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<document ID-DOI="10.1093/isd/ixy015" ID-ISSN="2399-3421" ID-Zenodo-Dep="7168382" approvalRequired="4" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="3" checkinTime="1665267159722" checkinUser="felipe" docAuthor="Mikó, István, Trietsch, Carolyn, Kamp, Thomas van de, Masner, Lubomír, Ulmer, Jonah M., Yoder, Matthew Jon, Zuber, Marcus, Sandall, Emily L., Baumbach, Tilo &amp; Deans, Andrew R." docDate="2018" docId="03DD3922C55CC72CFCA104ADFEC9F9DE" docLanguage="en" docName="InsectSystDivers.2.6.4.1-29.pdf" docOrigin="Insect Systematics and Diversity (AIFB) 2 (6), No. 4" docSource="http://dx.doi.org/10.1093/isd/ixy015" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.3:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="3" docTitle="Trassedia yanegai Miko and Trietsch 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="FFE4415AC54AC73BFF93014AFFEFFF8E" masterDocTitle="Revision of Trassedia (Hymenoptera: Ceraphronidae), an Evolutionary Relict With an Unusual Distribution" masterLastPageNumber="29" masterPageNumber="1" pageNumber="23" updateTime="1665277179783" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Revision of Trassedia (Hymenoptera: Ceraphronidae), an Evolutionary Relict With an Unusual Distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mikó, István</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
<mods:nameIdentifier type="email">istvan.miko@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Trietsch, Carolyn</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kamp, Thomas van de</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Masner, Lubomír</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Central Experimental Farm, Ottawa, ON, Canada,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ulmer, Jonah M.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoder, Matthew Jon</mods:namePart>
<mods:affiliation>Illinois Natural History Survey, University of Illinois at Urbana-Champaign, Champaign, IL,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zuber, Marcus</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sandall, Emily L.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumbach, Tilo</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deans, Andrew R.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
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<mods:date>2018</mods:date>
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<mods:title>AIFB</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2018-12-04</mods:number>
</mods:detail>
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<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
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<mods:number>4</mods:number>
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<mods:identifier type="DOI">10.1093/isd/ixy015</mods:identifier>
<mods:identifier type="ISSN">2399-3421</mods:identifier>
<mods:identifier type="Zenodo-Dep">7168382</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DD3922C55CC72CFCA104ADFEC9F9DE" httpUri="http://treatment.plazi.org/id/03DD3922C55CC72CFCA104ADFEC9F9DE" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection pageId="22" pageNumber="23" type="nomenclature">
<paragraph blockId="22.[818,1317,1511,1534]" box="[818,1317,1511,1534]" pageId="22" pageNumber="23">
<heading box="[818,1317,1511,1534]" fontSize="9" level="1" pageId="22" pageNumber="23" reason="6">
<taxonomicName authority="Miko and Trietsch" authorityName="Miko and Trietsch" authorityYear="2018" box="[818,1222,1511,1534]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="yanegai" status="sp. nov.">
<emphasis box="[818,1020,1511,1534]" italics="true" pageId="22" pageNumber="23">Trassedia yanegai</emphasis>
Mikó and Trietsch
</taxonomicName>
<taxonomicNameLabel box="[1229,1317,1511,1534]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="22.[818,1443,1545,1595]" pageId="22" pageNumber="23">
(Zoobank LSID:
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urn:lsid:zoobank. org:act:
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)
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="diagnosis">
<paragraph blockId="22.[818,1443,1639,1952]" box="[818,908,1639,1659]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[818,908,1639,1659]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="22.[818,1443,1639,1952]" lastBlockId="23.[146,770,148,227]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<taxonomicName authorityName="Miko and Trietsch" authorityYear="2018" box="[818,979,1668,1688]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="yanegai">
<emphasis box="[818,979,1668,1688]" italics="true" pageId="22" pageNumber="23">Trassedia yanegai</emphasis>
</taxonomicName>
shares the short setae on the dorsal region of the cranium (shorter than the diameter of the median ocellus;
<figureCitation captionStart="Fig" captionStartId="4.[146,173,1312,1329]" captionTargetBox="[314,1274,147,1277]" captionTargetId="figure-323@4.[314,1274,147,1277]" captionTargetPageId="4" captionText="Fig. 3. Brightfield images of Trassedia species. (A) Female antenna of Trassedia brasiliensis Masner and Mikó sp. nov. (OSUC 372988). (B) Male antenna of Trassedia nigrorufus Mikó and Masner sp. nov. (NCSU 53672). (C) Head of Trassedia yanegai Mikó and Trietsch sp. nov., anterior view (UCRCENT 392055). (D) Head of Trassedia luapi Cancemi 1996, anterior view (CASENT 2044510). (E) Head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., dorsal view (UCRC_ENT 392055). (F) Head and mesosoma of Trassedia luapi Cancemi 1996, dorsal view (CASENT 2002185) (aal = antero-admedian line, io = inner orbit, not = notaulus, pof = preocellar furrow, pro = pronope, ver = vertex)." figureDoi="http://doi.org/10.5281/zenodo.7168388" httpUri="https://zenodo.org/record/7168388/files/figure.png" pageId="22" pageNumber="23">Fig. 3C and D</figureCitation>
), the reduced anterior ocellar fovea that does not extend to the antennal scrobe (
<figureCitation box="[1010,1142,1756,1776]" captionStart="Fig" captionStartId="4.[146,173,1312,1329]" captionTargetBox="[314,1274,147,1277]" captionTargetId="figure-323@4.[314,1274,147,1277]" captionTargetPageId="4" captionText="Fig. 3. Brightfield images of Trassedia species. (A) Female antenna of Trassedia brasiliensis Masner and Mikó sp. nov. (OSUC 372988). (B) Male antenna of Trassedia nigrorufus Mikó and Masner sp. nov. (NCSU 53672). (C) Head of Trassedia yanegai Mikó and Trietsch sp. nov., anterior view (UCRCENT 392055). (D) Head of Trassedia luapi Cancemi 1996, anterior view (CASENT 2044510). (E) Head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., dorsal view (UCRC_ENT 392055). (F) Head and mesosoma of Trassedia luapi Cancemi 1996, dorsal view (CASENT 2002185) (aal = antero-admedian line, io = inner orbit, not = notaulus, pof = preocellar furrow, pro = pronope, ver = vertex)." figureDoi="http://doi.org/10.5281/zenodo.7168388" httpUri="https://zenodo.org/record/7168388/files/figure.png" pageId="22" pageNumber="23">Fig. 3C and D</figureCitation>
), the punctate ocular impression along the inner orbit (io:
<figureCitation box="[1065,1204,1785,1805]" captionStart="Fig" captionStartId="4.[146,173,1312,1329]" captionTargetBox="[314,1274,147,1277]" captionTargetId="figure-323@4.[314,1274,147,1277]" captionTargetPageId="4" captionText="Fig. 3. Brightfield images of Trassedia species. (A) Female antenna of Trassedia brasiliensis Masner and Mikó sp. nov. (OSUC 372988). (B) Male antenna of Trassedia nigrorufus Mikó and Masner sp. nov. (NCSU 53672). (C) Head of Trassedia yanegai Mikó and Trietsch sp. nov., anterior view (UCRCENT 392055). (D) Head of Trassedia luapi Cancemi 1996, anterior view (CASENT 2044510). (E) Head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., dorsal view (UCRC_ENT 392055). (F) Head and mesosoma of Trassedia luapi Cancemi 1996, dorsal view (CASENT 2002185) (aal = antero-admedian line, io = inner orbit, not = notaulus, pof = preocellar furrow, pro = pronope, ver = vertex)." figureDoi="http://doi.org/10.5281/zenodo.7168388" httpUri="https://zenodo.org/record/7168388/files/figure.png" pageId="22" pageNumber="23">Fig. 3C and D</figureCitation>
) and a wide Waterstons evaporatorium that reaches the lateral 13 of the tergite (wop:
<figureCitation box="[818,949,1844,1864]" captionStart="Fig" captionStartId="5.[146,173,1433,1450]" captionTargetBox="[314,1274,154,1398]" captionTargetId="figure-234@5.[314,1274,147,1398]" captionTargetPageId="5" captionText="Fig. 4. Brightfield images of Trassedia species. (A, B) Metasoma of Trassedia yanegai Mikó andTrietsch sp. nov. showing Waterstons evaporatoria, dorsal view (UCRCENT 392055). (C) Head of Trassedia brasiliensis Masner and Mikó sp. nov., anterolateral view (OSUC 372988). (D) Head and mesosoma of Trassedia angustioculus Mikó and Masner sp. nov., anterolateral view (PSUC_FEM 17569). (E) Head and mesosoma of Trassedia angustioculus Mikó and Masner sp. nov., lateral view (PSUC_FEM 17569).(F) Head and mesosoma of Trassedia guianensis Mikó and Masner sp.nov., dorsal view (PSUC_FEM 96179) (aal = anteroadmedian line, ala = anterolateral edge of mesoscutum, io = inner orbit, not = notaulus, pcs = anterior ocellar fovea, pof = preoccipital furrow, sc = antennal scrobe, wop = Waterstons evaporatorium)." figureDoi="http://doi.org/10.5281/zenodo.7168390" httpUri="https://zenodo.org/record/7168390/files/figure.png" pageId="22" pageNumber="23">Fig. 4A and B</figureCitation>
) with
<taxonomicName authorityName="Cancemi" authorityYear="1996" box="[1011,1082,1844,1864]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="luapi">
<emphasis box="[1011,1082,1844,1864]" italics="true" pageId="22" pageNumber="23">T. luapi</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[1131,1268,1844,1864]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis box="[1131,1268,1844,1864]" italics="true" pageId="22" pageNumber="23">T. australiensis</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Miko and Trietsch" authorityYear="2018" box="[1280,1441,1844,1864]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="yanegai">
<emphasis box="[1280,1441,1844,1864]" italics="true" pageId="22" pageNumber="23">Trassedia yanegai</emphasis>
</taxonomicName>
differs from these species in the lack of the notaulus, the lack of scutes on the vertex, the anteriorly gradually widening preoccipital sulcus (pof:
<figureCitation box="[929,997,1932,1952]" captionStart="Fig" captionStartId="4.[146,173,1312,1329]" captionTargetBox="[314,1274,147,1277]" captionTargetId="figure-323@4.[314,1274,147,1277]" captionTargetPageId="4" captionText="Fig. 3. Brightfield images of Trassedia species. (A) Female antenna of Trassedia brasiliensis Masner and Mikó sp. nov. (OSUC 372988). (B) Male antenna of Trassedia nigrorufus Mikó and Masner sp. nov. (NCSU 53672). (C) Head of Trassedia yanegai Mikó and Trietsch sp. nov., anterior view (UCRCENT 392055). (D) Head of Trassedia luapi Cancemi 1996, anterior view (CASENT 2044510). (E) Head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., dorsal view (UCRC_ENT 392055). (F) Head and mesosoma of Trassedia luapi Cancemi 1996, dorsal view (CASENT 2002185) (aal = antero-admedian line, io = inner orbit, not = notaulus, pof = preocellar furrow, pro = pronope, ver = vertex)." figureDoi="http://doi.org/10.5281/zenodo.7168388" httpUri="https://zenodo.org/record/7168388/files/figure.png" pageId="22" pageNumber="23">Fig. 3C</figureCitation>
), the foveolate sternaulus (ster:
<figureCitation box="[1292,1359,1932,1952]" captionStart="Fig" captionStartId="7.[146,173,1316,1333]" captionTargetBox="[314,1274,152,1281]" captionTargetId="figure-334@7.[314,1274,147,1281]" captionTargetPageId="7" captionText="Fig. 5. Brightfield images of Trassedia species. (A, B) Female metasoma and Waterstons evaporatoria of Trassedia brasiliensis Masner and Mikó sp.nov. dorsal view (OSUC 372988). (C) Female head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., lateral view (UCRCENT 392055). (D) Female antenna of Trassedia luapi Cancemi 1996 (NCSU 71196) (ster = sternaulus, wop = Waterstons evaporatorium)." figureDoi="http://doi.org/10.5281/zenodo.7168392" httpUri="https://zenodo.org/record/7168392/files/figure.png" pageId="22" pageNumber="23">Fig. 5C</figureCitation>
) and the large eyes (HW/IOS = 3.7,
<figureCitation box="[399,469,148,168]" captionStart="Fig" captionStartId="4.[146,173,1312,1329]" captionTargetBox="[314,1274,147,1277]" captionTargetId="figure-323@4.[314,1274,147,1277]" captionTargetPageId="4" captionText="Fig. 3. Brightfield images of Trassedia species. (A) Female antenna of Trassedia brasiliensis Masner and Mikó sp. nov. (OSUC 372988). (B) Male antenna of Trassedia nigrorufus Mikó and Masner sp. nov. (NCSU 53672). (C) Head of Trassedia yanegai Mikó and Trietsch sp. nov., anterior view (UCRCENT 392055). (D) Head of Trassedia luapi Cancemi 1996, anterior view (CASENT 2044510). (E) Head and mesosoma of Trassedia yanegai Mikó andTrietsch sp. nov., dorsal view (UCRC_ENT 392055). (F) Head and mesosoma of Trassedia luapi Cancemi 1996, dorsal view (CASENT 2002185) (aal = antero-admedian line, io = inner orbit, not = notaulus, pof = preocellar furrow, pro = pronope, ver = vertex)." figureDoi="http://doi.org/10.5281/zenodo.7168388" httpUri="https://zenodo.org/record/7168388/files/figure.png" pageId="23" pageNumber="24">Fig. 3C</figureCitation>
). The female pedicel and F1 are brown and F2F4 are whitish in
<taxonomicName authorityName="Miko and Trietsch" authorityYear="2018" box="[447,539,178,196]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="yanegai">
<emphasis box="[447,539,178,196]" italics="true" pageId="23" pageNumber="24">T. yanegai</emphasis>
</taxonomicName>
, while the female pedicel and F1F3 are yellow and F4F9 are dark brown in
<taxonomicName authorityName="Cancemi" authorityYear="1996" box="[628,698,207,227]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="luapi">
<emphasis box="[628,698,207,227]" italics="true" pageId="23" pageNumber="24">T. luapi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="description">
<paragraph blockId="23.[146,771,271,1435]" box="[146,253,271,291]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[146,253,271,291]" pageId="23" pageNumber="24">Description</emphasis>
</paragraph>
<paragraph blockId="23.[146,771,271,1435]" pageId="23" pageNumber="24">
Body length:
<quantity box="[273,361,300,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="23" pageNumber="24" unit="mm" value="3.95">3.95 mm</quantity>
. Color hue pattern female: cranium, mesosomablack, F5F9 black, F2F4 white, scape, legs, pedicel, F1 ochre. Color intensity pattern female: pedicel, F1, fore leg, mesocoxa darker than rest of middle leg, hind leg, scape. Structure of scutes on head and mesosoma: scute surface on head and mesosoma flat, scutes indistinct.
<emphasis bold="true" box="[402,458,447,467]" pageId="23" pageNumber="24">Head:</emphasis>
HW:HH = 1.1. HW/IOS Female: 3.8. Maximum eye diameter versus minimum eye diameter: 1.2. Interommatidial seta length: interommatidial seta length less than facet diameter. Occipital carina medially: continuous medially. Seta length on dorsal region of cranium versus diameter of median ocellus: shorter. Scutes on vertex count: absent. Preoccipital furrow anterior extension: adjacent anteriorly to the posterior margin of the median ocellus. Preoccipital furrow anterior region versus posterior region sculpture: posterior region crenulate, anterior region smooth or finely reticulate. Preoccipital furrow anterior region width versus posterior region width: wider anteriorly than posteriorly. Female OOL: POL: LOL: 0.1:0.5:1.0. Preocellar pit count: absent. Carina delimiting antennal scrobe count: absent. Transverse striation on upper face count: present. Anterior ocellar fovea shape: fovea not extended ventrally into facial sulcus. Supraclypeal depression count: present. Ocular impression sculpture: punctate (fovea of ocellar impression are well separated from each other).
<emphasis bold="true" box="[611,696,916,936]" pageId="23" pageNumber="24">Antenna:</emphasis>
Female scape length versus pedicel length: 2.4. Female F1 length versus pedicel length: 1.1. Female ninth flagellomere length: F9 less than F7+F8.
<emphasis bold="true" box="[146,251,1004,1024]" pageId="23" pageNumber="24">Mesosoma:</emphasis>
Mesosoma shape: not compressed laterally, as wide as high or wider than high. Pronope count: absent. Anterior slope of mesonotum shape: Anterior slope of mesonotum at obtuse angle to dorsal surface of mesonotum in lateral view. Antero-admedian line count: present. Notaulus count: absent. Notaulus anterior origin versus anterolateral angle of mesoscutum (ball-and-socket articulation between pronotum and mesoscutum): Notaulus arises medially of anterolateral angle of the mesoscutum. Scutes on posterior region of mesoscutum and dorsal region of mesoscutellum count: present. Epicnemial carina count: complete. Sternaulus count: present. Sternaulus sculpture: scalloped. Anterior metapleural carina count: present. Carina limiting posteriorly antecosta count: present. Lateral propodeal carina count: present.
<emphasis bold="true" box="[523,587,1356,1376]" pageId="23" pageNumber="24">Wings:</emphasis>
Stigmal vein length versus pterostigma marginal length: stigmal vein longer than the pterostigma marginal length.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="etymology">
<paragraph blockId="23.[146,770,1479,1616]" box="[146,245,1479,1499]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[146,245,1479,1499]" pageId="23" pageNumber="24">Etymology</emphasis>
</paragraph>
<paragraph blockId="23.[146,770,1479,1616]" pageId="23" pageNumber="24">
The species epithet refers to Douglas Yanega (Senior Museum Scientist of the Entomology Research Museum, University of
<collectingRegion box="[146,241,1567,1587]" country="United States of America" name="California" pageId="23" pageNumber="24">California</collectingRegion>
, Riverside), who drew our attention to the
<typeStatus box="[661,742,1567,1587]" pageId="23" pageNumber="24">holotype</typeStatus>
of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>