340 lines
47 KiB
XML
340 lines
47 KiB
XML
<document id="AC66213CCB503189FBA5223A31D5EB98" ID-CLB-Dataset="28123" ID-DOI="10.11646/zootaxa.4568.3.2" ID-GBIF-Dataset="d40eb95b-5a98-418a-b607-7c2cbec7c897" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601607" ID-ZooBank="8FD0AC89-424E-4CAC-92A5-A5600A481140" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172834442" checkinUser="plazi" docAuthor="Kozłowska, Anna, Bates, Denis, Zalasiewicz, Jan & Radzevičius, Sigitas" docDate="2019" docId="03A687FCFFF9985B8FA2D165FCA7FE0C" docLanguage="en" docName="zootaxa.4568.3.2.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gothograptus domeyki Kozłowska & Bates & Zalasiewicz & Radzevičius 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="457" masterDocId="FF9FFF84FFEA984D8F35D37CFD67FFA1" masterDocTitle="Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania" masterLastPageNumber="469" masterPageNumber="435" pageNumber="454" updateTime="1698703580205" updateUser="ExternalLinkService">
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<mods:titleInfo id="854346013A8330A7BBC24A720662A388">
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<mods:title id="7102D64F1C5F487B49796C84E8840B01">Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania</mods:title>
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<mods:namePart id="F1152837264AEF84AE85D1574E122CE7">Kozłowska, Anna</mods:namePart>
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<mods:namePart id="CF7D106100C6E3C1927B1EC81145E1A9">Bates, Denis</mods:namePart>
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<mods:namePart id="F57BABE7525A4D2D29B310E3590749FF">Zalasiewicz, Jan</mods:namePart>
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<mods:namePart id="4F8BB261DA7933BD68AA63963BCBA8C2">Radzevičius, Sigitas</mods:namePart>
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<mods:title id="1A8D0CCBBBB4360BB9D1B5AD336692BA">Zootaxa</mods:title>
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<mods:date id="4FB12E3E0E128A2C9DAF99D5CABCD94C">2019</mods:date>
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<mods:number id="E9A28E4880560D7D6BCA1D116F254F21">4568</mods:number>
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<treatment id="03A687FCFFF9985B8FA2D165FCA7FE0C" ID-DOI="http://doi.org/10.5281/zenodo.5935882" ID-GBIF-Taxon="154474979" ID-Zenodo-Dep="5935882" LSID="urn:lsid:plazi:treatment:03A687FCFFF9985B8FA2D165FCA7FE0C" httpUri="http://treatment.plazi.org/id/03A687FCFFF9985B8FA2D165FCA7FE0C" lastPageId="22" lastPageNumber="457" pageId="19" pageNumber="454">
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<subSubSection id="C3156561FFF9985E8FA2D165FC94FD92" box="[151,499,537,563]" pageId="19" pageNumber="454" type="nomenclature">
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<paragraph id="8BB036EAFFF9985E8FA2D165FC94FD92" blockId="19.[151,499,537,597]" box="[151,499,537,563]" pageId="19" pageNumber="454">
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<heading id="D0F88186FFF9985E8FA2D165FC94FD92" bold="true" box="[151,499,537,563]" fontSize="11" level="1" pageId="19" pageNumber="454" reason="1">
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<emphasis id="B97BEAF8FFF9985E8FA2D165FC94FD92" bold="true" box="[151,499,537,563]" pageId="19" pageNumber="454">
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<taxonomicName id="4C0F4D69FFF9985E8FA2D165FCCEFD92" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[151,425,537,563]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
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<emphasis id="B97BEAF8FFF9985E8FA2D165FCCEFD92" bold="true" box="[151,425,537,563]" italics="true" pageId="19" pageNumber="454">Gothograptus domeyki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2485783FFF9985E8E85D165FC94FD92" box="[432,499,537,563]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156561FFF9985E8FA2D140FE28FD3B" pageId="19" pageNumber="454" type="description">
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<paragraph id="8BB036EAFFF9985E8FA2D140FC72FDF4" blockId="19.[151,499,537,597]" box="[151,277,572,597]" pageId="19" pageNumber="454">
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<figureCitation id="13342A6FFFF9985E8FA2D140FC72FDF4" box="[151,277,572,597]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStartId-0="20.[151,250,1878,1900]" captionStartId-1="21.[151,260,1819,1844]" captionTargetBox-0="[223,1370,193,1878]" captionTargetBox-1="[260,1334,184,1819]" captionTargetId-0="figure@20.[223,1370,193,1868]" captionTargetId-1="figure@21.[260,1334,184,1804]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURE 15. Gothograptus domeyki n. sp. tubaria of young colonies, Šiupyliai-69 core, depth 1009.4 m, lundgreni Biozone, Lithuania. A, E, F. tubarium with five pairs of thecae, ventro-lateral view, VU RET-10; A. whole specimen; E. enlargement of orifice of th4 1; F. enlargement of orifice of th2 1. B. tubarium with six pairs of thecae, ventro-lateral view, VU RET-11. C, D. tubarium with seven pairs of thecae, VU RET-12; C. whole tubarium, lateral view; D. enlargement of orifice of th1 1." captionText-1="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri-0="https://zenodo.org/record/2601649/files/figure.png" httpUri-1="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Figs 15–16</figureCitation>
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</paragraph>
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<paragraph id="8BB036EAFFF9985E8FA2D1F8FE28FD3B" blockId="19.[151,847,644,666]" box="[151,847,644,666]" pageId="19" pageNumber="454">
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?1952
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<taxonomicName id="4C0F4D69FFF9985E8FEED1F8FF0BFD3B" authority=", Boucek & Munch" authorityName="Boucek & Munch" box="[219,620,644,666]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
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<emphasis id="B97BEAF8FFF9985E8FEED1F8FCCFFD3B" box="[219,424,644,666]" italics="true" pageId="19" pageNumber="454">Gothograptus nassa</emphasis>
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, Bouček & Münch
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</taxonomicName>
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, pp. 11–15, fig. 2, a–i.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3156561FFF9985E8FA2D1BAFED9FC86" pageId="19" pageNumber="454" type="materials_examined">
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<paragraph id="8BB036EAFFF9985E8FA2D1BAFED9FC86" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
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<emphasis id="B97BEAF8FFF9985E8FA2D1BAFC26FD7E" bold="true" box="[151,321,710,735]" pageId="19" pageNumber="454">Type material</emphasis>
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.
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<materialsCitation id="3B673CB7FFF9985E8E64D1BAFED9FC86" ID-GBIF-Occurrence="2013965661" country="Lithuania" location="Biozone" pageId="19" pageNumber="454" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54B48848FFF9985E8E64D1BAFCDEFD7E" box="[337,441,710,735]" pageId="19" pageNumber="454" type="holotype">Holotype</typeStatus>
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number VU RET-13, mature specimen with seven pairs of thecae and appendix (
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<figureCitation id="13342A6FFFF9985E8A5BD1BAFDD5FCA2" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Fig. 16</figureCitation>
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A–B), Šiupyliai-69 core, 1009,1 m,
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<emphasis id="B97BEAF8FFF9985E8D7ED197FFD0FCA2" box="[587,695,747,771]" italics="true" pageId="19" pageNumber="454">lundgreni</emphasis>
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Biozone,
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<emphasis id="B97BEAF8FFF9985E8C1ED196FEF1FCA2" bold="true" box="[811,918,746,771]" pageId="19" pageNumber="454">
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<typeStatus id="54B48848FFF9985E8C1ED196FEF1FCA2" box="[811,918,746,771]" pageId="19" pageNumber="454" type="paratype">paratype</typeStatus>
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</emphasis>
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number VU RET-14, finite tubarium (
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<figureCitation id="13342A6FFFF9985E8A7FD196F8FBFCA2" box="[1354,1436,746,771]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Fig. 16</figureCitation>
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C–E), Šiupyliai-69 core, depth 1010.0 m,
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<emphasis id="B97BEAF8FFF9985E8D59D072FFBFFC87" box="[620,728,782,806]" italics="true" pageId="19" pageNumber="454">lundgreni</emphasis>
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<location id="8ED06031FFF9985E8DEAD072FE58FC86" LSID="urn:lsid:plazi:treatment:03A687FCFFF9985B8FA2D165FCA7FE0C:8ED06031FFF9985E8DEAD072FE58FC86" box="[735,831,782,807]" country="Lithuania" name="Biozone" pageId="19" pageNumber="454">Biozone</location>
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,
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<collectingCountry id="F318767AFFF9985E8C7ED072FEDDFC86" box="[843,954,782,807]" name="Lithuania" pageId="19" pageNumber="454">Lithuania</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156561FFF9985E8FF2D04EFC0BFCCE" pageId="19" pageNumber="454" type="etymology">
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<paragraph id="8BB036EAFFF9985E8FF2D04EFC0BFCCE" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
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<emphasis id="B97BEAF8FFF9985E8FF2D04EFC29FCEA" bold="true" box="[199,334,818,843]" pageId="19" pageNumber="454">Etymology.</emphasis>
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Ignacy Domeyko (
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<date id="FFB1102AFFF9985E8D13D04FFFC0FCEA" box="[550,679,819,843]" pageId="19" pageNumber="454" value="1802" valueMax="1889">1802–1889</date>
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) was a Polish geologist, mineralogist and educator; most of his life was spent in
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<collectingCountry id="F318767AFFF9985E8E1DD02AFC0FFCCE" box="[296,360,854,879]" name="Chile" pageId="19" pageNumber="454">Chile</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156561FFF9985E8FF2D006FF01FB82" pageId="19" pageNumber="454" type="diagnosis">
|
||
<paragraph id="8BB036EAFFF9985E8FF2D006FF01FB82" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
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<emphasis id="B97BEAF8FFF9985E8FF2D006FC5DFC32" bold="true" box="[199,314,890,915]" pageId="19" pageNumber="454">Diagnosis</emphasis>
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.
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<taxonomicName id="4C0F4D69FFF9985E8E7DD007FC83FC32" authorityName=", Frech" authorityYear="1897" box="[328,484,891,915]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="genus">
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<emphasis id="B97BEAF8FFF9985E8E7DD007FC83FC32" box="[328,484,891,915]" italics="true" pageId="19" pageNumber="454">Gothograptus</emphasis>
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</taxonomicName>
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with genicular hoods of
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<taxonomicName id="4C0F4D69FFF9985E8DCED007FE5CFC33" box="[763,827,891,914]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
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<emphasis id="B97BEAF8FFF9985E8DCED007FE5CFC33" box="[763,827,891,914]" italics="true" pageId="19" pageNumber="454">nassa</emphasis>
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||
</taxonomicName>
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<typeStatus id="54B48848FFF9985E8C77D007FE15FC32" box="[834,882,891,915]" pageId="19" pageNumber="454">type</typeStatus>
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developed on the first pair of thecae; second pair of thecae may have paired spines with seams or remnants of reticulum between them; subsequent thecae lacking genicular processes except for the last theca with a reticulated hood. Distinctively thicker thecal lips than pleural and genicular lists. G
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<emphasis id="B97BEAF8FFF9985E8EADD09AFFEDFC5F" box="[408,650,998,1022]" italics="true" pageId="19" pageNumber="454">
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othograptus
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<taxonomicName id="4C0F4D69FFF9985E8D18D09AFFEDFC5F" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[557,650,998,1022]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki">domeyki</taxonomicName>
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</emphasis>
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differs from other
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<taxonomicName id="4C0F4D69FFF9985E8C4CD09AF972FC5F" authorityName=", Frech" authorityYear="1897" box="[889,1045,998,1022]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="genus">
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<emphasis id="B97BEAF8FFF9985E8C4CD09AF972FC5F" box="[889,1045,998,1022]" italics="true" pageId="19" pageNumber="454">Gothograptus</emphasis>
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</taxonomicName>
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species in the lack of genicular structures in the medial and distal thecae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3156561FFF9985E8FF2D752F80FFABE" pageId="19" pageNumber="454" type="discussion">
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<paragraph id="8BB036EAFFF9985E8FF2D752F9BAFB12" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
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||
<emphasis id="B97BEAF8FFF9985E8FF2D752FC5CFBE6" bold="true" box="[199,315,1070,1095]" pageId="19" pageNumber="454">Remarks.</emphasis>
|
||
<taxonomicName id="4C0F4D69FFF9985E8E76D752FF22FBE7" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[323,581,1070,1094]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFF9985E8E76D752FF22FBE7" box="[323,581,1070,1094]" italics="true" pageId="19" pageNumber="454">Gothograptus domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFF9985E8D79D752FFEAFBE6" bold="true" box="[588,653,1070,1095]" pageId="19" pageNumber="454">
|
||
<taxonomicNameLabel id="A2485783FFF9985E8D79D752FFEAFBE6" box="[588,653,1070,1095]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from other
|
||
<taxonomicName id="4C0F4D69FFF9985E8C52D752F964FBE7" authorityName=", Frech" authorityYear="1897" box="[871,1027,1070,1094]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="genus">
|
||
<emphasis id="B97BEAF8FFF9985E8C52D752F964FBE7" box="[871,1027,1070,1094]" italics="true" pageId="19" pageNumber="454">Gothograptus</emphasis>
|
||
</taxonomicName>
|
||
species in the lack of any genicular processes in most thecae as well as in having thick lips.
|
||
<taxonomicName id="4C0F4D69FFF9985E8C31D72FFE18FBCA" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[772,895,1107,1131]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFF9985E8C31D72FFE18FBCA" box="[772,895,1107,1131]" italics="true" pageId="19" pageNumber="454">G. domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFF9985E8CB3D72EFEA3FBCA" bold="true" box="[902,964,1106,1131]" pageId="19" pageNumber="454">
|
||
<taxonomicNameLabel id="A2485783FFF9985E8CB3D72EFEA3FBCA" box="[902,964,1106,1131]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C0F4D69FFF9985E8B61D72FF980FBCA" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[1108,1255,1107,1131]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="kozlowskii">
|
||
<emphasis id="B97BEAF8FFF9985E8B61D72FF980FBCA" box="[1108,1255,1107,1131]" italics="true" pageId="19" pageNumber="454">G. kozlowskii</emphasis>
|
||
</taxonomicName>
|
||
in having
|
||
<taxonomicName id="4C0F4D69FFF9985E8A69D72FF8FBFBCB" box="[1372,1436,1107,1130]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
|
||
<emphasis id="B97BEAF8FFF9985E8A69D72FF8FBFBCB" box="[1372,1436,1107,1130]" italics="true" pageId="19" pageNumber="454">nassa</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54B48848FFF9985E8FA2D70BFDA0FB2E" box="[151,199,1143,1167]" pageId="19" pageNumber="454">type</typeStatus>
|
||
hoods on the first pair of thecae and reticulated hoods on the last pair of thecae. Similar reticulated hoods on the last thecae are also found in
|
||
<taxonomicName id="4C0F4D69FFF9985E8ECAD7E7FF0EFB12" authorityName="Lenz & Kozlowska" authorityYear="2006" box="[511,617,1179,1203]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="436" phylum="Hemichordata" rank="species" species="storchi">
|
||
G.
|
||
<emphasis id="B97BEAF8FFF9985E8D28D7E7FF0EFB12" box="[541,617,1179,1203]" italics="true" pageId="19" pageNumber="454">storchi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C0F4D69FFF9985E8D4DD7E7FF93FB12" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[632,756,1179,1203]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFF9985E8D4DD7E7FF93FB12" box="[632,756,1179,1203]" italics="true" pageId="19" pageNumber="454">G. domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFF9985E8DCED7E6FE5DFB12" bold="true" box="[763,826,1178,1203]" pageId="19" pageNumber="454">
|
||
<taxonomicNameLabel id="A2485783FFF9985E8DCED7E6FE5DFB12" box="[763,826,1178,1203]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is so far known only from
|
||
<collectingCountry id="F318767AFFF9985E8B5FD7E6F9BEFB12" box="[1130,1241,1178,1203]" name="Lithuania" pageId="19" pageNumber="454">Lithuania</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF9985E8FF2D7C2F80FFABE" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
<bibRefCitation id="EF9E4B1BFFF9985E8FF2D7C2FCBCFB76" author="Boucek, B. & Munch, A." box="[199,475,1214,1239]" pageId="19" pageNumber="454" pagination="1 - 151" refId="ref10591" refString="Boucek, B. & Munch, A. (1952) Retioliti stredoevropskeho svrchniho wenloku a ludlowu. Sbornik Ustredniho Ustavu geologickeho 19, oddil paleontologicky, 1 - 151." type="journal article" year="1952">Bouček & Münch (1952)</bibRefCitation>
|
||
described some retiolitines as
|
||
<taxonomicName id="4C0F4D69FFF9985E8C1CD7C2F961FB77" box="[809,1030,1214,1238]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
|
||
<emphasis id="B97BEAF8FFF9985E8C1CD7C2F961FB77" box="[809,1030,1214,1238]" italics="true" pageId="19" pageNumber="454">Gothograptus nassa</emphasis>
|
||
</taxonomicName>
|
||
, from the
|
||
<collectingCountry id="F318767AFFF9985E8B42D7C2F84EFB76" box="[1143,1321,1214,1239]" name="Czech Republic" pageId="19" pageNumber="454">Czech Republic</collectingCountry>
|
||
,
|
||
<emphasis id="B97BEAF8FFF9985E8A06D7C2F8FBFB77" box="[1331,1436,1214,1238]" italics="true" pageId="19" pageNumber="454">lundgreni</emphasis>
|
||
Biozone. Their drawings show some tubaria with thick lips, and without genicular
|
||
<taxonomicName id="4C0F4D69FFF9985E8B7AD79FF9EAFB5B" box="[1103,1165,1251,1274]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
|
||
<emphasis id="B97BEAF8FFF9985E8B7AD79FF9EAFB5B" box="[1103,1165,1251,1274]" italics="true" pageId="19" pageNumber="454">nassa</emphasis>
|
||
</taxonomicName>
|
||
hoods, similar to those described herein for the first time in
|
||
<taxonomicName id="4C0F4D69FFF9985E8D10D67BFFF9FABF" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[549,670,1286,1310]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFF9985E8D10D67BFFF9FABF" box="[549,670,1286,1310]" italics="true" pageId="19" pageNumber="454">G. domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFF9985E8D90D67AFF84FABE" bold="true" box="[677,739,1286,1311]" pageId="19" pageNumber="454">
|
||
<taxonomicNameLabel id="A2485783FFF9985E8D90D67AFF84FABE" box="[677,739,1286,1311]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Proper recognition of this form needs some detailed study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156561FFF9985E8FF2D656FC08FA2A" pageId="19" pageNumber="454" type="materials_examined">
|
||
<paragraph id="8BB036EAFFF9985E8FF2D656FC08FA2A" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
<emphasis id="B97BEAF8FFF9985E8FF2D656FC48FAE2" bold="true" box="[199,303,1322,1347]" pageId="19" pageNumber="454">Material</emphasis>
|
||
. Two cores from
|
||
<collectingCountry id="F318767AFFF9985E8ECCD656FF0EFAE2" box="[505,617,1322,1347]" name="Lithuania" pageId="19" pageNumber="454">Lithuania</collectingCountry>
|
||
: Šiupyliai-69 core, depths:
|
||
<quantity id="4CF79B0FFFF9985E8CAAD657F96DFAE3" box="[927,1034,1323,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0097" pageId="19" pageNumber="454" unit="m" value="1009.7">1009.7 m</quantity>
|
||
30 young, 12 mature;
|
||
<quantity id="4CF79B0FFFF9985E8A38D657F81FFAE3" box="[1293,1400,1323,1347]" metricMagnitude="3" metricUnit="m" metricValue="1.0090999999999999" pageId="19" pageNumber="454" unit="m" value="1009.1">1009.1 m</quantity>
|
||
50 young, 31 mature; Vilkaviškis-131 core, depth
|
||
<quantity id="4CF79B0FFFF9985E8DF0D632FE53FAC7" box="[709,820,1358,1383]" metricMagnitude="3" metricUnit="m" metricValue="1.0958" pageId="19" pageNumber="454" unit="m" value="1095.8">1095.8 m</quantity>
|
||
four fragments of mature specimens. Upper part of
|
||
<emphasis id="B97BEAF8FFF9985E8FA2D60FFC64FA2A" box="[151,259,1395,1419]" italics="true" pageId="19" pageNumber="454">lundgreni</emphasis>
|
||
Biozone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156561FFF9985B8FF2D6EAFCA7FE0C" lastPageId="22" lastPageNumber="457" pageId="19" pageNumber="454" type="description">
|
||
<paragraph id="8BB036EAFFF9985E8FF2D6EAF9D0FA56" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
<emphasis id="B97BEAF8FFF9985E8FF2D6EAFC3EFA0E" bold="true" box="[199,345,1430,1455]" pageId="19" pageNumber="454">Description.</emphasis>
|
||
Mature tubaria usually have seven or eight pairs of thecae and appendix. The total length of the tubarium is about 7.0 mm, together with the appendix. The tubarium is almost parallel-sided, widest at
|
||
<quantity id="4CF79B0FFFF9985E8A28D6C7F8E6FA73" box="[1309,1409,1467,1491]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="19" pageNumber="454" unit="mm" value="0.83">0.83 mm</quantity>
|
||
at the level of the first thecal pair, and then it slowly narrows to
|
||
<quantity id="4CF79B0FFFF9985E8C7DD6A3FEF9FA57" box="[840,926,1503,1527]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="454" unit="mm" value="0.4">0.4 mm</quantity>
|
||
just below the appendix.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF9985E8FF2D57FFE63F926" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
In some mature specimens the prongs of the ancora, as well as the sicular rim and lists above the proximal lateral orifices, are extremely thick ((
|
||
<figureCitation id="13342A6FFFF9985E8D02D55AFE73F99E" box="[567,788,1574,1599]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Fig. 16A, B, E, H, I</figureCitation>
|
||
). An outer ancora is well developed in mature specimens. The proximal ventral orifices are overgrown by reticulum in mature colonies. Mid-ventral lists are
|
||
<quantity id="4CF79B0FFFF9985E8BC5D536F821F9C3" box="[1264,1350,1610,1635]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="454" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long in the first and last thecae, the rest are about
|
||
<quantity id="4CF79B0FFFF9985E8D5BD513FFA3F927" box="[622,708,1647,1671]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="454" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF9985E8FF2D5EEF9B3F952" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
Genicular structures start to develop when the colony reaches about five thecal pairs. There are three
|
||
<typeStatus id="54B48848FFF9985E8A77D5EFF81AF90A" box="[1346,1405,1683,1707]" pageId="19" pageNumber="454">types</typeStatus>
|
||
of genicular structures in mature tubaria of
|
||
<taxonomicName id="4C0F4D69FFF9985E8D51D5CBFF86F96F" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[612,737,1718,1742]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFF9985E8D51D5CBFF86F96F" box="[612,737,1718,1742]" italics="true" pageId="19" pageNumber="454">G. domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFF9985E8DDCD5CAFE4EF96E" bold="true" box="[745,809,1718,1743]" pageId="19" pageNumber="454">
|
||
<taxonomicNameLabel id="A2485783FFF9985E8DDCD5CAFE4EF96E" box="[745,809,1718,1743]" pageId="19" pageNumber="454" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The first pair of thecae have
|
||
<taxonomicName id="4C0F4D69FFF9985E8B4DD5CBF9DFF96F" box="[1144,1208,1719,1742]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="nassa">
|
||
<emphasis id="B97BEAF8FFF9985E8B4DD5CBF9DFF96F" box="[1144,1208,1719,1742]" italics="true" pageId="19" pageNumber="454">nassa</emphasis>
|
||
</taxonomicName>
|
||
hoods; usually they are small and do not cover the orifices of thecae or they are slightly larger than the thecal orifice.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF9985E8FF2D582F935F822" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
The second and third thecal pairs have a pair of curved spines grown on the pleural lists, extending horizontally. Seams on the inner sides of the spines indicate that there was a thin membrane between them, so that the spines form the sides of a hood. The bases of the spines in the mature tubaria are of triangular shape (
|
||
<figureCitation id="13342A6FFFF9985E8A1DD43AF8F7F8FE" box="[1320,1424,1862,1887]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Fig. 16G</figureCitation>
|
||
). Similar spines forming hoods are also found in
|
||
<taxonomicName id="4C0F4D69FFF9985E8D9ED417FEA5F822" authorityName=", Kozlowska-Dawidziuk" authorityYear="1990" box="[683,962,1899,1923]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="19" pageNumber="454" phylum="Hemichordata" rank="species" species="kozlowskii">
|
||
<emphasis id="B97BEAF8FFF9985E8D9ED417FEA5F822" box="[683,962,1899,1923]" italics="true" pageId="19" pageNumber="454">Gothograptus kozlowskii</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFF9985E8FF2D4F2F82CF84E" blockId="19.[151,1437,710,2031]" pageId="19" pageNumber="454">
|
||
The next thecae have no hoods; their lips are thick, about twice as thick as the geniculum list. The genicular lists became significantly thinner in the last thecae. The last pair of thecae have reticulated hoods covering orifices; the genicular list is not developed. The reticulum of these hoods is connected with the appendix (
|
||
<figureCitation id="13342A6FFFF9985E8BE3D4AAF859F84E" box="[1238,1342,2006,2031]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="19" pageNumber="454">Fig. 16A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF706662FFFE98598FA2D42AF848F86C" httpUri="https://zenodo.org/record/2601649/files/figure.png" pageId="20" pageNumber="455" startId="20.[151,250,1878,1900]" targetBox="[223,1370,193,1878]" targetPageId="20">
|
||
<paragraph id="8BB036EAFFFE98598FA2D42AF848F86C" blockId="20.[151,1436,1878,1997]" pageId="20" pageNumber="455">
|
||
<emphasis id="B97BEAF8FFFE98598FA2D42AFC47F8CD" bold="true" box="[151,288,1878,1900]" pageId="20" pageNumber="455">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C0F4D69FFFE98598E1DD42AFF77F8CD" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[296,528,1878,1900]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="20" pageNumber="455" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFFE98598E1DD42AFF77F8CD" box="[296,528,1878,1900]" italics="true" pageId="20" pageNumber="455">Gothograptus domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFFE98598D22D42AFF36F8CD" bold="true" box="[535,593,1878,1900]" pageId="20" pageNumber="455">
|
||
<taxonomicNameLabel id="A2485783FFFE98598D22D42AFF36F8CD" box="[535,593,1878,1900]" pageId="20" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
tubaria of young colonies, Šiupyliai-69 core, depth 1009.4 m,
|
||
<emphasis id="B97BEAF8FFFE98598BEDD42AF85EF8CD" box="[1240,1337,1878,1900]" italics="true" pageId="20" pageNumber="455">lundgreni</emphasis>
|
||
Biozone, Lithuania. A, E, F. tubarium with five pairs of thecae, ventro-lateral view, VU RET-10; A. whole specimen; E. enlargement of orifice of th4
|
||
<superScript id="7C7A9BA2FFFE98598E2BD4EFFC42F83E" attach="none" box="[286,293,1939,1951]" fontSize="5" pageId="20" pageNumber="455">1</superScript>
|
||
; F. enlargement of orifice of th2
|
||
<superScript id="7C7A9BA2FFFE98598D42D4EFFF19F83E" attach="right" box="[631,638,1939,1951]" fontSize="5" pageId="20" pageNumber="455">1</superScript>
|
||
. B. tubarium with six pairs of thecae, ventro-lateral view, VU RET-11. C, D. tubarium with seven pairs of thecae, VU RET-12; C. whole tubarium, lateral view; D. enlargement of orifice of th1
|
||
<superScript id="7C7A9BA2FFFE98598A17D4C8F84EF861" attach="none" box="[1314,1321,1972,1984]" fontSize="5" pageId="20" pageNumber="455">1</superScript>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF706662FFFF98588FA2D467F8FCF86C" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="21" pageNumber="456" startId="21.[151,260,1819,1844]" targetBox="[260,1334,184,1819]" targetPageId="21">
|
||
<paragraph id="8BB036EAFFFF98588FA2D467F8FCF86C" blockId="21.[151,1437,1819,2033]" pageId="21" pageNumber="456">
|
||
<emphasis id="B97BEAF8FFFF98588FA2D467FC49F895" bold="true" box="[151,302,1819,1844]" pageId="21" pageNumber="456">FIGURE 16.</emphasis>
|
||
Tubaria of mature colonies of
|
||
<taxonomicName id="4C0F4D69FFFF98588D51D461FE2CF892" authority="Kozłowska & Bates & Zalasiewicz & Radzevičius, 2019" authorityName="Kozłowska & Bates & Zalasiewicz & Radzevičius" authorityYear="2019" box="[612,843,1821,1843]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="21" pageNumber="456" phylum="Hemichordata" rank="species" species="domeyki" status="sp. nov.">
|
||
<emphasis id="B97BEAF8FFFF98588D51D461FE2CF892" box="[612,843,1821,1843]" italics="true" pageId="21" pageNumber="456">Gothograptus domeyki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97BEAF8FFFF98588C67D461FEEDF892" bold="true" box="[850,906,1821,1843]" pageId="21" pageNumber="456">
|
||
<taxonomicNameLabel id="A2485783FFFF98588C67D461FEEDF892" box="[850,906,1821,1843]" pageId="21" pageNumber="456" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Šiupyliai-69 core, upper part of
|
||
<emphasis id="B97BEAF8FFFF98588BEFD461F85BF892" box="[1242,1340,1821,1843]" italics="true" pageId="21" pageNumber="456">lundgreni</emphasis>
|
||
Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix,
|
||
<emphasis id="B97BEAF8FFFF98588CBAD441FE8BF8F2" bold="true" box="[911,1004,1853,1875]" pageId="21" pageNumber="456">holotype</emphasis>
|
||
, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium,
|
||
<emphasis id="B97BEAF8FFFF98588FA2D4E5FD9FF80E" bold="true" box="[151,248,1945,1967]" pageId="21" pageNumber="456">paratype</emphasis>
|
||
, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB036EAFFFF98588FA2D4A7FCE3F850" blockId="21.[151,1437,1819,2033]" box="[151,388,2011,2033]" pageId="21" pageNumber="456">umbrella, outside view.</paragraph>
|
||
<paragraph id="8BB036EAFFFC985B8FF2D3E4F9F1FEBC" blockId="22.[151,1437,151,429]" pageId="22" pageNumber="457">
|
||
Some young tubaria with five pairs of thecae have reticulum up to the third pair of thecae, small
|
||
<taxonomicName id="4C0F4D69FFFC985B8A24D3E5F836FF11" box="[1297,1361,153,176]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="22" pageNumber="457" phylum="Hemichordata" rank="species" species="nassa">
|
||
<emphasis id="B97BEAF8FFFC985B8A24D3E5F836FF11" box="[1297,1361,153,176]" italics="true" pageId="22" pageNumber="457">nassa</emphasis>
|
||
</taxonomicName>
|
||
hoods on the first thecae, the two next pairs of thecae having thin genicular spines. A specimen with seven pairs of thecae has a well-developed reticulum, except on the two last thecae. The lists of young colonies are thin; the thickest is the nema (
|
||
<figureCitation id="13342A6FFFFC985B8E39D278FCF1FEBD" box="[268,406,260,285]" captionStart="FIGURE 15" captionStartId="20.[151,250,1878,1900]" captionTargetBox="[223,1370,193,1878]" captionTargetId="figure@20.[223,1370,193,1868]" captionTargetPageId="20" captionText="FIGURE 15. Gothograptus domeyki n. sp. tubaria of young colonies, Šiupyliai-69 core, depth 1009.4 m, lundgreni Biozone, Lithuania. A, E, F. tubarium with five pairs of thecae, ventro-lateral view, VU RET-10; A. whole specimen; E. enlargement of orifice of th4 1; F. enlargement of orifice of th2 1. B. tubarium with six pairs of thecae, ventro-lateral view, VU RET-11. C, D. tubarium with seven pairs of thecae, VU RET-12; C. whole tubarium, lateral view; D. enlargement of orifice of th1 1." httpUri="https://zenodo.org/record/2601649/files/figure.png" pageId="22" pageNumber="457">Fig. 15A, D</figureCitation>
|
||
). During astogeny the next thicker lists are the lateral apertural rods.
|
||
</paragraph>
|
||
<paragraph id="8BB036EAFFFC985B8FF2D25BFCA7FE0C" blockId="22.[151,1437,151,429]" pageId="22" pageNumber="457">
|
||
The lips and genicular lists are so thick that the thecal orifice is very narrow (
|
||
<figureCitation id="13342A6FFFFC985B8B56D25BF9ACFEE1" box="[1123,1227,295,320]" captionStart="FIGURE 15" captionStartId="20.[151,250,1878,1900]" captionTargetBox="[223,1370,193,1878]" captionTargetId="figure@20.[223,1370,193,1868]" captionTargetPageId="20" captionText="FIGURE 15. Gothograptus domeyki n. sp. tubaria of young colonies, Šiupyliai-69 core, depth 1009.4 m, lundgreni Biozone, Lithuania. A, E, F. tubarium with five pairs of thecae, ventro-lateral view, VU RET-10; A. whole specimen; E. enlargement of orifice of th4 1; F. enlargement of orifice of th2 1. B. tubarium with six pairs of thecae, ventro-lateral view, VU RET-11. C, D. tubarium with seven pairs of thecae, VU RET-12; C. whole tubarium, lateral view; D. enlargement of orifice of th1 1." httpUri="https://zenodo.org/record/2601649/files/figure.png" pageId="22" pageNumber="457">Fig. 15E</figureCitation>
|
||
,
|
||
<figureCitation id="13342A6FFFFC985B8BEFD25BF86BFEE1" box="[1242,1292,295,320]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="22" pageNumber="457">16E</figureCitation>
|
||
). The thecal orifices vary in shape from nearly rectangular to pyramidal with rounded corners (
|
||
<figureCitation id="13342A6FFFFC985B8B58D230F9ABFEC5" box="[1133,1228,332,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,894,916]" captionTargetBox="[151,1435,280,787]" captionTargetId="figure@9.[151,1435,280,787]" captionTargetPageId="9" captionText="FIGURE 7. Thecal orifices and hoods of Gothograptus. A. Pyramidal orifices in Gothograptus domeyki n. sp., VU RET-13, Šiupyliai-69 core, Lithuania, depth 1010.0 m, upper part of lundgreni Biozone. B. rectangular orifice in Gothograptus nassa (Holm, 1890), Bartoszyce IG-1 core, depth 1656.8 m, nassa Biozone, ZPAL G.55/13." httpUri="https://zenodo.org/record/2601627/files/figure.png" pageId="22" pageNumber="457">Fig. 7A</figureCitation>
|
||
,
|
||
<figureCitation id="13342A6FFFFC985B8BE9D230F854FEC5" box="[1244,1331,332,357]" captionStart="FIGURE 16" captionStartId="21.[151,260,1819,1844]" captionTargetBox="[260,1334,184,1819]" captionTargetId="figure@21.[260,1334,184,1804]" captionTargetPageId="21" captionText="FIGURE 16. Tubaria of mature colonies of Gothograptus domeyki n. sp., Šiupyliai-69 core, upper part of lundgreni Biozone, Lithuania. A–B. finite tubarium with seven pairs of thecae and appendix, holotype, depth 1009,1 m, VU RET-13; A. ventrolateral view of specimen; B. enlargement of thick ancora and broken outer ancora, oblique view. C, H. finite tubarium, depth 1010.0 m, VU RET-14; C. enlargement of two thecae; H. enlargement of proximal end. D–E. fragment of finite tubarium, paratype, depth 1010.0 m, VU RET-15; D. whole specimen, ventro-lateral view; E. proximal thecae with spines. F–G. proximal end of young tubarium, depth 1010.0 m, VU RET-16; F. enlargement of proximal theca; G. enlargement of ancora" httpUri="https://zenodo.org/record/2601651/files/figure.png" pageId="22" pageNumber="457">16C, E</figureCitation>
|
||
). Mature colonies have a dense reticulum of similar thickness to the clathrial lists, thus the structures formed by clathrial lists are difficult to distinguish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |