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<document id="2A512534B9B6A6476A0B3140BD49DA82" ID-DOI="10.5281/zenodo.212710" ID-GBIF-Dataset="cc4e0f3b-19df-4bca-b666-e737d285210a" ID-ISSN="1175-5326" ID-Zenodo-Dep="212710" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460477492253" checkinUser="plazi" docAuthor="Ament, Danilo César" docDate="2012" docId="306CBA7B3F34FFBA2DA2F9C2BD93F9F5" docLanguage="en" docName="zt03298p061.pdf" docOrigin="Zootaxa 3298" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mannheimsia conica Ament, 2012, sp. nov." docType="treatment" docVersion="11" lastPageNumber="57" masterDocId="CC55C2033F37FFBE2D35FF8FBC6BFF80" masterDocTitle="Mannheimsia Beyer (Diptera: Phoridae): description of a new species, redescription of Mannheimsia stricta and studies on hypopygial morphology" masterLastPageNumber="61" masterPageNumber="53" pageNumber="56" updateTime="1698307012574" updateUser="plazi">
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<mods:title id="B7CFC7F35B96C518D659D00026DF5700">Mannheimsia Beyer (Diptera: Phoridae): description of a new species, redescription of Mannheimsia stricta and studies on hypopygial morphology</mods:title>
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<mods:namePart id="033707D53592D7460E14AF1C5D2E865F">Ament, Danilo César</mods:namePart>
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<mods:date id="ED80EFC1DB8D7CE866B8EBD3F6C84146">2012</mods:date>
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<treatment id="306CBA7B3F34FFBA2DA2F9C2BD93F9F5" ID-DOI="http://doi.org/10.5281/zenodo.5672333" ID-GBIF-Taxon="119516621" ID-Zenodo-Dep="5672333" LSID="urn:lsid:plazi:treatment:306CBA7B3F34FFBA2DA2F9C2BD93F9F5" httpUri="http://treatment.plazi.org/id/306CBA7B3F34FFBA2DA2F9C2BD93F9F5" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<subSubSection id="F0DF58E63F34FFBD2DA2F9C2BDACF908" pageId="3" pageNumber="56" type="nomenclature">
<paragraph id="B87A0B6D3F34FFBD2DA2F9C2BE69F9E7" blockId="3.[151,514,1613,1672]" box="[151,514,1613,1639]" pageId="3" pageNumber="56">
<heading id="E332BC013F34FFBD2DA2F9C2BE69F9E7" bold="true" box="[151,514,1613,1639]" fontSize="11" level="1" pageId="3" pageNumber="56" reason="1">
<emphasis id="8AB1D77F3F34FFBD2DA2F9C2BE69F9E7" bold="true" box="[151,514,1613,1639]" pageId="3" pageNumber="56">
<emphasis id="8AB1D77F3F34FFBD2DA2F9C2BDF6F9E7" bold="true" box="[151,413,1613,1639]" italics="true" pageId="3" pageNumber="56">
<taxonomicName id="7FC570EE3F34FFBD2DA2F9C2BDFCF9E7" ID-CoL="9J2LG" box="[151,407,1613,1639]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">Mannheimsia conica</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="91826A043F34FFBD2C91F9C2BE69F9E7" box="[420,514,1613,1639]" pageId="3" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B87A0B6D3F34FFBD2DA2F9FFBDACF908" blockId="3.[151,514,1613,1672]" box="[151,455,1648,1672]" pageId="3" pageNumber="56">
(
<figureCitation id="20FE17E83F34FFBD2DAAF9FFBD60F908" box="[159,267,1648,1672]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[159,1413,196,1831]" captionTargetId="figure@2.[151,1436,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 3. Mannheimsia conica, sp. nov. and Mannheimsia stricta. 1. M. conica, sp. nov. habitus; 2. M. conica, sp. nov. head in lateral view; 3. M. stricta abdomen in terminal-ventral view." httpUri="https://zenodo.org/record/212711/files/figure.png" pageId="3" pageNumber="56">Figs. 12</figureCitation>
,
<figureCitation id="20FE17E83F34FFBD2C22F9FFBD4CF908" box="[279,295,1648,1672]" captionStart="FIGURES 4 7" captionStartId="3.[151,264,1511,1534]" captionTargetBox="[164,1436,844,1474]" captionTargetId="figure@3.[151,1436,831,1489]" captionTargetPageId="3" captionText="FIGURES 4 7. Mannheimsia wings. 4. M. conica, sp. nov.; 5. M. stricta; 6. M. stylodactyla; 7. M. tianzena." httpUri="https://zenodo.org/record/212712/files/figure.png" pageId="3" pageNumber="56">4</figureCitation>
,
<figureCitation id="20FE17E83F34FFBD2C07F9FFBD07F908" box="[306,364,1648,1672]" captionStart="FIGURES 8 23" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[178,1404,339,1817]" captionTargetId="figure@5.[151,1436,313,1841]" captionTargetPageId="5" captionText="FIGURES 8 23. Mannheimsia hypopygium. 8 11. Mannheimsia conica, sp. nov., 8. Dorsal, 9. Ventral, 10. Left lateral, 11. Right lateral; 12 15. Mannheimsia stricta, 12. Dorsal, 13. Ventral, 14. Left lateral, 15. Right lateral; 16 19. Mannheimsia stylodactyla, 16. Dorsal, 17. Ventral, 18. Left lateral, 19. Right lateral; 20 23. Mannheimsia tianzena, 20. Dorsal, 21. Ventral, 22. Left lateral, 23. Right lateral. Colors indicating homology: blue, hypandrium left lobe; yellow, hypandrium right lobe; red, hypandrium right lobe medial process. Abbreviations: cerc, cercus; lep, left epandrial process (fused surstyli); hyprct, hypoproct; rsp, right subepandrial process." httpUri="https://zenodo.org/record/212713/files/figure.png" pageId="3" pageNumber="56">811</figureCitation>
,
<figureCitation id="20FE17E83F34FFBD2C42F9FFBDD4F908" box="[375,447,1648,1672]" captionStart="FIGURES 24 31" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[256,1286,195,1887]" captionTargetId="figure@6.[256,1331,157,1923]" captionTargetPageId="6" captionText="FIGURES 24 31. Mannheimsia phallus. 24 25. M. conica, sp. nov., 24. Dorsal, 25. Left lateral; 26 27. M. stricta, 26. Dorsal, 27. Left lateral; 28 29. M. stylodactyla, 28. Dorsal; 29. Left lateral; 30 31. M. tianzena, 30. Dorsal, 31. Left lateral. Abbreviations: bs, basiphallus; epi, epiphallus; epl, epiphallus posterior lobe; p 1 and p 2, processes 1 and 2, respectively; ra, right arm. Core plate represented in yellow." httpUri="https://zenodo.org/record/212714/files/figure.png" pageId="3" pageNumber="56">2425</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F34FFBD2DA2F937BD7BF974" pageId="3" pageNumber="56" type="diagnosis">
<paragraph id="B87A0B6D3F34FFBD2DA2F937BD7BF974" blockId="3.[151,1436,1720,2032]" pageId="3" pageNumber="56">
<emphasis id="8AB1D77F3F34FFBD2DA2F937BD61F951" bold="true" box="[151,266,1720,1745]" pageId="3" pageNumber="56">Diagnosis</emphasis>
(male). Flagellomere 1 elongate-conical. Two setae on hindtibia. Right lobe of hypandrium clearly bifurcated.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F34FFBD2DFCF88FBD36F8BC" pageId="3" pageNumber="56" type="materials_examined">
<paragraph id="B87A0B6D3F34FFBD2DFCF88FBD36F8BC" blockId="3.[151,1436,1720,2032]" pageId="3" pageNumber="56">
<emphasis id="8AB1D77F3F34FFBD2DFCF88FBDC7F899" bold="true" box="[201,428,1792,1817]" pageId="3" pageNumber="56">Material examined</emphasis>
.
<typeStatus id="677EB5CF3F34FFBD2C8EF88FBE43F898" box="[443,552,1792,1816]" pageId="3" pageNumber="56" type="holotype">Holotype</typeStatus>
: ɗ,
<emphasis id="8AB1D77F3F34FFBD2F60F88FBE87F899" bold="true" box="[597,748,1792,1817]" pageId="3" pageNumber="56">
<collectingCountry id="C0D24BFD3F34FFBD2F60F88FBE87F899" box="[597,748,1792,1817]" name="Thailand" pageId="3" pageNumber="56">THAILAND</collectingCountry>
</emphasis>
: Chiang Mai,
<geoCoordinate id="DDF16DAA3F34FFBD2EA4F88FBF85F898" box="[913,1006,1792,1816]" direction="north" orientation="latitude" pageId="3" pageNumber="56" precision="555" value="18.7">18.70ºN</geoCoordinate>
<geoCoordinate id="DDF16DAA3F34FFBD2EC3F88FB83AF898" box="[1014,1105,1792,1816]" direction="east" orientation="longitude" pageId="3" pageNumber="56" precision="555" value="98.8">98.80ºE</geoCoordinate>
,
<date id="CC7B2DAD3F34FFBD2968F88FB897F898" box="[1117,1276,1792,1816]" pageId="3" pageNumber="56" value="1997-12-01" valueMax="1997-12-31" valueMin="1997-12-01">131.xii.1997</date>
, Malaise trap, S. Sonthichai col.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F34FFBA2DFCF8C7BFC7FE2F" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
<paragraph id="B87A0B6D3F34FFBA2DFCF8C7B8B2FE0A" blockId="3.[151,1436,1720,2032]" lastBlockId="4.[151,1437,151,576]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<emphasis id="8AB1D77F3F34FFBD2DFCF8C7BD3FF8E1" bold="true" box="[201,340,1864,1889]" pageId="3" pageNumber="56">Description</emphasis>
.
<emphasis id="8AB1D77F3F34FFBD2C54F8C6BDF6F8E1" bold="true" box="[353,413,1865,1889]" pageId="3" pageNumber="56">Male</emphasis>
. Body length,
<quantity id="7F3DA6883F34FFBD2F71F8C7BEF5F8E0" box="[580,670,1864,1888]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="56" unit="mm" value="1.9">1.9 mm</quantity>
.
<emphasis id="8AB1D77F3F34FFBD2F9CF8C6BE82F8E1" bold="true" box="[681,745,1865,1889]" pageId="3" pageNumber="56">Head</emphasis>
. Frons brown, pubescent, without median furrow. Flagellomere 1 light brown, pubescent, conical-elongated. Arista apical, pubescent. Palpus light brown; four medium size genal setae (
<figureCitation id="20FE17E83F34FFBD2C1FF81FBD18F828" box="[298,371,1936,1960]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[159,1413,196,1831]" captionTargetId="figure@2.[151,1436,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 3. Mannheimsia conica, sp. nov. and Mannheimsia stricta. 1. M. conica, sp. nov. habitus; 2. M. conica, sp. nov. head in lateral view; 3. M. stricta abdomen in terminal-ventral view." httpUri="https://zenodo.org/record/212711/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
).
<emphasis id="8AB1D77F3F34FFBD2CBEF81EBD8FF829" bold="true" box="[395,484,1937,1961]" pageId="3" pageNumber="56">Thorax</emphasis>
. Scutum, scutellum and pleural regions brown; anepisternum setulose. Legs light brown. Foretibia with one dorsal seta at basal third and anterodorsal row of strong setae. Midtibia with one anterodorsal and one posterodorsal setae.
<collectingCountry id="C0D24BFD3F34FFBD2F17F857BE31F870" box="[546,602,2008,2032]" name="India" pageId="3" pageNumber="56">Hind</collectingCountry>
femur not swollen.
<collectingCountry id="C0D24BFD3F34FFBD2E0CF857BF1AF870" box="[825,881,2008,2032]" name="India" pageId="3" pageNumber="56">Hind</collectingCountry>
tibia with one anterodorsal and one posterodorsal setae near midlength.
<emphasis id="8AB1D77F3F33FFBA2CBAFF18BDA4FF30" bold="true" box="[399,463,151,176]" pageId="4" pageNumber="57">Wing</emphasis>
. (
<figureCitation id="20FE17E83F33FFBA2CD2FF18BE40FF2F" box="[487,555,151,175]" captionStart="FIGURES 4 7" captionStartId="3.[151,264,1511,1534]" captionTargetBox="[164,1436,844,1474]" captionTargetId="figure@3.[151,1436,831,1489]" captionTargetPageId="3" captionText="FIGURES 4 7. Mannheimsia wings. 4. M. conica, sp. nov.; 5. M. stricta; 6. M. stylodactyla; 7. M. tianzena." httpUri="https://zenodo.org/record/212712/files/figure.png" pageId="4" pageNumber="57">Fig. 4</figureCitation>
) Costa 0.36 of the wing length, M2 sinuous. Halter gray with apical black circle.
<emphasis id="8AB1D77F3F33FFBA2DF0FF31BD52FF57" bold="true" box="[197,313,190,215]" pageId="4" pageNumber="57">Abdomen</emphasis>
. Tergites brown. Hypopygium yellowish-brown (
<figureCitation id="20FE17E83F33FFBA2E53FF30BFB4FF57" box="[870,991,191,215]" captionStart="FIGURES 8 23" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[178,1404,339,1817]" captionTargetId="figure@5.[151,1436,313,1841]" captionTargetPageId="5" captionText="FIGURES 8 23. Mannheimsia hypopygium. 8 11. Mannheimsia conica, sp. nov., 8. Dorsal, 9. Ventral, 10. Left lateral, 11. Right lateral; 12 15. Mannheimsia stricta, 12. Dorsal, 13. Ventral, 14. Left lateral, 15. Right lateral; 16 19. Mannheimsia stylodactyla, 16. Dorsal, 17. Ventral, 18. Left lateral, 19. Right lateral; 20 23. Mannheimsia tianzena, 20. Dorsal, 21. Ventral, 22. Left lateral, 23. Right lateral. Colors indicating homology: blue, hypandrium left lobe; yellow, hypandrium right lobe; red, hypandrium right lobe medial process. Abbreviations: cerc, cercus; lep, left epandrial process (fused surstyli); hyprct, hypoproct; rsp, right subepandrial process." httpUri="https://zenodo.org/record/212713/files/figure.png" pageId="4" pageNumber="57">Figs. 811</figureCitation>
). Left epandrial process (fused surstyli) with a medially-directed tooth-like projection near apex. Right epandrial margin narrowing ventrally, with subepandrial right process bearing setae. Hypandrium left lobe bifurcated in ventral bare extension and dorsal microtrichose extension; right lobe apically microtrichose, with medially-directed elongated process. Hypoproct with two setae. Cerci large.
<emphasis id="8AB1D77F3F33FFBA2CFBFEC0BE4CFEE7" bold="true" box="[462,551,335,359]" pageId="4" pageNumber="57">Phallus</emphasis>
(
<figureCitation id="20FE17E83F33FFBA2F03FEC0BEABFEE7" box="[566,704,335,359]" captionStart="FIGURES 24 31" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[256,1286,195,1887]" captionTargetId="figure@6.[256,1331,157,1923]" captionTargetPageId="6" captionText="FIGURES 24 31. Mannheimsia phallus. 24 25. M. conica, sp. nov., 24. Dorsal, 25. Left lateral; 26 27. M. stricta, 26. Dorsal, 27. Left lateral; 28 29. M. stylodactyla, 28. Dorsal; 29. Left lateral; 30 31. M. tianzena, 30. Dorsal, 31. Left lateral. Abbreviations: bs, basiphallus; epi, epiphallus; epl, epiphallus posterior lobe; p 1 and p 2, processes 1 and 2, respectively; ra, right arm. Core plate represented in yellow." httpUri="https://zenodo.org/record/212714/files/figure.png" pageId="4" pageNumber="57">Figs. 2425</figureCitation>
). Right arm with dorsal projection. Epiphallus mostly absent, just posterior lobe present. Core plate hairy, with three dorsally curved processes.
<emphasis id="8AB1D77F3F33FFBA2EC8FEFCB83FFE0B" bold="true" box="[1021,1108,371,395]" pageId="4" pageNumber="57">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="B87A0B6D3F33FFBA2DFCFE19BFC7FE2F" blockId="4.[151,1437,151,576]" box="[201,940,406,431]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFE19BD82FE2F" bold="true" box="[201,489,406,431]" pageId="4" pageNumber="57">Geographic distribution</emphasis>
. Known from only one site in
<collectingCountry id="C0D24BFD3F33FFBA2E74FE18BFC3FE2F" box="[833,936,407,431]" name="Thailand" pageId="4" pageNumber="57">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DFCFE34B88FFE52" box="[201,1252,442,467]" pageId="4" pageNumber="57" type="etymology">
<paragraph id="B87A0B6D3F33FFBA2DFCFE34B88FFE52" blockId="4.[151,1437,151,576]" box="[201,1252,442,467]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFE34BD23FE53" bold="true" box="[201,328,443,467]" pageId="4" pageNumber="57">Etymology</emphasis>
.
<taxonomicName id="7FC570EE3F33FFBA2C63FE34BE2BFE53" box="[342,576,443,467]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="conica">
<emphasis id="8AB1D77F3F33FFBA2C63FE34BE2BFE53" box="[342,576,443,467]" italics="true" pageId="4" pageNumber="57">Mannheimsia conica</emphasis>
</taxonomicName>
was named after its characteristic conical first flagellomere.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DFCFE51B89CFDBF" pageId="4" pageNumber="57" type="discussion">
<paragraph id="B87A0B6D3F33FFBA2DFCFE51B89CFDBF" blockId="4.[151,1437,151,576]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFE51BD21FE77" bold="true" box="[201,330,478,503]" pageId="4" pageNumber="57">Comments</emphasis>
. In Browns key for
<taxonomicName id="7FC570EE3F33FFBA2F74FE50BEB2FE77" box="[577,729,479,503]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="8AB1D77F3F33FFBA2F74FE50BEB2FE77" box="[577,729,479,503]" italics="true" pageId="4" pageNumber="57">Mannheimsia</emphasis>
</taxonomicName>
species (
<bibRefCitation id="DC54769C3F33FFBA2E73FE50BFBDFE77" author="Brown" box="[838,982,479,503]" pageId="4" pageNumber="57" refString="Brown, B. V. (2005) Classification of two poorly known genera of African Phoridae (Diptera). African Invertebrates, 46, 133 - 140." type="journal article" year="2005">Brown 2005</bibRefCitation>
)
<taxonomicName id="7FC570EE3F33FFBA2EDDFE6FB835FE77" box="[1000,1118,480,503]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="8AB1D77F3F33FFBA2EDDFE6FBF94FE77" box="[1000,1023,480,503]" italics="true" pageId="4" pageNumber="57">M</emphasis>
.
<emphasis id="8AB1D77F3F33FFBA2925FE6FB835FE77" box="[1040,1118,480,503]" italics="true" pageId="4" pageNumber="57">conica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="91826A043F33FFBA2959FE6FB8AEFE77" box="[1132,1221,480,503]" pageId="4" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
keys in couplet 2, together with
<taxonomicName id="7FC570EE3F33FFBA2C01FD8BBDCAFD9B" box="[308,417,516,539]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="stricta">
<emphasis id="8AB1D77F3F33FFBA2C01FD8BBDCAFD9B" box="[308,417,516,539]" italics="true" pageId="4" pageNumber="57">M. stricta</emphasis>
</taxonomicName>
.
<taxonomicName id="7FC570EE3F33FFBA2C9AFD8CBEF1FD9B" box="[431,666,515,539]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="8AB1D77F3F33FFBA2C9AFD8CBEF1FD9B" box="[431,666,515,539]" italics="true" pageId="4" pageNumber="57">Mannheimsia conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91826A043F33FFBA2F97FD8CBE92FD9A" box="[674,761,515,538]" pageId="4" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from this species by its elongate-conical flagellomere 1, two setae on the hindtibia, and many differences in the hypopygium and phallus.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DA2FD1DBEB0FC94" pageId="4" pageNumber="57" type="reference_group">
<paragraph id="B87A0B6D3F33FFBA2DA2FD1DBDD1FD4F" blockId="4.[151,532,658,719]" pageId="4" pageNumber="57">
<taxonomicName id="7FC570EE3F33FFBA2DA2FD1DBE7FFD2A" ID-CoL="3XVN6" authority="Beyer, 1965" authorityName="Beyer" authorityYear="1965" box="[151,532,658,683]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="stricta">
<emphasis id="8AB1D77F3F33FFBA2DA2FD1DBDEEFD2B" bold="true" box="[151,389,658,683]" italics="true" pageId="4" pageNumber="57">Mannheimsia stricta</emphasis>
<bibRefCitation id="DC54769C3F33FFBA2CB9FD1CBE7FFD2A" author="Beyer" box="[396,532,658,683]" pageId="4" pageNumber="57" refString="Beyer, E. M. (1965) Phoridae. Exploration Parc National Albert, Mission G. F. de Witte (1933 - 1935), 99, 1 - 211." type="journal article" year="1965">Beyer, 1965</bibRefCitation>
</taxonomicName>
(
<figureCitation id="20FE17E83F33FFBA2DAAFD38BC85FD4F" box="[159,238,695,719]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[159,1413,196,1831]" captionTargetId="figure@2.[151,1436,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 3. Mannheimsia conica, sp. nov. and Mannheimsia stricta. 1. M. conica, sp. nov. habitus; 2. M. conica, sp. nov. head in lateral view; 3. M. stricta abdomen in terminal-ventral view." httpUri="https://zenodo.org/record/212711/files/figure.png" pageId="4" pageNumber="57">Figs. 3</figureCitation>
,
<figureCitation id="20FE17E83F33FFBA2DCFFD38BD61FD4F" box="[250,266,695,719]" captionStart="FIGURES 4 7" captionStartId="3.[151,264,1511,1534]" captionTargetBox="[164,1436,844,1474]" captionTargetId="figure@3.[151,1436,831,1489]" captionTargetPageId="3" captionText="FIGURES 4 7. Mannheimsia wings. 4. M. conica, sp. nov.; 5. M. stricta; 6. M. stylodactyla; 7. M. tianzena." httpUri="https://zenodo.org/record/212712/files/figure.png" pageId="4" pageNumber="57">5</figureCitation>
,
<figureCitation id="20FE17E83F33FFBA2C22FD38BD35FD4F" box="[279,350,695,719]" captionStart="FIGURES 8 23" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[178,1404,339,1817]" captionTargetId="figure@5.[151,1436,313,1841]" captionTargetPageId="5" captionText="FIGURES 8 23. Mannheimsia hypopygium. 8 11. Mannheimsia conica, sp. nov., 8. Dorsal, 9. Ventral, 10. Left lateral, 11. Right lateral; 12 15. Mannheimsia stricta, 12. Dorsal, 13. Ventral, 14. Left lateral, 15. Right lateral; 16 19. Mannheimsia stylodactyla, 16. Dorsal, 17. Ventral, 18. Left lateral, 19. Right lateral; 20 23. Mannheimsia tianzena, 20. Dorsal, 21. Ventral, 22. Left lateral, 23. Right lateral. Colors indicating homology: blue, hypandrium left lobe; yellow, hypandrium right lobe; red, hypandrium right lobe medial process. Abbreviations: cerc, cercus; lep, left epandrial process (fused surstyli); hyprct, hypoproct; rsp, right subepandrial process." httpUri="https://zenodo.org/record/212713/files/figure.png" pageId="4" pageNumber="57">1215</figureCitation>
,
<figureCitation id="20FE17E83F33FFBA2C5FFD38BDD9FD4F" box="[362,434,695,719]" captionStart="FIGURES 24 31" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[256,1286,195,1887]" captionTargetId="figure@6.[256,1331,157,1923]" captionTargetPageId="6" captionText="FIGURES 24 31. Mannheimsia phallus. 24 25. M. conica, sp. nov., 24. Dorsal, 25. Left lateral; 26 27. M. stricta, 26. Dorsal, 27. Left lateral; 28 29. M. stylodactyla, 28. Dorsal; 29. Left lateral; 30 31. M. tianzena, 30. Dorsal, 31. Left lateral. Abbreviations: bs, basiphallus; epi, epiphallus; epl, epiphallus posterior lobe; p 1 and p 2, processes 1 and 2, respectively; ra, right arm. Core plate represented in yellow." httpUri="https://zenodo.org/record/212714/files/figure.png" pageId="4" pageNumber="57">2627</figureCitation>
)
</paragraph>
<paragraph id="B87A0B6D3F33FFBA2DA2FD70BEB0FC94" blockId="4.[151,731,766,789]" box="[151,731,766,789]" pageId="4" pageNumber="57">
<treatmentCitationGroup id="98D52C433F33FFBA2DA2FD70BEB0FC94" box="[151,731,766,789]" pageId="4" pageNumber="57">
<taxonomicName id="7FC570EE3F33FFBA2DA2FD70BE79FC94" ID-CoL="3XVN6" authority="Beyer, 1965: 29" authorityName="Beyer" authorityPageNumber="29" authorityYear="1965" box="[151,530,766,789]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="stricta">
<emphasis id="8AB1D77F3F33FFBA2DA2FD70BD0CFC95" box="[151,359,767,789]" italics="true" pageId="4" pageNumber="57">Mannheimsia stricta</emphasis>
<treatmentCitation id="39642D7C3F33FFBA2C5BFD70BE79FC94" author="Beyer" box="[366,530,766,788]" page="29" pageId="4" pageNumber="57" year="1965">
<bibRefCitation id="DC54769C3F33FFBA2C5BFD70BD87FC94" author="Beyer" box="[366,492,766,788]" pageId="4" pageNumber="57" refString="Beyer, E. M. (1965) Phoridae. Exploration Parc National Albert, Mission G. F. de Witte (1933 - 1935), 99, 1 - 211." type="journal article" year="1965">Beyer, 1965</bibRefCitation>
: 29
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="39642D7C3F33FFBA2F28FD70BEBCFC94" ID-DOI="http://doi.org/10.5281/zenodo.7667293" ID-Zenodo-Dep="7667293" author="Brown" box="[541,727,766,788]" httpUri="http://treatment.plazi.org/id/3E7AF841323DEB5B5691FB0434D5AFAC" page="136" pageId="4" pageNumber="57" year="2005">
<bibRefCitation id="DC54769C3F33FFBA2F28FD70BECEFC94" author="Brown" box="[541,677,766,788]" pageId="4" pageNumber="57" refString="Brown, B. V. (2005) Classification of two poorly known genera of African Phoridae (Diptera). African Invertebrates, 46, 133 - 140." type="journal article" year="2005">Brown, 2005</bibRefCitation>
: 136
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DA2FCCEB965FC21" pageId="4" pageNumber="57" type="materials_examined">
<paragraph id="B87A0B6D3F33FFBA2DA2FCCEB965FC21" blockId="4.[151,1437,833,1653]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DA2FCCEBD10FCDA" bold="true" box="[151,379,833,858]" pageId="4" pageNumber="57">Material examined</emphasis>
. 1ɗ,
<emphasis id="8AB1D77F3F33FFBA2C88FCCEBE4BFCDA" bold="true" box="[445,544,833,858]" pageId="4" pageNumber="57">
<collectingCountry id="C0D24BFD3F33FFBA2C88FCCEBE4BFCDA" box="[445,544,833,858]" name="Kenya" pageId="4" pageNumber="57">KENYA</collectingCountry>
</emphasis>
: Kakamega: Isecheno Nature Reserve,
<geoCoordinate id="DDF16DAA3F33FFBA2EC1FCCEB829FCD9" box="[1012,1090,833,857]" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="555" value="0.24">0.24ºN</geoCoordinate>
<geoCoordinate id="DDF16DAA3F33FFBA2979FCCEB8C3FCD9" box="[1100,1192,833,857]" direction="east" orientation="longitude" pageId="4" pageNumber="57" precision="555" value="34.86">34.86ºE</geoCoordinate>
,
<date id="CC7B2DAD3F33FFBA2980FCCEB93CFCD9" box="[1205,1367,833,857]" pageId="4" pageNumber="57" value="2001-04-19" valueMax="2001-04-30" valueMin="2001-04-19">1930.iv.2001</date>
, Malaise trap,
<quantity id="7F3DA6883F33FFBA2C31FCEABD3FFCFD" box="[260,340,869,893]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="57" unit="m" value="1800.0">1800m</quantity>
, R. Snelling col; 2ɗ, same data but
<date id="CC7B2DAD3F33FFBA2FDDFCEABF12FCFD" box="[744,889,869,893]" pageId="4" pageNumber="57" value="2002-02-01" valueMax="2002-02-10" valueMin="2002-02-01">110.ii.2002</date>
; 1ɗ, same data but
<date id="CC7B2DAD3F33FFBA2960FCEAB89BFCFD" box="[1109,1264,869,893]" pageId="4" pageNumber="57" value="2002-05-11" valueMax="2002-05-20" valueMin="2002-05-11">1120.v.2002</date>
, W. Okeka col; 1ɗ,
<emphasis id="8AB1D77F3F33FFBA2DF0FC06BD2BFC22" bold="true" box="[197,320,905,930]" pageId="4" pageNumber="57">
<collectingCountry id="C0D24BFD3F33FFBA2DF0FC06BD2BFC22" box="[197,320,905,930]" name="Nigeria" pageId="4" pageNumber="57">NIGERIA</collectingCountry>
</emphasis>
: Onde Idanre Hills,
<geoCoordinate id="DDF16DAA3F33FFBA2F18FC06BE11FC21" box="[557,634,905,929]" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="555" value="7.09">7.09ºN</geoCoordinate>
<geoCoordinate id="DDF16DAA3F33FFBA2FB7FC06BEA4FC21" box="[642,719,905,929]" direction="east" orientation="longitude" pageId="4" pageNumber="57" precision="555" value="5.4">5.40ºE</geoCoordinate>
,
<date id="CC7B2DAD3F33FFBA2FEFFC06BFECFC21" box="[730,903,905,929]" pageId="4" pageNumber="57" value="2008-07-16" valueMax="2008-07-19" valueMin="2008-07-16">1619.vii.2008</date>
, Malaise trap,
<quantity id="7F3DA6883F33FFBA291EFC06B805FC21" box="[1067,1134,905,929]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="57" unit="m" value="440.0">440m</quantity>
, J. Heraty col.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DFCFC22BF40FA64" pageId="4" pageNumber="57" type="description">
<paragraph id="B87A0B6D3F33FFBA2DFCFC22BFE4FA1C" blockId="4.[151,1437,833,1653]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFC22BD04FC46" bold="true" box="[201,367,941,966]" pageId="4" pageNumber="57">Redescription</emphasis>
.
<emphasis id="8AB1D77F3F33FFBA2CB7FC22BDD5FC45" bold="true" box="[386,446,941,965]" pageId="4" pageNumber="57">Male</emphasis>
. Body length,
<quantity id="7F3DA6883F33FFBA2F47FC22BF6AFC45" box="[626,769,941,965]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="4" pageNumber="57" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1 mm</quantity>
.
<emphasis id="8AB1D77F3F33FFBA2E25FC22BF3BFC45" bold="true" box="[784,848,941,965]" pageId="4" pageNumber="57">Head</emphasis>
. Frons brown, pubescent, without median furrow. Flagellomere 1 light brown, pubescent, globose. Arista dorsal, pubescent. Palpus light brown; 46 medium size genal setae.
<emphasis id="8AB1D77F3F33FFBA2C10FC7ABD16FB8D" bold="true" box="[293,381,1013,1037]" pageId="4" pageNumber="57">Thorax</emphasis>
. Scutum, scutellum and pleural regions yellowish-brown; anepisternum setulose. Legs light brown, midcoxa brown. Foretibia with one dorsal seta at basal third and anterodorsal row of strong setae. Midtibia with one anterodorsal and one posterodorsal setae.
<collectingCountry id="C0D24BFD3F33FFBA2FE9FBB2BF7FFBD5" box="[732,788,1085,1109]" name="India" pageId="4" pageNumber="57">Hind</collectingCountry>
femur not swollen.
<collectingCountry id="C0D24BFD3F33FFBA2ECCFBB2B85AFBD5" box="[1017,1073,1085,1109]" name="India" pageId="4" pageNumber="57">Hind</collectingCountry>
tibia with one anterodorsal seta near midlength.
<emphasis id="8AB1D77F3F33FFBA2C78FBEEBDE6FBFA" bold="true" box="[333,397,1121,1146]" pageId="4" pageNumber="57">Wing</emphasis>
. (
<figureCitation id="20FE17E83F33FFBA2C90FBEEBD8CFBF9" box="[421,487,1121,1145]" captionStart="FIGURES 4 7" captionStartId="3.[151,264,1511,1534]" captionTargetBox="[164,1436,844,1474]" captionTargetId="figure@3.[151,1436,831,1489]" captionTargetPageId="3" captionText="FIGURES 4 7. Mannheimsia wings. 4. M. conica, sp. nov.; 5. M. stricta; 6. M. stylodactyla; 7. M. tianzena." httpUri="https://zenodo.org/record/212712/files/figure.png" pageId="4" pageNumber="57">Fig. 5</figureCitation>
) Costa 0.37 of the wing length; M2 proximal half slightly concave, distal half almost straight. Halter yellowish-brown with apical black circle.
<emphasis id="8AB1D77F3F33FFBA2E2DFB08BFE7FB20" bold="true" box="[792,908,1159,1184]" pageId="4" pageNumber="57">Abdomen</emphasis>
. Tergites brown. Ventral pair of gland-like projections near the hypopygium, connected to the internal wall of abdomen (
<figureCitation id="20FE17E83F33FFBA2EC1FB23B851FB44" box="[1012,1082,1196,1220]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[159,1413,196,1831]" captionTargetId="figure@2.[151,1436,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 3. Mannheimsia conica, sp. nov. and Mannheimsia stricta. 1. M. conica, sp. nov. habitus; 2. M. conica, sp. nov. head in lateral view; 3. M. stricta abdomen in terminal-ventral view." httpUri="https://zenodo.org/record/212711/files/figure.png" pageId="4" pageNumber="57">Fig. 3</figureCitation>
). Venter of abdomen with two pairs of rounded elevations near hypopygium bearing few strong setae. Hypopygium yellowish-brown (
<figureCitation id="20FE17E83F33FFBA2804FB5FBCDDFA8C" captionStart="FIGURES 8 23" captionStartId="5.[151,264,1862,1885]" captionTargetBox="[178,1404,339,1817]" captionTargetId="figure@5.[151,1436,313,1841]" captionTargetPageId="5" captionText="FIGURES 8 23. Mannheimsia hypopygium. 8 11. Mannheimsia conica, sp. nov., 8. Dorsal, 9. Ventral, 10. Left lateral, 11. Right lateral; 12 15. Mannheimsia stricta, 12. Dorsal, 13. Ventral, 14. Left lateral, 15. Right lateral; 16 19. Mannheimsia stylodactyla, 16. Dorsal, 17. Ventral, 18. Left lateral, 19. Right lateral; 20 23. Mannheimsia tianzena, 20. Dorsal, 21. Ventral, 22. Left lateral, 23. Right lateral. Colors indicating homology: blue, hypandrium left lobe; yellow, hypandrium right lobe; red, hypandrium right lobe medial process. Abbreviations: cerc, cercus; lep, left epandrial process (fused surstyli); hyprct, hypoproct; rsp, right subepandrial process." httpUri="https://zenodo.org/record/212713/files/figure.png" pageId="4" pageNumber="57">Figs. 12 15</figureCitation>
). Left epandrial process (fused surstyli) with median finger-like extension. Right epandrial posterior margin narrowing ventrally, with subepandrial right process bearing setae. Hypandrium with two similar lobes; left lobe sometimes with a suture, with a medially-directed process. Hypoproct with two setae. Cerci small.
<emphasis id="8AB1D77F3F33FFBA29CEFAB2B93FFAD5" bold="true" box="[1275,1364,1341,1365]" pageId="4" pageNumber="57">Phallus</emphasis>
(
<figureCitation id="20FE17E83F33FFBA2851FAB3BC8AFAF8" captionStart="FIGURES 24 31" captionStartId="6.[151,264,1888,1911]" captionTargetBox="[256,1286,195,1887]" captionTargetId="figure@6.[256,1331,157,1923]" captionTargetPageId="6" captionText="FIGURES 24 31. Mannheimsia phallus. 24 25. M. conica, sp. nov., 24. Dorsal, 25. Left lateral; 26 27. M. stricta, 26. Dorsal, 27. Left lateral; 28 29. M. stylodactyla, 28. Dorsal; 29. Left lateral; 30 31. M. tianzena, 30. Dorsal, 31. Left lateral. Abbreviations: bs, basiphallus; epi, epiphallus; epl, epiphallus posterior lobe; p 1 and p 2, processes 1 and 2, respectively; ra, right arm. Core plate represented in yellow." httpUri="https://zenodo.org/record/212714/files/figure.png" pageId="4" pageNumber="57">Figs. 2627</figureCitation>
). Right arm with dorsal projection. Epiphallus large, almost reaching the posterior part of the phallus. Core plate large, bare and flattened, with three dorsally curved processes.
</paragraph>
<paragraph id="B87A0B6D3F33FFBA2DFCFA27BDCFFA40" blockId="4.[151,1437,833,1653]" box="[201,420,1448,1472]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFA27BD4BFA40" bold="true" box="[201,288,1448,1472]" pageId="4" pageNumber="57">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="B87A0B6D3F33FFBA2DFCFA43BF40FA64" blockId="4.[151,1437,833,1653]" box="[201,811,1484,1509]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFA43BD82FA65" bold="true" box="[201,489,1484,1509]" pageId="4" pageNumber="57">Geographic distribution</emphasis>
.
<collectingCountry id="C0D24BFD3F33FFBA2CC2FA42BE28FA65" box="[503,579,1485,1509]" name="Kenya" pageId="4" pageNumber="57">Kenya</collectingCountry>
,
<collectingCountry id="C0D24BFD3F33FFBA2F7AFA43BEF2FA64" box="[591,665,1484,1508]" name="Republic of the Congo" pageId="4" pageNumber="57">Congo</collectingCountry>
and
<collectingCountry id="C0D24BFD3F33FFBA2FFAFA43BF4CFA64" box="[719,807,1484,1508]" name="Nigeria" pageId="4" pageNumber="57">Nigeria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F0DF58E63F33FFBA2DFCFA60BD93F9F5" pageId="4" pageNumber="57" type="discussion">
<paragraph id="B87A0B6D3F33FFBA2DFCFA60BD93F9F5" blockId="4.[151,1437,833,1653]" pageId="4" pageNumber="57">
<emphasis id="8AB1D77F3F33FFBA2DFCFA60BD21F988" bold="true" box="[201,330,1519,1544]" pageId="4" pageNumber="57">Comments</emphasis>
. The recognition of gland-like structures and the rounded setose elevations in the venter of the abdomen of
<taxonomicName id="7FC570EE3F33FFBA2C11F99ABDFAF9AC" box="[292,401,1557,1580]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="stricta">
<emphasis id="8AB1D77F3F33FFBA2C11F99ABDFAF9AC" box="[292,401,1557,1580]" italics="true" pageId="4" pageNumber="57">M. stricta</emphasis>
</taxonomicName>
are the main complements to the original description (
<bibRefCitation id="DC54769C3F33FFBA2ECDF99BB816F9AC" author="Beyer" box="[1016,1149,1556,1580]" pageId="4" pageNumber="57" refString="Beyer, E. M. (1965) Phoridae. Exploration Parc National Albert, Mission G. F. de Witte (1933 - 1935), 99, 1 - 211." type="journal article" year="1965">Beyer 1965</bibRefCitation>
) and Browns revision of this species (
<bibRefCitation id="DC54769C3F33FFBA2C10F9B7BDDDF9D0" author="Brown" box="[293,438,1592,1616]" pageId="4" pageNumber="57" refString="Brown, B. V. (2005) Classification of two poorly known genera of African Phoridae (Diptera). African Invertebrates, 46, 133 - 140." type="journal article" year="2005">Brown 2005</bibRefCitation>
).
<taxonomicName id="7FC570EE3F33FFBA2CFDF9B7BEC5F9D0" box="[456,686,1592,1616]" class="Insecta" family="Phoridae" genus="Mannheimsia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="stricta">
<emphasis id="8AB1D77F3F33FFBA2CFDF9B7BEC5F9D0" box="[456,686,1592,1616]" italics="true" pageId="4" pageNumber="57">Mannheimsia stricta</emphasis>
</taxonomicName>
is the only species in the genus with relatively small cerci, a condition present in some outgroups.
</paragraph>
</subSubSection>
</treatment>
</document>