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<document id="AC0312A286D8733F40425B901980D1F8" ID-DOI="10.5281/zenodo.190536" ID-GBIF-Dataset="d7ff8abd-bb96-4ea4-b61a-66aa70982a9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="190536" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461114760079" checkinUser="plazi" docAuthor="Pesic, Vladimir &amp; Smit, Harry" docDate="2009" docId="03DD801AE54DFFB8FF43D25AFCD8FE5D" docLanguage="en" docName="zt02240p030.pdf" docOrigin="Zootaxa 2240" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractides (Atractides) putihi Wiles 1991" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FFE4F862E555FFA1FFD4D17BFF87FFBA" masterDocTitle="New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="25" updateTime="1698594892522" updateUser="plazi">
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<mods:title id="82A430CBC4DFA5A7A4E3787C5553D45C">New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species</mods:title>
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<mods:namePart id="DFC77CDBB3B17A0CE363EF07A8EB614F">Pesic, Vladimir</mods:namePart>
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<mods:namePart id="F46F0B9CC8DC459CC67B112A749EB5FA">Smit, Harry</mods:namePart>
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<mods:date id="9D8E83C6F26947479514744FD2722462">2009</mods:date>
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<subSubSection id="C36E6287E54DFFB9FF43D25AFE22FCE4" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8BCB310CE54DFFB9FF43D25AFD0CFC80" blockId="24.[151,651,800,862]" box="[151,651,800,827]" pageId="24" pageNumber="25">
<heading id="D0838660E54DFFB9FF43D25AFD0CFC80" bold="true" box="[151,651,800,827]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<taxonomicName id="4C744A8FE54DFFB9FF43D25AFD0CFC80" authority="Wiles, 1991" authorityName="Wiles" authorityYear="1991" box="[151,651,800,827]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="putihi" subGenus="Atractides">
<emphasis id="B900ED1EE54DFFB9FF43D25AFD0CFC80" bold="true" box="[151,651,800,827]" pageId="24" pageNumber="25">
<emphasis id="B900ED1EE54DFFB9FF43D25AFE96FC81" bold="true" box="[151,273,801,827]" italics="true" pageId="24" pageNumber="25">Atractides</emphasis>
(
<emphasis id="B900ED1EE54DFFB9FEF6D25AFE1BFC81" bold="true" box="[290,412,801,827]" italics="true" pageId="24" pageNumber="25">Atractides</emphasis>
)
<emphasis id="B900ED1EE54DFFB9FE79D25AFE72FC81" bold="true" box="[429,501,801,827]" italics="true" pageId="24" pageNumber="25">putihi</emphasis>
<bibRefCitation id="EFE54CFDE54DFFB9FE28D25BFD0CFC80" author="Wiles" box="[508,651,800,826]" pageId="24" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF43D23FFE22FCE4" blockId="24.[151,651,800,862]" box="[151,421,836,862]" pageId="24" pageNumber="25">
(
<figureCitation id="134F2D89E54DFFB9FF4BD23FFEBDFCE4" box="[159,314,836,862]" captionStart="FIGURE 18 A D" captionStartId="25.[151,255,1883,1907]" captionTargetBox="[187,1421,562,1827]" captionTargetId="figure@25.[151,1436,542,1853]" captionTargetPageId="25" captionText="FIGURE 18 A D. Atractides putihi Wiles, 1991, male, from unnamed creek crossing track from watchtower, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190554/files/figure.png" pageId="24" pageNumber="25">Figs. 18AD</figureCitation>
,
<figureCitation id="134F2D89E54DFFB9FE93D23FFE1AFCE4" box="[327,413,836,862]" captionStart="FIGURE 19 A C" captionStartId="26.[151,258,1700,1724]" captionTargetBox="[171,1416,192,1669]" captionTargetId="figure@26.[151,1436,163,1692]" captionTargetPageId="26" captionText="FIGURE 19 A C. Atractides putihi Wiles, 1991, female, stream near Haewsai Waterfall, Thailand: A = idiosoma, ventral view; B = I-Leg- 5 and - 6; C = IV-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190555/files/figure.png" pageId="24" pageNumber="25">19AC</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E54DFFB9FF43D2EBFA07F89E" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BCB310CE54DFFB9FF43D2EBFC4AFBA5" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
<emphasis id="B900ED1EE54DFFB9FF43D2EBFE83FC10" bold="true" box="[151,260,912,938]" pageId="24" pageNumber="25">Material</emphasis>
.
<collectingCountry id="F363719CE54DFFB9FEC6D2EAFE29FC11" box="[274,430,913,939]" name="Thailand" pageId="24" pageNumber="25">THAILAND</collectingCountry>
: unnamed creek crossing track from watchtower, Khao Yai NP
<date id="FFCA17CCE54DFFB9FB45D2EAFA97FC11" box="[1169,1296,913,939]" pageId="24" pageNumber="25" value="2007-11-11">11.xi.2007</date>
, 14°
<geoCoordinate id="EE4057CBE54DFFB9FA9FD2EAFF2BFC68" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="27.533">27.533 N</geoCoordinate>
101°
<geoCoordinate id="EE4057CBE54DFFB9FF20D2C3FEE7FC68" box="[244,352,952,978]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="20.386">20.386 E</geoCoordinate>
, leg. Smit, 1/0/0 (1/0/0 mounted); stream near Haewsai Waterfall, Nam Nao NP, alt.
<quantity id="4C8C9CE9E54DFFB9FA9ED2C3FA1FFC68" box="[1354,1432,952,978]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="24" pageNumber="25" unit="m" value="425.0">425 m</quantity>
,
<date id="FFCA17CCE54DFFB9FF43D2A5FE90FC42" box="[151,279,990,1016]" pageId="24" pageNumber="25" value="2007-11-16">16.xi.2007</date>
, 16°
<geoCoordinate id="EE4057CBE54DFFB9FE86D2A5FE39FC42" box="[338,446,990,1016]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="40.668">40.668 N</geoCoordinate>
101°
<geoCoordinate id="EE4057CBE54DFFB9FDD7D2A5FDE9FC42" box="[515,622,990,1016]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="41.856">41.856 E</geoCoordinate>
, leg. Smit, 0/1/0 (0/1/0 mounted); slow flowing stream, Nam Nao NP, alt.
<quantity id="4C8C9CE9E54DFFB9FF17D57EFE9AFBA5" box="[195,285,1029,1055]" metricMagnitude="3" metricUnit="m" metricValue="1.19" pageId="24" pageNumber="25" unit="m" value="1190.0">1190 m</quantity>
,
<date id="FFCA17CCE54DFFB9FEFDD57EFE2EFBA5" box="[297,425,1029,1055]" pageId="24" pageNumber="25" value="2007-11-15">15.xi.2007</date>
, 15°
<geoCoordinate id="EE4057CBE54DFFB9FE31D57EFDD5FBA5" box="[485,594,1029,1055]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="14.442">14.442 N</geoCoordinate>
101°
<geoCoordinate id="EE4057CBE54DFFB9FD4DD57EFC83FBA5" box="[665,772,1029,1055]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="34.774">34.774 E</geoCoordinate>
, leg. Smit, 0/1/0.
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF12D550FBF4FABD" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
<emphasis id="B900ED1EE54DFFB9FF12D550FEE0FBFF" bold="true" box="[198,359,1067,1093]" pageId="24" pageNumber="25">Morphology.</emphasis>
General features. Integument dorsally striated. Muscle attachment plates unsclerotized. Coxal field with the medioposterior margin of Cx-1+2 sharply curved, lateral and posterior margins of Cx- 3+4 with extended secondary sclerotization, distinctly projecting posteriorly to lateral margins. Excretory pore smooth; Vgl-1 separated from Vgl-2. Palp with a weak sexual dimorphism, P-4 with a strong sword seta, inserting proximally from proximoventral hair; I-Leg: setae S-1 and S-2 similar in length, close to each other; IV-Leg-5 (
<figureCitation id="134F2D89E54DFFB9FEC7D596FEFBFABD" box="[275,380,1261,1287]" captionStart="FIGURE 19 A C" captionStartId="26.[151,258,1700,1724]" captionTargetBox="[171,1416,192,1669]" captionTargetId="figure@26.[151,1436,163,1692]" captionTargetPageId="26" captionText="FIGURE 19 A C. Atractides putihi Wiles, 1991, female, stream near Haewsai Waterfall, Thailand: A = idiosoma, ventral view; B = I-Leg- 5 and - 6; C = IV-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190555/files/figure.png" pageId="24" pageNumber="25">Fig. 19C</figureCitation>
) with two swimming setae at distal and nearly long as IV-Leg-6.
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF11D46FFF3BFA72" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
Male (from unnamed creek crossing track from watchtower,
<collectingCountry id="F363719CE54DFFB9FC71D46FFB91FA94" box="[933,1046,1300,1326]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
): Idiosoma L/W 431/309. Coxal field (
<figureCitation id="134F2D89E54DFFB9FF0AD441FECAFAEE" box="[222,333,1338,1364]" captionStart="FIGURE 18 A D" captionStartId="25.[151,255,1883,1907]" captionTargetBox="[187,1421,562,1827]" captionTargetId="figure@25.[151,1436,542,1853]" captionTargetPageId="25" captionText="FIGURE 18 A D. Atractides putihi Wiles, 1991, male, from unnamed creek crossing track from watchtower, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190554/files/figure.png" pageId="24" pageNumber="25">Fig. 18A</figureCitation>
): L 234, Cx-1+2 mL 112, Cx-3 W 242, Cx-1+2 lL 178. Palp (
<figureCitation id="134F2D89E54DFFB9FBE6D441FB27FAEE" box="[1074,1184,1338,1364]" captionStart="FIGURE 18 A D" captionStartId="25.[151,255,1883,1907]" captionTargetBox="[187,1421,562,1827]" captionTargetId="figure@25.[151,1436,542,1853]" captionTargetPageId="25" captionText="FIGURE 18 A D. Atractides putihi Wiles, 1991, male, from unnamed creek crossing track from watchtower, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190554/files/figure.png" pageId="24" pageNumber="25">Fig. 18C</figureCitation>
): total L 195, dL and %L (in parentheses): P-1 18 (9.2), P-2 49 (25.1), P-3 34 (17.4), P-4 68 (34.9), P-5 26 (13.3); L P-2/P-4 0.72. Genital field (
<figureCitation id="134F2D89E54DFFB9FEEED4F3FE21FA18" box="[314,422,1416,1442]" captionStart="FIGURE 18 A D" captionStartId="25.[151,255,1883,1907]" captionTargetBox="[187,1421,562,1827]" captionTargetId="figure@25.[151,1436,542,1853]" captionTargetPageId="25" captionText="FIGURE 18 A D. Atractides putihi Wiles, 1991, male, from unnamed creek crossing track from watchtower, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190554/files/figure.png" pageId="24" pageNumber="25">Fig. 18B</figureCitation>
): with nearly straight or slightly convex anterior margin, L/W 65/86, L Ac-1-3: 21-21- 27.
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF11D4AEFAF9F9AC" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
I-Leg (
<figureCitation id="134F2D89E54DFFB9FEC2D4AEFE02FA55" box="[278,389,1493,1519]" captionStart="FIGURE 18 A D" captionStartId="25.[151,255,1883,1907]" captionTargetBox="[187,1421,562,1827]" captionTargetId="figure@25.[151,1436,542,1853]" captionTargetPageId="25" captionText="FIGURE 18 A D. Atractides putihi Wiles, 1991, male, from unnamed creek crossing track from watchtower, Thailand: A = coxal and genital field; B = genital field; C = palp, medial view; D = I-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190554/files/figure.png" pageId="24" pageNumber="25">Fig. 18D</figureCitation>
): I-Leg-5 dL 94, vL 73, dL/vL 1.29, HB 33, dL/HB 2.9, S-1 L 46, L/
<geoCoordinate id="EE4057CBE54DFFB9FB67D4AEFB79FA55" box="[1203,1278,1493,1519]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5555" value="-6.8">W 6.8</geoCoordinate>
, S-2 L 46, L/
<geoCoordinate id="EE4057CBE54DFFB9FF43D487FF66F9AC" box="[151,225,1532,1558]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5555" value="-6.1">W 6.1</geoCoordinate>
, distance of sword setae at I-Leg-5 6, L S-1/2 1.0; I-Leg-6 L 57, HB 11, L/HB 5.0; L I-Leg-5/6 1.65.
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF11D759FA04F96D" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
Female (from stream near Haewsai Waterfall,
<collectingCountry id="F363719CE54DFFB9FD33D759FCD1F986" box="[743,854,1570,1596]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
): Idiosoma L/W 480/359. Coxal field (
<figureCitation id="134F2D89E54DFFB9FAF4D759FA08F986" box="[1312,1423,1570,1596]" captionStart="FIGURE 19 A C" captionStartId="26.[151,258,1700,1724]" captionTargetBox="[171,1416,192,1669]" captionTargetId="figure@26.[151,1436,163,1692]" captionTargetPageId="26" captionText="FIGURE 19 A C. Atractides putihi Wiles, 1991, female, stream near Haewsai Waterfall, Thailand: A = idiosoma, ventral view; B = I-Leg- 5 and - 6; C = IV-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190555/files/figure.png" pageId="24" pageNumber="25">Fig. 19A</figureCitation>
): L 221, Cx-3 W 245, Cx-1+2 mL 85, Cx-1+2 lL 156. Palp: total L 210, dL and %L (in parentheses): P-1 21 (10.0), P-2 52 (24.8), P-3 36 (17.1), P-4 70 (33.3), P-5 31 (14.8); L P-2/P-4 0.74; capitulum ventral L 84. Genital field (
<figureCitation id="134F2D89E54DFFB9FEE9D7EDFE2AF90A" box="[317,429,1686,1712]" captionStart="FIGURE 19 A C" captionStartId="26.[151,258,1700,1724]" captionTargetBox="[171,1416,192,1669]" captionTargetId="figure@26.[151,1436,163,1692]" captionTargetPageId="26" captionText="FIGURE 19 A C. Atractides putihi Wiles, 1991, female, stream near Haewsai Waterfall, Thailand: A = idiosoma, ventral view; B = I-Leg- 5 and - 6; C = IV-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190555/files/figure.png" pageId="24" pageNumber="25">Fig. 19A</figureCitation>
): gonopore long, genital plates in a right angle, restricted to the level of the postgen, Ac in triangular position, L/W 111/113, genital plate L 52, L Ac-1-3: 22-21-21; egg diameter (n=2) 97-100.
</paragraph>
<paragraph id="8BCB310CE54DFFB9FF11D79FFA07F89E" blockId="24.[151,1437,912,2022]" pageId="24" pageNumber="25">
I-Leg (
<figureCitation id="134F2D89E54DFFB9FEC2D79FFE06F944" box="[278,385,1764,1790]" captionStart="FIGURE 19 A C" captionStartId="26.[151,258,1700,1724]" captionTargetBox="[171,1416,192,1669]" captionTargetId="figure@26.[151,1436,163,1692]" captionTargetPageId="26" captionText="FIGURE 19 A C. Atractides putihi Wiles, 1991, female, stream near Haewsai Waterfall, Thailand: A = idiosoma, ventral view; B = I-Leg- 5 and - 6; C = IV-Leg- 5 and - 6. Scale bars = 100 μm." httpUri="https://zenodo.org/record/190555/files/figure.png" pageId="24" pageNumber="25">Fig. 19B</figureCitation>
): I-Leg-5 dL 92, vL 77, dL/vL 1.2, HB 32, dL/HB 2.9, S-1 L 45, L/
<geoCoordinate id="EE4057CBE54DFFB9FB4ED79FFB63F944" box="[1178,1252,1764,1790]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5555" value="-7.3">W 7.3</geoCoordinate>
, S-2 L 44, L/
<geoCoordinate id="EE4057CBE54DFFB9FA54D79FFF38F89E" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="5555" value="-6.0">W 6.0</geoCoordinate>
, distance of sword setae at I-Leg-5 5.0, L S-1/2 1.02; I-Leg-6 L 59, HB 11, L/HB 5.2; L I-Leg-5/6 1.56.
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E54DFFB8FF12D64BFBB8FE7A" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BCB310CE54DFFB8FF12D64BFBB8FE7A" blockId="24.[151,1437,912,2022]" lastBlockId="25.[151,1437,152,487]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B900ED1EE54DFFB9FF12D64BFEBFF8F0" bold="true" box="[198,312,1840,1866]" pageId="24" pageNumber="25">Remarks</emphasis>
. In the original description
<bibRefCitation id="EFE54CFDE54DFFB9FD42D64AFCB2F8F1" author="Wiles" box="[662,821,1841,1867]" pageId="24" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles (1991)</bibRefCitation>
mentioned the similarity with
<taxonomicName id="4C744A8FE54DFFB9FB61D64AFF50F8C8" authority="Cook" authorityName="Cook" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B900ED1EE54DFFB9FB61D64AFA1BF8F0" box="[1205,1436,1841,1866]" italics="true" pageId="24" pageNumber="25">Atractides diversus</emphasis>
Cook
</taxonomicName>
and
<taxonomicName id="4C744A8FE54DFFB9FEDBD623FE64F8C8" authority="Walter" authorityName="Walter" box="[271,483,1880,1906]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B900ED1EE54DFFB9FEDBD623FE0AF8CB" box="[271,397,1880,1905]" italics="true" pageId="24" pageNumber="25">A. minutus</emphasis>
Walter
</taxonomicName>
, two polyacetabulate members of the subgenus
<taxonomicName id="4C744A8FE54DFFB9FBC0D623FB35F8CB" box="[1044,1202,1880,1905]" class="Arachnida" family="Hygrobatidae" genus="Polymegapus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B900ED1EE54DFFB9FBC0D623FB35F8CB" box="[1044,1202,1880,1905]" italics="true" pageId="24" pageNumber="25">Polymegapus</emphasis>
</taxonomicName>
, known from
<collectingCountry id="F363719CE54DFFB9FA8CD623FA1FF8C8" box="[1368,1432,1880,1906]" name="India" pageId="24" pageNumber="25">India</collectingCountry>
.
<taxonomicName id="4C744A8FE54DFFB9FF43D604FEDDF822" box="[151,346,1919,1944]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B900ED1EE54DFFB9FF43D604FEDDF822" box="[151,346,1919,1944]" italics="true" pageId="24" pageNumber="25">Atractides putihi</emphasis>
</taxonomicName>
resemble the species of the later subgenus in the shape of the palp (sword seta on P-4 fingerlike, inserting proximal to proximoventral hair), but differs in the following features (in parentheses diagnostic features of the subgenus
<taxonomicName id="4C744A8FE54DFFB9FD97D6B7FD64F85F" box="[579,739,1996,2021]" class="Arachnida" family="Hygrobatidae" genus="Polymegapus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B900ED1EE54DFFB9FD97D6B7FD64F85F" box="[579,739,1996,2021]" italics="true" pageId="24" pageNumber="25">Polymegapus</emphasis>
</taxonomicName>
, from
<bibRefCitation id="EFE54CFDE54DFFB9FCE0D6B7FC58F85C" author="Gerecke" box="[820,991,1996,2022]" pageId="24" pageNumber="30" refString="Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke 2003</bibRefCitation>
): I-Leg-5 with ventral margin shorter than dorsal margin (the ventrodistal edge of I-Leg-5 extends clearly over the dorsodistal one), S-1 not extremely inflated (S-1 strongly enlarged in the proximal part, distally narrowed to a rather fine trip) and idiosoma with unsclerotized muscle attachment plates (idiosoma with strong sclerification due to sclerotized muscle attachments). Its possible that
<taxonomicName id="4C744A8FE54CFFB8FDBED077FCB1FE9F" box="[618,822,268,293]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B900ED1EE54CFFB8FDBED077FCB1FE9F" box="[618,822,268,293]" italics="true" pageId="25" pageNumber="26">Atractides putihi</emphasis>
</taxonomicName>
within
<taxonomicName id="4C744A8FE54CFFB8FC41D077FB95FE9F" box="[917,1042,268,293]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B900ED1EE54CFFB8FC41D077FB95FE9F" box="[917,1042,268,293]" italics="true" pageId="25" pageNumber="26">Atractides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2335065E54CFFB8FBCED077FBD4FE9C" box="[1050,1107,268,294]" pageId="25" pageNumber="26" sensu="stricto">s.str.</taxonomicNameLabel>
represents an intermediate species on the bridge to
<emphasis id="B900ED1EE54CFFB8FE1CD049FDE8FEF1" box="[456,623,306,331]" italics="true" pageId="25" pageNumber="26">Polymagapus</emphasis>
. To address this interesting question, additional studies on larval morphology and the application of molecular genetics techniques are necessery. Only the female of this species is known (
<bibRefCitation id="EFE54CFDE54CFFB8FEB9D0FBFE70FE20" author="Wiles" box="[365,503,384,410]" pageId="25" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles 1991</bibRefCitation>
), and this is the first description of the male. According to
<bibRefCitation id="EFE54CFDE54CFFB8FB7DD0FBFAC5FE20" author="Wiles" box="[1193,1346,384,410]" pageId="25" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles (1991)</bibRefCitation>
several specimens of this species, including a male, have been collected from Sulawesi.
</paragraph>
</subSubSection>
<subSubSection id="C36E6287E54CFFB8FF11D0B6FCD8FE5D" box="[197,863,461,487]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BCB310CE54CFFB8FF11D0B6FCD8FE5D" blockId="25.[151,1437,152,487]" box="[197,863,461,487]" pageId="25" pageNumber="26">
<emphasis id="B900ED1EE54CFFB8FF11D0B6FED9FE5D" bold="true" box="[197,350,461,487]" pageId="25" pageNumber="26">Distribution</emphasis>
.
<collectingCountry id="F363719CE54CFFB8FEB9D0B6FE5FFE5D" box="[365,472,461,487]" name="Malaysia" pageId="25" pageNumber="26">Malaysia</collectingCountry>
(
<bibRefCitation id="EFE54CFDE54CFFB8FE33D0B6FDF5FE5D" author="Wiles" box="[487,626,461,487]" pageId="25" pageNumber="30" refString="Wiles, P. R. (1991) Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia. Acarologia, 32, 41 - 56." type="journal article" year="1991">Wiles 1991</bibRefCitation>
). New for
<collectingCountry id="F363719CE54CFFB8FD3BD0B6FCDCFE5D" box="[751,859,461,487]" name="Thailand" pageId="25" pageNumber="26">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>