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<document id="D1BC20205184E86F98431E092E0DAB3A" ID-CLB-Dataset="28123" ID-DOI="10.11646/zootaxa.4568.3.2" ID-GBIF-Dataset="d40eb95b-5a98-418a-b607-7c2cbec7c897" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601607" ID-ZooBank="8FD0AC89-424E-4CAC-92A5-A5600A481140" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172834442" checkinUser="plazi" docAuthor="Kozłowska, Anna, Bates, Denis, Zalasiewicz, Jan &amp; Radzevičius, Sigitas" docDate="2019" docId="03A687FCFFF598528FA2D230FF8FFA86" docLanguage="en" docName="zootaxa.4568.3.2.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gothograptus diminutus Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius 2019, n. sp." docType="treatment" docVersion="9" lastPageNumber="466" masterDocId="FF9FFF84FFEA984D8F35D37CFD67FFA1" masterDocTitle="Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania" masterLastPageNumber="469" masterPageNumber="435" pageNumber="466" updateTime="1698703580205" updateUser="ExternalLinkService">
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<mods:title id="F07DAFF3DF4EA5DB9F8C99438EF050E9">Evolutionary significance of the retiolitine Gothograptus (Graptolithina) with four new species from the Silurian of the East European Platform (Baltica), Poland and Lithuania</mods:title>
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<mods:namePart id="9F3A458569AD88B2EA7DF6680C70B5D0">Kozłowska, Anna</mods:namePart>
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<mods:namePart id="969B5BB68B387203BE25413E4CBEBFF0">Bates, Denis</mods:namePart>
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<mods:namePart id="10D76B6307FA29EAB44F6F3BE1716A46">Zalasiewicz, Jan</mods:namePart>
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<treatment id="03A687FCFFF598528FA2D230FF8FFA86" ID-DOI="http://doi.org/10.5281/zenodo.5935890" ID-GBIF-Taxon="154474980" ID-Zenodo-Dep="5935890" LSID="urn:lsid:plazi:treatment:03A687FCFFF598528FA2D230FF8FFA86" httpUri="http://treatment.plazi.org/id/03A687FCFFF598528FA2D230FF8FFA86" lastPageNumber="466" pageId="31" pageNumber="466">
<subSubSection id="C3156561FFF598528FA2D230FF6FFEC7" box="[151,520,332,358]" pageId="31" pageNumber="466" type="nomenclature">
<paragraph id="8BB036EAFFF598528FA2D230FF6FFEC7" blockId="31.[151,520,332,391]" box="[151,520,332,358]" pageId="31" pageNumber="466">
<heading id="D0F88186FFF598528FA2D230FF6FFEC7" bold="true" box="[151,520,332,358]" fontSize="11" level="1" pageId="31" pageNumber="466" reason="1">
<emphasis id="B97BEAF8FFF598528FA2D230FF6FFEC7" bold="true" box="[151,520,332,358]" pageId="31" pageNumber="466">
<taxonomicName id="4C0F4D69FFF598528FA2D230FCD9FEC7" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[151,446,332,358]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="diminutus" status="sp. nov.">
<emphasis id="B97BEAF8FFF598528FA2D230FCD9FEC7" bold="true" box="[151,446,332,358]" italics="true" pageId="31" pageNumber="466">Gothograptus diminutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2485783FFF598528EF0D230FF6FFEC7" box="[453,520,332,358]" pageId="31" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3156561FFF598528FA2D212FD81FE26" box="[151,230,366,391]" pageId="31" pageNumber="466" type="description">
<paragraph id="8BB036EAFFF598528FA2D212FD81FE26" blockId="31.[151,520,332,391]" box="[151,230,366,391]" pageId="31" pageNumber="466">
<figureCitation id="13342A6FFFF598528FA2D212FD81FE26" box="[151,230,366,391]" captionStart="FIGURE 23" captionStartId="30.[151,250,1862,1884]" captionTargetBox="[227,1367,193,1853]" captionTargetId="figure@30.[227,1367,193,1853]" captionTargetPageId="30" captionText="FIGURE 23. Gothograptus diminutus n. sp. Bartoszyce IG-1core, Poland. A. lateral view of tubarium, almost mature, with seven pairs of thecae, 1659.9 m, ZPAL G.55/32. B, C, E. mature tubarium with six pairs of thecae and appendix, holotype, 1659.9 m, ZPAL G.29/21; B. ventro-lateral view; C, fragment of third pair of theca; E. proximal end. D. finite tubarium with six pairs of thecae and appendix, paratype, depth 1659.9 m, ZPAL G.55/34. F. mature tubarium with seven pairs of thecae, ventrolateral view, depth 1658.8 m, ZPAL G.54/8. G. young tubarium with four pairs of thecae, depth 1659.9 m, ZPAL G.55/35." httpUri="https://zenodo.org/record/2601665/files/figure.png" pageId="31" pageNumber="466">Fig. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FA2D2CAFE2BFE4A" pageId="31" pageNumber="466" type="reference_group">
<paragraph id="8BB036EAFFF598528FA2D2CAFEAFFE6D" blockId="31.[151,968,438,491]" box="[151,968,438,461]" pageId="31" pageNumber="466">
1999
<taxonomicName id="4C0F4D69FFF598528FE5D2CAFF49FE6D" authority="(Holm, 1890)" baseAuthorityName="Holm" baseAuthorityYear="1890" box="[208,558,438,461]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528FE5D2CAFCFAFE6C" box="[208,413,438,461]" italics="true" pageId="31" pageNumber="466">Gothograptus nassa</emphasis>
(
<bibRefCitation id="EF9E4B1BFFF598528E9FD2CAFF40FE6D" author="Holm, G." box="[426,551,438,460]" pageId="31" pageNumber="466" pagination="1 - 34" refId="ref10823" refString="Holm, G. (1890) Gotlands graptoliter. Svenska Vetenskap-Akademiens handlingar, 16, 1 - 34." type="journal article" year="1890">Holm, 1890</bibRefCitation>
)
</taxonomicName>
, Kozłowska-Dawidziuk, p. 156, fig. 3B.
</paragraph>
<paragraph id="8BB036EAFFF598528FA2D2A9FE2BFE4A" blockId="31.[151,968,438,491]" box="[151,844,469,491]" pageId="31" pageNumber="466">
2016
<taxonomicName id="4C0F4D69FFF598528FE5D2A9FF49FE4A" authority="(Holm, 1890)" baseAuthorityName="Holm" baseAuthorityYear="1890" box="[208,558,469,491]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528FE5D2A9FCFAFE4A" box="[208,413,469,491]" italics="true" pageId="31" pageNumber="466">Gothograptus nassa</emphasis>
(
<bibRefCitation id="EF9E4B1BFFF598528E9FD2A9FF40FE4A" author="Holm, G." box="[426,551,469,491]" pageId="31" pageNumber="466" pagination="1 - 34" refId="ref10823" refString="Holm, G. (1890) Gotlands graptoliter. Svenska Vetenskap-Akademiens handlingar, 16, 1 - 34." type="journal article" year="1890">Holm, 1890</bibRefCitation>
)
</taxonomicName>
, Kozłowska, p. 535, fig. 1b.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FA2D16BFF0DFDD9" pageId="31" pageNumber="466" type="materials_examined">
<paragraph id="8BB036EAFFF598528FA2D16BFF0DFDD9" blockId="31.[151,1437,535,1320]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FA2D16BFC26FD91" bold="true" box="[151,321,535,560]" pageId="31" pageNumber="466">Type material</emphasis>
.
<typeStatus id="54B48848FFF598528E66D16BFCDCFD91" box="[339,443,535,560]" pageId="31" pageNumber="466" type="holotype">Holotype</typeStatus>
ZPAL G.29/21, mature specimen with six pairs of thecae (
<figureCitation id="13342A6FFFF598528B47D16BF844FD91" box="[1138,1315,535,560]" captionStart="FIGURE 23" captionStartId="30.[151,250,1862,1884]" captionTargetBox="[227,1367,193,1853]" captionTargetId="figure@30.[227,1367,193,1853]" captionTargetPageId="30" captionText="FIGURE 23. Gothograptus diminutus n. sp. Bartoszyce IG-1core, Poland. A. lateral view of tubarium, almost mature, with seven pairs of thecae, 1659.9 m, ZPAL G.55/32. B, C, E. mature tubarium with six pairs of thecae and appendix, holotype, 1659.9 m, ZPAL G.29/21; B. ventro-lateral view; C, fragment of third pair of theca; E. proximal end. D. finite tubarium with six pairs of thecae and appendix, paratype, depth 1659.9 m, ZPAL G.55/34. F. mature tubarium with seven pairs of thecae, ventrolateral view, depth 1658.8 m, ZPAL G.54/8. G. young tubarium with four pairs of thecae, depth 1659.9 m, ZPAL G.55/35." httpUri="https://zenodo.org/record/2601665/files/figure.png" pageId="31" pageNumber="466">Fig. 23B, C, E</figureCitation>
);
<typeStatus id="54B48848FFF598528A09D164F8FBFD91" box="[1340,1436,536,560]" pageId="31" pageNumber="466" type="paratype">paratype</typeStatus>
ZPAL G.55/34, mature tubarium with six pairs of thecae and appendix (
<figureCitation id="13342A6FFFF598528CEBD140F92DFDF5" box="[990,1098,572,597]" captionStart="FIGURE 23" captionStartId="30.[151,250,1862,1884]" captionTargetBox="[227,1367,193,1853]" captionTargetId="figure@30.[227,1367,193,1853]" captionTargetPageId="30" captionText="FIGURE 23. Gothograptus diminutus n. sp. Bartoszyce IG-1core, Poland. A. lateral view of tubarium, almost mature, with seven pairs of thecae, 1659.9 m, ZPAL G.55/32. B, C, E. mature tubarium with six pairs of thecae and appendix, holotype, 1659.9 m, ZPAL G.29/21; B. ventro-lateral view; C, fragment of third pair of theca; E. proximal end. D. finite tubarium with six pairs of thecae and appendix, paratype, depth 1659.9 m, ZPAL G.55/34. F. mature tubarium with seven pairs of thecae, ventrolateral view, depth 1658.8 m, ZPAL G.54/8. G. young tubarium with four pairs of thecae, depth 1659.9 m, ZPAL G.55/35." httpUri="https://zenodo.org/record/2601665/files/figure.png" pageId="31" pageNumber="466">Fig. 23D</figureCitation>
). Bartoszyce IG-1well, depth
<quantity id="4CF79B0FFFF598528FA2D123FC64FDD6" box="[151,259,607,632]" metricMagnitude="3" metricUnit="m" metricValue="1.6599" pageId="31" pageNumber="466" unit="m" value="1659.9">1659.9 m</quantity>
,
<collectingCountry id="F318767AFFF598528E3BD123FC38FDD9" box="[270,351,607,632]" name="Poland" pageId="31" pageNumber="466">Poland</collectingCountry>
,
<emphasis id="B97BEAF8FFF598528E5FD11CFCA4FDD6" box="[362,451,608,631]" italics="true" pageId="31" pageNumber="466">
<taxonomicName id="4C0F4D69FFF598528E5FD11CFCD2FDD6" box="[362,437,608,631]" class="Graptolithina" family="Rhabdinoporidae" genus="Pristiograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="parvus">parvus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C0F4D69FFF598528EF7D11CFF0DFDD9" authority="Biozone." authorityName="Biozone." box="[450,618,607,632]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528EF7D11CFC99FDD6" box="[450,510,608,631]" italics="true" pageId="31" pageNumber="466">nassa</emphasis>
Biozone.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FF2D1F8F94AFD3C" box="[199,1069,644,669]" pageId="31" pageNumber="466" type="etymology">
<paragraph id="8BB036EAFFF598528FF2D1F8F94AFD3C" blockId="31.[151,1437,535,1320]" box="[199,1069,644,669]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FF2D1F8FC20FD3C" bold="true" box="[199,327,644,669]" pageId="31" pageNumber="466">Etymology</emphasis>
. Latin
<emphasis id="B97BEAF8FFF598528EA3D1F8FF7AFD3D" box="[406,541,644,668]" italics="true" pageId="31" pageNumber="466">diminutus—</emphasis>
relating to the small size of the finite tubarium.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FF2D1DBFE67FCA9" pageId="31" pageNumber="466" type="diagnosis">
<paragraph id="8BB036EAFFF598528FF2D1DBFE67FCA9" blockId="31.[151,1437,535,1320]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FF2D1DBFC5DFD61" bold="true" box="[199,314,679,704]" pageId="31" pageNumber="466">Diagnosis</emphasis>
. Small finite tubarium reaching six or seven pairs of thecae and appendix. Genicular structures of the
<taxonomicName id="4C0F4D69FFF598528FF1D1B1FC63FD45" box="[196,260,717,740]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528FF1D1B1FC63FD45" box="[196,260,717,740]" italics="true" pageId="31" pageNumber="466">nassa</emphasis>
</taxonomicName>
<typeStatus id="54B48848FFF598528E3AD1B1FC58FD44" box="[271,319,717,741]" pageId="31" pageNumber="466">type</typeStatus>
of hood are usually developed on medial and distal thecae. It differs from other
<taxonomicName id="4C0F4D69FFF598528A35D1B0F8FBFD45" authorityName=", Frech" authorityYear="1897" box="[1280,1436,716,740]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="genus">
<emphasis id="B97BEAF8FFF598528A35D1B0F8FBFD45" box="[1280,1436,716,740]" italics="true" pageId="31" pageNumber="466">Gothograptus</emphasis>
</taxonomicName>
species in the small hoods and small number of thecae.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FF2D068FE10FCF1" pageId="31" pageNumber="466" type="materials_examined">
<paragraph id="8BB036EAFFF598528FF2D068FE10FCF1" blockId="31.[151,1437,535,1320]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FF2D068FC48FC8C" bold="true" box="[199,303,788,813]" pageId="31" pageNumber="466">Material</emphasis>
. Bartoszyce IG-1 core,
<collectingCountry id="F318767AFFF598528D09D068FFEAFC8C" box="[572,653,788,813]" name="Poland" pageId="31" pageNumber="466">Poland</collectingCountry>
,
<emphasis id="B97BEAF8FFF598528DADD068FF97FC8A" box="[664,752,788,811]" italics="true" pageId="31" pageNumber="466">
<taxonomicName id="4C0F4D69FFF598528DADD068FF84FC8A" box="[664,739,788,811]" class="Graptolithina" family="Rhabdinoporidae" genus="Pristiograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="parvus">parvus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C0F4D69FFF598528DC5D068FEF4FC8C" authority="Biozone" authorityName="Biozone" box="[752,915,788,813]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528DC5D068FE4BFC8A" box="[752,812,788,811]" italics="true" pageId="31" pageNumber="466">nassa</emphasis>
Biozone
</taxonomicName>
, depths
<quantity id="4CF79B0FFFF598528CDAD068F93BFC8D" box="[1007,1116,788,813]" metricMagnitude="3" metricUnit="m" metricValue="1.6599" pageId="31" pageNumber="466" unit="m" value="1659.9">1659.9 m</quantity>
; five mature specimens, and about 60 young tubaria or fragments;
<quantity id="4CF79B0FFFF598528D0BD04BFFCDFCEE" box="[574,682,823,848]" metricMagnitude="3" metricUnit="m" metricValue="1.6588" pageId="31" pageNumber="466" unit="m" value="1658.8">1658.8 m</quantity>
, a few specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FF2D020FE7EFBD2" pageId="31" pageNumber="466" type="description">
<paragraph id="8BB036EAFFF598528FF2D020FE7EFBD2" blockId="31.[151,1437,535,1320]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FF2D020FC36FCD4" bold="true" box="[199,337,860,885]" pageId="31" pageNumber="466">Description</emphasis>
. Tubaria of
<taxonomicName id="4C0F4D69FFF598528EEBD020FF0CFCD5" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[478,619,860,884]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="diminutus" status="sp. nov.">
<emphasis id="B97BEAF8FFF598528EEBD020FF0CFCD5" box="[478,619,860,884]" italics="true" pageId="31" pageNumber="466">G. diminutus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFF598528D41D020FFD4FCD4" bold="true" box="[628,691,860,885]" pageId="31" pageNumber="466">
<taxonomicNameLabel id="A2485783FFF598528D41D020FFD4FCD4" box="[628,691,860,885]" pageId="31" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are small; length of mature tubarium with six pairs of thecae and appendix is about 5.0
<quantity id="4CF79B0FFFF598528EA0D003FC8AFC36" box="[405,493,895,920]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="466" unit="mm" value="5.5">5.5 mm</quantity>
. Nema short. Length of appendix is about
<quantity id="4CF79B0FFFF598528CFDD003F946FC36" box="[968,1057,895,920]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="466" unit="mm" value="0.6">0.6 mm</quantity>
. Width of proximal end above the ancora umbrella rim is
<quantity id="4CF79B0FFFF598528EAFD0D8FC95FC1D" box="[410,498,932,956]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="466" unit="mm" value="0.7">0.7 mm</quantity>
. Maximum width of tubarium reaches about
<quantity id="4CF79B0FFFF598528CDDD0D8F959FC1D" box="[1000,1086,932,956]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="466" unit="mm" value="0.9">0.9 mm</quantity>
at the level of second and third pairs of thecae. The tubarium slowly tapers distally; below the appendix it is
<quantity id="4CF79B0FFFF598528B3FD0BBF9F4FC7E" box="[1034,1171,967,992]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="31" pageNumber="466" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40.5 mm</quantity>
wide. Mid-ventral lists are slightly longer in medial thecae, about
<quantity id="4CF79B0FFFF598528DB5D093FFBCFBA6" box="[640,731,1007,1031]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="466" unit="mm" value="0.7">0.7 mm</quantity>
. Length of mid-ventral list of th1
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is
<quantity id="4CF79B0FFFF598528BB5D092F98FFBA6" box="[1152,1256,1006,1031]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="466" unit="mm" value="0.35">0.35 mm</quantity>
. Mature tubaria have
<taxonomicName id="4C0F4D69FFF598528FE2D76FFC70FB8B" box="[215,279,1043,1066]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528FE2D76FFC70FB8B" box="[215,279,1043,1066]" italics="true" pageId="31" pageNumber="466">nassa</emphasis>
</taxonomicName>
-
<typeStatus id="54B48848FFF598528E2BD76FFC37FB8A" box="[286,336,1043,1067]" pageId="31" pageNumber="466">type</typeStatus>
hoods developed on medial and distal thecae. The biggest hoods are in distal thecae. Mature tubaria usually have thin lists and poorly developed reticulum (
<figureCitation id="13342A6FFFF598528C6BD74AFE8EFBEE" box="[862,1001,1078,1103]" captionStart="FIGURE 23" captionStartId="30.[151,250,1862,1884]" captionTargetBox="[227,1367,193,1853]" captionTargetId="figure@30.[227,1367,193,1853]" captionTargetPageId="30" captionText="FIGURE 23. Gothograptus diminutus n. sp. Bartoszyce IG-1core, Poland. A. lateral view of tubarium, almost mature, with seven pairs of thecae, 1659.9 m, ZPAL G.55/32. B, C, E. mature tubarium with six pairs of thecae and appendix, holotype, 1659.9 m, ZPAL G.29/21; B. ventro-lateral view; C, fragment of third pair of theca; E. proximal end. D. finite tubarium with six pairs of thecae and appendix, paratype, depth 1659.9 m, ZPAL G.55/34. F. mature tubarium with seven pairs of thecae, ventrolateral view, depth 1658.8 m, ZPAL G.54/8. G. young tubarium with four pairs of thecae, depth 1659.9 m, ZPAL G.55/35." httpUri="https://zenodo.org/record/2601665/files/figure.png" pageId="31" pageNumber="466">Fig. 23A, D</figureCitation>
). Some membrane of the metasicula is observed (
<figureCitation id="13342A6FFFF598528E3ED726FCE8FBD2" box="[267,399,1114,1139]" captionStart="FIGURE 23" captionStartId="30.[151,250,1862,1884]" captionTargetBox="[227,1367,193,1853]" captionTargetId="figure@30.[227,1367,193,1853]" captionTargetPageId="30" captionText="FIGURE 23. Gothograptus diminutus n. sp. Bartoszyce IG-1core, Poland. A. lateral view of tubarium, almost mature, with seven pairs of thecae, 1659.9 m, ZPAL G.55/32. B, C, E. mature tubarium with six pairs of thecae and appendix, holotype, 1659.9 m, ZPAL G.29/21; B. ventro-lateral view; C, fragment of third pair of theca; E. proximal end. D. finite tubarium with six pairs of thecae and appendix, paratype, depth 1659.9 m, ZPAL G.55/34. F. mature tubarium with seven pairs of thecae, ventrolateral view, depth 1658.8 m, ZPAL G.54/8. G. young tubarium with four pairs of thecae, depth 1659.9 m, ZPAL G.55/35." httpUri="https://zenodo.org/record/2601665/files/figure.png" pageId="31" pageNumber="466">Fig. 23B, F</figureCitation>
). The outer ancora is not observed.
</paragraph>
</subSubSection>
<subSubSection id="C3156561FFF598528FF2D702FF8FFA86" pageId="31" pageNumber="466" type="discussion">
<paragraph id="8BB036EAFFF598528FF2D702FF8FFA86" blockId="31.[151,1437,535,1320]" pageId="31" pageNumber="466">
<emphasis id="B97BEAF8FFF598528FF2D702FC53FB36" bold="true" box="[199,308,1150,1175]" pageId="31" pageNumber="466">Remarks</emphasis>
.
<taxonomicName id="4C0F4D69FFF598528E7FD703FF05FB36" authority="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius, 2019" authorityName="Kozłowska &amp; Bates &amp; Zalasiewicz &amp; Radzevičius" authorityYear="2019" box="[330,610,1151,1175]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="diminutus" status="sp. nov.">
<emphasis id="B97BEAF8FFF598528E7FD703FF05FB36" box="[330,610,1151,1175]" italics="true" pageId="31" pageNumber="466">Gothograptus diminutus</emphasis>
</taxonomicName>
<emphasis id="B97BEAF8FFF598528D47D702FFDEFB36" bold="true" box="[626,697,1150,1175]" pageId="31" pageNumber="466">
<taxonomicNameLabel id="A2485783FFF598528D47D702FFDEFB36" box="[626,697,1150,1175]" pageId="31" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the smallest form among the
<taxonomicName id="4C0F4D69FFF598528B6DD703F993FB36" authorityName=", Frech" authorityYear="1897" box="[1112,1268,1151,1175]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="genus">
<emphasis id="B97BEAF8FFF598528B6DD703F993FB36" box="[1112,1268,1151,1175]" italics="true" pageId="31" pageNumber="466">Gothograptus</emphasis>
</taxonomicName>
species. It is distinctive in that its mature tubaria do not have thick lists and dense reticulum, characteristic of mature colonies of all other
<taxonomicName id="4C0F4D69FFF598528FCCD7BBFCF2FB7E" authorityName=", Frech" authorityYear="1897" box="[249,405,1223,1247]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="genus">
<emphasis id="B97BEAF8FFF598528FCCD7BBFCF2FB7E" box="[249,405,1223,1247]" italics="true" pageId="31" pageNumber="466">Gothograptus</emphasis>
</taxonomicName>
species. This form is comparable in size to the dwarf monograptid,
<taxonomicName id="4C0F4D69FFF598528BBCD7B4F81AFB7E" box="[1161,1405,1224,1247]" class="Graptolithina" family="Rhabdinoporidae" genus="Pristiograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="parvus">
<emphasis id="B97BEAF8FFF598528BBCD7B4F81AFB7E" box="[1161,1405,1224,1247]" italics="true" pageId="31" pageNumber="466">Pristiograptus parvus</emphasis>
</taxonomicName>
. It is noteworthy that both species occur at the same stratigraphical level, in the
<emphasis id="B97BEAF8FFF598528CCAD797F930FAA3" box="[1023,1111,1259,1282]" italics="true" pageId="31" pageNumber="466">
<taxonomicName id="4C0F4D69FFF598528CCAD797F92DFAA3" box="[1023,1098,1259,1282]" class="Graptolithina" family="Rhabdinoporidae" genus="Pristiograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="parvus">parvus</taxonomicName>
/
</emphasis>
<taxonomicName id="4C0F4D69FFF598528B62D797F85FFAA2" authority="Biozone. This" authorityName="Biozone. This" box="[1111,1336,1258,1283]" class="Graptolithina" family="Retiolitidae" genus="Gothograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Graptoloidea" pageId="31" pageNumber="466" phylum="Hemichordata" rank="species" species="nassa">
<emphasis id="B97BEAF8FFF598528B62D797F9F4FAA3" box="[1111,1171,1259,1282]" italics="true" pageId="31" pageNumber="466">nassa</emphasis>
Biozone. This
</taxonomicName>
material comes only from the Bartoszyce IG-1core of
<collectingCountry id="F318767AFFF598528DA6D672FF83FA86" box="[659,740,1294,1319]" name="Poland" pageId="31" pageNumber="466">Poland</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>