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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.179.21253" ID-GBIF-Dataset="144aa3b4-ee51-4c26-8436-aaa65ae3b22b" ID-PMC="PMC5740450" ID-Pensoft-Pub="1313-2970-179-75" ID-PubMed="29290697" ID-ZBK="36C4E2C8E07D4CC9A1D696B0FCE92CCF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-179-75" ModsDocOrigin="ZooKeys 179" ModsDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" checkinTime="1512688543562" checkinUser="pensoft" docAuthor="Burke, Alan &amp; Zolnerowich, Gregory" docDate="2017" docId="6908CA4FEB56AF339497A20DF6B761A8" docLanguage="en" docName="ZooKeys 719: 75-157" docOrigin="ZooKeys 179" docSource="http://dx.doi.org/10.3897/zookeys.179.21253" docTitle="Onychotillus Chapin 1945" docType="treatment" docVersion="5" lastPageNumber="137" masterDocId="8A60FFA894215C48ED67FF86FF8F3B5B" masterDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" masterLastPageNumber="157" masterPageNumber="75" pageNumber="135" updateTime="1668165163296" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Burke, Alan</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zolnerowich, Gregory</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>179</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>157</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.179.21253</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.179.21253</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-179-75</mods:identifier>
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<treatment ID-GBIF-Taxon="137663077" LSID="urn:lsid:plazi:treatment:6908CA4FEB56AF339497A20DF6B761A8" httpUri="http://treatment.plazi.org/id/6908CA4FEB56AF339497A20DF6B761A8" lastPageId="62" lastPageNumber="137" pageId="60" pageNumber="135">
<subSubSection pageId="60" pageNumber="135" type="nomenclature">
<paragraph pageId="60" pageNumber="135">
<taxonomicName authority="Chapin, 1945" authorityName="Chapin" authorityYear="1945" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="60" pageNumber="135" phylum="Arthropoda" rank="genus">Onychotillus Chapin, 1945</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="135" type="type species">
<paragraph pageId="60" pageNumber="135">Type species.</paragraph>
<paragraph pageId="60" pageNumber="135">
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="60" pageNumber="135" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
Chapin, original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="135" type="distribution">
<paragraph pageId="60" pageNumber="135">Distribution.</paragraph>
<paragraph pageId="60" pageNumber="135">Shown in Fig. 21K.</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="136" type="differential diagnosis">
<paragraph pageId="61" pageNumber="136">
<pageBreakToken pageId="61" pageNumber="136" start="start">Differential</pageBreakToken>
diagnosis.
</paragraph>
<paragraph pageId="61" pageNumber="136">
Species of
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
somewhat resemble small-sized members of
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
and species of
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
. These genera can be differentiated without difficulty based on the number of tarsal claws.
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
species have a single tarsal claw, while members of
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
and
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
have two tarsal claws (Fig. 7B). Additionally, the distribution of
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
does not overlap with that of
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
and
<taxonomicName class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
, with the former restricted to the West Indies, while the latter two have a continental distribution.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="136" type="redescription">
<paragraph pageId="61" pageNumber="136">Redescription.</paragraph>
<paragraph pageId="61" pageNumber="136">
Size: 3-12 mm. Color: Body ranging from light fuscous to ferrugineous; some individuals may possess one pale, median fascia on each elytron. Body: Winged species; short to moderately long, slightly robust, somewhat subparallel specimens (Fig. 5
<normalizedToken originalValue="AB">A-B</normalizedToken>
).
</paragraph>
<paragraph pageId="61" pageNumber="136">
Head: Including eye slightly narrower than pronotum; integument coarsely punctate; punctations vary from narrow to wide; eyes large, finely faceted, feebly emarginate posteriorly, bulging laterally; antennae with 11 antennomeres, extending beyond posterior margin of pronotum; last antennomere as long as or conspicuously longer than ninth antennomere (Fig. 10
<normalizedToken originalValue="EF">E-F</normalizedToken>
); frons feebly bi-impressed; terminal labial palpi securiform; terminal maxillary palpi cylindrical, distally compressed.
</paragraph>
<paragraph pageId="61" pageNumber="136">Thorax: Pronotum shiny, smooth to feebly punctate; punctation may range from narrow to slightly wide; lateral margins subparallel, slightly constricted anteriorly, somewhat wider posteriorly. Prosternum: Much wider than long, smooth to rugose, variously punctate. Mesoventrite: Wider than long, smooth, feebly to moderately punctate. Metaventrite: Convex, wider than long, surface conspicuously punctate to almost smooth. Metaventral process compressed anteriorly. Metepisterna largely exposed, the elytra do not coverer these plates in lateral view.</paragraph>
<paragraph pageId="61" pageNumber="136">Elytra: Short to elongate, subparallel, widest on middle half, surface moderately to coarsely punctate; punctations arranged in regular striae; punctations do not extend to apex; epipleural fold complete, narrowing toward apex.</paragraph>
<paragraph pageId="61" pageNumber="136">Legs: Femora smooth to rugose; rather swollen distally; tibiae rugose to rugulose; pulvillar formula 4-4-4; one tarsal denticle, tarsal denticle subtriangular in shape.</paragraph>
<paragraph pageId="61" pageNumber="136">Abdomen: Six visible ventrites. Ventrites 1-5 not impressed laterally. Sixth ventrite of males feebly differentiated from that of females; females with sixth ventrite broadly rounded, pygidium simple.</paragraph>
</subSubSection>
<subSubSection lastPageId="62" lastPageNumber="137" pageId="61" pageNumber="136" type="remarks">
<paragraph pageId="61" pageNumber="136">Remarks.</paragraph>
<paragraph lastPageId="62" lastPageNumber="137" pageId="61" pageNumber="136">
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
is currently composed of five species, all inhabiting the West Indies. Of the five valid species, three are thought to be restricted to Cuba (
<bibRefCitation pageId="61" pageNumber="136">De Zayas 1988</bibRefCitation>
). Based on the limited descriptive work given by
<bibRefCitation pageId="61" pageNumber="136">De Zayas (1988)</bibRefCitation>
, where no key for identification was provided, specimens of what appears to be
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus cubana" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="species" species="cubana">Onychotillus cubana</taxonomicName>
were examined (Fig. 5A); however, none of these individuals were collected in Cuba, so these specimens are tentatively assigned to
<taxonomicName lsidName="O. cubana" pageId="61" pageNumber="136" rank="species" species="cubana">O. cubana</taxonomicName>
, pending further material to be examined. It is important to note that De
<normalizedToken originalValue="Zayas">Zayas'</normalizedToken>
descriptions were based, in most cases, on a single specimen, and access to that material has been particularly difficult. Therefore, due to the lack of availability of material and the poorly detailed descriptions given by De Zayas, three
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
species are excluded from this revision,
<taxonomicName lsidName="O. trinitatis" pageId="61" pageNumber="136" rank="species" species="trinitatis">O. trinitatis</taxonomicName>
,
<taxonomicName lsidName="O. minuta" pageId="61" pageNumber="136" rank="species" species="minuta">O. minuta</taxonomicName>
and
<taxonomicName lsidName="O. dimidiata" pageId="61" pageNumber="136" rank="species" species="dimidiata">O. dimidiata</taxonomicName>
, all endemic to Cuba. On the other hand, we examined
<pageBreakToken pageId="62" pageNumber="137" start="start">material</pageBreakToken>
of
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
collected in the Dominican Republic and the Cayman Islands. These collecting localities represent new distribution records for
<taxonomicName lsidName="O. vittatus" pageId="62" pageNumber="137" rank="species" species="vittatus">O. vittatus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="137" type="key">
<paragraph pageId="62" pageNumber="137">
Key to species of
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
</paragraph>
<paragraph pageId="62" pageNumber="137">
<table pageId="62" pageNumber="137">
<tr pageId="62" pageNumber="137">
<td colspan="1" pageId="62" pageNumber="137" rowspan="1">5B10F</td>
<td colspan="1" pageId="62" pageNumber="137" rowspan="1">
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
</td>
</tr>
<tr pageId="62" pageNumber="137">
<td colspan="1" pageId="62" pageNumber="137" rowspan="1">5A10E</td>
<td colspan="1" pageId="62" pageNumber="137" rowspan="1">
<taxonomicName class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus cubana" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="cubana">Onychotillus cubana</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>