304 lines
41 KiB
XML
304 lines
41 KiB
XML
<document id="2929D107DB12EA3C833E393E90FD0CED" ID="10.11646/zootaxa.4189.1.8" ID-DOI="10.11646/zootaxa.4189.1.8" ID-GBIF-Dataset="b5e15f1c-e028-4aa6-ad2a-4cd8421e2d72" ID-ISSN="1175-5326" ID-Zenodo-Dep="165712" ID-ZooBank="61692FA8-DF4B-4098-8F4D-3B332677D5CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478678021184" checkinUser="plazi" docAuthor="Mjobo, Sifiso & Thandar, Ahmed S." docDate="2016" docId="C67F6425FFA52F37FF0BF710FF0C1F51" docLanguage="en" docName="zootaxa.4189.1.8.pdf" docOrigin="Zootaxa 4189 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemiocnus rubrobrunneus Mjobo & Thandar, 2016, n.sp." docType="treatment" docVersion="9" lastPageNumber="162" masterDocId="3A461C5DFFA12F30FF9CF22FFFE71E4C" masterDocTitle="A new genus and a new species in the sea cucumber subfamily Colochirinae (Echinodermata: Holothuroidea: Dendrochirotida: Cucumariidae) in the Mediterranean Sea" masterLastPageNumber="164" masterPageNumber="156" pageNumber="160" updateTime="1698392810130" updateUser="plazi">
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<mods:mods id="F6C8B03A9E3D45EA348AF24AB342D6EC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="75DCB50F20DDDA9AF64580FD91F7CBA9">
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<mods:title id="C2185087DB3794C5C233B2FEC6F9783F">A new genus and a new species in the sea cucumber subfamily Colochirinae (Echinodermata: Holothuroidea: Dendrochirotida: Cucumariidae) in the Mediterranean Sea</mods:title>
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</mods:titleInfo>
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<mods:name id="37DEA55AC9C8CC28F6975823B1E7A37D" type="personal">
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<mods:roleTerm id="FF2133874CEEE1295857E729FB1931B3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="14ABB61CCF7136F23F40A8A901D73DBB">Mjobo, Sifiso</mods:namePart>
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</mods:name>
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<mods:name id="E7FB9DE18BED6C4245365B4F8E94E061" type="personal">
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<mods:roleTerm id="3ADC825F9835421B8BF9A1DF868B77FB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E277A952391457FD96645258246D3DB3">Thandar, Ahmed S.</mods:namePart>
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<mods:typeOfResource id="AE218C25BA94F3231F54C23D8A367748">text</mods:typeOfResource>
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<mods:titleInfo id="F6F4C487B1F93E76411A0FB4582992BA">
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<mods:title id="4D765202C4C5CF3FF240674060696A9A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="963E70965E5E4EE916BEAC03CC19FC2D">
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<mods:date id="A27475B963FA49AC09529B93E0751F39">2016</mods:date>
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<mods:detail id="0EB8A42C4CF6933FA42B13BE251A5406" type="volume">
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<mods:number id="B38B058C78CACF2DFA619925F71E7CDF">4189</mods:number>
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</mods:detail>
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<mods:detail id="F103F37FDA84427669EEAD5E914E64BB" type="issue">
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<mods:number id="6C51FE7475D2BAB24A144FC27C3D2161">1</mods:number>
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<mods:classification id="0089282C2979A98276AB566E26B002AD">journal article</mods:classification>
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<mods:identifier id="8663C29BF91505DB0AFBA9BE1A2EB448" type="DOI">10.11646/zootaxa.4189.1.8</mods:identifier>
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<mods:identifier id="0CF1B9C7CB117ED0D28831853BCB9737" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="09A16A25E926F2D06CE572C18E9767CE" type="Zenodo-Dep">165712</mods:identifier>
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<treatment id="C67F6425FFA52F37FF0BF710FF0C1F51" ID-DOI="http://doi.org/10.5281/zenodo.6087193" ID-GBIF-Taxon="124658044" ID-Zenodo-Dep="6087193" LSID="urn:lsid:plazi:treatment:C67F6425FFA52F37FF0BF710FF0C1F51" httpUri="http://treatment.plazi.org/id/C67F6425FFA52F37FF0BF710FF0C1F51" lastPageId="7" lastPageNumber="162" pageId="4" pageNumber="160">
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<subSubSection id="06CC86B8FFA52F34FF0BF710FEC91B37" pageId="4" pageNumber="160" type="nomenclature">
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<paragraph id="4E69D533FFA52F34FF0BF710FDC41B16" blockId="4.[151,547,1343,1403]" box="[151,547,1343,1370]" pageId="4" pageNumber="160">
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<heading id="1521625FFFA52F34FF0BF710FDC41B16" bold="true" box="[151,547,1343,1370]" fontSize="11" level="1" pageId="4" pageNumber="160" reason="1">
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<emphasis id="7CA20921FFA52F34FF0BF710FDC41B16" bold="true" box="[151,547,1343,1370]" pageId="4" pageNumber="160">
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<taxonomicName id="89D6AEB0FFA52F34FF0BF710FE061B15" ID-CoL="6LP4B" box="[151,481,1343,1369]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="160" phylum="Echinodermata" rank="species" species="rubrobrunneus" status="sp. nov.">
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<emphasis id="7CA20921FFA52F34FF0BF710FE061B15" bold="true" box="[151,481,1343,1369]" italics="true" pageId="4" pageNumber="160">Hemiocnus rubrobrunneus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6791B45AFFA52F34FE74F76FFDC41B16" box="[488,547,1344,1370]" pageId="4" pageNumber="160" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4E69D533FFA52F34FF0BF74DFEC91B37" blockId="4.[151,547,1343,1403]" box="[151,302,1378,1403]" pageId="4" pageNumber="160">
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(
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<figureCitation id="D6EDC9B6FFA52F34FF02F74DFEC11B37" box="[158,294,1378,1403]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="5.[151,250,1702,1724]" captionStartId-1="6.[151,250,1591,1613]" captionStartId-2="7.[151,250,1089,1111]" captionStartId-3="7.[151,250,1872,1894]" captionTargetBox-0="[340,1246,701,1675]" captionTargetBox-1="[340,1246,553,1569]" captionTargetBox-3="[369,1224,1175,1843]" captionTargetId-0="figure@5.[340,1247,701,1678]" captionTargetId-1="figure@6.[340,1247,553,1569]" captionTargetId-3="figure@7.[340,1248,1161,1851]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-3="7" captionText-0="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." captionText-1="FIGURE 4. Hemiocnus rubrobrunneus n. sp. A. Rosettes from tentacles; B. Small rods from tentacles; C. Large rods from tentacles. D. Paratype (entire)." captionText-2="FIGURE 5. Hemiocnus rubrobrunneus n. sp. Plates from the anal region." captionText-3="FIGURE 6. Hemiocnus rubrobrunneus. A. Rods from tentacles; B. Rosettes from tentacles; C. Rods from tube feet; D. Rosettes from introvert; E. Rods from introvert; F. Round knobbed button from introvert; G. Part of calcareous ring. A, D, E & F = scale a." httpUri-0="https://zenodo.org/record/165715/files/figure.png" httpUri-1="https://zenodo.org/record/256162/files/figure.png" httpUri-3="https://zenodo.org/record/165717/files/figure.png" pageId="4" pageNumber="160">Figures 3–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="06CC86B8FFA52F34FF0BF785FAAA183B" pageId="4" pageNumber="160" type="diagnosis">
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<paragraph id="4E69D533FFA52F34FF0BF785FAAA183B" blockId="4.[151,1437,1450,2015]" pageId="4" pageNumber="160">
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<emphasis id="7CA20921FFA52F34FF0BF785FEF61B8F" bold="true" box="[151,273,1450,1475]" pageId="4" pageNumber="160">Diagnosis.</emphasis>
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Species soft, flaccid; body subcylindrical to barrel-shaped; colour in alcohol dark reddish-brown in bright light. Length up to
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<quantity id="892E78D6FFA52F34FE59F7E0FDFD1BAA" box="[453,538,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="160" unit="mm" value="32.0">32 mm</quantity>
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. Mouth and anus terminal. Anal teeth/papillae absent. Tube feet short, mostly non-retractile, two rows per ambulacrum, few also scattered in dorsal interambulacra. Body wall ossicles comprise a layer of complete and incomplete baskets, accompanied by rosettes, in addition to small, rounded, knobbed plates. Large smooth to knobbed plates sometimes present in anal region. Tube feet deposits comprise baskets, rods and rosettes. Tentacles and introvert with rods and rosettes; knobbed, rounded plates also occur in introvert.
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</paragraph>
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</subSubSection>
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<subSubSection id="06CC86B8FFA52F34FF5BF4ADFC2318D7" box="[199,964,1666,1691]" pageId="4" pageNumber="160" type="etymology">
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||
<paragraph id="4E69D533FFA52F34FF5BF4ADFC2318D7" blockId="4.[151,1437,1450,2015]" box="[199,964,1666,1691]" pageId="4" pageNumber="160">
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<emphasis id="7CA20921FFA52F34FF5BF4ADFEA918D7" bold="true" box="[199,334,1666,1691]" pageId="4" pageNumber="160">Etymology.</emphasis>
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The species is named for its reddish-brown colouration.
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</paragraph>
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</subSubSection>
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<subSubSection id="06CC86B8FFA52F34FF5BF489FBC4194B" pageId="4" pageNumber="160" type="materials_examined">
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<paragraph id="4E69D533FFA52F34FF5BF489FBC4194B" blockId="4.[151,1437,1450,2015]" pageId="4" pageNumber="160">
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<materialsCitation id="FEBEDF6EFFA52F34FF5BF489FDED18AE" ID-GBIF-Occurrence="1324949261" collectionCode="USNM" elevation="1" location="Jerba Island" pageId="4" pageNumber="160" specimenCount="1" typeStatus="holotype">
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<emphasis id="7CA20921FFA52F34FF5BF489FE5618F3" bold="true" box="[199,433,1702,1727]" pageId="4" pageNumber="160">Material examined.</emphasis>
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<typeStatus id="916D6B91FFA52F34FE24F489FDC218F3" box="[440,549,1702,1727]" pageId="4" pageNumber="160" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="28C74DF6FFA52F34FDB3F488FD6718F3" box="[559,640,1703,1727]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="160">USNM</collectionCode>
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(Smithsonian Institution),
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<accessionNumber id="518548D0FFA52F34FC33F489FBEE18F3" box="[943,1033,1702,1727]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E22590" pageId="4" pageNumber="160" type="EnaNcbi">E22590</accessionNumber>
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,
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<location id="4B0983E8FFA52F34FB89F489FB7918F3" LSID="urn:lsid:plazi:treatment:C67F6425FFA52F37FF0BF710FF0C1F51:4B0983E8FFA52F34FB89F489FB7918F3" box="[1045,1182,1702,1727]" name="Jerba Island" pageId="4" pageNumber="160">Jerba Island</location>
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,
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<locationDeviation id="5FDBCC11FFA52F34FB36F489FE2D18AF" pageId="4" pageNumber="160">between Houmt Sauq and Bordj Djillidj, Tunisia</locationDeviation>
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,
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||
<quantity id="892E78D6FFA52F34FE4BF4E4FDED18AE" box="[471,522,1739,1763]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="160" unit="m" value="1.0">
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||
<specimenCount id="58D01EBAFFA52F34FE4BF4E4FDED18AE" box="[471,522,1739,1763]" pageId="4" pageNumber="160" type="generic">
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<elevation id="C5FB3200FFA52F34FE4BF4E4FDED18AE" box="[471,522,1739,1763]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="160" unit="m" value="1.0">1 m</elevation>
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</specimenCount>
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||
</quantity>
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||
</materialsCitation>
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||
.,
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<materialsCitation id="FEBEDF6EFFA52F34FDBAF4E4FBC4194B" ID-GBIF-Occurrence="1324949259" collectionCode="USNM" collectorName="Jones, M. L. & C. Gust & C. Ahearn" elevation="1" location="Jerba Island" pageId="4" pageNumber="160" specimenCount="1" typeStatus="paratype">
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||
<collectorName id="E323B0E5FFA52F34FDBAF4E4FD4A18AE" box="[550,685,1739,1762]" pageId="4" pageNumber="160">Jones, M.L.</collectorName>
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1973;
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<typeStatus id="916D6B91FFA52F34FD63F4E4FC8818AF" box="[767,879,1739,1763]" pageId="4" pageNumber="160" type="paratype">Paratypes</typeStatus>
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, same data as holotype, 2 spec. (all specimens formerly determined as
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<taxonomicName id="89D6AEB0FFA52F34FE3FF4C1FD27194A" box="[419,704,1774,1798]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnella" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="160" phylum="Echinodermata" rank="species" species="syracusana">
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<emphasis id="7CA20921FFA52F34FE3FF4C1FD27194A" box="[419,704,1774,1798]" italics="true" pageId="4" pageNumber="160">Pseudocnella syracusana</emphasis>
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</taxonomicName>
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by
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<collectorName id="E323B0E5FFA52F34FD76F4C0FCA7194B" box="[746,832,1775,1799]" pageId="4" pageNumber="160">C. Gust</collectorName>
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||
(
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<collectorName id="E323B0E5FFA52F34FCCCF4C0FC2F194B" box="[848,968,1774,1799]" pageId="4" pageNumber="160">C. Ahearn</collectorName>
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), 1981).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="06CC86B8FFA52F35FF5BF53DFBD01CCA" lastPageId="5" lastPageNumber="161" pageId="4" pageNumber="160" type="description">
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<paragraph id="4E69D533FFA52F35FF5BF53DFC3C1EB6" blockId="4.[151,1437,1450,2015]" lastBlockId="5.[151,1437,151,646]" lastPageId="5" lastPageNumber="161" pageId="4" pageNumber="160">
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||
<emphasis id="7CA20921FFA52F34FF5BF53DFE091967" bold="true" box="[199,494,1810,1835]" pageId="4" pageNumber="160">
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Description of
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||
<typeStatus id="916D6B91FFA52F34FE1CF53DFE0E1967" box="[384,489,1810,1835]" pageId="4" pageNumber="160" type="holotype">holotype</typeStatus>
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||
.
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</emphasis>
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Form subcylindrical to barrel-shaped (
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<figureCitation id="D6EDC9B6FFA52F34FC25F53DFBF91967" box="[953,1054,1810,1835]" captionStart="FIGURE 4" captionStartId="6.[151,250,1591,1613]" captionTargetBox="[340,1246,553,1569]" captionTargetId="figure@6.[340,1247,553,1569]" captionTargetPageId="6" captionText="FIGURE 4. Hemiocnus rubrobrunneus n. sp. A. Rosettes from tentacles; B. Small rods from tentacles; C. Large rods from tentacles. D. Paratype (entire)." httpUri="https://zenodo.org/record/256162/files/figure.png" pageId="4" pageNumber="160">Figure 4</figureCitation>
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D),
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||
<quantity id="892E78D6FFA52F34FBD2F53CFB451966" box="[1102,1186,1811,1835]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="160" unit="mm" value="29.0">29 mm</quantity>
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||
in length,
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||
<quantity id="892E78D6FFA52F34FABBF53CFA9C1966" box="[1319,1403,1811,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="160" unit="mm" value="13.0">13 mm</quantity>
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in breadth in mid-body (
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<typeStatus id="916D6B91FFA52F34FE15F518FE111903" box="[393,502,1847,1871]" pageId="4" pageNumber="160" type="paratype">paratypes</typeStatus>
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||
<date id="3A68F3F3FFA52F34FE61F518FDB61903" box="[509,593,1847,1871]" pageId="4" pageNumber="160" value="2012-10-29">
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||
29 x
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||
<quantity id="892E78D6FFA52F34FDA9F518FDB61903" box="[565,593,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="160" unit="mm" value="12.0">12</quantity>
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</date>
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||
mm and
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||
<date id="3A68F3F3FFA52F34FD25F518FCEA1903" box="[697,781,1847,1871]" pageId="4" pageNumber="160" value="1932-10-12">
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||
32 x
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||
<quantity id="892E78D6FFA52F34FD6DF518FCEA1903" box="[753,781,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="160" unit="mm" value="12.0">12</quantity>
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||
</date>
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||
mm). Skin soft, colour in alcohol, a dark reddish brown in bright light. Mouth and anus terminal. Tube feet short, distributed in two distinct rows ventrally, and in somewhat zigzag rows dorsally with few also in dorsal interrambulacra. Tentacles retracted, bushy, 10, ventral-most two reduced. Calcareous ring (
|
||
<figureCitation id="D6EDC9B6FFA52F34FE5EF58DFDC319F7" box="[450,548,1954,1979]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[369,1224,1175,1843]" captionTargetId="figure@7.[340,1248,1161,1851]" captionTargetPageId="7" captionText="FIGURE 6. Hemiocnus rubrobrunneus. A. Rods from tentacles; B. Rosettes from tentacles; C. Rods from tube feet; D. Rosettes from introvert; E. Rods from introvert; F. Round knobbed button from introvert; G. Part of calcareous ring. A, D, E & F = scale a." httpUri="https://zenodo.org/record/165717/files/figure.png" pageId="4" pageNumber="160">Figure 6</figureCitation>
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G) typically cucumariid, simple, without posterior prolongations, radial plates bifid anteriorly, all plates deeply notched posteriorly. Madreporite bean-shaped, stone canal short; Polian vesicle single. Retractor muscles short, arising from longitudinal bands at about half body length from anterior end. Longitudinal muscles not paired. Respiratory trees well branched, about ¾ of body length from posterior end. Gonad in single tuft, tubules unbranched, partially full of developing eggs.
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||
</paragraph>
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<paragraph id="4E69D533FFA42F35FF5BF32AFBD01CCA" blockId="5.[151,1437,151,646]" pageId="5" pageNumber="161">
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||
Body wall deposits consist of baskets and knobbed, perforated buttons/plates and rosettes. Buttons (70–100 µm) with mostly 4(–6) holes, rarely more, most about 100 µm, with 1–3 central knobs, rarely more (
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||
<figureCitation id="D6EDC9B6FFA42F35FA8AF306FA901F0D" box="[1302,1399,297,322]" captionStart="FIGURE 3" captionStartId="5.[151,250,1702,1724]" captionTargetBox="[340,1246,701,1675]" captionTargetId="figure@5.[340,1247,701,1678]" captionTargetPageId="5" captionText="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161">Figure 3</figureCitation>
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A). Baskets numerous, both complete and incomplete (
|
||
<figureCitation id="D6EDC9B6FFA42F35FD7CF362FCAA1F2A" box="[736,845,333,358]" captionStart="FIGURE 3" captionStartId="5.[151,250,1702,1724]" captionTargetBox="[340,1246,701,1675]" captionTargetId="figure@5.[340,1247,701,1678]" captionTargetPageId="5" captionText="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161">Figures 3</figureCitation>
|
||
B & D), incomplete baskets appearing as X- or Yshaped simple, smooth deposits with dichotomous branchings, whereas complete baskets appear as shallow, flat cups with blunt spines around the rim. Rosettes appear as both open and closed
|
||
<typeStatus id="916D6B91FFA42F35FBE3F3B9FB5A1FE2" box="[1151,1213,406,430]" pageId="5" pageNumber="161">types</typeStatus>
|
||
. Anal region with multilocular, smooth and/or knobbed plates (
|
||
<figureCitation id="D6EDC9B6FFA42F35FD13F396FD161F9E" box="[655,753,441,466]" captionStart="FIGURE 5" captionStartId="7.[151,250,1089,1111]" captionText="FIGURE 5. Hemiocnus rubrobrunneus n. sp. Plates from the anal region." pageId="5" pageNumber="161">Figure 5</figureCitation>
|
||
). Tube feet deposits include rods, rosettes and baskets. Rods simple, mostly table-like, perforated at ends and with a spire-like medial extension (
|
||
<figureCitation id="D6EDC9B6FFA42F35FBC9F3F2FB501FBA" box="[1109,1207,477,502]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[369,1224,1175,1843]" captionTargetId="figure@7.[340,1248,1161,1851]" captionTargetPageId="7" captionText="FIGURE 6. Hemiocnus rubrobrunneus. A. Rods from tentacles; B. Rosettes from tentacles; C. Rods from tube feet; D. Rosettes from introvert; E. Rods from introvert; F. Round knobbed button from introvert; G. Part of calcareous ring. A, D, E & F = scale a." httpUri="https://zenodo.org/record/165717/files/figure.png" pageId="5" pageNumber="161">Figure 6</figureCitation>
|
||
C). Rosettes simple (
|
||
<figureCitation id="D6EDC9B6FFA42F35FF02F02EFEE41C55" box="[158,259,513,538]" captionStart="FIGURE 3" captionStartId="5.[151,250,1702,1724]" captionTargetBox="[340,1246,701,1675]" captionTargetId="figure@5.[340,1247,701,1678]" captionTargetPageId="5" captionText="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161">Figure 3</figureCitation>
|
||
E), or developing into complex, opened ones (
|
||
<figureCitation id="D6EDC9B6FFA42F35FCB7F02EFC771C55" box="[811,912,513,538]" captionStart="FIGURE 3" captionStartId="5.[151,250,1702,1724]" captionTargetBox="[340,1246,701,1675]" captionTargetId="figure@5.[340,1247,701,1678]" captionTargetPageId="5" captionText="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161">Figure 3</figureCitation>
|
||
F), and eventually into closed knobbed ones (
|
||
<figureCitation id="D6EDC9B6FFA42F35FF02F00AFF191C72" box="[158,254,549,574]" captionStart="FIGURE 3" captionStartId="5.[151,250,1702,1724]" captionTargetBox="[340,1246,701,1675]" captionTargetId="figure@5.[340,1247,701,1678]" captionTargetPageId="5" captionText="FIGURE 3. Hemiocnus rubrobrunneus n. sp. ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a)." httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161">Figure 3</figureCitation>
|
||
G). Baskets of tube feet complete and incomplete, similar to those of body wall. End-plates present, better developed ventrally. Tentacle and introvert ossicles include rosettes in addition to rods (
|
||
<figureCitation id="D6EDC9B6FFA42F35FBF2F066FB3D1C2D" box="[1134,1242,585,610]" captionStart="FIGURE 4" captionStartId="6.[151,250,1591,1613]" captionTargetBox="[340,1246,553,1569]" captionTargetId="figure@6.[340,1247,553,1569]" captionTargetPageId="6" captionText="FIGURE 4. Hemiocnus rubrobrunneus n. sp. A. Rosettes from tentacles; B. Small rods from tentacles; C. Large rods from tentacles. D. Paratype (entire)." httpUri="https://zenodo.org/record/256162/files/figure.png" pageId="5" pageNumber="161">Figures 4</figureCitation>
|
||
A, B&C; 6A,B,D & E); some simple knobbed rounded plates (
|
||
<figureCitation id="D6EDC9B6FFA42F35FD0EF042FD151CCA" box="[658,754,621,646]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[369,1224,1175,1843]" captionTargetId="figure@7.[340,1248,1161,1851]" captionTargetPageId="7" captionText="FIGURE 6. Hemiocnus rubrobrunneus. A. Rods from tentacles; B. Rosettes from tentacles; C. Rods from tube feet; D. Rosettes from introvert; E. Rods from introvert; F. Round knobbed button from introvert; G. Part of calcareous ring. A, D, E & F = scale a." httpUri="https://zenodo.org/record/165717/files/figure.png" pageId="5" pageNumber="161">Figure 6</figureCitation>
|
||
F) also observed in introvert.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1AA985BBFFA42F35FF0BF489FAA918B5" httpUri="https://zenodo.org/record/165715/files/figure.png" pageId="5" pageNumber="161" targetBox="[340,1246,701,1675]" targetPageId="5">
|
||
<paragraph id="4E69D533FFA42F35FF0BF489FAA918B5" blockId="5.[151,1436,1701,1785]" pageId="5" pageNumber="161">
|
||
<emphasis id="7CA20921FFA42F35FF0BF489FD6418F7" bold="true" box="[151,643,1701,1724]" pageId="5" pageNumber="161">
|
||
FIGURE 3.
|
||
<taxonomicName id="89D6AEB0FFA42F35FE80F489FDDD18F0" box="[284,570,1701,1724]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="161" phylum="Echinodermata" rank="species" species="rubrobrunneus" status="sp. nov.">
|
||
<emphasis id="7CA20921FFA42F35FE80F489FDDD18F0" bold="true" box="[284,570,1701,1724]" italics="true" pageId="5" pageNumber="161">Hemiocnus rubrobrunneus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6791B45AFFA42F35FDD5F489FD6418F7" box="[585,643,1702,1723]" pageId="5" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
ossicles. A. Buttons from body wall; B. Incomplete baskets from body wall and tube feet; C. Basket on knobbed button; D. Complete baskets from tube feet; E. Simple open rosettes from body wall; F. Complex open rosettes from body wall; G Closed rosette from body wall. (B, D, E, F & G = scale b; A & C = scale a).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="06CC86B8FFA42F35FF5BF50AFE40192A" pageId="5" pageNumber="161" type="distribution">
|
||
<paragraph id="4E69D533FFA42F35FF5BF50AFCE11972" blockId="5.[151,1436,1829,2038]" box="[199,774,1829,1854]" pageId="5" pageNumber="161">
|
||
<emphasis id="7CA20921FFA42F35FF5BF50AFE871972" bold="true" box="[199,352,1829,1854]" pageId="5" pageNumber="161">Distribution.</emphasis>
|
||
At present known only from
|
||
<collectingCountry id="36C195A3FFA42F35FD37F50AFCE51972" box="[683,770,1829,1854]" name="Tunisia" pageId="5" pageNumber="161">Tunisia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4E69D533FFA42F35FF5BF562FE40192A" blockId="5.[151,1436,1829,2038]" box="[199,423,1869,1894]" pageId="5" pageNumber="161">
|
||
<emphasis id="7CA20921FFA42F35FF5BF562FECD192A" bold="true" box="[199,298,1869,1894]" pageId="5" pageNumber="161">Habitat.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06CC86B8FFA42F37FF5BF55EFF0C1F51" lastPageId="7" lastPageNumber="163" pageId="5" pageNumber="161" type="discussion">
|
||
<paragraph id="4E69D533FFA42F36FF5BF55EFCEB1FB9" blockId="5.[151,1436,1829,2038]" lastBlockId="6.[151,1436,151,501]" lastPageId="6" lastPageNumber="162" pageId="5" pageNumber="161">
|
||
<materialsCitation id="FEBEDF6EFFA42F36FF5BF55EFBB51EE3" ID-GBIF-Occurrence="1324949264" collectionCode="USNM, NHMUK, AST" lastPageId="6" lastPageNumber="162" location="The" pageId="5" pageNumber="161" specimenCount="3" stateProvince="Hamburg">
|
||
<emphasis id="7CA20921FFA42F35FF5BF55EFEDC19C6" bold="true" box="[199,315,1905,1930]" pageId="5" pageNumber="161">Remarks.</emphasis>
|
||
The
|
||
<specimenCount id="58D01EBAFFA42F35FEEAF55DFDE719C6" box="[374,512,1905,1930]" pageId="5" pageNumber="161" type="generic">3 specimens</specimenCount>
|
||
from the
|
||
<collectionCode id="28C74DF6FFA42F35FDECF55DFD2319C6" box="[624,708,1906,1930]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="161">USNM</collectionCode>
|
||
, labelled as
|
||
<taxonomicName id="89D6AEB0FFA42F35FCD3F55DFB8B19C5" box="[847,1132,1906,1930]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnella" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="161" phylum="Echinodermata" rank="species" species="syracusana">
|
||
<emphasis id="7CA20921FFA42F35FCD3F55DFB8B19C5" box="[847,1132,1906,1930]" italics="true" pageId="5" pageNumber="161">Pseudocnella syracusana</emphasis>
|
||
</taxonomicName>
|
||
, are small (max. length
|
||
<quantity id="892E78D6FFA42F35FA1CF55DFF2319E2" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="161" unit="mm" value="32.0">32 mm</quantity>
|
||
) and very distinct from the single specimen from the
|
||
<collectingRegion id="8C121BD1FFA42F35FCB3F5BAFC7F19E2" box="[815,920,1941,1966]" country="Germany" name="Hamburg" pageId="5" pageNumber="161">Hamburg</collectingRegion>
|
||
Museum and those examined at
|
||
<collectionCode id="28C74DF6FFA42F35FA92F5B9FA9F19E1" box="[1294,1400,1942,1965]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="161">NHMUK</collectionCode>
|
||
by
|
||
<collectionCode id="28C74DF6FFA42F35FF0BF595FF2A199E" box="[151,205,1978,2002]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15510" name="University of Aston" pageId="5" pageNumber="161">AST</collectionCode>
|
||
, which are much larger (maximum length
|
||
<quantity id="892E78D6FFA42F35FD2EF596FCE3199E" box="[690,772,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="161" unit="mm" value="56.0">56 mm</quantity>
|
||
) and identified as
|
||
<taxonomicName id="89D6AEB0FFA42F35FC44F595FB17199D" box="[984,1264,1978,2002]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnus" kingdom="Animalia" order="Dendrochirotida" pageId="5" pageNumber="161" phylum="Echinodermata" rank="species" species="syracusanus">
|
||
<emphasis id="7CA20921FFA42F35FC44F595FB17199D" box="[984,1264,1978,2002]" italics="true" pageId="5" pageNumber="161">Pseudocnus syracusanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<location id="4B0983E8FFA42F35FA9DF596FACA199E" LSID="urn:lsid:plazi:treatment:C67F6425FFA52F37FF0BF710FF0C1F51:4B0983E8FFA42F35FA9DF596FACA199E" box="[1281,1325,1977,2002]" name="The" pageId="5" pageNumber="161" stateProvince="Hamburg">The</location>
|
||
colour of the preserved
|
||
<collectionCode id="28C74DF6FFA42F35FEA9F5F1FE6119BA" box="[309,390,2014,2038]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="161">USNM</collectionCode>
|
||
material, collected in 1973, is a dark reddish brown (
|
||
<figureCitation id="D6EDC9B6FFA42F35FC78F5F2FBA119BA" box="[996,1094,2013,2038]" captionStart="FIGURE 2" captionStartId="4.[151,250,1210,1232]" captionTargetBox="[340,1246,193,1188]" captionTargetId="figure@4.[340,1247,193,1189]" captionTargetPageId="4" captionText="FIGURE 2. Hemiocnus syracusanus. A. Fir-cone-shaped plates from body wall; B. Large round plates from body wall; C. Buttons from body wall; D. Small rods from tentacles; E. Large rods from tentacles; F. Specimen entire." httpUri="https://zenodo.org/record/256161/files/figure.png" pageId="5" pageNumber="161">Figure 2</figureCitation>
|
||
D), while that of the
|
||
<collectingRegion id="8C121BD1FFA42F35FAAFF5F2FA7B19BA" box="[1331,1436,2013,2038]" country="Germany" name="Hamburg" pageId="5" pageNumber="161">Hamburg</collectingRegion>
|
||
specimen, collected in 1982, is light brown (
|
||
<figureCitation id="D6EDC9B6FFA72F36FD0CF2B8FD161EFC" box="[656,753,151,176]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[369,1224,1175,1843]" captionTargetId="figure@7.[340,1248,1161,1851]" captionTargetPageId="7" captionText="FIGURE 6. Hemiocnus rubrobrunneus. A. Rods from tentacles; B. Rosettes from tentacles; C. Rods from tube feet; D. Rosettes from introvert; E. Rods from introvert; F. Round knobbed button from introvert; G. Part of calcareous ring. A, D, E & F = scale a." httpUri="https://zenodo.org/record/165717/files/figure.png" pageId="6" pageNumber="162">Figure 6</figureCitation>
|
||
F) similar to those at
|
||
<collectionCode id="28C74DF6FFA72F36FC79F2B7FBB51EE3" box="[997,1106,152,175]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="162">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="FEBEDF6EFFA72F36FBC2F2B8FCEB1FB9" ID-GBIF-Occurrence="1324949262" collectionCode="USNM, NHMUK" county="Both" location="While" municipality="The" pageId="6" pageNumber="162" specimenCount="1" stateProvince="Hamburg">
|
||
<collectingCounty id="A708ADBFFFA72F36FBC2F2B8FB731EFC" box="[1118,1172,151,176]" pageId="6" pageNumber="162">Both</collectingCounty>
|
||
materials also differ in their size and tube feet distribution.
|
||
<collectingMunicipality id="AE0D4F49FFA72F36FDB2F293FDBD1E99" box="[558,602,188,213]" pageId="6" pageNumber="162">The</collectingMunicipality>
|
||
maximum length recorded for a true
|
||
<taxonomicName id="89D6AEB0FFA72F36FC63F292FB7C1E98" box="[1023,1179,189,212]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnella" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="species" species="syracusana">
|
||
<emphasis id="7CA20921FFA72F36FC63F292FB7C1E98" box="[1023,1179,189,212]" italics="true" pageId="6" pageNumber="162">P. syracusana</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<quantity id="892E78D6FFA72F36FB21F293FAEB1E98" box="[1213,1292,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="162" unit="mm" value="58.0">58 mm</quantity>
|
||
(by
|
||
<bibRefCitation id="2A47A8C2FFA72F36FADCF293FF341EB4" author="Panning" pageId="6" pageNumber="162" refString="Panning, A. (1962) Bermerkungen uber die Holothurien - Familie Cucumariidae (Ordnung Dendrochirota) 3 Teil. Die Gattung Pseudocnus (Panning 1949). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 60,57 - 80." type="journal article" year="1962">Panning 1962</bibRefCitation>
|
||
).
|
||
<location id="4B0983E8FFA72F36FF7AF2F0FECD1EB4" LSID="urn:lsid:plazi:treatment:C67F6425FFA52F37FF0BF710FF0C1F51:4B0983E8FFA72F36FF7AF2F0FECD1EB4" box="[230,298,223,248]" county="Both" municipality="The" name="While" pageId="6" pageNumber="162" stateProvince="Hamburg">While</location>
|
||
the
|
||
<collectingRegion id="8C121BD1FFA72F36FEC5F2F0FE251EB4" box="[345,450,223,248]" country="Germany" name="Hamburg" pageId="6" pageNumber="162">Hamburg</collectingRegion>
|
||
specimen is hard and has, in addition to scattered tube feet, distinct interradial ‘papulaelike’ extensions, reminiscent of those of
|
||
<taxonomicName id="89D6AEB0FFA72F36FDC1F32BFD131F50" box="[605,756,260,284]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnella" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7CA20921FFA72F36FDC1F32BFD131F50" box="[605,756,260,284]" italics="true" pageId="6" pageNumber="162">Pseudocnella</emphasis>
|
||
</taxonomicName>
|
||
spp., the
|
||
<collectionCode id="28C74DF6FFA72F36FCC3F32BFC571F50" box="[863,944,260,284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="162">USNM</collectionCode>
|
||
specimens are soft, with tube feet restricted in two distinct rows per ambulacrum, especially ventrally, and lack ‘papulae-like’ structures of any sort. As far as ossicles are concerned the new species has in its body wall knobbed rounded plates and both complete and incomplete baskets whereas the
|
||
<collectingRegion id="8C121BD1FFA72F36FD98F340FD8A1FC4" box="[516,621,367,392]" country="Germany" name="Hamburg" pageId="6" pageNumber="162">Hamburg</collectingRegion>
|
||
and
|
||
<collectionCode id="28C74DF6FFA72F36FD3AF35FFCF71FCB" box="[678,784,368,391]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="162">NHMUK</collectionCode>
|
||
specimen have large fir-cone shaped plates in addition to round plates and a few cross-shaped incomplete baskets but the latter were absent in most specimens from
|
||
<collectionCode id="28C74DF6FFA72F36FF0BF397FEE21F83" box="[151,261,440,463]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="162">NHMUK</collectionCode>
|
||
. The new species has also rosettes in the body wall and rods, rosettes and baskets in the tube feet. Hence, there is no doubt that both materials are not conspecific.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="1AA985BBFFA72F36FF0BF418FE2C1827" httpUri="https://zenodo.org/record/256162/files/figure.png" pageId="6" pageNumber="162" targetBox="[340,1246,553,1569]" targetPageId="6">
|
||
<paragraph id="4E69D533FFA72F36FF0BF418FE2C1827" blockId="6.[151,1436,1590,1643]" pageId="6" pageNumber="162">
|
||
<emphasis id="7CA20921FFA72F36FF0BF418FD9D1800" bold="true" box="[151,634,1590,1613]" pageId="6" pageNumber="162">
|
||
FIGURE 4.
|
||
<taxonomicName id="89D6AEB0FFA72F36FE87F418FDDE1801" box="[283,569,1590,1613]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="species" species="rubrobrunneus" status="sp. nov.">
|
||
<emphasis id="7CA20921FFA72F36FE87F418FDDE1801" bold="true" box="[283,569,1590,1613]" italics="true" pageId="6" pageNumber="162">Hemiocnus rubrobrunneus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6791B45AFFA72F36FDDBF418FD9D1800" box="[583,634,1591,1612]" pageId="6" pageNumber="162" rank="species">n. sp</taxonomicNameLabel>
|
||
</emphasis>
|
||
. A. Rosettes from tentacles; B. Small rods from tentacles; C. Large rods from tentacles. D. Paratype (entire).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4E69D533FFA72F37FF5BF4BCFF0C1F51" blockId="6.[151,1436,1683,2032]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="163" pageId="6" pageNumber="162">
|
||
At first we were of the opinion that the locality of the species and form of the ossicles, except for the presence of quadrilocular baskets, clearly places the new species in the genus
|
||
<taxonomicName id="89D6AEB0FFA72F36FC27F498FBE41882" box="[955,1027,1719,1742]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7CA20921FFA72F36FC27F498FBE41882" box="[955,1027,1719,1742]" italics="true" pageId="6" pageNumber="162">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
and we described it as such in the manuscript. However, Dr Rowe who is most familiar with the Mediterranean holothuroids and especially the genus
|
||
<taxonomicName id="89D6AEB0FFA72F36FF0BF4D0FF38195A" box="[151,223,1791,1814]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7CA20921FFA72F36FF0BF4D0FF38195A" box="[151,223,1791,1814]" italics="true" pageId="6" pageNumber="162">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
, first reviewed the manuscript and is of the opinion that the baskets are not reminiscent of species of
|
||
<taxonomicName id="89D6AEB0FFA72F36FAC8F4D0FA7C195A" box="[1364,1435,1791,1814]" class="Holothuroidea" family="Cucumariidae" genus="Ocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7CA20921FFA72F36FAC8F4D0FA7C195A" box="[1364,1435,1791,1814]" italics="true" pageId="6" pageNumber="162">Ocnus</emphasis>
|
||
</taxonomicName>
|
||
and the incomplete ones suggest that the specimens perhaps represent juvenile of the preceding species or at best a new species within
|
||
<taxonomicName id="89D6AEB0FFA72F36FEEFF567FE161913" box="[371,497,1864,1887]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="157" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="7CA20921FFA72F36FEEFF567FE161913" box="[371,497,1864,1887]" italics="true" pageId="6" pageNumber="162">Hemiocnus</emphasis>
|
||
</taxonomicName>
|
||
. We contest the former viewpoint as the gonadal tubules of at least one specimen is in the process of maturity, the specimens lack the complex plates of
|
||
<emphasis id="7CA20921FFA72F36FC09F543FC5619CF" box="[917,945,1900,1923]" italics="true" pageId="6" pageNumber="162">H.</emphasis>
|
||
<taxonomicName id="89D6AEB0FFA72F36FC24F543FBD819C8" box="[952,1087,1900,1924]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="162" phylum="Echinodermata" rank="species" species="syracusanus">syracusanus</taxonomicName>
|
||
and those of the anal region of at least one specimen are rather simple, there are rosettes present together with complete baskets, and the colour and texture of the body wall are quite different. We therefore agree with Dr Rowe’s second point and here describe the new species in
|
||
<taxonomicName id="89D6AEB0FFA72F36FEF1F5F7FE0C19A3" box="[365,491,2008,2031]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" kingdom="Animalia" order="Dendrochirotida" pageId="6" pageNumber="157" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="7CA20921FFA72F36FEF1F5F7FE0C19A3" box="[365,491,2008,2031]" italics="true" pageId="6" pageNumber="162">Hemiocnus</emphasis>
|
||
</taxonomicName>
|
||
as no other genus exists to accommodate it. It is possible that the Tunisian fauna is quite different from that of the western Mediterranean as quite recently, Stohr (2015) (in
|
||
<emphasis id="7CA20921FFA62F37FB65F2B7FA741EFC" box="[1273,1427,152,176]" italics="true" pageId="7" pageNumber="163">Echinoderm-l</emphasis>
|
||
) commented, with reference to the ophiuroids, that
|
||
<collectingCountry id="36C195A3FFA62F37FD49F293FCCF1E99" box="[725,808,188,213]" name="Tunisia" pageId="7" pageNumber="163">Tunisia</collectingCountry>
|
||
is at the western border of the Levant and thus it has a somewhat different fauna from the northern and western Med. Hence this may also apply to other echinoderm groups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |