treatments-xml/data/03/D2/95/03D295485E13551480A4C04A6B9CFA85.xml
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<document id="12D07CAA4A849464D2A75380B15FD095" ID-CLB-Dataset="25012" ID-DOI="10.11646/zootaxa.4691.2.11" ID-GBIF-Dataset="d9b9e6f4-c8af-4b94-818b-72061226085e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3527111" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1572857388071" checkinUser="plazi" docAuthor="Peng, Yunfei, Ji, Lanzhu &amp; Bian, Dongju" docDate="2019" docId="03D295485E13551480A4C04A6B9CFA85" docLanguage="en" docName="zootaxa.4691.2.11.pdf" docOrigin="Zootaxa 4691 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius shengshanensis Peng &amp; Ji &amp; Bian 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="200" masterDocId="FFEBED305E1355178033C5096E00FF8D" masterDocTitle="Description of Laccobius (s. str.) shengshanensis sp. n. from China (Coleoptera: Hydrophilidae)" masterLastPageNumber="200" masterPageNumber="197" pageNumber="197" updateTime="1698761649523" updateUser="ExternalLinkService">
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<mods:title id="6A9FE01C5A4002C358FD43F3080B1700">Description of Laccobius (s. str.) shengshanensis sp. n. from China (Coleoptera: Hydrophilidae)</mods:title>
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<mods:namePart id="B4598C08DBC01073314FFE801921098D">Peng, Yunfei</mods:namePart>
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<mods:namePart id="56D98C31BC94731ED16B3A3C1CBDE9AB">Ji, Lanzhu</mods:namePart>
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<mods:namePart id="BB5486534C46991EA41539C02C2455C7">Bian, Dongju</mods:namePart>
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<mods:date id="E03BEDC11DDB8B4537C79C908942A3AD">2019</mods:date>
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(s. str.)
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locality.
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, Leg. Peng&quot;, &quot;
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&quot; [red printed label]
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labeled: &quot;
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, Leg. Peng&quot;
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;
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,
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<elevation id="0056C36D5E1355178290C3496CE2F9D5" box="[675,738,1600,1624]" metricMagnitude="2" metricUnit="m" metricValue="3.53" pageId="0" pageNumber="197" unit="m" value="353.0">353m</elevation>
</quantity>
,
<date id="FFC5029E5E13551782DEC3486D68F9D4" box="[749,872,1601,1625]" pageId="0" pageNumber="197" value="2017-09-03">
<collectingDate id="EF81FB765E13551782DEC3486D68F9D4" box="[749,872,1601,1625]" pageId="0" pageNumber="197" value="2017-09-03">2017.09.03</collectingDate>
</date>
, Leg. Peng&quot;.
<collectorName id="268E41885E13551783C5C3486A18F9D4" box="[1014,1048,1601,1625]" pageId="0" pageNumber="197">All</collectorName>
<typeStatus id="54C09AFC5E135517842CC3486A85F9D4" box="[1055,1157,1601,1625]" pageId="0" pageNumber="197" type="paratype">paratypes</typeStatus>
are labeled with a respective red printed
<typeStatus id="54C09AFC5E1355178172C36C6F9DF9F0" box="[321,413,1637,1661]" pageId="0" pageNumber="197" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36177D55E13551480F4C3816ADBFD7C" lastPageId="3" lastPageNumber="200" pageId="0" pageNumber="197" type="description">
<paragraph id="8BC4245E5E13551780F4C3816FA0F964" blockId="0.[151,1437,1347,2021]" pageId="0" pageNumber="197">
<emphasis id="B90FF84C5E13551780F4C3816F4BF92D" bold="true" box="[199,331,1672,1696]" pageId="0" pageNumber="197">Description</emphasis>
. Total length
<quantity id="4C8389BB5E13551781D7C3816C65F92D" box="[484,613,1672,1696]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="0" pageNumber="197" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
(
<typeStatus id="54C09AFC5E1355178246C3806CD4F92C" box="[629,724,1673,1697]" pageId="0" pageNumber="197" type="holotype">holotype</typeStatus>
<quantity id="4C8389BB5E13551782EFC3806D34F92C" box="[732,820,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="0" pageNumber="197" unit="mm" value="2.4">2.4 mm</quantity>
); maximum width
<quantity id="4C8389BB5E1355178431C3806A84F92D" box="[1026,1156,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="0" pageNumber="197" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.21.4 mm</quantity>
(
<typeStatus id="54C09AFC5E13551784A7C3806AF3F92C" box="[1172,1267,1673,1697]" pageId="0" pageNumber="197" type="holotype">holotype</typeStatus>
<quantity id="4C8389BB5E13551784C8C3806B53F92C" box="[1275,1363,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="0" pageNumber="197" unit="mm" value="1.3">1.3 mm</quantity>
). Total length / total width ratio = 1.8. Body oval, moderately convex, with broadest width in anterior third of elytra, evenly narrowed posteriad (
<figureCitation id="134038DB5E1355178162C3D86F90F964" box="[337,400,1745,1769]" captionStart="FIGURE 1" captionStartId="1.[151,244,1980,2002]" captionTargetBox="[161,1428,492,1956]" captionTargetId="figure@1.[151,1436,481,1959]" captionTargetPageId="1" captionText="FIGURE 1. Laccobius (s. str.) shengshanensis sp. n. ac: general habitus (a: dorsal view; b: lateral view; c: ventral view). d: meso- and metaventrites, e and f: aedeagus (e: dorsal view; f: lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3527113" httpUri="https://zenodo.org/record/3527113/files/figure.png" pageId="0" pageNumber="197">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BC4245E5E13551680F4C3FC6FE7FE90" blockId="0.[151,1437,1347,2021]" lastBlockId="1.[151,1437,154,466]" lastPageId="1" lastPageNumber="198" pageId="0" pageNumber="197">
<emphasis id="B90FF84C5E13551780F4C3FC6F04F880" bold="true" box="[199,260,1781,1805]" pageId="0" pageNumber="197">Head</emphasis>
. Labrum blackish brown, slightly convex at anterior margin, closely adhering to clypeus; surface densely punctured, anteriorly with stiff white setae. Clypeus and frons shiny black, preocular pale spots completely absent, surface with irregular, sparsely arranged fine punctures, punctures slightly impressed but larger than labral punctures, each puncture with decumbent white seta; microsculpture absent. Clypeus nearly 2.5× as wide as long, deeply emarginated anteriorly; frontoclypeal suture scarcely apparent. Eyes oval, moderately large, occupying 1/3 side margin of head. Inter-ocular distance ca. 3.1x diameters of eye. Ventral surface black, except the brown maxillae. Mentum slightly rugulose with fine punctures. Postmentum smooth, gula granulate. Maxillary palpi yellowish; palpomere 3 inflated, palpomere 4 darker at apex, asymmetrical, straight medially and convex laterally. Labial palpi yellowish. With 8 antennomeres, antennomeres 15 yellowish, antennal club (antennomeres 68) dark brown; antennomere 1 longer than antennomeres 24 combined; antennomere 5 asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, bearing densely arranged setae.
</paragraph>
<paragraph id="8BC4245E5E12551680F4C4236C5FFE5F" blockId="1.[151,1437,154,466]" pageId="1" pageNumber="198">
<emphasis id="B90FF84C5E12551680F4C4236F23FECF" bold="true" box="[199,291,298,322]" pageId="1" pageNumber="198">Thorax.</emphasis>
Pronotum transverse, slightly impressed sublaterally, 0.44× as long as wide; black with yellowish lateral margins; yellow area about 0.5× as wide as posterior margin of eye along base, sometimes yellow edge vaguely defined from black disc; surface densely punctured, without micro-sculpture; punctures coarse and dense. Scutellum shield equilaterally triangular, black, with few punctures. Prosternum black, tectiform, with longitudinal keel. Mesoventrite with longitudinal, anteriorly arrow-shaped keel.
</paragraph>
<caption id="DF0474D65E12551680A4C2B56D00F87B" ID-DOI="http://doi.org/10.5281/zenodo.3527113" ID-Zenodo-Dep="3527113" httpUri="https://zenodo.org/record/3527113/files/figure.png" pageId="1" pageNumber="198" startId="1.[151,244,1980,2002]" targetBox="[161,1428,492,1956]" targetPageId="1">
<paragraph id="8BC4245E5E12551680A4C2B56D00F87B" blockId="1.[151,1437,1980,2038]" pageId="1" pageNumber="198">
<emphasis id="B90FF84C5E12551680A4C2B56F0CF85C" bold="true" box="[151,268,1980,2002]" pageId="1" pageNumber="198">FIGURE 1.</emphasis>
<taxonomicName id="4C7B5FDD5E1255168122C2B56C54F85C" authority="Peng &amp; Ji &amp; Bian, 2019" authorityName="Peng &amp; Ji &amp; Bian" authorityYear="2019" box="[273,596,1980,2002]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="shengshanensis" status="sp. nov.">
<emphasis id="B90FF84C5E1255168122C2B56F74F85C" box="[273,372,1980,2001]" italics="true" pageId="1" pageNumber="198">Laccobius</emphasis>
(s. str.)
<emphasis id="B90FF84C5E125516818CC2B56C54F85C" box="[447,596,1980,2001]" italics="true" pageId="1" pageNumber="198">shengshanensis</emphasis>
</taxonomicName>
<emphasis id="B90FF84C5E125516826AC2B46C8EF85F" bold="true" box="[601,654,1981,2002]" pageId="1" pageNumber="198">
<taxonomicNameLabel id="A23C45375E125516826AC2B46C8EF85F" box="[601,654,1981,2002]" pageId="1" pageNumber="198" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
ac: general habitus (a: dorsal view; b: lateral view; c: ventral view). d: meso- and metaventrites, e and f: aedeagus (e: dorsal view; f: lateral view).
</paragraph>
</caption>
<caption id="DF0474D65E11551580A4C2B56F33F87B" ID-DOI="http://doi.org/10.5281/zenodo.3527115" ID-Zenodo-Dep="3527115" httpUri="https://zenodo.org/record/3527115/files/figure.png" pageId="2" pageNumber="199" startId="2.[151,244,1980,2002]" targetBox="[151,1436,198,1912]" targetPageId="2">
<paragraph id="8BC4245E5E11551580A4C2B56F33F87B" blockId="2.[151,1437,1980,2038]" pageId="2" pageNumber="199">
<emphasis id="B90FF84C5E11551580A4C2B56F11F85C" bold="true" box="[151,273,1980,2002]" pageId="2" pageNumber="199">FIGURE 2:</emphasis>
Type locality of
<taxonomicName id="4C7B5FDD5E11551581F3C2B56D0EF85C" authority="Peng &amp; Ji &amp; Bian, 2019" authorityName="Peng &amp; Ji &amp; Bian" authorityYear="2019" box="[448,782,1980,2002]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="shengshanensis" status="sp. nov.">
<emphasis id="B90FF84C5E11551581F3C2B56C23F85C" box="[448,547,1980,2001]" italics="true" pageId="2" pageNumber="199">Laccobius</emphasis>
(s. str.)
<emphasis id="B90FF84C5E115515824AC2B56D0EF85C" box="[633,782,1980,2001]" italics="true" pageId="2" pageNumber="199">shengshanensis</emphasis>
</taxonomicName>
<emphasis id="B90FF84C5E1155158324C2B46D50F85F" bold="true" box="[791,848,1981,2002]" pageId="2" pageNumber="199">
<taxonomicNameLabel id="A23C45375E1155158324C2B46D50F85F" box="[791,848,1981,2002]" pageId="2" pageNumber="199" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Heilongjiang Province, Heihe County, Shengshan National Nature Reserve.
</paragraph>
</caption>
<paragraph id="8BC4245E5E10551480F4C5906DC7FECC" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C5906F17FF3C" bold="true" box="[199,279,153,177]" pageId="3" pageNumber="200">Elytra.</emphasis>
Black on disc, yellowish along lateral margins, lateral pale stripe slightly widening anteriorly, very narrow at middle, then widening posteriorly, ca. 1.2× as long as wide, Elytral punctation consisting of fine setiferous punctures (some bearing a white seta) arranged in ca. twenty longitudinal rows, all punctures alike, with almost equally broad intervals. Parasutural furrow absent. Microsculpture absent. Epipleura ending at level of metafemora. Metaventrite with decumbent pubescence, which are absent from a small medio-posterior area.
</paragraph>
<paragraph id="8BC4245E5E10551480F4C4446AAAFD94" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C4446F03FEE8" bold="true" box="[199,259,333,357]" pageId="3" pageNumber="200">Legs.</emphasis>
Coxae and trochanters dark brown, femora dark brown except apical part, tibiae and tarsi yellowish brown. Femora with thin punctures, profemora pubescent at base 1/3. Tibiae with ranges of spines more developed on outer margin, and the spines on mesotibiae more developed; metatibiae slightly curved, distinctly longer than pretibiae and mesotibae. Tarsal natatory setae nearly absent. Tarsi with five tarsomeres; protarsomere 5 long, nearly 3× as long as other protarsomeres; mesotarsomere 1 is very short, mesotarsomeres 2 and 5 longer than other mesotarsomeres; metatarsomeres similar to mesotarsomeres. Claws shortly arcuate. Male with basal 2 protarsomeres dilated.
</paragraph>
<paragraph id="8BC4245E5E10551480F4C72C6EDDFDEC" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C72C6F36FDB0" bold="true" box="[199,310,549,573]" pageId="3" pageNumber="200">Abdomen</emphasis>
. Venerates wrinkled, sparsely punctured and pubescent; pubescence denser on ventrites 15 than on ventrite 6.
</paragraph>
<paragraph id="8BC4245E5E10551480F4C7646ADBFD7C" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C7646F35FD08" bold="true" box="[199,309,621,645]" pageId="3" pageNumber="200">Aedeagus</emphasis>
. Total length
<quantity id="4C8389BB5E10551481E3C7646C34FD08" box="[464,564,621,645]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="200" unit="mm" value="0.65">0.65 mm</quantity>
. Parameres nearly 1.8x as long as phallobase. Phallobase 1.2x as long as wide. Median lobe (without apophyses) nearly as long as parameres, slim at basal 2/3, then gradually broad, apices truncate, with one small angle at each side. Parameres gradually narrowing from base to basal 2/3, outer edge almost parallel, then slightly constricted, distal 1/3 extended outwards, inner-edge of apices with an indentation (
<figureCitation id="134038DB5E1055148459C7D06AA5FD7C" box="[1130,1189,729,753]" captionStart="FIGURE 1" captionStartId="1.[151,244,1980,2002]" captionTargetBox="[161,1428,492,1956]" captionTargetId="figure@1.[151,1436,481,1959]" captionTargetPageId="1" captionText="FIGURE 1. Laccobius (s. str.) shengshanensis sp. n. ac: general habitus (a: dorsal view; b: lateral view; c: ventral view). d: meso- and metaventrites, e and f: aedeagus (e: dorsal view; f: lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3527113" httpUri="https://zenodo.org/record/3527113/files/figure.png" pageId="3" pageNumber="200">Fig. 1</figureCitation>
ef).
</paragraph>
</subSubSection>
<subSubSection id="C36177D55E10551480F4C7F46B6DFC44" pageId="3" pageNumber="200" type="biology_ecology">
<paragraph id="8BC4245E5E10551480F4C7F46F24FCB4" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C7F46F23FC98" bold="true" box="[199,291,765,789]" pageId="3" pageNumber="200">Biology.</emphasis>
All known specimens were collected in an aquatic situation, living underwater in the sand at the edge of a pool (
<figureCitation id="134038DB5E10551480E6C6286F17FCB4" box="[213,279,801,825]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[151,1436,198,1912]" captionTargetId="figure@2.[151,1436,198,1912]" captionTargetPageId="2" captionText="FIGURE 2: Type locality of Laccobius (s. str.) shengshanensis sp. n. Heilongjiang Province, Heihe County, Shengshan National Nature Reserve." figureDoi="http://doi.org/10.5281/zenodo.3527115" httpUri="https://zenodo.org/record/3527115/files/figure.png" pageId="3" pageNumber="200">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BC4245E5E10551480F4C64C6B6DFC44" blockId="3.[151,1437,153,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C64C6FC0FCD0" bold="true" box="[199,448,837,861]" pageId="3" pageNumber="200">Differential diagnosis.</emphasis>
This new species is very similar to
<taxonomicName id="4C7B5FDD5E1055148303C64F6A71FCD0" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[816,1137,837,861]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="cinereus">
<emphasis id="B90FF84C5E1055148303C64F6DA3FCD0" box="[816,931,838,861]" italics="true" pageId="3" pageNumber="200">L. cinereus</emphasis>
Motschulsky,
<quantity id="4C8389BB5E105514840FC64C6A71FCD0" box="[1084,1137,837,861]" metricMagnitude="1" metricUnit="m" metricValue="4.724399999999999" pageId="3" pageNumber="200" unit="in" value="1860.0">1860</quantity>
</taxonomicName>
in habitus from
<collectingCountry id="F36C64CE5E1055148528C64F6B98FCD3" box="[1307,1432,838,862]" name="Kyrgyzstan" pageId="3" pageNumber="200">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="F36C64CE5E10551480A4C6606EDEFC0C" box="[151,222,873,897]" name="Russia" pageId="3" pageNumber="200">Russia</collectingCountry>
and
<collectingCountry id="F36C64CE5E1055148126C6606F7DFC0C" box="[277,381,873,897]" name="Mongolia" pageId="3" pageNumber="200">Mongolia</collectingCountry>
and
<collectingRegion id="49BFEABC5E1055148180C6606C78FC0C" box="[435,632,873,897]" country="China" name="Liaoning" pageId="3" pageNumber="200">Liaoning province</collectingRegion>
in
<collectingCountry id="F36C64CE5E10551482AEC6606CDFFC0C" box="[669,735,873,897]" name="China" pageId="3" pageNumber="200">China</collectingCountry>
, but it can be distinguished by the apex of median lobe truncate, distal third of parameres oriented externally, and apices of parameres with an indention at inner edge, while in
<taxonomicName id="4C7B5FDD5E1055148512C6876B95FC28" authorityName="Motschulsky" authorityYear="1860" box="[1313,1429,910,933]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="cinereus">
<emphasis id="B90FF84C5E1055148512C6876B95FC28" box="[1313,1429,910,933]" italics="true" pageId="3" pageNumber="200">L. cinereus</emphasis>
</taxonomicName>
, the apex of median lobe round, the parameres are nearly parallel to median lobe, and apices of parameres expanded.
</paragraph>
</subSubSection>
<subSubSection id="C36177D55E10551480F4C6DC6B8CFC60" box="[199,1420,981,1005]" pageId="3" pageNumber="200" type="etymology">
<paragraph id="8BC4245E5E10551480F4C6DC6B8CFC60" blockId="3.[151,1437,153,1077]" box="[199,1420,981,1005]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C6DC6F48FC60" bold="true" box="[199,328,981,1005]" pageId="3" pageNumber="200">Etymology.</emphasis>
This species name refers to Shengshan National Nature Reserve in which this new species was found.
</paragraph>
</subSubSection>
<subSubSection id="C36177D55E10551480F4C6F06A7AFB9C" box="[199,1146,1017,1042]" pageId="3" pageNumber="200" type="distribution">
<paragraph id="8BC4245E5E10551480F4C6F06A7AFB9C" blockId="3.[151,1437,153,1077]" box="[199,1146,1017,1042]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C6F06F5AFB9C" bold="true" box="[199,346,1017,1041]" pageId="3" pageNumber="200">Distribution.</emphasis>
So far known only from the
<typeStatus id="54C09AFC5E10551482BDC6F36CBCFB9F" box="[654,700,1018,1042]" pageId="3" pageNumber="200">type</typeStatus>
locality in
<collectingRegion id="49BFEABC5E1055148306C6F06A29FB9C" box="[821,1065,1017,1041]" country="China" name="Heilongjiang" pageId="3" pageNumber="200">Heilongjiang Province</collectingRegion>
,
<collectingCountry id="F36C64CE5E1055148400C6F06A76FB9C" box="[1075,1142,1017,1041]" name="China" pageId="3" pageNumber="200">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36177D55E10551480F4C1146B9CFA85" pageId="3" pageNumber="200" type="discussion">
<paragraph id="8BC4245E5E10551480F4C1146B95FBB8" blockId="3.[151,1437,153,1077]" box="[199,1429,1053,1077]" pageId="3" pageNumber="200">
<emphasis id="B90FF84C5E10551480F4C1146F2FFBB8" bold="true" box="[199,303,1053,1077]" pageId="3" pageNumber="200">Remarks</emphasis>
: the key to the species of the subgenus
<taxonomicName id="4C7B5FDD5E10551482E5C1146D44FBB8" authorityName="Erichson" authorityYear="1837" box="[726,836,1053,1077]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="B90FF84C5E10551482E5C1146D44FBB8" box="[726,836,1053,1077]" italics="true" pageId="3" pageNumber="200">Laccobius</emphasis>
</taxonomicName>
(s. str.) (
<bibRefCitation id="EFEA59AF5E1055148392C1146A31FBB8" author="Gentili, E." box="[929,1073,1053,1077]" pageId="3" pageNumber="200" pagination="245 - 286" refId="ref1933" refString="Gentili, E. (1995) Hydrophilidae 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). In: Jach, M. A. &amp; Ji, L. (Eds.), Water Beetles of China. Vol. 1. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, pp. 245 - 286." type="book chapter" year="1995">Gentili, 1995</bibRefCitation>
) could be modified as following:
</paragraph>
<paragraph id="8BC4245E5E10551480A4C1686B9CFBFB" blockId="3.[151,1437,1120,1289]" box="[151,1436,1120,1142]" pageId="3" pageNumber="200">7 Pronotum and basal part of elytra strongly shagreened, elytra pointed at tips...................................... 7a</paragraph>
<paragraph id="8BC4245E5E10551480A4C1776B9CFB1E" blockId="3.[151,1437,1120,1289]" box="[151,1436,1149,1171]" pageId="3" pageNumber="200">7 Pronotum more weakly shagreened, elytra not shagreened; broadened, oval; elytra usually rounded apically............. 8</paragraph>
<paragraph id="8BC4245E5E10551480A4C1926B9CFB43" blockId="3.[151,1437,1120,1289]" pageId="3" pageNumber="200">
7a Pronotum and basal part of elytra narrower and parallel-sided, the apex of median lobe round, the parameres are nearly parallel to median lobe, and apices of parameres expanded........................................
<taxonomicName id="4C7B5FDD5E1055148483C1B06B9CFB43" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1860" box="[1200,1436,1208,1230]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="cinereus">
<emphasis id="B90FF84C5E1055148483C1B06B1AFB43" box="[1200,1306,1209,1230]" italics="true" pageId="3" pageNumber="200">L. cinereus</emphasis>
Motschulsky
</taxonomicName>
</paragraph>
<paragraph id="8BC4245E5E10551480A4C1DC6B9CFA85" blockId="3.[151,1437,1120,1289]" pageId="3" pageNumber="200">
7b Pronotum and basal part of elytra not narrower and parallel-sided, the apex of median lobe truncate, distal third of parameres oriented externally, and apices of parameres with an indention at inner edge....................
<emphasis id="B90FF84C5E105514849AC1FA6B9CFA85" bold="true" box="[1193,1436,1266,1288]" pageId="3" pageNumber="200">
<taxonomicName id="4C7B5FDD5E105514849AC1FA6B60FA85" authority="Peng &amp; Ji &amp; Bian, 2019" authorityName="Peng &amp; Ji &amp; Bian" authorityYear="2019" box="[1193,1376,1266,1288]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="shengshanensis" status="sp. nov.">
<emphasis id="B90FF84C5E105514849AC1FA6B60FA85" bold="true" box="[1193,1376,1266,1288]" italics="true" pageId="3" pageNumber="200">L. shengshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C45375E1055148555C1FA6B9CFA85" box="[1382,1436,1267,1288]" pageId="3" pageNumber="200" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>