7175098" ID-ZooBank="D3356439-3DE5-4DF8-87C0-70F6046BB1CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665292877568" checkinUser="felipe" docAuthor="Dworschak, Peter C. &amp; Anker, Arthur" docDate="2022" docId="D65A2A57FFD7FF825CE0FC51FA3BFB5E" docLanguage="en" docName="RafflesBZool.70.108133.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Upogebia singaporensis Dworschak &amp; Anker 2022, new species" docType="treatment" docUuid="1623DBDB-55ED-4F7D-BF6D-481F2760AF1C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="126" masterDocId="2A63522FFFD8FF905F54FFACFF90FFD3" masterDocTitle="Axiidea (Crustacea: Callianassidae, Callichiridae and Ctenochelidae) and Gebiidea (Upogebiidae) collected during the Comprehensive Marine Biodiversity Survey of Singapore" masterLastPageNumber="133" masterPageNumber="108" pageNumber="123" updateTime="1699380736798" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1FCEA044C0AE19341DFCB50FC6A973F8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="640D1FA71F47DF4B221FC295AF2B9FCD">Axiidea (Crustacea: Callianassidae, Callichiridae and Ctenochelidae) and Gebiidea (Upogebiidae) collected during the Comprehensive Marine Biodiversity Survey of Singapore</mods:title>
</mods:titleInfo>
<mods:name id="1292ABC33ACDF3359CEA686EDD32DBA0" type="personal">
<mods:role id="A5CBF30EDB45BE5E7FA4A65E4CD8AB6D">
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</mods:role>
<mods:namePart id="468BE2CBC8238AB5062302B0F4F26C93">Dworschak, Peter C.</mods:namePart>
</mods:name>
<mods:name id="029B9A42A3B3A54583DEB33BCA43606E" type="personal">
<mods:role id="C196990C442DD260AE2AD04D1E821105">
<mods:roleTerm id="8F9C395F1747FEBA8C3BB668AB94A77C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5888E347C9D7DE2DC0FD19BF64C4BD25">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="9C359E983F8C6CCF2A7FC2622958B50F">text</mods:typeOfResource>
<mods:relatedItem id="4D1163F6CCE10D03BBC0582F4DA3383D" type="host">
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<mods:title id="38CB71AF5C360EC08F07FED8DD327B79">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="8B95E94A74A8969112CD07EF41E8C4B3">
<mods:date id="543920854193C3831BC602083F2D3897">2022</mods:date>
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<mods:number id="BDF2B2C897F3C4660EA366BC0F7EAD4D">2022-03-23</mods:number>
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<mods:number id="12E3EE1211BADABB7AB227C80AC05860">70</mods:number>
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<mods:classification id="FB7E6B2CA9A6164D2EF0D5839B56E480">journal article</mods:classification>
<mods:identifier id="A3C1D9253B74BE679A5D03625EE50313" type="DOI">10.26107/RBZ-2022-0008</mods:identifier>
<mods:identifier id="1F3AC6BC6A0B1E4DD96405B7A6622216" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="BEFA100BF6B00F692D245EFDDF81AB10" type="Zenodo-Dep">7175098</mods:identifier>
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</mods:mods>
<treatment id="D65A2A57FFD7FF825CE0FC51FA3BFB5E" LSID="urn:lsid:zoobank.org:act:1623DBDB-55ED-4F7D-BF6D-481F2760AF1C" httpUri="http://treatment.plazi.org/id/D65A2A57FFD7FF825CE0FC51FA3BFB5E" lastPageId="18" lastPageNumber="126" pageId="15" pageNumber="123">
<subSubSection id="16E9C8CAFFD7FF9F5CE0FC51FAC2FBC6" box="[948,1362,1021,1045]" pageId="15" pageNumber="123" type="nomenclature">
<paragraph id="5E4C9B41FFD7FF9F5CE0FC51FAC2FBC6" blockId="15.[948,1362,1021,1077]" box="[948,1362,1021,1045]" pageId="15" pageNumber="123">
<heading id="05042C2DFFD7FF9F5CE0FC51FAC2FBC6" bold="true" box="[948,1362,1021,1045]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="123" reason="2">
<emphasis id="6C874753FFD7FF9F5CE0FC51FAC2FBC6" bold="true" box="[948,1362,1021,1045]" pageId="15" pageNumber="123">
<taxonomicName id="99F3E0C2FFD7FF9F5CE0FC51FB2DFBC6" ID-CoL="d3f01234-3354-4903-9f49-c313f1b1c44c" authority="Dworschak &amp; Anker, 2022" authorityName="Dworschak &amp; Anker" authorityYear="2022" box="[948,1213,1021,1045]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov.">
<emphasis id="6C874753FFD7FF9F5CE0FC51FB2DFBC6" bold="true" box="[948,1213,1021,1045]" italics="true" pageId="15" pageNumber="123">Upogebia singaporensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="77B4FA28FFD7FF9F5B98FC51FAC2FBC6" box="[1228,1362,1021,1045]" pageId="15" pageNumber="123" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFD7FF9F5B70FBB1FB71FBE6" box="[1060,1249,1053,1077]" pageId="15" pageNumber="123" type="description">
<paragraph id="5E4C9B41FFD7FF9F5B70FBB1FB71FBE6" blockId="15.[948,1362,1021,1077]" box="[1060,1249,1053,1077]" pageId="15" pageNumber="123">
(
<figureCitation id="C6C887C4FFD7FF9F5B78FBB1FB16FBE6" box="[1068,1158,1053,1077]" captionStart="Fig" captionStartId="1.[136,171,1856,1878]" captionTargetBox="[136,1474,311,1840]" captionTargetId="figure-7@1.[136,1474,311,1840]" captionTargetPageId="1" captionText="Fig. 1. Specimens, habitus (dorsal and/or ventral views), showing colouration in life: a, Gourretia sinica Liu &amp; Liu, 2010, female (23/5.3) (ZRC 2017.0947); b, Karumballichirus karumba (Poore &amp; Griffin, 1979), female (32/7.6) (ZRC 2018.0526); c, Aqaballianassa brevirostris Sakai, 2002, ovigerous female (28/5.6) (ZRC 2017.0948); d, Upogebia singaporensis, new species, female holotype (16/5.4) (ZRC 2017.0951); e, f, Neogebicula johorensis, new species, male holotype (18/5.6) (ZRC 2017.0956); g, h, same, ovigerous female paratype (21/6.3) (ZRC 2017.0955); i, j, Wolffogebia phuketensis Sakai, 1982, male (37/10.3) (NHMW 26036). Not to scale. [Photographs by AA]." figureDoi="http://doi.org/10.5281/zenodo.7175102" httpUri="https://zenodo.org/record/7175102/files/figure.png" pageId="15" pageNumber="123">Figs. 1d</figureCitation>
, 11, 12)
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFD7FF9F5C17FBF1FB9CF9E6" pageId="15" pageNumber="123" type="materials_examined">
<paragraph id="5E4C9B41FFD7FF9F5C17FBF1FAF8FA46" blockId="15.[835,1475,1117,1430]" pageId="15" pageNumber="123">
<materialsCitation id="EE9B911CFFD7FF9F5C17FBF1FAF5FA46" collectingDate="2013-05-25" collectingDateMax="2013-06-05" collectingDateMin="2013-05-25" collectionCode="CMBS" collectorName="Pulau Hantu &amp; KS Koh &amp; Koh, TS &amp; Ahyong, K &amp; HH Tan, S &amp; S De Grave &amp; B Richer de Forges" country="Singapore" county="Richer de Forges" elevation="4" latitude="1.2030001" location="Kusu Island" longLatPrecision="1" longitude="103.86875" municipality="Kusu Island" pageId="15" pageNumber="123" specimenCode="ZRC 2017.0951, ZRC 2017.0952, ZRC 2017.0950, ZRC 2017.0953" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="2" typeStatus="holotype">
<emphasis id="6C874753FFD7FF9F5C17FBF1FB8AFBA6" bold="true" box="[835,1050,1117,1141]" pageId="15" pageNumber="123">
<collectionCode id="38E20384FFD7FF9F5C17FBF1FC06FBA6" box="[835,918,1117,1141]" pageId="15" pageNumber="123">CMBS</collectionCode>
material.
</emphasis>
<typeStatus id="814825E3FFD7FF9F5B7CFBF1FB0EFBA6" box="[1064,1182,1117,1141]" pageId="15" pageNumber="123" type="holotype">Holotype</typeStatus>
: female (16/5.4) (
<specimenCode id="0E55333AFFD7FF9F5AD3FBF1FC2DFB46" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="123">ZRC 2017.0951</specimenCode>
), sta. IT120,
<collectorName id="F306FE97FFD7FF9F5B03FBD1FB74FB46" box="[1111,1252,1149,1173]" pageId="15" pageNumber="123">Pulau Hantu</collectorName>
, intertidal, coll.
<collectorName id="F306FE97FFD7FF9F5AF5FBD1FCE5FB66" pageId="15" pageNumber="123">KS Koh</collectorName>
, TS
<collectorName id="F306FE97FFD7FF9F5CF6FB31FB87FB66" box="[930,1047,1181,1205]" pageId="15" pageNumber="123">Ahyong, K</collectorName>
Tilbrook et al.,
<date id="2A4DBD81FFD7FF9F5BE9FB31FAD9FB66" box="[1213,1353,1181,1206]" pageId="15" pageNumber="123" value="2013-05-31">
<collectingDate id="3A094469FFD7FF9F5BE9FB31FAD9FB66" box="[1213,1353,1181,1206]" pageId="15" pageNumber="123" value="2013-05-31">31 May 2013</collectingDate>
</date>
(SS-3982).
<typeStatus id="814825E3FFD7FF9F5C17FB12FC3FFB05" box="[835,943,1214,1238]" pageId="15" pageNumber="123" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="48F550C8FFD7FF9F5CE8FB11FB8CFB06" box="[956,1052,1213,1237]" pageId="15" pageNumber="123" type="female">1 female</specimenCount>
(13/4.8) (
<specimenCode id="0E55333AFFD7FF9F5BDFFB11FAD1FB06" box="[1163,1345,1213,1237]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="123">ZRC 2017.0952</specimenCode>
),
<specimenCount id="48F550C8FFD7FF9F5A01FB11FA52FB06" box="[1365,1474,1213,1237]" pageId="15" pageNumber="123" type="juvenile">1 juvenile</specimenCount>
(12/3.9) (
<specimenCode id="0E55333AFFD7FF9F5CFDFB71FBCEFB26" box="[937,1118,1245,1269]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="123">ZRC 2017.0950</specimenCode>
), sta. SB055,
<location id="5B2CCD9AFFD7FF9F5BADFB72FAEFFB26" LSID="urn:lsid:plazi:treatment:D65A2A57FFD7FF825CE0FC51FA3BFB5E:5B2CCD9AFFD7FF9F5BADFB72FAEFFB26" box="[1273,1407,1245,1269]" country="Singapore" county="Richer de Forges" latitude="1.2030001" longLatPrecision="1" longitude="103.86875" municipality="Kusu Island" name="Kusu Island" pageId="15" pageNumber="123">Kusu Island</location>
, coral rubble brushing, ~
<quantity id="990B36A4FFD7FF9F5B5DFB51FBA5FAC6" box="[1033,1077,1277,1301]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="123" unit="m" value="4.0">
<elevation id="D5DE7C72FFD7FF9F5B5DFB51FBA5FAC6" box="[1033,1077,1277,1301]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="123" unit="m" value="4.0">4 m</elevation>
</quantity>
, coll.
<collectorName id="F306FE97FFD7FF9F5B2CFB52FB7DFAC6" box="[1144,1261,1277,1301]" pageId="15" pageNumber="123">HH Tan, S</collectorName>
De Grave et al.,
<date id="2A4DBD81FFD7FF9F5AF3FB51FC21FAE6" pageId="15" pageNumber="123" value="2013-05-25">
<collectingDate id="3A094469FFD7FF9F5AF3FB51FC21FAE6" pageId="15" pageNumber="123" value="2013-05-25">25 May 2013</collectingDate>
</date>
(SS-1631);
<specimenCount id="48F550C8FFD7FF9F5B67FAB1FB0DFAE6" box="[1075,1181,1309,1333]" pageId="15" pageNumber="123" type="juvenile">1 juvenile</specimenCount>
(11/3.6) (
<specimenCode id="0E55333AFFD7FF9F5A52FAB1FA28FAE6" box="[1286,1464,1309,1333]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="15" pageNumber="123">ZRC 2017.0953</specimenCode>
), sta. TB172,
<location id="5B2CCD9AFFD7FF9F5C94FA91FB93FA86" LSID="urn:lsid:plazi:treatment:D65A2A57FFD7FF825CE0FC51FA3BFB5E:5B2CCD9AFFD7FF9F5C94FA91FB93FA86" box="[960,1027,1341,1365]" country="Singapore" county="Richer de Forges" latitude="1.2030001" longLatPrecision="1" longitude="103.86875" municipality="Kusu Island" name="Straits" pageId="15" pageNumber="123">Straits</location>
of
<collectingCountry id="26E4DBD1FFD7FF9F5B76FA91FB1CFA86" box="[1058,1164,1341,1365]" name="Singapore" pageId="15" pageNumber="123">Singapore</collectingCountry>
near
<collectingMunicipality id="BE28013BFFD7FF9F5B97FA92FAD1FA86" box="[1219,1345,1341,1365]" pageId="15" pageNumber="123">Kusu Island</collectingMunicipality>
,
<geoCoordinate id="3BC7FD86FFD7FF9F5A1EFA90FA52FA86" box="[1354,1474,1340,1365]" degrees="1" direction="north" minutes="12.180" orientation="latitude" pageId="15" pageNumber="123" precision="1" value="1.2030001">1°12.180N</geoCoordinate>
<geoCoordinate id="3BC7FD86FFD7FF9F5C17FAF0FC48FAA6" box="[835,984,1372,1397]" degrees="103" direction="east" minutes="52.125" orientation="longitude" pageId="15" pageNumber="123" precision="1" value="103.86875">103°52.125E</geoCoordinate>
, consolidated marine clay,
<quantity id="990B36A4FFD7FF9F5A56FAF1FA16FAA6" box="[1282,1414,1373,1397]" metricMagnitude="1" metricUnit="m" metricValue="9.725" metricValueMax="10.0" metricValueMin="9.45" pageId="15" pageNumber="123" unit="m" value="97.25" valueMax="100.0" valueMin="94.5">
<elevation id="D5DE7C72FFD7FF9F5A56FAF1FA16FAA6" box="[1282,1414,1373,1397]" metricMagnitude="1" metricUnit="m" metricValue="9.725" metricValueMax="10.0" metricValueMin="9.45" pageId="15" pageNumber="123" unit="m" value="97.25" valueMax="100.0" valueMin="94.5">94.5100 m</elevation>
</quantity>
, coll.
<collectorName id="F306FE97FFD7FF9F5C17FAD2FB88FA45" box="[835,1048,1405,1430]" pageId="15" pageNumber="123">
B
<collectingCounty id="B72DE3CDFFD7FF9F5C0AFAD1FB88FA45" box="[862,1048,1405,1430]" pageId="15" pageNumber="123">Richer de Forges</collectingCounty>
</collectorName>
et al.,
<date id="2A4DBD81FFD7FF9F5B32FAD1FB7AFA46" box="[1126,1258,1405,1429]" pageId="15" pageNumber="123" value="2013-06-05">
<collectingDate id="3A094469FFD7FF9F5B32FAD1FB7AFA46" box="[1126,1258,1405,1429]" pageId="15" pageNumber="123" value="2013-06-05">5 June 2013</collectingDate>
</date>
(SS-4522)
</materialsCitation>
.
</paragraph>
<paragraph id="5E4C9B41FFD7FF9F5C17FA11FB9CF9E6" blockId="15.[834,1474,1469,1589]" pageId="15" pageNumber="123">
<emphasis id="6C874753FFD7FF9F5C17FA11FBD1FA06" bold="true" box="[835,1089,1469,1493]" pageId="15" pageNumber="123">Comparative material.</emphasis>
<taxonomicName id="99F3E0C2FFD7FF9F5B12FA11FA2EFA06" authority="Sakai, 2006" authorityName="Sakai" authorityYear="2006" box="[1094,1470,1469,1493]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="123" phylum="Arthropoda" rank="species" species="srilankaensis">
<emphasis id="6C874753FFD7FF9F5B12FA11FAACFA06" box="[1094,1340,1469,1493]" italics="true" pageId="15" pageNumber="123">Upogebia srilankaensis</emphasis>
<materialsCitation id="EE9B911CFFD7FF9F5A15FA11FA2EFA06" accessionNumber="SMF 30213" box="[1345,1470,1469,1493]" collectingDate="1912-12-03" collectionCode="SMF" collectorName="O. Low-Beer" country="Sri Lanka" latitude="5.9845667" location="Matara" longLatPrecision="1" longitude="80.32825" municipality="Weligama Bay" pageId="15" pageNumber="123" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="3A62E6B0FFD7FF9F5A15FA11FA2EFA06" author="Sakai K" box="[1345,1470,1469,1493]" pageId="15" pageNumber="123" pagination="1 - 185" refId="ref16901" refString="Sakai K (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6: 1 - 185." type="journal article" year="2006">Sakai, 2006</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<typeStatus id="814825E3FFD7FF9F5C16FA71FC32FA26" box="[834,930,1501,1525]" pageId="15" pageNumber="123" type="holotype">holotype</typeStatus>
, male (24/7.4) (
<collectionCode id="38E20384FFD7FF9F5B00FA71FB1BFA26" box="[1108,1163,1501,1525]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="15" pageNumber="123">SMF</collectionCode>
30213),
<collectingCountry id="26E4DBD1FFD7FF9F5BA6FA71FAF0FA26" box="[1266,1376,1501,1525]" name="Sri Lanka" pageId="15" pageNumber="123">Sri Lanka</collectingCountry>
,
<location id="5B2CCD9AFFD7FF9F5A3AFA72FA2CFA26" LSID="urn:lsid:plazi:treatment:D65A2A57FFD7FF825CE0FC51FA3BFB5E:5B2CCD9AFFD7FF9F5A3AFA72FA2CFA26" box="[1390,1468,1502,1525]" country="Sri Lanka" latitude="5.9845667" longLatPrecision="1" longitude="80.32825" municipality="Weligama Bay" name="Matara" pageId="15" pageNumber="123">Matara</location>
,
<collectingMunicipality id="BE28013BFFD7FF9F5C17FA51FC4EF9C5" box="[835,990,1533,1558]" pageId="15" pageNumber="123">Weligama Bay</collectingMunicipality>
(
<geoCoordinate id="3BC7FD86FFD7FF9F5CBFFA50FBF5F9C6" box="[1003,1125,1532,1557]" degrees="5" direction="north" minutes="59.074" orientation="latitude" pageId="15" pageNumber="123" precision="1" value="5.9845667">5°59.074N</geoCoordinate>
<geoCoordinate id="3BC7FD86FFD7FF9F5B3DFA50FB7FF9C6" box="[1129,1263,1532,1557]" degrees="80" direction="east" minutes="19.695" orientation="longitude" pageId="15" pageNumber="123" precision="1" value="80.32825">80°19.695E</geoCoordinate>
), coll.
<collectorName id="F306FE97FFD7FF9F5A67FA51FA2EF9C6" box="[1331,1470,1533,1557]" pageId="15" pageNumber="123">O. Löw-Beer</collectorName>
,
<date id="2A4DBD81FFD7FF9F5C17F9B1FB98F9E6" box="[835,1032,1565,1589]" pageId="15" pageNumber="123" value="1912-12-03">
<collectingDate id="3A094469FFD7FF9F5C17F9B1FB98F9E6" box="[835,1032,1565,1589]" pageId="15" pageNumber="123" value="1912-12-03">3 December 1912</collectingDate>
</date>
.
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFD7FF9F5C17F9F1FA53F886" pageId="15" pageNumber="123" type="diagnosis">
<paragraph id="5E4C9B41FFD7FF9F5C17F9F1FA53F886" blockId="15.[835,1475,1629,1877]" pageId="15" pageNumber="123">
<emphasis id="6C874753FFD7FF9F5C17F9F1FC28F9A6" bold="true" box="[835,952,1629,1653]" pageId="15" pageNumber="123">Diagnosis.</emphasis>
Rostrum elongate, with four strong dorsal spines, without infrarostral spine. Lateral ridges of gastric region weakly developed, with obtuse distal tooth and posterior to that tooth 911 tubercles. Postocular spine absent. Pleonal sternites unarmed. First pereopod chelate, merus lacking subdistal dorsal spine and bearing row of obtuse ventral teeth; carpus bearing 1 strong distoventral and 1 strong distodorsal spine. Meri of third to fifth pereopods unarmed.
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFD7FF815C17F8D1FA19F796" lastPageId="17" lastPageNumber="125" pageId="15" pageNumber="123" type="description">
<paragraph id="5E4C9B41FFD7FF815C17F8D1FE32FAD7" blockId="15.[835,1475,1917,2101]" lastBlockId="17.[136,775,1165,1285]" lastPageId="17" lastPageNumber="125" pageId="15" pageNumber="123">
<emphasis id="6C874753FFD7FF9F5C17F8D1FB2EF846" bold="true" box="[835,1214,1917,1941]" pageId="15" pageNumber="123">
Description of female
<typeStatus id="814825E3FFD7FF9F5B05F8D1FB28F846" box="[1105,1208,1917,1941]" pageId="15" pageNumber="123" type="holotype">holotype</typeStatus>
.
</emphasis>
Rostrum (Fig. 11a, b) elongate, twice as long as broad at base, setose on dorsal surface, bearing 1 pair of distal and 1 pair of subdistal dorsal spines, lacking infrarostral spine; lateral ridges of gastric region slightly protruding laterally, bearing row of 89 unequal tubercles; mid-dorsal area of gastric region with scattered tubercles. Cervical groove lacking spines. Postocular spine absent. Eyestalk stout, reaching to midlength of rostrum, unarmed. Linea thalassinica reaching to posterior end of carapace.
</paragraph>
<paragraph id="5E4C9B41FFC9FF815FDCFA81FEF6FA76" blockId="17.[136,775,1325,1445]" pageId="17" pageNumber="125">Antennular peduncle (Figs. 11a, 12a) shorter than antennal peduncle; articles unarmed, second article short, third article elongate; flagella of same length, dorsal thicker than ventral, as long as peduncle.</paragraph>
<paragraph id="5E4C9B41FFC9FF815FDCFA61FE71F9F6" blockId="17.[136,775,1485,1573]" pageId="17" pageNumber="125">Antennal peduncle (Figs. 11a, 12b) overreaching rostrum by distal margin of penultimate article; articles unarmed; scaphocerite weakly developed.</paragraph>
<paragraph id="5E4C9B41FFC9FF815FDCF9E1FEF7F8F7" blockId="17.[136,776,1613,1829]" pageId="17" pageNumber="125">
Epistome (Fig. 11a) rounded, unarmed. Mandible (Fig. 12c, d) without mesio-anterior tooth. First and second maxilla as illustrated (Fig. 12e, f). First maxilliped (Fig.
<quantity id="990B36A4FFC9FF815FDCF901FF22F916" box="[136,178,1709,1733]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="17" pageNumber="125" unit="g" value="12.0">12g</quantity>
) without epipod. Second maxilliped (Fig. 12h) with epipod. Third maxilliped (Fig. 12i) without epipod; exopod simple (without flagellum) distally, reaching almost to end of endopodal merus.
</paragraph>
<paragraph id="5E4C9B41FFC9FF815FDCF8E1FB44FAB6" blockId="17.[136,776,1869,2117]" lastBlockId="17.[835,1474,1165,1381]" pageId="17" pageNumber="125">First pereopod (Fig. 11cf) chelate; coxa and basis unarmed. Ischium bearing 1 subterminal spine on ventral margin. Merus bearing row of 1314 obtuse thick spines on ventral margin, increasing in size to mid-length of merus, distal ones diminishing in size; dorsal margin unarmed. Carpus triangular, lateral surface bearing longitudinal ventral carina; 1 strong, mesially directed dorsal spine, 1 small median spine, and 1 prominent dorsal spine present on distal margin; 6 small spines present on mesial surface. Palm 2.1 times as long as high; upper border unarmed, with several low tubercles on inner dorsal surface; lower border keeled with 78 tubercles proximally; 3 tubercles present on lower inner surface proximally. Fixed finger 0.3 times length of palm, with low denticles on cutting edge. Dactylus (Fig. 11d, f) with proximal tooth on cutting edge.</paragraph>
<paragraph id="5E4C9B41FFC9FF815C17FA21FB45F996" blockId="17.[835,1475,1421,1605]" pageId="17" pageNumber="125">
Second pereopod (Fig. 11h) unarmed; merus 5.3 times as long as wide; propodus 2.8 times as long as wide. Third and fourth pereopods (Fig. 11i, k) simple; dactylus with comb on lower border and 5 spaced corneous spines on upper border (Fig. 11j, l). Fifth pereopod (Fig.
<quantity id="990B36A4FFC9FF815BFFF9A1FB4CF9F6" box="[1195,1244,1549,1573]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="125" unit="m" value="11.0">11m</quantity>
) subchelate; dactylus curved, with corneous tip (Fig. 11n).
</paragraph>
<paragraph id="5E4C9B41FFC9FF815C17F9C1FA3EF8F6" blockId="17.[835,1474,1645,1829]" pageId="17" pageNumber="125">Telson (Fig. 11p) slightly longer than maximal width; dorsal surface with U-shaped concavity, latter with denticulate transverse carina. Uropodal endopod broadened, about as long as telson, with somewhat truncate, slightly convex distal margin; uropodal exopod broadened, about as long as endopod, with prominent spine proximally (Fig. 11o).</paragraph>
<paragraph id="5E4C9B41FFC9FF815C17F8E1FB11F856" blockId="17.[835,1474,1869,1925]" pageId="17" pageNumber="125">
Arthrobranchs of
<typeStatus id="814825E3FFC9FF815B56F8E1FBA0F8B6" box="[1026,1072,1869,1893]" pageId="17" pageNumber="125">type</typeStatus>
A, one pair each on third maxilliped, and first to fourth pereopods.
</paragraph>
<paragraph id="5E4C9B41FFC9FF815C17F801FA19F796" blockId="17.[834,1475,1965,2117]" pageId="17" pageNumber="125">
<emphasis id="6C874753FFC9FF815C17F801FC52F816" bold="true" box="[835,962,1965,1989]" pageId="17" pageNumber="125">Variations.</emphasis>
The number of spines on the lower border of the first pereopod merus ranges from 9 (juvenile 11/3.6) to 15 (female 13/4.8) on the right side and between 9 and 14 on the left side. The teeth on the lower border of first pereopod propodus differ between sides and range from 3 to 8.
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFCAFF825F25FC79FF75FBDE" pageId="18" pageNumber="126" type="etymology">
<paragraph id="5E4C9B41FFCAFF825F25FC79FF75FBDE" blockId="18.[113,751,981,1037]" pageId="18" pageNumber="126">
<emphasis id="6C874753FFCAFF825F25FC79FF66FC3E" bold="true" box="[113,246,981,1005]" pageId="18" pageNumber="126">Etymology.</emphasis>
Named after the
<typeStatus id="814825E3FFCAFF825E95FC7AFE60FC3D" box="[449,496,982,1006]" pageId="18" pageNumber="126">type</typeStatus>
locality, the Straits of
<collectingCountry id="26E4DBD1FFCAFF825F25FC59FF71FBDE" box="[113,225,1013,1037]" name="Singapore" pageId="18" pageNumber="126">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFCAFF825F25FB99FE1DFAFE" pageId="18" pageNumber="126" type="distribution">
<paragraph id="5E4C9B41FFCAFF825F25FB99FD70FB9E" blockId="18.[113,736,1077,1101]" box="[113,736,1077,1101]" pageId="18" pageNumber="126">
<emphasis id="6C874753FFCAFF825F25FB99FE94FB9E" bold="true" box="[113,260,1077,1101]" pageId="18" pageNumber="126">Distribution.</emphasis>
Only known from the Straits of
<collectingCountry id="26E4DBD1FFCAFF825D39FB99FD4DFB9E" box="[621,733,1077,1101]" name="Singapore" pageId="18" pageNumber="126">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="5E4C9B41FFCAFF825F25FBD9FE1DFAFE" blockId="18.[113,754,1141,1325]" pageId="18" pageNumber="126">
<emphasis id="6C874753FFCAFF825F25FBD9FF40FB5E" bold="true" box="[113,208,1141,1165]" pageId="18" pageNumber="126">Habitat.</emphasis>
<taxonomicName id="99F3E0C2FFCAFF825F8CFBD9FE4DFB5E" authority="Dworschak &amp; Anker, 2022" authorityName="Dworschak &amp; Anker" authorityYear="2022" box="[216,477,1141,1165]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov.">
<emphasis id="6C874753FFCAFF825F8CFBD9FE4DFB5E" box="[216,477,1141,1165]" italics="true" pageId="18" pageNumber="126">Upogebia singaporensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="77B4FA28FFCAFF825EB8FBDAFDE1FB5E" box="[492,625,1141,1165]" pageId="18" pageNumber="126" rank="species">new species</taxonomicNameLabel>
, has a wide bathymetric distribution:
<specimenCount id="48F550C8FFCAFF825E2CFB3AFE79FB7E" box="[376,489,1173,1197]" pageId="18" pageNumber="126" type="female">one female</specimenCount>
(16.1/5.42) was collected in the intertidal zone at Pulau Hantu (together with
<taxonomicName id="99F3E0C2FFCAFF825D82FB1AFE97FB3E" authorityName="de Man" authorityYear="1905" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="ancylodactyla">
<emphasis id="6C874753FFCAFF825D82FB1AFE97FB3E" italics="true" pageId="18" pageNumber="126">U. ancylodactyla</emphasis>
</taxonomicName>
);
<specimenCount id="48F550C8FFCAFF825E75FB7AFE51FB3E" box="[289,449,1237,1261]" pageId="18" pageNumber="126" type="generic">two specimens</specimenCount>
came from coral brushings at
<quantity id="990B36A4FFCAFF825FDFFB59FF24FADE" box="[139,180,1269,1293]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="126" unit="m" value="4.0">4 m</quantity>
depth at Kusu Island, and one from consolidated clay at a depth of
<quantity id="990B36A4FFCAFF825E50FAB9FE19FAFE" box="[260,393,1301,1325]" metricMagnitude="1" metricUnit="m" metricValue="9.725" metricValueMax="10.0" metricValueMin="9.45" pageId="18" pageNumber="126" unit="m" value="97.25" valueMax="100.0" valueMin="94.5">
94.5
<specimenCount id="48F550C8FFCAFF825E16FAB9FE19FAFE" box="[322,393,1301,1325]" pageId="18" pageNumber="126" type="generic">100 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="16E9C8CAFFCAFF825F25FAF9FA3BFB5E" pageId="18" pageNumber="126" type="discussion">
<paragraph id="5E4C9B41FFCAFF825F25FAF9FDEBF89E" blockId="18.[112,754,1365,1869]" pageId="18" pageNumber="126">
<emphasis id="6C874753FFCAFF825F25FAF9FF7BFABE" bold="true" box="[113,235,1365,1389]" pageId="18" pageNumber="126">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="99F3E0C2FFCAFF825D2FFAF9FE13FA5E" authority="Sakai, 2006" authorityName="Sakai" authorityYear="2006" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="srilankaensis">
<emphasis id="6C874753FFCAFF825D2FFAF9FF6EFA5E" italics="true" pageId="18" pageNumber="126">Upogebia srilankaensis</emphasis>
<bibRefCitation id="3A62E6B0FFCAFF825E51FAD9FE13FA5E" author="Sakai K" box="[261,387,1397,1421]" pageId="18" pageNumber="126" pagination="1 - 185" refId="ref16901" refString="Sakai K (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6: 1 - 185." type="journal article" year="2006">
Sakai,
<quantity id="990B36A4FFCAFF825E1AFAD9FE13FA5E" box="[334,387,1397,1421]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="18" pageNumber="126" unit="in" value="2006.0">2006</quantity>
</bibRefCitation>
</taxonomicName>
in possessing a long rostrum with two pairs of prominent dorsal spines (1 distal and 1 subdistal pair); in the shape of the spines on the lower border of the first pereopod merus; in having upper and lower spines on the first pereopod carpus; and in the general shape of telson and uropods. However,
<taxonomicName id="99F3E0C2FFCAFF825E33F9BAFD87F9FE" authority="Dworschak &amp; Anker, 2022" authorityName="Dworschak &amp; Anker" authorityYear="2022" box="[359,535,1558,1581]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov.">
<emphasis id="6C874753FFCAFF825E33F9BAFD87F9FE" box="[359,535,1558,1581]" italics="true" pageId="18" pageNumber="126">U. singaporensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="77B4FA28FFCAFF825D70F9BAFD33F9FE" box="[548,675,1557,1581]" pageId="18" pageNumber="126" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="99F3E0C2FFCAFF825FFAF99AFECDF99E" authorityName="Sakai" authorityYear="2006" box="[174,349,1589,1613]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="srilankaensis">
<emphasis id="6C874753FFCAFF825FFAF99AFECDF99E" box="[174,349,1589,1613]" italics="true" pageId="18" pageNumber="126">U. srilankaensis</emphasis>
</taxonomicName>
(characters in parentheses) by 1) the absence of a ventral spine on the first article of the antennal peduncle (present); 2) the slightly produced lateral ridges of the gastric carapace region (not protruding); 3) the dorsal spine on the mesial surface of the first pereopod carpus prominent (much smaller); 4) the ventral margin of the first pereopod propodus with several tubercles proximally (without tubercles); and 5) the fixed finger of the first pereopod entire (with a deep concavity on the occlusal margin).
</paragraph>
<paragraph id="5E4C9B41FFCAFF825F25F8D9FA3BFB5E" blockId="18.[113,753,1909,2093]" lastBlockId="18.[812,1452,981,1165]" pageId="18" pageNumber="126">
A re-examination of the
<typeStatus id="814825E3FFCAFF825E39F8D9FE59F85E" box="[365,457,1909,1933]" pageId="18" pageNumber="126" type="holotype">holotype</typeStatus>
of
<taxonomicName id="99F3E0C2FFCAFF825EBCF8DAFD01F85E" authorityName="Sakai" authorityYear="2006" box="[488,657,1909,1933]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="126" phylum="Arthropoda" rank="species" species="srilankaensis">
<emphasis id="6C874753FFCAFF825EBCF8DAFD01F85E" box="[488,657,1909,1933]" italics="true" pageId="18" pageNumber="126">U. srilankaensis</emphasis>
</taxonomicName>
(Fig. 13) revealed some inconsistencies and omissions in the original description by
<bibRefCitation id="3A62E6B0FFCAFF825E46F819FE30F81E" author="Sakai K" box="[274,416,1973,1997]" pageId="18" pageNumber="126" pagination="1 - 185" refId="ref16901" refString="Sakai K (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6: 1 - 185." type="journal article" year="2006">Sakai (2006)</bibRefCitation>
: 1) the antennal peduncle has a distinct, acute scaphocerite; 2) the first pereopod palm has a row of three tubercles proximally on the lower mesial surface (only one visible in lateral view in
<bibRefCitation id="3A62E6B0FFCAFF825D14F7B9FD52F7FE" author="Sakai K" box="[576,706,2069,2093]" pageId="18" pageNumber="126" pagination="1 - 185" refId="ref16901" refString="Sakai K (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6: 1 - 185." type="journal article" year="2006">Sakai, 2006</bibRefCitation>
: fig. 23D); 3) there are two spines on the upper mesial face of the first pereopod carpus; 4) the left first pereopod has two spines ventrally on the ischium; and 5) the third and fourth pereopod dactyli have a comb on the lower border and a row of corneous spines on the upper border, as in the present new species (not illustrated nor mentioned by
<bibRefCitation id="3A62E6B0FFCAFF825A48FBD9FA30FB5E" author="Sakai K" box="[1308,1440,1141,1165]" pageId="18" pageNumber="126" pagination="1 - 185" refId="ref16901" refString="Sakai K (2006) Upogebiidae of the world (Decapoda, Thalassinidea). Crustaceana Monographs, 6: 1 - 185." type="journal article" year="2006">Sakai, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>