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<document id="DD1E63F28F8F35128864BD5C73862797" ID-DOI="10.1111/j.1096-3642.2008.00510.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443607" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630455996773" checkinUser="felipe" docAuthor="Prathapan, K. D. &amp; Viraktamath, C. A." docDate="2009" docId="03E7375EFFD7FF98FC119D5C38F89D8B" docLanguage="en" docName="j.1096-3642.2008.00510.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00510.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Tegyrius anupama Prathapan &amp; Viraktamath 2009, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="335" masterDocId="FFDE4F26FFD1FF91FF80996E3A70983B" masterDocTitle="Revision of Tegyrius (Coleoptera: Chrysomelidae: Galerucinae: Alticini), with descriptions of eight new species" masterLastPageNumber="358" masterPageNumber="326" pageNumber="332" updateTime="1699277305514" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="78298F3734B056D681C071076FE4F48F">Revision of Tegyrius (Coleoptera: Chrysomelidae: Galerucinae: Alticini), with descriptions of eight new species</mods:title>
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<mods:namePart id="E1A19C923D783472C3640B8A2727D3FE">Prathapan, K. D.</mods:namePart>
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<mods:date id="2436F32D0E2EFDB1105539DED9316B61">2009</mods:date>
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<treatment id="03E7375EFFD7FF98FC119D5C38F89D8B" ID-DOI="http://doi.org/10.5281/zenodo.5492289" ID-Zenodo-Dep="5492289" LSID="urn:lsid:plazi:treatment:03E7375EFFD7FF98FC119D5C38F89D8B" httpUri="http://treatment.plazi.org/id/03E7375EFFD7FF98FC119D5C38F89D8B" lastPageId="9" lastPageNumber="335" pageId="6" pageNumber="332">
<subSubSection id="C354D5C3FFD7FF97FC119D5C3F3A9C72" box="[913,1354,1073,1098]" pageId="6" pageNumber="332" type="nomenclature">
<paragraph id="8BF18648FFD7FF97FC119D5C3F3A9C72" blockId="6.[913,1354,1073,1130]" box="[913,1354,1073,1098]" pageId="6" pageNumber="332">
<heading id="D0B93124FFD7FF97FC119D5C3F3A9C72" allCaps="true" box="[913,1354,1073,1098]" centered="true" fontSize="10" level="1" pageId="6" pageNumber="332" reason="1">
<emphasis id="B93A5A5AFFD7FF97FC119D5C3F3A9C72" bold="true" box="[913,1354,1073,1098]" pageId="6" pageNumber="332">
<taxonomicName id="4C4EFDCBFFD7FF97FC119D5C3EB69C71" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[913,1222,1074,1098]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="343" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">
<emphasis id="B93A5A5AFFD7FF97FC119D5C3EB69C71" bold="true" box="[913,1222,1074,1098]" italics="true" pageId="6" pageNumber="332">TEGYRIUS ANUPAMA</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD7FF97FB519D5F3F3A9C72" box="[1233,1354,1073,1097]" pageId="6" pageNumber="332" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C354D5C3FFD7FF98FB9B9D3C38F89D8B" lastPageId="9" lastPageNumber="335" pageId="6" pageNumber="332" type="description">
<paragraph id="8BF18648FFD7FF97FB9B9D3C3EB09C52" blockId="6.[913,1354,1073,1130]" box="[1051,1216,1105,1130]" pageId="6" pageNumber="332">
(
<figureCitation id="13759ACDFFD7FF97FBA39D3C3EC89C52" box="[1059,1208,1105,1130]" captionStart-0="Figures 1521" captionStart-1="Figures 2224" captionStartId-0="7.[144,234,1590,1609]" captionStartId-1="8.[164,254,1270,1289]" captionTargetBox-0="[308,1263,197,1560]" captionTargetBox-1="[326,1280,201,1238]" captionTargetId-0="figure-69@7.[303,1263,194,1560]" captionTargetId-1="figure-250@8.[323,1283,196,1241]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." captionText-1="Figures 2224. Tegyrius anupama sp. nov.: 22, spermatheca; 23, vaginal palpi; 24, tignum." figureDoi-0="http://doi.org/10.5281/zenodo.5443613" figureDoi-1="http://doi.org/10.5281/zenodo.5443617" httpUri-0="https://zenodo.org/record/5443613/files/figure.png" httpUri-1="https://zenodo.org/record/5443617/files/figure.png" pageId="6" pageNumber="332">FIGS 1524</figureCitation>
)
</paragraph>
<paragraph id="8BF18648FFD7FF97FCBA9DFD3E239C92" blockId="6.[826,1107,1171,1193]" box="[826,1107,1171,1193]" pageId="6" pageNumber="332">
<emphasis id="B93A5A5AFFD7FF97FCBA9DFD39BC9C93" box="[826,972,1171,1192]" italics="true" pageId="6" pageNumber="332">Distribution:</emphasis>
<collectingCountry id="F359C6D8FFD7FF97FC569DFD3E3F9C92" box="[982,1103,1171,1193]" name="Sri Lanka" pageId="6" pageNumber="332">
Sri
<taxonomicName id="4C4EFDCBFFD7FF97FB829DFD3E3F9C92" box="[1026,1103,1171,1193]" class="Insecta" family="Chrysomelidae" genus="Lanka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="358" phylum="Arthropoda" rank="genus">Lanka</taxonomicName>
</collectingCountry>
.
</paragraph>
<paragraph id="8BF18648FFD7FF97FCBA9DBA3EF19CD2" blockId="6.[826,1153,1235,1257]" box="[826,1153,1235,1257]" pageId="6" pageNumber="332">
<emphasis id="B93A5A5AFFD7FF97FCBA9DBA3E109CD2" box="[826,1120,1235,1257]" italics="true" pageId="6" pageNumber="332">
Host plant:
<taxonomicName id="4C4EFDCBFFD7FF97FC459DBA3E109CD2" box="[965,1120,1236,1257]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="6" pageNumber="332" phylum="Tracheophyta" rank="species" species="nigrum">Piper nigrum</taxonomicName>
</emphasis>
L.
</paragraph>
<paragraph id="8BF18648FFD7FF97FCBA9C7A3E529DDB" blockId="6.[826,1441,1299,1903]" pageId="6" pageNumber="332">
<emphasis id="B93A5A5AFFD7FF97FCBA9C7A39B29D12" box="[826,962,1300,1321]" italics="true" pageId="6" pageNumber="332">Description:</emphasis>
General colour shiny black. Head piceous, with brownish tinge. Pronotum and elytra shiny black. Sternites dark brown to black. Antennae and legs yellowish brown, except pro- and mesofemora, which are dark brown in the proximal two-thirds, and metafemora, which are dark brown with a lighter apex.
</paragraph>
<paragraph id="8BF18648FFD7FF97FCD39C843E189EEE" blockId="6.[826,1441,1299,1903]" pageId="6" pageNumber="332">Supraorbital sulcus weak, but distinct. Third antennomere slightly shorter than fourth; fifth slightly longer than fourth; sixth shorter than fifth; seventh longer than sixth; eighth to tenth each slightly shorter than preceding one; last subequal to or slightly shorter than first. Distal five antennomeres slightly thickened. Antennae reach half of the elytra over the pronotum.</paragraph>
<paragraph id="8BF18648FFD7FF96FCD39FB03BE49F0A" blockId="6.[826,1441,1299,1903]" lastBlockId="7.[144,759,1758,1903]" lastPageId="7" lastPageNumber="333" pageId="6" pageNumber="332">Antebasal transverse impression absent. Apex of prosternal intercoxal process convex, preapical depressions indistinct. Prosternal intercoxal process 4.3 times wider than distance between anterior margin of prosternum to coxal cavity; distance from anterior margin of prosternum to end of intercoxal prosternal process 2.0 times the minimum width of the prosternal intercoxal process.</paragraph>
<caption id="DF31D6C0FFD6FF96FF109F583B1F9EBF" ID-DOI="http://doi.org/10.5281/zenodo.5443613" ID-Zenodo-Dep="5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="7" pageNumber="333" startId="7.[144,234,1590,1609]" targetBox="[308,1263,197,1560]" targetPageId="7">
<paragraph id="8BF18648FFD6FF96FF109F583B1F9EBF" blockId="7.[144,1421,1590,1668]" pageId="7" pageNumber="333">
<emphasis id="B93A5A5AFFD6FF96FF109F5838FB9E72" bold="true" box="[144,651,1590,1609]" pageId="7" pageNumber="333">
Figures 1521.
<taxonomicName id="4C4EFDCBFFD6FF96FEC29F5838519E72" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[322,545,1590,1609]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="333" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">
<emphasis id="B93A5A5AFFD6FF96FEC29F5838519E72" bold="true" box="[322,545,1590,1609]" italics="true" pageId="7" pageNumber="333">Tegyrius anupama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD6FF96FDAE9F5838FB9E72" box="[558,651,1590,1609]" pageId="7" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen.
</paragraph>
</caption>
<paragraph id="8BF18648FFD6FF99FF289E543B699D9A" blockId="7.[144,759,1758,1903]" lastBlockId="8.[164,779,1358,1900]" lastPageId="8" lastPageNumber="334" pageId="7" pageNumber="333">Mesoscutellum broader than long, with broadly rounded apex. Mesosternal intercoxal process depressed on top, posterior margin concave. Elytra with maximum width at anterior two-thirds. Elytral apex convex, and moderately broad. Elytral punctures minute, tend to form rows, with the distance between adjacent punctures 1.53.5 times the diameter of a puncture in the anterior half of the elytron. First metatarsomere distinctly longer than half of metatibia.</paragraph>
<caption id="DF31D6C0FFD9FF99FF249D983EEF9D31" ID-DOI="http://doi.org/10.5281/zenodo.5443617" ID-Zenodo-Dep="5443617" box="[164,1183,1270,1290]" httpUri="https://zenodo.org/record/5443617/files/figure.png" pageId="8" pageNumber="334" startId="8.[164,254,1270,1289]" targetBox="[326,1280,201,1238]" targetPageId="8">
<paragraph id="8BF18648FFD9FF99FF249D983EEF9D31" blockId="8.[164,1183,1270,1290]" box="[164,1183,1270,1290]" pageId="8" pageNumber="334">
<emphasis id="B93A5A5AFFD9FF99FF249D9838E39D31" bold="true" box="[164,659,1270,1290]" pageId="8" pageNumber="334">
Figures 2224.
<taxonomicName id="4C4EFDCBFFD9FF99FED69D9938419D31" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[342,561,1271,1290]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="334" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">
<emphasis id="B93A5A5AFFD9FF99FED69D9938419D31" bold="true" box="[342,561,1271,1290]" italics="true" pageId="8" pageNumber="334">Tegyrius anupama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD9FF99FDBB9D9938E39D31" box="[571,659,1271,1290]" pageId="8" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 22, spermatheca; 23, vaginal palpi; 24, tignum.
</paragraph>
</caption>
<paragraph id="8BF18648FFD9FF99FF3C9CC438B49E4C" blockId="8.[164,779,1358,1900]" pageId="8" pageNumber="334">
Subparalllel ridges along first abdominal ventrite well developed, extending beyond two-thirds of ventrite, nearly parallel with converging ends (
<figureCitation id="13759ACDFFD9FF99FD2D9C89388D9DC6" box="[685,765,1511,1533]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 15</figureCitation>
). Last visible tergite of females with a short groove. Last visible ventrite in males with a group of microsetae on each side laterally, lobe in middle of last male ventrite with a convex apical margin (
<figureCitation id="13759ACDFFD9FF99FDE69F0F38C79E4C" box="[614,695,1633,1655]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BF18648FFD9FF99FF3C9FEE38DB9F57" blockId="8.[164,779,1358,1900]" pageId="8" pageNumber="334">
Aedeagus laterally compressed, moderately curved in lateral view, with a transverse depression preapically (
<figureCitation id="13759ACDFFD9FF99FF699FD33B4A9EE8" box="[233,314,1725,1747]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 19</figureCitation>
). In ventral view, apex without angulate extremities, with a narrow transparent window along the middle, sharply raised along the middle, and forming an acute preapical ridge (
<figureCitation id="13759ACDFFD9FF99FDD79E7738D79F14" box="[599,679,1817,1839]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 18</figureCitation>
). Distal opening partially covered with lamina (
<figureCitation id="13759ACDFFD9FF99FDF29E5938B49F75" box="[626,708,1847,1870]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 20</figureCitation>
). Arm of tegmen much shorter than stem (
<figureCitation id="13759ACDFFD9FF99FDCE9E3838EF9F57" box="[590,671,1878,1900]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BF18648FFD9FF99FCD39C2039D09E4C" blockId="8.[826,1441,1358,1655]" pageId="8" pageNumber="334">
Spermatheca with horizontal part of pump shorter than vertical part, receptacle nearly three times longer than wide, outer and inner sides slighty convex in middle, and narrowed towards duct; duct folded three or four times (
<figureCitation id="13759ACDFFD9FF99FB439CA63F659DE5" box="[1219,1301,1480,1502]" captionStart="Figures 2224" captionStartId="8.[164,254,1270,1289]" captionTargetBox="[326,1280,201,1238]" captionTargetId="figure-250@8.[323,1283,196,1241]" captionTargetPageId="8" captionText="Figures 2224. Tegyrius anupama sp. nov.: 22, spermatheca; 23, vaginal palpi; 24, tignum." figureDoi="http://doi.org/10.5281/zenodo.5443617" httpUri="https://zenodo.org/record/5443617/files/figure.png" pageId="8" pageNumber="334">Fig. 22</figureCitation>
). Proximal sclerotization of vaginal palpus subequal to distal sclerotization, with each sclerotization being shorter than the unsclerotized lateral area (
<figureCitation id="13759ACDFFD9FF99FAC39F4A3FE59E01" box="[1347,1429,1572,1594]" captionStart="Figures 2224" captionStartId="8.[164,254,1270,1289]" captionTargetBox="[326,1280,201,1238]" captionTargetId="figure-250@8.[323,1283,196,1241]" captionTargetPageId="8" captionText="Figures 2224. Tegyrius anupama sp. nov.: 22, spermatheca; 23, vaginal palpi; 24, tignum." figureDoi="http://doi.org/10.5281/zenodo.5443617" httpUri="https://zenodo.org/record/5443617/files/figure.png" pageId="8" pageNumber="334">Fig. 23</figureCitation>
). Tignum curved with broad distal sclerotization (
<figureCitation id="13759ACDFFD9FF99FCC29F0F39E49E4C" box="[834,916,1633,1655]" captionStart="Figures 2224" captionStartId="8.[164,254,1270,1289]" captionTargetBox="[326,1280,201,1238]" captionTargetId="figure-250@8.[323,1283,196,1241]" captionTargetPageId="8" captionText="Figures 2224. Tegyrius anupama sp. nov.: 22, spermatheca; 23, vaginal palpi; 24, tignum." figureDoi="http://doi.org/10.5281/zenodo.5443617" httpUri="https://zenodo.org/record/5443617/files/figure.png" pageId="8" pageNumber="334">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="8BF18648FFD9FF99FCBA9FD03E9C9F55" blockId="8.[826,1441,1726,1902]" pageId="8" pageNumber="334">
<emphasis id="B93A5A5AFFD9FF99FCBA9FD03E559EE8" box="[826,1061,1726,1747]" italics="true" pageId="8" pageNumber="334">Sexual dimorphism:</emphasis>
First pro- and mesotarsomeres slightly enlarged in males. First metatarsomere of males modified as in
<taxonomicName id="4C4EFDCBFFD9FF99FBB19F923EBE9F2A" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" baseAuthorityName="Medvedev" baseAuthorityYear="2001" box="[1073,1230,1788,1809]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="334" phylum="Arthropoda" rank="species" species="antennatus" status="comb. nov.">
<emphasis id="B93A5A5AFFD9FF99FBB19F923EBE9F2A" box="[1073,1230,1788,1809]" italics="true" pageId="8" pageNumber="334">T. antennatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD9FF99FB569F923F209F2A" box="[1238,1360,1788,1810]" pageId="8" pageNumber="334" rank="species">comb. nov.</taxonomicNameLabel>
, but to a lesser extent. Posterior margin of last visible ventrite in females deeply incised, the incision with an acute process in the middle (
<figureCitation id="13759ACDFFD9FF99FB0E9E393EAF9F55" box="[1166,1247,1879,1902]" captionStart="Figures 1521" captionStartId="7.[144,234,1590,1609]" captionTargetBox="[308,1263,197,1560]" captionTargetId="figure-69@7.[303,1263,194,1560]" captionTargetPageId="7" captionText="Figures 1521. Tegyrius anupama sp. nov.: 15, intercoxal ridges on the first abdominal ventrite; 16, male last abdominal ventrite; 17, female last abdominal ventrite; 18, aedeagus, ventral view; 19, aedeagus, lateral view; 20, distal opening; 21, tegmen." figureDoi="http://doi.org/10.5281/zenodo.5443613" httpUri="https://zenodo.org/record/5443613/files/figure.png" pageId="8" pageNumber="334">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BF18648FFD8FF98FF1099A93B549A36" blockId="9.[144,759,198,526]" pageId="9" pageNumber="335">
<emphasis id="B93A5A5AFFD8FF98FF1099A93BEC98E0" box="[144,412,198,220]" italics="true" pageId="9" pageNumber="335">Measurements (n = 4):</emphasis>
Length, 2.002.21 (2.10); width, 1.101.30 (1.24); length of pronotum, 0.490.62 (0.57); width of pronotum, 0.740.82 (0.79); width of head across eyes, 0.550.60 (0.57); transverse diameter of one eye, 0.210.24 (0.22); distance between eyes, 0.210.26 (0.23); distance between eye and antennal socket, 0.020.03 (0.03); distance between antennal sockets, 0.070.08 (0.07); diameter of one antennal socket, 0.060.07 (0.07); length of aedeagus, 0.90; length of receptacle, 0.25; length of vaginal palpus, 0.39.
</paragraph>
<paragraph id="8BF18648FFD8FF98FF109B5438E59A90" blockId="9.[144,759,569,928]" pageId="9" pageNumber="335">
<emphasis id="B93A5A5AFFD8FF98FF109B543AA59A74" box="[144,213,570,591]" italics="true" pageId="9" pageNumber="335">Types:</emphasis>
<typeStatus id="54F538EAFFD8FF98FF619B573B379A74" box="[225,327,569,591]" pageId="9" pageNumber="335" type="holotype">Holotype</typeStatus>
<specimenCount id="9D484DC1FFD8FF98FED29B523B1B9A75" box="[338,363,572,590]" pageId="9" pageNumber="335" type="male"></specimenCount>
. Labels: (1)
<collectingCountry id="F359C6D8FFD8FF98FD839B57380D9A74" box="[515,637,569,591]" name="Sri Lanka" pageId="9" pageNumber="335">
Sri
<taxonomicName id="4C4EFDCBFFD8FF98FDB19B57380D9A74" authority=", Knuckles Range, Atanwala" authorityName="Knuckles Range, Atanwala" box="[561,637,569,591]" class="Insecta" family="Chrysomelidae" genus="Lanka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="genus">Lanka</taxonomicName>
</collectingCountry>
, Knuckles Range, Atanwala,
<date id="FFF0A088FFD8FF98FEFC9B3638229A55" box="[380,594,600,622]" pageId="9" pageNumber="335" value="2006-03-07" valueMax="2006-03-08" valueMin="2006-03-07">78 March 2006</date>
, coll. K. D. Prathapan; (2)
<taxonomicName id="4C4EFDCBFFD8FF98FEC09B1938659AB7" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[320,533,631,652]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">
<emphasis id="B93A5A5AFFD8FF98FEC09B1938659AB7" box="[320,533,631,652]" italics="true" pageId="9" pageNumber="335">Tegyrius anupama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD8FF98FD9E9B1938049AB7" box="[542,628,631,652]" pageId="9" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
Prathapan &amp; Viraktamath, 2007; (3)
<typeStatus id="54F538EAFFD8FF98FE419BFB38529A90" box="[449,546,661,683]" pageId="9" pageNumber="335" type="holotype">holotype</typeStatus>
(BMNH).
</paragraph>
<paragraph id="8BF18648FFD8FF98FF289BDA3B769B9B" blockId="9.[144,759,569,928]" pageId="9" pageNumber="335">
<typeStatus id="54F538EAFFD8FF98FF289BDA3B6B9AF2" box="[168,283,692,713]" pageId="9" pageNumber="335" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D484DC1FFD8FF98FEB29BDA3BA49AF1" box="[306,468,692,714]" pageId="9" pageNumber="335" type="generic">17 specimens</specimenCount>
):
<materialsCitation id="3B268C15FFD8FF98FE6D9BDA3A949B1E" collectingDate="2006-03-06" pageId="9" pageNumber="335" specimenCount="15" specimenCount-female="9" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D484DC1FFD8FF98FE6D9BDA38639AF3" box="[493,531,692,714]" pageId="9" pageNumber="335" type="male">6♂</specimenCount>
and
<specimenCount id="9D484DC1FFD8FF98FDD99BDA380C9AF3" box="[601,636,692,713]" pageId="9" pageNumber="335" type="female">9♀</specimenCount>
, with the same data as for the holotype, except the locality was Kandy and the date was
<date id="FFF0A088FFD8FF98FE459B9F381C9B3C" box="[453,620,753,775]" pageId="9" pageNumber="335" value="2006-03-06">
<collectingDate id="EFB45960FFD8FF98FE459B9F381C9B3C" box="[453,620,753,775]" pageId="9" pageNumber="335" value="2006-03-06">6 March 2006</collectingDate>
</date>
from
<taxonomicName id="4C4EFDCBFFD8FF98FD3A9B9F3A949B1E" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="9" pageNumber="335" phylum="Tracheophyta" rank="species" species="nigrum">
<emphasis id="B93A5A5AFFD8FF98FD3A9B9F3A949B1E" italics="true" pageId="9" pageNumber="335">Piper nigrum</emphasis>
</taxonomicName>
</materialsCitation>
;
<materialsCitation id="3B268C15FFD8FF98FF769A7E3B729B9B" collectingDate="2006-03-17" collectionCode="BMNH, NHMB, NPC, UASB, USNM" pageId="9" pageNumber="335" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D484DC1FFD8FF98FF769A7E3B6C9B1F" box="[246,284,784,805]" pageId="9" pageNumber="335" type="male">1♂</specimenCount>
and
<specimenCount id="9D484DC1FFD8FF98FEDD9A7E3B0F9B1F" box="[349,383,784,805]" pageId="9" pageNumber="335" type="female">1♀</specimenCount>
, with the same data as for the holotype, except the locality was Siripagama and the date was
<date id="FFF0A088FFD8FF98FE839A233BC59B58" box="[259,437,845,867]" pageId="9" pageNumber="335" value="2006-03-17">
<collectingDate id="EFB45960FFD8FF98FE839A233BC59B58" box="[259,437,845,867]" pageId="9" pageNumber="335" value="2006-03-17">17 March 2006</collectingDate>
</date>
(3
<collectionCode id="ED5F1E8DFFD8FF98FE619A2338459B59" box="[481,565,845,866]" pageId="9" pageNumber="335">BMNH</collectionCode>
, 2 EUJ, 2 KUJ, 1 LMC, 2
<collectionCode id="ED5F1E8DFFD8FF98FE939A023B179BBA" box="[275,359,876,897]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="9" pageNumber="335">NHMB</collectionCode>
, 3
<collectionCode id="ED5F1E8DFFD8FF98FE149A053BBD9BBB" box="[404,461,875,896]" country="India" httpUri="http://grbio.org/cool/nwky-2b63" name="National Pusa Collection" pageId="9" pageNumber="335">NPC</collectionCode>
, 1 PKDC, 1
<collectionCode id="ED5F1E8DFFD8FF98FDF09A0538CA9BBB" box="[624,698,875,896]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="9" pageNumber="335">UASB</collectionCode>
, and 2
<collectionCode id="ED5F1E8DFFD8FF98FF279AE43A8A9B9B" box="[167,250,906,928]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="335" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF18648FFD8FF98FF109AA538549C25" blockId="9.[144,759,971,1054]" pageId="9" pageNumber="335">
<emphasis id="B93A5A5AFFD8FF98FF109AA53BF69BDB" box="[144,390,971,992]" italics="true" pageId="9" pageNumber="335">Etymology: Anupama</emphasis>
literally means unique in Sanskrit, and refers to the unique features of the species. The name is a noun in apposition.
</paragraph>
<paragraph id="8BF18648FFD8FF98FF109D2438F89D8B" blockId="9.[144,759,1098,1456]" pageId="9" pageNumber="335">
<emphasis id="B93A5A5AFFD8FF98FF109D243BA69C64" box="[144,470,1098,1119]" italics="true" pageId="9" pageNumber="335">
Remarks:
<taxonomicName id="4C4EFDCBFFD8FF98FE839D243BA69C64" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[259,470,1098,1119]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">Tegyrius anupama</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A209E721FFD8FF98FE5E9D2438439C64" box="[478,563,1098,1119]" pageId="9" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
is unique among the species of the genus in having the last female ventrite with a deep incision bearing an acute process in the middle. Among the Oriental flea beetles,
<taxonomicName id="4C4EFDCBFFD8FF98FD469DC838C29CE1" authority="Chen &amp; Wang (1980)" authorityName="Chen &amp; Wang" authorityYear="1980" class="Insecta" family="Chrysomelidae" genus="Trachyaphthona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="B93A5A5AFFD8FF98FD469DC83BED9CE2" italics="true" pageId="9" pageNumber="335">Trachyaphthona bidentata</emphasis>
Chen &amp; Wang (1980)
</taxonomicName>
from
<collectingCountry id="F359C6D8FFD8FF98FF109D8D3AA59CC2" box="[144,213,1251,1273]" name="China" pageId="9" pageNumber="335">China</collectingCountry>
exhibits a similar sexual dimorphism. A laterally compressed aedeagus, with a transverse preapical depression in lateral view, is another unique feature of this species.
<taxonomicName id="4C4EFDCBFFD8FF98FE179C51381A9D6F" authority="Prathapan &amp; Viraktamath, 2009" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" box="[407,618,1343,1364]" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="species" species="anupama" status="sp. nov.">
<emphasis id="B93A5A5AFFD8FF98FE179C51381A9D6F" box="[407,618,1343,1364]" italics="true" pageId="9" pageNumber="335">Tegyrius anupama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD8FF98FDF29C5138B89D6F" box="[626,712,1343,1364]" pageId="9" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
is a common pest of black pepper,
<taxonomicName id="4C4EFDCBFFD8FF98FE639C3038229D48" box="[483,594,1374,1395]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="9" pageNumber="335" phylum="Tracheophyta" rank="species" species="nigrum">
<emphasis id="B93A5A5AFFD8FF98FE639C3038229D48" box="[483,594,1374,1395]" italics="true" pageId="9" pageNumber="335">P. nigrum</emphasis>
</taxonomicName>
, in
<collectingCountry id="F359C6D8FFD8FF98FDFD9C3338829D48" box="[637,754,1373,1395]" name="Sri Lanka" pageId="9" pageNumber="335">
Sri
<taxonomicName id="4C4EFDCBFFD8FF98FD269C3338829D48" box="[678,754,1373,1395]" class="Insecta" family="Chrysomelidae" genus="Lanka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="358" phylum="Arthropoda" rank="genus">Lanka</taxonomicName>
</collectingCountry>
, feeding on the underside of mature leaves as
<taxonomicName id="4C4EFDCBFFD8FF98FD2C9C123A9C9D94" authorityName="Prathapan &amp; Viraktamath" authorityYear="2009" baseAuthorityName="Doeberl" baseAuthorityYear="2003" class="Insecta" family="Chrysomelidae" genus="Tegyrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="335" phylum="Arthropoda" rank="species" species="keralaensis" status="comb. nov.">
<emphasis id="B93A5A5AFFD8FF98FD2C9C123A9C9D94" italics="true" pageId="9" pageNumber="335">T. keralaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209E721FFD8FF98FF769CF43B1F9D8B" box="[246,367,1434,1456]" pageId="9" pageNumber="335" rank="species">comb. nov.</taxonomicNameLabel>
does in southern
<collectingCountry id="F359C6D8FFD8FF98FDC49CF438F49D8B" box="[580,644,1434,1456]" name="India" pageId="9" pageNumber="335">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>