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<document ID-DOI="10.1093/isd/ixy015" ID-ISSN="2399-3421" approvalRequired="4" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="3" checkinTime="1665267159722" checkinUser="felipe" docAuthor="Mikó, István, Trietsch, Carolyn, Kamp, Thomas van de, Masner, Lubomír, Ulmer, Jonah M., Yoder, Matthew Jon, Zuber, Marcus, Sandall, Emily L., Baumbach, Tilo &amp; Deans, Andrew R." docDate="2018" docId="03DD3922C55BC729FCA107C5FC65FC05" docLanguage="en" docName="InsectSystDivers.2.6.4.1-29.pdf" docOrigin="Insect Systematics and Diversity (AIFB) 2 (6), No. 4" docSource="http://dx.doi.org/10.1093/isd/ixy015" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.3:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="3" docTitle="Trassedia gauldi Miko and Masner 2018, sp. nov." docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="FFE4415AC54AC73BFF93014AFFEFFF8E" masterDocTitle="Revision of Trassedia (Hymenoptera: Ceraphronidae), an Evolutionary Relict With an Unusual Distribution" masterLastPageNumber="29" masterPageNumber="1" pageNumber="18" updateTime="1665271147411" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Revision of Trassedia (Hymenoptera: Ceraphronidae), an Evolutionary Relict With an Unusual Distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mikó, István</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
<mods:nameIdentifier type="email">istvan.miko@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Trietsch, Carolyn</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kamp, Thomas van de</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Masner, Lubomír</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Central Experimental Farm, Ottawa, ON, Canada,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ulmer, Jonah M.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoder, Matthew Jon</mods:namePart>
<mods:affiliation>Illinois Natural History Survey, University of Illinois at Urbana-Champaign, Champaign, IL,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zuber, Marcus</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sandall, Emily L.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumbach, Tilo</mods:namePart>
<mods:affiliation>Laboratory for Applications of Synchrotron Radiation, Karlsruhe Institute of Technology (KIT), Kaiserstr. 12, 76131 Karlsruhe, Germany,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deans, Andrew R.</mods:namePart>
<mods:affiliation>Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA,</mods:affiliation>
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<mods:title>Insect Systematics and Diversity</mods:title>
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<mods:date>2018</mods:date>
<mods:detail type="series">
<mods:title>AIFB</mods:title>
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<mods:detail type="pubDate">
<mods:number>2018-12-04</mods:number>
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<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
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<mods:number>4</mods:number>
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<mods:url>http://dx.doi.org/10.1093/isd/ixy015</mods:url>
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<mods:identifier type="DOI">10.1093/isd/ixy015</mods:identifier>
<mods:identifier type="ISSN">2399-3421</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03DD3922C55BC729FCA107C5FC65FC05" httpUri="http://treatment.plazi.org/id/03DD3922C55BC729FCA107C5FC65FC05" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection pageId="17" pageNumber="18" type="nomenclature">
<paragraph blockId="17.[818,1301,1679,1702]" box="[818,1301,1679,1702]" pageId="17" pageNumber="18">
<heading box="[818,1301,1679,1702]" fontSize="9" level="1" pageId="17" pageNumber="18" reason="6">
<taxonomicName authority="Miko and Masner" authorityName="Miko and Masner" authorityYear="2018" box="[818,1205,1679,1702]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gauldi" status="sp. nov.">
<emphasis box="[818,1001,1679,1702]" italics="true" pageId="17" pageNumber="18">Trassedia gauldi</emphasis>
Mikó and Masner
</taxonomicName>
<taxonomicNameLabel box="[1212,1301,1679,1702]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="17.[818,1441,1714,1763]" pageId="17" pageNumber="18">
(Zoobank LSID:
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urn:lsid:zoobank.org:act:
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)
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph blockId="17.[818,1442,1810,1947]" box="[818,908,1810,1830]" pageId="17" pageNumber="18">
<emphasis bold="true" box="[818,908,1810,1830]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="17.[818,1442,1810,1947]" lastBlockId="18.[146,770,148,549]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[818,968,1839,1859]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[818,968,1839,1859]" italics="true" pageId="17" pageNumber="18">Trassedia gauldi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[1147,1335,1839,1859]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[1147,1335,1839,1859]" italics="true" pageId="17" pageNumber="18">Trassedia guianensis</emphasis>
</taxonomicName>
in the lack of elongate interommatidial setae (in both species the setae are less than 1.5× as long as the diameter of the facets) and the lack of convex scutes on the dorsal cranium and mesosoma. Female specimens differ in POL:OOL ratio (POL as long as OOL in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[617,698,148,168]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[617,698,148,168]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
and 2× as long as OOL in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[326,447,178,196]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[326,447,178,196]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
), the completeness of the occipital carina (the carina is incomplete medially in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[543,662,208,226]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[543,662,208,226]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
), the length of the antennomeres (F1 shorter than the pedicel in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[644,727,236,256]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[644,727,236,256]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
and longer than the pedicel in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[401,523,266,284]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[401,523,266,284]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
) and the body coloration (head and mesosoma black and metasoma brown in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[644,764,296,314]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[644,764,296,314]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
; head, mesosoma, and metasoma brown in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[549,631,324,344]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[549,631,324,344]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
). Males of the two species differ in the length of setae on the flagellomeres (setae shorter than the width of the flagellomeres in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[580,662,383,403]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[580,662,383,403]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
and longer than the width of the flagellomeres in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[489,607,413,431]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[489,607,413,431]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
), POL:OOL ratio (POL shorter than OOL in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[398,479,441,461]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[398,479,441,461]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
and as long as OOL in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
) and the length/width ratio of the gonostyle/volsella complex (gonostyle/volsella complex wider than long in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[603,726,501,519]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis box="[603,726,501,519]" italics="true" pageId="18" pageNumber="19">T. guianensis</emphasis>
</taxonomicName>
and longer than wide in
<taxonomicName authorityName="Miko and Masner" authorityYear="2018" box="[331,411,529,549]" class="Insecta" family="Megaspilidae" genus="Trassedia" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis box="[331,411,529,549]" italics="true" pageId="18" pageNumber="19">T. gauldi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="description">
<paragraph blockId="18.[146,771,588,1957]" box="[146,253,588,608]" pageId="18" pageNumber="19">
<emphasis bold="true" box="[146,253,588,608]" pageId="18" pageNumber="19">Description</emphasis>
</paragraph>
<paragraph blockId="18.[146,771,588,1957]" lastBlockId="18.[818,1443,148,784]" pageId="18" pageNumber="19">
Body length:
<quantity box="[268,406,617,637]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.6" metricValueMin="1.6" pageId="18" pageNumber="19" unit="mm" value="2.1" valueMax="2.6" valueMin="1.6">1.602.60 mm</quantity>
. Color hue pattern female: brown, legs, scape, pedicel, and F1F4 yellowish, forewing brownish with a transverse discolored band at level of stigmal vein. Color intensity pattern female: head lighter than mesosoma and F5F9, metasoma lighter than head, hind femur lighter than the rest of legs. Color hue pattern male: brown, legs yellowish, forewing brownish, with a transverse discolored band at level of stigmal vein. Structure of scutes on head and mesosoma: Scute surface on head and mesosoma flat, scutes indistinct.
<emphasis bold="true" box="[307,357,852,872]" pageId="18" pageNumber="19">Head</emphasis>
: HW:HH = 1.11.2. HW/IOS Female: 2.5 2.7. HW/IOS male: 1.9. Maximum eye diameter versus minimum eye diameter: 1.11.3. Interommatidial seta length: Interommatidial seta length less than facet diameter. Occipital carina medially: continuous medially. Seta length on dorsal region of cranium versus diameter of median ocellus: longer. Scutes on vertex count: absent. Preoccipital furrow anterior extension: extending anteriorly of the anatomical line connecting the two lateral ocelli and not adjacent to anterior ocellus. Preoccipital furrow anterior region versus posterior region sculpture: posterior region crenulate, anterior region smooth or finely reticulate. Preoccipital furrow anterior region width versus posterior region width: wider anteriorly than posteriorly. Female OOL: POL: LOL: 0.40.6:0.50.6:1.0. Male OOL: POL: LOL: 1.4:1.0:1.0. Preocellar pit count: present. Carina delimiting antennal scrobe count: present. Transverse striation on upper face count: absent. Anterior ocellar fovea shape: fovea extended ventrally into facial sulcus, reaching dorsal margin of antennal scrobe. Supraclypeal depression count: present. Ocular impression sculpture: scalloped (foveae composing ocellar impression adjacent, sometimes not separated from each other).
<emphasis bold="true" box="[541,625,1409,1429]" pageId="18" pageNumber="19">Antenna:</emphasis>
Length of setae on male flagellomere versus male flagellomere width: setae shorter than width of flagellomeres. Male scape length versus pedicel length: 1.7. Male F1 length versus pedicel length: 1.2. Male scape length versus combined length of F1+F2: shorter. Male F6 length versus combined length of F7+F8: Shorter than length of flagellomere 7 + 8. Number of flagellomeres with male-specific ventral sensilla: F6-9. Female scape length versus pedicel length: 2.02.6. Female F1 length versus pedicel length: 0.81.0. Female ninth flagellomere length: F9 less than F7+F8.
<emphasis bold="true" box="[300,406,1673,1693]" pageId="18" pageNumber="19">Mesosoma:</emphasis>
Mesosoma shape: compressed laterally, distinctly higher than wide. Pronope count: present. Anterior slope of mesonotum shape: Anterior slope of mesonotum at obtuse angle to dorsal surface of mesonotum in lateral view. Antero-admedian line count: present. Notaulus count: present. Notaulus anterior origin versus anterolateral angle of mesoscutum (ball-and-socket articulation between pronotum and mesoscutum): notaulus arises at anterolateral angle of the mesoscutum. Posterior end of notaulus versus posterior end of antero-admedian line location: anteroadmedian line extends more posteriorly than notaulus. Scutes on posterior region of mesoscutum and dorsal region of mesoscutellum count: absent. Epicnemial carina count: complete. Sternaulus count: present. Sternaulus sculpture: smooth. Anterior metapleural carina count: absent. Carina limiting posteriorly antecosta count: present. Lateral propodeal carina count: present. Wings: Stigmal vein length versus pterostigma marginal length: stigmal vein longer than the pterostigma marginal length. Metasoma: Waterstons evaporatorium shape medially male: paired, left and right evaporatoria are not continuous medially.
<emphasis bold="true" box="[980,1118,383,403]" pageId="18" pageNumber="19">Male genitalia:</emphasis>
Anterior margin shape of male S9: convex. Male S9 distal setal line/setal patch count: distal setae composing transverse setiferous line(s). Distal margin of male S9 shape: concave. Proximolateral corner of male S9 shape: blunt. Gonostyle/ volsella complex length: longer than wide. Gonostyle/volsella complex proximodorsal margin shape: straight or slightly concave. Distal sensilla on gonossiculus versus proximal margin of gonossiculus location: closer to distal margin than to proximal margin of gonossiculus. Distal margin of harpe in lateral view: shape: acute. Lateral setae of harpe count: present. Lateral margin of harpe shape: widest point of harpe is at its articulation site with gonostyle-volsella complex. Dorsomedial cupulo-gonostipal muscles orientation: parallel. Ventral gonostipo-penisvalval muscles site of origin—proximal extension: extends distally on parossiculus.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="etymology">
<paragraph blockId="18.[818,1441,828,907]" box="[818,917,828,848]" pageId="18" pageNumber="19">
<emphasis bold="true" box="[818,917,828,848]" pageId="18" pageNumber="19">Etymology</emphasis>
</paragraph>
<paragraph blockId="18.[818,1441,828,907]" pageId="18" pageNumber="19">
The species epithet refers to Ian David Gauld, the collector of the
<typeStatus box="[818,902,887,907]" pageId="18" pageNumber="19">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>