treatments-xml/data/03/A6/87/03A687CCCA2F4770FF11F948FB4910D2.xml

<document id="4C781D4EC1BA912C30C175F996675772" ID-DOI="10.11646/zootaxa.4803.3.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3920609" ID-ZooBank="E8A09DAA-8E40-47AD-B268-F36FF7529BBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593414574084" checkinUser="plazi" docAuthor="Katwate, Unmesh, Kumkar, Pradeep, Raghavan, Rajeev &amp; Dahanukar, Neelesh" docDate="2020" docId="03A687CCCA2F4770FF11F948FB4910D2" docLanguage="en" docName="zootaxa.4803.3.9.pdf" docOrigin="Zootaxa 4803 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Waikhomia hira Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="553" masterDocId="FF9FFFB4CA2B4779FF86FFAFFF831612" masterDocTitle="Taxonomy and systematics of the ‘ Maharaja Barbs’ (Teleostei: Cyprinidae), with the description of a new genus and species from the Western Ghats, India" masterLastPageNumber="560" masterPageNumber="544" pageNumber="548" updateTime="1700277268978" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3BF06494B4FDC2DA551936017CBB051A">Taxonomy and systematics of the ‘ Maharaja Barbs’ (Teleostei: Cyprinidae), with the description of a new genus and species from the Western Ghats, India</mods:title>
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<mods:name id="4A4133D0567EB94B8DBD5C2DB58C10AC" type="personal">
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<mods:namePart id="6411D3A43C876CC2862DB2A32E0C0CE7">Katwate, Unmesh</mods:namePart>
<mods:affiliation id="6EEEE2A65869418F775A0BE13330DCDA">Freshwater Research Unit, Bombay Natural History Society (BNHS), Mumbai 400 001, Maharashtra, India. https: // orcid. org / 0000 - 0001 - 8924 - 4156 &amp; School of Ocean Science and Technology, Kerala University of Fisheries and Ocean Studies (KUFOS), Kochi 682 506, Kerala, India</mods:affiliation>
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<mods:namePart id="180CF4F435273BD06D87ADBD11E75C3A">Kumkar, Pradeep</mods:namePart>
<mods:affiliation id="2B89F10668935A885CEB557F8323F312">Department of Zoology, Modern College of Arts, Science and Commerce, Pune 411 007, Maharashtra, India. https: // orcid. org / 0000 - 0002 - 5443 - 1899</mods:affiliation>
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<mods:namePart id="FAAAA7BC1BA587F3BE062C4AFF6CE3AF">Raghavan, Rajeev</mods:namePart>
<mods:affiliation id="7BD4D5D18108868B60A3B7F5682109D5">Department of Fisheries Resource Management, Kerala University of Fisheries and Ocean Studies (KUFOS), Kochi 682 506, Kerala,</mods:affiliation>
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<mods:namePart id="49E8B44C2548D6E4E5043742B474DD0E">Dahanukar, Neelesh</mods:namePart>
<mods:affiliation id="95F5DB9ABA6A931F97BD12E46857C209">Indian Institute of Science Education and Research (IISER), Pune 411 008, Maharashtra, India. https: // orcid. org / 0000 - 0001 - 7162 - 9023 &amp; Systematics, Ecology and Conservation Laboratory, Zoo Outreach Organization (ZOO), Coimbatore, Tamil Nadu 641 035, India</mods:affiliation>
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<mods:date id="639E6086450ECFCFE0EBF1377A516317">2020</mods:date>
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<heading id="D0F881B6CA2F477DFF11F948FE441110" bold="true" box="[151,455,1767,1794]" fontSize="11" level="1" pageId="4" pageNumber="548" reason="1">
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<taxonomicName id="4C0F4D59CA2F477DFF11F948FEDF1110" authority="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar, 2020" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[151,348,1767,1794]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="548" phylum="Chordata" rank="species" species="hira" status="sp. nov.">
<emphasis id="B97BEAC8CA2F477DFF11F948FEDF1110" bold="true" box="[151,348,1767,1794]" italics="true" pageId="4" pageNumber="548">Waikhomia hira</emphasis>
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, 
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(
<figureCitation id="13342A5FCA2F477DFF19F8A4FE831134" box="[159,256,1803,1830]" captionStart="FIGURE 3" captionStartId="7.[151,250,1625,1650]" captionTargetBox="[151,1435,237,1600]" captionTargetId="figure@7.[151,1435,237,1600]" captionTargetPageId="7" captionText="FIGURE 3. Waikhomia hira holotype (male, BNHS FWF 602, 42.9 mm SL) in (A) life and (B) preserved." figureDoi="http://doi.org/10.5281/zenodo.3920623" httpUri="https://zenodo.org/record/3920623/files/figure.png" pageId="4" pageNumber="548">Figure 3</figureCitation>
, 
<figureCitation id="13342A5FCA2F477DFE8BF8A3FE9F1134" box="[269,284,1804,1830]" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="4" pageNumber="548">4</figureCitation>
)
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<subSubSection id="C3156551CA2F477DFF11F8E1FAE611EA" pageId="4" pageNumber="548" type="materials_examined">
<paragraph id="8BB036DACA2F477DFF11F8E1FDBE11A2" blockId="4.[151,1437,1870,2041]" pageId="4" pageNumber="548">
<materialsCitation id="3B673C87CA2F477DFF11F8E1FDBE11A2" collectingDate="2016-02-05" collectionCode="BNHS, R" collectorName="U. Katwate &amp; P. Kumkar &amp; N. Dahanukar &amp; N. Sawant &amp; N. Sood &amp; Britz" country="India" county="Uttara Kannada District" elevation="592" latitude="15.370501" location="Chandewadi" longLatPrecision="1" longitude="74.41" municipality="Kali River" pageId="4" pageNumber="548" specimenCode="FWF 602" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<emphasis id="B97BEAC8CA2F477DFF11F8E1FE88117A" bold="true" box="[151,267,1870,1896]" pageId="4" pageNumber="548">
<typeStatus id="54B48878CA2F477DFF11F8E1FE85117A" box="[151,262,1870,1896]" pageId="4" pageNumber="548" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED1EAE1FCA2F477DFE97F8E1FEDF117A" box="[273,348,1870,1896]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">BNHS</collectionCode>
<specimenCode id="DBA99EA1CA2F477DFEE7F8E0FE4E117A" box="[353,461,1870,1897]" collectionCode="FWF" pageId="4" pageNumber="548">FWF 602</specimenCode>
, 
<quantity id="4CF79B3FCA2F477DFE51F8E1FDB9117B" box="[471,570,1870,1897]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="4" pageNumber="548" unit="mm" value="42.9">42.9 mm</quantity>
SL, male; 
<collectingCountry id="F318764ACA2F477DFD36F8E1FD6D117A" box="[688,750,1870,1896]" name="India" pageId="4" pageNumber="548">India</collectingCountry>
: 
<collectingRegion id="49CBF838CA2F477DFD71F8E1FCEC117A" box="[759,879,1870,1896]" country="India" name="Karnataka" pageId="4" pageNumber="548">Karnataka</collectingRegion>
: 
<collectingCounty id="62D14E56CA2F477DFCFEF8E0FB07117A" box="[888,1156,1870,1897]" pageId="4" pageNumber="548">Uttara Kannada District</collectingCounty>
: 
<collectingMunicipality id="6BD4ACA0CA2F477DFB08F8E1FA81117A" box="[1166,1282,1870,1896]" pageId="4" pageNumber="548">Kali River</collectingMunicipality>
near 
<location id="8ED06001CA2F477DFABBF8E1FF53119E" LSID="urn:lsid:plazi:treatment:03A687CCCA2F4770FF11F948FB4910D2:8ED06001CA2F477DFABBF8E1FF53119E" country="India" county="Uttara Kannada District" latitude="15.370501" longLatPrecision="1" longitude="74.41" municipality="Kali River" name="Chandewadi" pageId="4" pageNumber="548" stateProvince="Karnataka">Chandewadi</location>
, 
<location id="8ED06001CA2F477DFF58F8DCFEAE119F" LSID="urn:lsid:plazi:treatment:03A687CCCA2F4770FF11F948FB4910D2:8ED06001CA2F477DFF58F8DCFEAE119F" box="[222,301,1907,1933]" country="India" county="Uttara Kannada District" latitude="15.370501" longLatPrecision="1" longitude="74.41" municipality="Kali River" name="Kamra" pageId="4" pageNumber="548" stateProvince="Karnataka">Kamra</location>
, 
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, 
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, 
<elevation id="0022D1E9CA2F477DFD1AF8DEFCA7119E" box="[668,804,1905,1933]" metricMagnitude="2" metricUnit="m" metricValue="5.92" pageId="4" pageNumber="548" unit="m" value="592.0">
<quantity id="4CF79B3FCA2F477DFD1AF8DEFD66119F" box="[668,741,1905,1933]" metricMagnitude="2" metricUnit="m" metricValue="5.92" pageId="4" pageNumber="548" unit="m" value="592.0">592 m</quantity>
a.s.l.
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, 
<collectorName id="26FA530CCA2F477DFCB4F8DCFC34119F" box="[818,951,1907,1933]" pageId="4" pageNumber="548">U. Katwate</collectorName>
, 
<collectorName id="26FA530CCA2F477DFC42F8DCFBBC119E" box="[964,1087,1906,1933]" pageId="4" pageNumber="548">P. Kumkar</collectorName>
, 
<collectorName id="26FA530CCA2F477DFBCAF8DCFB6E119E" box="[1100,1261,1906,1933]" pageId="4" pageNumber="548">N. Dahanukar</collectorName>
, 
<collectorName id="26FA530CCA2F477DFB7CF8DCFAF0119E" box="[1274,1395,1906,1933]" pageId="4" pageNumber="548">N. Sawant</collectorName>
, 
<collectorName id="26FA530CCA2F477DFA07F8DCFF5211A2" pageId="4" pageNumber="548">N. Sood</collectorName>
and 
<collectionCode id="ED1EAE1FCA2F477DFE81F838FE9E11A3" box="[263,285,1943,1969]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="548">R</collectionCode>
. 
<collectorName id="26FA530CCA2F477DFEAEF839FEE011A2" box="[296,355,1942,1968]" pageId="4" pageNumber="548">Britz</collectorName>
, 
<date id="FFB1101ACA2F477DFEE8F83AFDBA11A2" box="[366,569,1941,1968]" pageId="4" pageNumber="548" value="2016-02-05">
<collectingDate id="EFF5E9F2CA2F477DFEE8F83AFDBA11A2" box="[366,569,1941,1968]" pageId="4" pageNumber="548" value="2016-02-05">05 February 2016</collectingDate>
</date>
.
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</paragraph>
<paragraph id="8BB036DACA2F477DFF41F815FAE611EA" blockId="4.[151,1437,1870,2041]" pageId="4" pageNumber="548">
<materialsCitation id="3B673C87CA2F477DFF41F815FB5111C6" box="[199,1234,1977,2005]" collectingDate="2016-02-05" collectionCode="BNHS" collectorName="U. Katwate &amp; P. Kumkar &amp; N. Dahanukar &amp; N. Sawant &amp; N. Sood &amp; Britz" country="India" county="Uttara Kannada District" elevation="592" latitude="15.370501" location="Chandewadi" longLatPrecision="1" longitude="74.41" municipality="Kali River" pageId="4" pageNumber="548" specimenCode="FWF 603, 604, 607, 1002" specimenCount="4" stateProvince="Karnataka" typeStatus="paratype">
<emphasis id="B97BEAC8CA2F477DFF41F815FEC511C6" bold="true" box="[199,326,1978,2004]" pageId="4" pageNumber="548">
<typeStatus id="54B48878CA2F477DFF41F815FEC211C6" box="[199,321,1978,2004]" pageId="4" pageNumber="548" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED1EAE1FCA2F477DFECAF815FE1411C6" box="[332,407,1978,2004]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">BNHS</collectionCode>
<specimenCode id="DBA99EA1CA2F477DFE1AF814FD8B11C6" box="[412,520,1978,2005]" collectionCode="FWF" pageId="4" pageNumber="548">FWF 603</specimenCode>
, 
<specimenCode id="DBA99EA1CA2F477DFD94F815FDBC11C6" box="[530,575,1978,2004]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">604</specimenCode>
, 
<specimenCode id="DBA99EA1CA2F477DFDCFF815FDF511C6" box="[585,630,1978,2004]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">607</specimenCode>
, 
<specimenCode id="DBA99EA1CA2F477DFD06F815FD3811C6" box="[640,699,1978,2004]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">1002</specimenCode>
, 
<specimenCount id="9D09FD53CA2F477DFD43F815FD7F11C7" box="[709,764,1978,2005]" pageId="4" pageNumber="548" type="generic">4 ex.</specimenCount>
, 
<quantity id="4CF79B3FCA2F477DFC81F815FC2B11C7" box="[775,936,1977,2005]" metricMagnitude="-2" metricUnit="m" metricValue="4.175" metricValueMax="4.52" metricValueMin="3.8299999999999996" pageId="4" pageNumber="548" unit="mm" value="41.75" valueMax="45.2" valueMin="38.3">38.3–45.2 mm</quantity>
SL, same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B673C87CA2F477DFB5DF815FAE111EA" collectingDate="2014-03-27" collectionCode="BNHS" collectorName="P. Kumkar &amp; N. Sawant" country="India" county="Uttara Kannada District" elevation="592" latitude="15.370501" location="Chandewadi" longLatPrecision="1" longitude="74.41" municipality="Kali River" pageId="4" pageNumber="548" specimenCode="FWF 605, 997" specimenCount="2" stateProvince="Karnataka" typeStatus="paratype">
<collectionCode id="ED1EAE1FCA2F477DFB5DF815FAA511C6" box="[1243,1318,1978,2004]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">BNHS</collectionCode>
<specimenCode id="DBA99EA1CA2F477DFAAAF814FA1B11C6" box="[1324,1432,1977,2005]" collectionCode="FWF" pageId="4" pageNumber="548">FWF 605</specimenCode>
, 
<specimenCode id="DBA99EA1CA2F477DFF11F871FF4711EA" box="[151,196,2014,2040]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="548">997</specimenCode>
, 
<specimenCount id="9D09FD53CA2F477DFF49F871FE8B11EB" box="[207,264,2014,2041]" pageId="4" pageNumber="548" type="generic">2 ex.</specimenCount>
, 
<quantity id="4CF79B3FCA2F477DFE92F871FE3411EB" box="[276,439,2014,2041]" metricMagnitude="-2" metricUnit="m" metricValue="2.845" metricValueMax="3.62" metricValueMin="2.07" pageId="4" pageNumber="548" unit="mm" value="28.45" valueMax="36.2" valueMin="20.7">20.7–36.2 mm</quantity>
SL, same coll. locality data as holotype, 
<collectorName id="26FA530CCA2F477DFC05F870FC7911EA" box="[899,1018,2014,2041]" pageId="4" pageNumber="548">P. Kumkar</collectorName>
and 
<collectorName id="26FA530CCA2F477DFBB6F870FB2411EA" box="[1072,1191,2014,2041]" pageId="4" pageNumber="548">N. Sawant</collectorName>
, 
<date id="FFB1101ACA2F477DFB35F871FAE111EA" box="[1203,1378,2014,2040]" pageId="4" pageNumber="548" value="2014-03-27">
<collectingDate id="EFF5E9F2CA2F477DFB35F871FAE111EA" box="[1203,1378,2014,2040]" pageId="4" pageNumber="548" value="2014-03-27">27 March 2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3156551CA2E477CFF41FF37FC9317C1" pageId="5" pageNumber="549" type="diagnosis">
<paragraph id="8BB036DACA2E477CFF41FF37FC9317C1" blockId="5.[151,1437,152,539]" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF41FF37FEC116A1" bold="true" box="[199,322,152,179]" pageId="5" pageNumber="549">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D59CA2E477CFECAFF37FD8116A0" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[332,514,152,178]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2E477CFECAFF37FD8116A0" box="[332,514,152,178]" italics="true" pageId="5" pageNumber="549">Waikhomia hira</emphasis>
</taxonomicName>
is distinguished from its only congener, 
<emphasis id="B97BEAC8CA2E477CFC66FF36FB2416A0" box="[992,1191,152,178]" italics="true" pageId="5" pageNumber="549">
<taxonomicName id="4C0F4D59CA2E477CFC66FF36FB2016A0" box="[992,1187,152,178]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="sahyadriensis">W. sahyadriensis</taxonomicName>
,
</emphasis>
by a combination of characters that includes small size (maximum recorded standard length 
<quantity id="4CF79B3FCA2E477CFC38FF13FBA416C5" box="[958,1063,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="5" pageNumber="549" unit="mm" value="45.2">45.2 mm</quantity>
); lateral line straight, with 23–25 perforated scales; pre-dorsal and pre-pelvic scales 10; 4–½4 scales between dorsal-fin origin and lateral-line scale row, 3½ between lateral line row and pelvic-fin origin. Furthermore, 
<taxonomicName id="4C0F4D59CA2E477CFC2FFEAAFC7F170C" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[937,1020,260,286]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2E477CFC2FFEAAFC7F170C" box="[937,1020,260,286]" italics="true" pageId="5" pageNumber="549">W. hira</emphasis>
</taxonomicName>
is distinguished by having a distinct body coloration consisting of: small rhomboidal spots (7–8) largely restricted to lateral-line scale row; nape and dorsal-base spot absent; cleithral spot small, confined to the first scale of lateral line scale row; post-pectoral and super-anal spots small; a lateral spot present on or between 8 
<superScript id="7C7A9B92CA2E477CFCB8FEC0FCD91799" attach="none" box="[830,858,367,395]" fontSize="6" pageId="5" pageNumber="549">th–</superScript>
9 
<superScript id="7C7A9B92CA2E477CFCE1FEC0FCF7176F" attach="left" box="[871,884,367,381]" fontSize="6" pageId="5" pageNumber="549">th</superScript>
lateral-line scales; two spots on caudal peduncle; no spots coalescing into blotches, bars or bands. Gill rakers 2–3 on distal end of epibranchial 1; infraorbital 2 narrow; supraneural 3 broad, separated from neural spine 4.
</paragraph>
</subSubSection>
<subSubSection id="C3156551CA2E4770FF41FE73FD241529" lastPageId="9" lastPageNumber="553" pageId="5" pageNumber="549" type="description">
<paragraph id="8BB036DACA2E477CFF41FE73FB321409" blockId="5.[151,1437,152,539]" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF41FE73FEDA17E5" bold="true" box="[199,345,476,503]" pageId="5" pageNumber="549">Description.</emphasis>
A small cyprinid, maximum adult size 
<quantity id="4CF79B3FCA2E477CFC96FE73FCF717E5" box="[784,884,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="5" pageNumber="549" unit="mm" value="45.2">45.2 mm</quantity>
SL. For general shape and appearance see 
<figureCitation id="13342A5FCA2E477CFAD5FE73FF241409" captionStart="FIGURE 3" captionStartId="7.[151,250,1625,1650]" captionTargetBox="[151,1435,237,1600]" captionTargetId="figure@7.[151,1435,237,1600]" captionTargetPageId="7" captionText="FIGURE 3. Waikhomia hira holotype (male, BNHS FWF 602, 42.9 mm SL) in (A) life and (B) preserved." figureDoi="http://doi.org/10.5281/zenodo.3920623" httpUri="https://zenodo.org/record/3920623/files/figure.png" pageId="5" pageNumber="549">Figure 3</figureCitation>
. Morphometric and meristic data of the 
<typeStatus id="54B48878CA2E477CFDECFDAFFD4F1408" box="[618,716,512,538]" pageId="5" pageNumber="549" type="holotype">holotype</typeStatus>
and five 
<typeStatus id="54B48878CA2E477CFCB3FDAEFC231409" box="[821,928,513,539]" pageId="5" pageNumber="549" type="paratype">paratypes</typeStatus>
are provided in 
<tableCitation id="C68D0361CA2E477CFBD1FDAFFB2E1409" box="[1111,1197,512,539]" captionStart="TABLE 2" captionStartId="5.[151,239,586,610]" captionText="TABLE 2. Biometric data of Waikhomia hira type material (holotype, BNHS FWF 602 and paratypes, BNHS FWF 603– 605, 607, 1002 and 997) and comparative material of Waikhomia sahyadriensis (BNHS FWF 582–597 and 599–601)." pageId="5" pageNumber="549">Table 2</tableCitation>
.
</paragraph>
<caption id="DF706652CA2E477CFF11FDE5FAFC1494" pageId="5" pageNumber="549">
<paragraph id="8BB036DACA2E477CFF11FDE5FAFC1494" blockId="5.[151,1436,586,646]" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF11FDE5FE8A1470" bold="true" box="[151,265,586,610]" pageId="5" pageNumber="549">TABLE 2.</emphasis>
Biometric data of 
<taxonomicName id="4C0F4D59CA2E477CFE56FDE5FDFA1470" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[464,633,586,610]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2E477CFE56FDE5FDFA1470" box="[464,633,586,610]" italics="true" pageId="5" pageNumber="549">Waikhomia hira</emphasis>
</taxonomicName>
type material (holotype, BNHS FWF 602 and paratypes, BNHS FWF 603– 605, 607, 1002 and 997) and comparative material of 
<taxonomicName id="4C0F4D59CA2E477CFD53FDC1FC651494" box="[725,998,622,646]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA2E477CFD53FDC1FC651494" box="[725,998,622,646]" italics="true" pageId="5" pageNumber="549">Waikhomia sahyadriensis</emphasis>
</taxonomicName>
(BNHS FWF 582–597 and 599–601).
</paragraph>
</caption>
<paragraph id="8BB036DACA2E477CFF25FD39FAE211AC" pageId="5" pageNumber="549">
<table id="F90FC47ACA2EB886FF25FD39FA1211AC" box="[163,1425,662,1982]" colsContinueIn="6.[163,1425,204,1133]" gridcols="7" gridrows="31" pageId="5" pageNumber="549">
<tr id="353F3498CA2EB886FF25FD39FA1214BD" box="[163,1425,662,687]" gridrow="0" pageId="5" pageNumber="549" rowspan-3="1" rowspan-4="1">
<th id="76EE5DE4CA2EB886FF25FD39FE5D14BD" box="[163,478,662,687]" gridcol="0" gridrow="0" pageId="5" pageNumber="549">Characters</th>
<th id="76EE5DE4CA2EB886FD87FD39FD7614BD" box="[513,757,662,687]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="5" pageNumber="549">
<taxonomicName id="4C0F4D59CA2E477CFD87FD39FD2114BC" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[513,674,662,686]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2E477CFD87FD39FD2114BC" box="[513,674,662,686]" italics="true" pageId="5" pageNumber="549">Waikhomia hira</emphasis>
</taxonomicName>
</th>
<th id="76EE5DE4CA2EB886FBC8FD39FA1214BD" box="[1102,1425,662,687]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="5" pageNumber="549">
<taxonomicName id="4C0F4D59CA2E477CFBC8FD39FAD314BC" box="[1102,1360,662,686]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="549" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA2E477CFBC8FD39FAD314BC" box="[1102,1360,662,686]" italics="true" pageId="5" pageNumber="549">Waikhomia sahyadriensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F3498CA2EB886FF25FD11FA121505" box="[163,1425,702,791]" gridrow="1" pageId="5" pageNumber="549" rowspan-0="1">
<td id="76EE5DE4CA2EB886FD87FD11FDDC1505" box="[513,607,702,791]" gridcol="1" gridrow="1" pageId="5" pageNumber="549">Holotype (602)</td>
<td id="76EE5DE4CA2EB886FD0BFD11FC091505" box="[653,906,702,791]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="5" pageNumber="549">Paratypes (n= 5) (603–605, 607, 1002)</td>
<td id="76EE5DE4CA2EB886FC53FD11FBAD1505" box="[981,1070,702,791]" gridcol="4" gridrow="1" pageId="5" pageNumber="549">Paratype (997)</td>
<td id="76EE5DE4CA2EB886FBC8FD11FA121505" box="[1102,1425,702,791]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="5" pageNumber="549">Topotypes + additional material (n=19) (582–597, 599–601)</td>
</tr>
<tr id="353F3498CA2EB886FF25FC8EFA121528" box="[163,1425,801,826]" gridrow="2" pageId="5" pageNumber="549" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="76EE5DE4CA2EB886FD0BFC8EFD761528" box="[653,757,801,826]" gridcol="2" gridrow="2" pageId="5" pageNumber="549">Mean (sd)</td>
<td id="76EE5DE4CA2EB886FCA2FC8EFC091528" box="[804,906,801,826]" gridcol="3" gridrow="2" pageId="5" pageNumber="549">Range</td>
<td id="76EE5DE4CA2EB886FBC8FC8EFB3E1528" box="[1102,1213,801,826]" gridcol="5" gridrow="2" pageId="5" pageNumber="549">Mean (sd)</td>
<td id="76EE5DE4CA2EB886FB7DFC8EFA121528" box="[1275,1425,801,826]" gridcol="6" gridrow="2" pageId="5" pageNumber="549">Range</td>
</tr>
<tr id="353F3498CA2EB886FF25FCE6FA121570" box="[163,1425,841,866]" gridrow="3" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FCE6FA121570" box="[163,1425,841,866]" colspan="7" colspanRight="6" gridcol="0" gridrow="3" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF25FCE6FEC01570" bold="true" box="[163,323,841,866]" italics="true" pageId="5" pageNumber="549">Morphometrics</emphasis>
</th>
</tr>
<tr id="353F3498CA2EB886FF25FCDEFA121598" box="[163,1425,881,906]" gridrow="4" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FCDEFE5D1598" box="[163,478,881,906]" gridcol="0" gridrow="4" pageId="5" pageNumber="549">Total length (mm)</th>
<td id="76EE5DE4CA2EB886FD87FCDEFDDC1598" box="[513,607,881,906]" gridcol="1" gridrow="4" pageId="5" pageNumber="549">55.8</td>
<td id="76EE5DE4CA2EB886FD0BFCDEFD761598" box="[653,757,881,906]" gridcol="2" gridrow="4" pageId="5" pageNumber="549">53.0 (5.2)</td>
<td id="76EE5DE4CA2EB886FCA2FCDEFC091598" box="[804,906,881,906]" gridcol="3" gridrow="4" pageId="5" pageNumber="549">46.2–60.6</td>
<td id="76EE5DE4CA2EB886FC53FCDEFBAD1598" box="[981,1070,881,906]" gridcol="4" gridrow="4" pageId="5" pageNumber="549">27.1</td>
<td id="76EE5DE4CA2EB886FBC8FCDEFB3E1598" box="[1102,1213,881,906]" gridcol="5" gridrow="4" pageId="5" pageNumber="549">53.8 (11.5)</td>
<td id="76EE5DE4CA2EB886FB7DFCDEFA121598" box="[1275,1425,881,906]" gridcol="6" gridrow="4" pageId="5" pageNumber="549">36.1–75.1</td>
</tr>
<tr id="353F3498CA2EB886FF25FC35FA1215A1" box="[163,1425,922,947]" gridrow="5" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FC35FE5D15A1" box="[163,478,922,947]" gridcol="0" gridrow="5" pageId="5" pageNumber="549">Standard length (SL, mm)</th>
<td id="76EE5DE4CA2EB886FD87FC35FDDC15A1" box="[513,607,922,947]" gridcol="1" gridrow="5" pageId="5" pageNumber="549">42.9</td>
<td id="76EE5DE4CA2EB886FD0BFC35FD7615A1" box="[653,757,922,947]" gridcol="2" gridrow="5" pageId="5" pageNumber="549">40.1 (3.4)</td>
<td id="76EE5DE4CA2EB886FCA2FC35FC0915A1" box="[804,906,922,947]" gridcol="3" gridrow="5" pageId="5" pageNumber="549">36.2–45.2</td>
<td id="76EE5DE4CA2EB886FC53FC35FBAD15A1" box="[981,1070,922,947]" gridcol="4" gridrow="5" pageId="5" pageNumber="549">20.7</td>
<td id="76EE5DE4CA2EB886FBC8FC35FB3E15A1" box="[1102,1213,922,947]" gridcol="5" gridrow="5" pageId="5" pageNumber="549">41.8 (9.1)</td>
<td id="76EE5DE4CA2EB886FB7DFC35FA1215A1" box="[1275,1425,922,947]" gridcol="6" gridrow="5" pageId="5" pageNumber="549">28.7–58.9</td>
</tr>
<tr id="353F3498CA2EB886FF25FC6EFA1215C8" box="[163,1425,961,986]" gridrow="6" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FC6EFA1215C8" box="[163,1425,961,986]" colspan="7" colspanRight="6" gridcol="0" gridrow="6" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF25FC6EFF5815C8" bold="true" box="[163,219,961,986]" pageId="5" pageNumber="549">%SL</emphasis>
</th>
</tr>
<tr id="353F3498CA2EB886FF25FC45FA121211" box="[163,1425,1002,1027]" gridrow="7" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FC45FE5D1211" box="[163,478,1002,1027]" gridcol="0" gridrow="7" pageId="5" pageNumber="549">Head length (HL)</th>
<td id="76EE5DE4CA2EB886FD87FC45FDDC1211" box="[513,607,1002,1027]" gridcol="1" gridrow="7" pageId="5" pageNumber="549">28.9</td>
<td id="76EE5DE4CA2EB886FD0BFC45FD761211" box="[653,757,1002,1027]" gridcol="2" gridrow="7" pageId="5" pageNumber="549">28.5 (0.7)</td>
<td id="76EE5DE4CA2EB886FCA2FC45FC091211" box="[804,906,1002,1027]" gridcol="3" gridrow="7" pageId="5" pageNumber="549">27.6–29.2</td>
<td id="76EE5DE4CA2EB886FC53FC45FBAD1211" box="[981,1070,1002,1027]" gridcol="4" gridrow="7" pageId="5" pageNumber="549">30.0</td>
<td id="76EE5DE4CA2EB886FBC8FC45FB3E1211" box="[1102,1213,1002,1027]" gridcol="5" gridrow="7" pageId="5" pageNumber="549">28.1 (2.2)</td>
<td id="76EE5DE4CA2EB886FB7DFC45FA121211" box="[1275,1425,1002,1027]" gridcol="6" gridrow="7" pageId="5" pageNumber="549">24.1–31.7</td>
</tr>
<tr id="353F3498CA2EB886FF25FBBDFA121239" box="[163,1425,1042,1067]" gridrow="8" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FBBDFE5D1239" box="[163,478,1042,1067]" gridcol="0" gridrow="8" pageId="5" pageNumber="549">Post-orbital head length</th>
<td id="76EE5DE4CA2EB886FD87FBBDFDDC1239" box="[513,607,1042,1067]" gridcol="1" gridrow="8" pageId="5" pageNumber="549">11.2</td>
<td id="76EE5DE4CA2EB886FD0BFBBDFD761239" box="[653,757,1042,1067]" gridcol="2" gridrow="8" pageId="5" pageNumber="549">11.3 (0.3)</td>
<td id="76EE5DE4CA2EB886FCA2FBBDFC091239" box="[804,906,1042,1067]" gridcol="3" gridrow="8" pageId="5" pageNumber="549">11.0–11.6</td>
<td id="76EE5DE4CA2EB886FC53FBBDFBAD1239" box="[981,1070,1042,1067]" gridcol="4" gridrow="8" pageId="5" pageNumber="549">11.1</td>
<td id="76EE5DE4CA2EB886FBC8FBBDFB3E1239" box="[1102,1213,1042,1067]" gridcol="5" gridrow="8" pageId="5" pageNumber="549">10.8 (0.9)</td>
<td id="76EE5DE4CA2EB886FB7DFBBDFA121239" box="[1275,1425,1042,1067]" gridcol="6" gridrow="8" pageId="5" pageNumber="549">8.7–12.4</td>
</tr>
<tr id="353F3498CA2EB886FF25FB95FA121241" box="[163,1425,1082,1107]" gridrow="9" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FB95FE5D1241" box="[163,478,1082,1107]" gridcol="0" gridrow="9" pageId="5" pageNumber="549">Head depth</th>
<td id="76EE5DE4CA2EB886FD87FB95FDDC1241" box="[513,607,1082,1107]" gridcol="1" gridrow="9" pageId="5" pageNumber="549">21.2</td>
<td id="76EE5DE4CA2EB886FD0BFB95FD761241" box="[653,757,1082,1107]" gridcol="2" gridrow="9" pageId="5" pageNumber="549">23.0 (0.6)</td>
<td id="76EE5DE4CA2EB886FCA2FB95FC091241" box="[804,906,1082,1107]" gridcol="3" gridrow="9" pageId="5" pageNumber="549">22.3–23.5</td>
<td id="76EE5DE4CA2EB886FC53FB95FBAD1241" box="[981,1070,1082,1107]" gridcol="4" gridrow="9" pageId="5" pageNumber="549">21.3</td>
<td id="76EE5DE4CA2EB886FBC8FB95FB3E1241" box="[1102,1213,1082,1107]" gridcol="5" gridrow="9" pageId="5" pageNumber="549">23.5 (1.6)</td>
<td id="76EE5DE4CA2EB886FB7DFB95FA121241" box="[1275,1425,1082,1107]" gridcol="6" gridrow="9" pageId="5" pageNumber="549">20.5–26.1</td>
</tr>
<tr id="353F3498CA2EB886FF25FBCCFA12126E" box="[163,1425,1123,1148]" gridrow="10" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FBCCFE5D126E" box="[163,478,1123,1148]" gridcol="0" gridrow="10" pageId="5" pageNumber="549">Head width</th>
<td id="76EE5DE4CA2EB886FD87FBCCFDDC126E" box="[513,607,1123,1148]" gridcol="1" gridrow="10" pageId="5" pageNumber="549">15.9</td>
<td id="76EE5DE4CA2EB886FD0BFBCCFD76126E" box="[653,757,1123,1148]" gridcol="2" gridrow="10" pageId="5" pageNumber="549">17.1 (0.7)</td>
<td id="76EE5DE4CA2EB886FCA2FBCCFC09126E" box="[804,906,1123,1148]" gridcol="3" gridrow="10" pageId="5" pageNumber="549">16.2–18.0</td>
<td id="76EE5DE4CA2EB886FC53FBCCFBAD126E" box="[981,1070,1123,1148]" gridcol="4" gridrow="10" pageId="5" pageNumber="549">15.0</td>
<td id="76EE5DE4CA2EB886FBC8FBCCFB3E126E" box="[1102,1213,1123,1148]" gridcol="5" gridrow="10" pageId="5" pageNumber="549">16.9 (1.1)</td>
<td id="76EE5DE4CA2EB886FB7DFBCCFA12126E" box="[1275,1425,1123,1148]" gridcol="6" gridrow="10" pageId="5" pageNumber="549">14.9–18.4</td>
</tr>
<tr id="353F3498CA2EB886FF25FB24FA1212B6" box="[163,1425,1163,1188]" gridrow="11" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FB24FE5D12B6" box="[163,478,1163,1188]" gridcol="0" gridrow="11" pageId="5" pageNumber="549">Body depth</th>
<td id="76EE5DE4CA2EB886FD87FB24FDDC12B6" box="[513,607,1163,1188]" gridcol="1" gridrow="11" pageId="5" pageNumber="549">34.0</td>
<td id="76EE5DE4CA2EB886FD0BFB24FD7612B6" box="[653,757,1163,1188]" gridcol="2" gridrow="11" pageId="5" pageNumber="549">36.4 (1.4)</td>
<td id="76EE5DE4CA2EB886FCA2FB24FC0912B6" box="[804,906,1163,1188]" gridcol="3" gridrow="11" pageId="5" pageNumber="549">34.5–37.8</td>
<td id="76EE5DE4CA2EB886FC53FB24FBAD12B6" box="[981,1070,1163,1188]" gridcol="4" gridrow="11" pageId="5" pageNumber="549">30.0</td>
<td id="76EE5DE4CA2EB886FBC8FB24FB3E12B6" box="[1102,1213,1163,1188]" gridcol="5" gridrow="11" pageId="5" pageNumber="549">36.7 (1.3)</td>
<td id="76EE5DE4CA2EB886FB7DFB24FA1212B6" box="[1275,1425,1163,1188]" gridcol="6" gridrow="11" pageId="5" pageNumber="549">34.3–39.0</td>
</tr>
<tr id="353F3498CA2EB886FF25FB1CFA1212DE" box="[163,1425,1203,1228]" gridrow="12" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FB1CFE5D12DE" box="[163,478,1203,1228]" gridcol="0" gridrow="12" pageId="5" pageNumber="549">Body width at dorsal-fin origin</th>
<td id="76EE5DE4CA2EB886FD87FB1CFDDC12DE" box="[513,607,1203,1228]" gridcol="1" gridrow="12" pageId="5" pageNumber="549">15.9</td>
<td id="76EE5DE4CA2EB886FD0BFB1CFD7612DE" box="[653,757,1203,1228]" gridcol="2" gridrow="12" pageId="5" pageNumber="549">15.0 (0.6)</td>
<td id="76EE5DE4CA2EB886FCA2FB1CFC0912DE" box="[804,906,1203,1228]" gridcol="3" gridrow="12" pageId="5" pageNumber="549">14.6–15.7</td>
<td id="76EE5DE4CA2EB886FC53FB1CFBAD12DE" box="[981,1070,1203,1228]" gridcol="4" gridrow="12" pageId="5" pageNumber="549">11.6</td>
<td id="76EE5DE4CA2EB886FBC8FB1CFB3E12DE" box="[1102,1213,1203,1228]" gridcol="5" gridrow="12" pageId="5" pageNumber="549">15.2 (1.4)</td>
<td id="76EE5DE4CA2EB886FB7DFB1CFA1212DE" box="[1275,1425,1203,1228]" gridcol="6" gridrow="12" pageId="5" pageNumber="549">12.7–17.4</td>
</tr>
<tr id="353F3498CA2EB886FF25FB74FA1212E6" box="[163,1425,1243,1268]" gridrow="13" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FB74FE5D12E6" box="[163,478,1243,1268]" gridcol="0" gridrow="13" pageId="5" pageNumber="549">Body width at anal-fin origin</th>
<td id="76EE5DE4CA2EB886FD87FB74FDDC12E6" box="[513,607,1243,1268]" gridcol="1" gridrow="13" pageId="5" pageNumber="549">12.6</td>
<td id="76EE5DE4CA2EB886FD0BFB74FD7612E6" box="[653,757,1243,1268]" gridcol="2" gridrow="13" pageId="5" pageNumber="549">10.0 (1.4)</td>
<td id="76EE5DE4CA2EB886FCA2FB74FC0912E6" box="[804,906,1243,1268]" gridcol="3" gridrow="13" pageId="5" pageNumber="549">8.8–11.5</td>
<td id="76EE5DE4CA2EB886FC53FB74FBAD12E6" box="[981,1070,1243,1268]" gridcol="4" gridrow="13" pageId="5" pageNumber="549">7.7</td>
<td id="76EE5DE4CA2EB886FBC8FB74FB3E12E6" box="[1102,1213,1243,1268]" gridcol="5" gridrow="13" pageId="5" pageNumber="549">10.5 (2.2)</td>
<td id="76EE5DE4CA2EB886FB7DFB74FA1212E6" box="[1275,1425,1243,1268]" gridcol="6" gridrow="13" pageId="5" pageNumber="549">6.9–15.2</td>
</tr>
<tr id="353F3498CA2EB886FF25FAABFA12130F" box="[163,1425,1284,1309]" gridrow="14" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FAABFE5D130F" box="[163,478,1284,1309]" gridcol="0" gridrow="14" pageId="5" pageNumber="549">Pre-dorsal distance</th>
<td id="76EE5DE4CA2EB886FD87FAABFDDC130F" box="[513,607,1284,1309]" gridcol="1" gridrow="14" pageId="5" pageNumber="549">49.2</td>
<td id="76EE5DE4CA2EB886FD0BFAABFD76130F" box="[653,757,1284,1309]" gridcol="2" gridrow="14" pageId="5" pageNumber="549">47.5 (1.6)</td>
<td id="76EE5DE4CA2EB886FCA2FAABFC09130F" box="[804,906,1284,1309]" gridcol="3" gridrow="14" pageId="5" pageNumber="549">45.2–49.4</td>
<td id="76EE5DE4CA2EB886FC53FAABFBAD130F" box="[981,1070,1284,1309]" gridcol="4" gridrow="14" pageId="5" pageNumber="549">51.2</td>
<td id="76EE5DE4CA2EB886FBC8FAABFB3E130F" box="[1102,1213,1284,1309]" gridcol="5" gridrow="14" pageId="5" pageNumber="549">47.2 (2.5)</td>
<td id="76EE5DE4CA2EB886FB7DFAABFA12130F" box="[1275,1425,1284,1309]" gridcol="6" gridrow="14" pageId="5" pageNumber="549">42.6–52.0</td>
</tr>
<tr id="353F3498CA2EB886FF25FA83FA121357" box="[163,1425,1324,1349]" gridrow="15" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FA83FE5D1357" box="[163,478,1324,1349]" gridcol="0" gridrow="15" pageId="5" pageNumber="549">Post-dorsal distance</th>
<td id="76EE5DE4CA2EB886FD87FA83FDDC1357" box="[513,607,1324,1349]" gridcol="1" gridrow="15" pageId="5" pageNumber="549">83.4</td>
<td id="76EE5DE4CA2EB886FD0BFA83FD761357" box="[653,757,1324,1349]" gridcol="2" gridrow="15" pageId="5" pageNumber="549">88.6 (3.9)</td>
<td id="76EE5DE4CA2EB886FCA2FA83FC091357" box="[804,906,1324,1349]" gridcol="3" gridrow="15" pageId="5" pageNumber="549">82.0–91.6</td>
<td id="76EE5DE4CA2EB886FC53FA83FBAD1357" box="[981,1070,1324,1349]" gridcol="4" gridrow="15" pageId="5" pageNumber="549">83.1</td>
<td id="76EE5DE4CA2EB886FBC8FA83FB3E1357" box="[1102,1213,1324,1349]" gridcol="5" gridrow="15" pageId="5" pageNumber="549">83.4 (3.4)</td>
<td id="76EE5DE4CA2EB886FB7DFA83FA121357" box="[1275,1425,1324,1349]" gridcol="6" gridrow="15" pageId="5" pageNumber="549">76.5–91.0</td>
</tr>
<tr id="353F3498CA2EB886FF25FAFBFA12137F" box="[163,1425,1364,1389]" gridrow="16" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FAFBFE5D137F" box="[163,478,1364,1389]" gridcol="0" gridrow="16" pageId="5" pageNumber="549">Dorsal to hypural distance</th>
<td id="76EE5DE4CA2EB886FD87FAFBFDDC137F" box="[513,607,1364,1389]" gridcol="1" gridrow="16" pageId="5" pageNumber="549">55.5</td>
<td id="76EE5DE4CA2EB886FD0BFAFBFD76137F" box="[653,757,1364,1389]" gridcol="2" gridrow="16" pageId="5" pageNumber="549">55.2 (1.9)</td>
<td id="76EE5DE4CA2EB886FCA2FAFBFC09137F" box="[804,906,1364,1389]" gridcol="3" gridrow="16" pageId="5" pageNumber="549">52.5–57.2</td>
<td id="76EE5DE4CA2EB886FC53FAFBFBAD137F" box="[981,1070,1364,1389]" gridcol="4" gridrow="16" pageId="5" pageNumber="549">53.6</td>
<td id="76EE5DE4CA2EB886FBC8FAFBFB3E137F" box="[1102,1213,1364,1389]" gridcol="5" gridrow="16" pageId="5" pageNumber="549">56.2 (2.9)</td>
<td id="76EE5DE4CA2EB886FB7DFAFBFA12137F" box="[1275,1425,1364,1389]" gridcol="6" gridrow="16" pageId="5" pageNumber="549">51.8–64.1</td>
</tr>
<tr id="353F3498CA2EB886FF25FAD3FA121387" box="[163,1425,1404,1429]" gridrow="17" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FAD3FE5D1387" box="[163,478,1404,1429]" gridcol="0" gridrow="17" pageId="5" pageNumber="549">Pre-pelvic distance</th>
<td id="76EE5DE4CA2EB886FD87FAD3FDDC1387" box="[513,607,1404,1429]" gridcol="1" gridrow="17" pageId="5" pageNumber="549">48.0</td>
<td id="76EE5DE4CA2EB886FD0BFAD3FD761387" box="[653,757,1404,1429]" gridcol="2" gridrow="17" pageId="5" pageNumber="549">50.2 (1.1)</td>
<td id="76EE5DE4CA2EB886FCA2FAD3FC091387" box="[804,906,1404,1429]" gridcol="3" gridrow="17" pageId="5" pageNumber="549">48.5–51.1</td>
<td id="76EE5DE4CA2EB886FC53FAD3FBAD1387" box="[981,1070,1404,1429]" gridcol="4" gridrow="17" pageId="5" pageNumber="549">49.3</td>
<td id="76EE5DE4CA2EB886FBC8FAD3FB3E1387" box="[1102,1213,1404,1429]" gridcol="5" gridrow="17" pageId="5" pageNumber="549">48.8 (1.9)</td>
<td id="76EE5DE4CA2EB886FB7DFAD3FA121387" box="[1275,1425,1404,1429]" gridcol="6" gridrow="17" pageId="5" pageNumber="549">46.3–52.4</td>
</tr>
<tr id="353F3498CA2EB886FF25FA0AFA1213AC" box="[163,1425,1445,1470]" gridrow="18" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FA0AFE5D13AC" box="[163,478,1445,1470]" gridcol="0" gridrow="18" pageId="5" pageNumber="549">Pre-anal distance</th>
<td id="76EE5DE4CA2EB886FD87FA0AFDDC13AC" box="[513,607,1445,1470]" gridcol="1" gridrow="18" pageId="5" pageNumber="549">69.2</td>
<td id="76EE5DE4CA2EB886FD0BFA0AFD7613AC" box="[653,757,1445,1470]" gridcol="2" gridrow="18" pageId="5" pageNumber="549">71.2 (1.2)</td>
<td id="76EE5DE4CA2EB886FCA2FA0AFC0913AC" box="[804,906,1445,1470]" gridcol="3" gridrow="18" pageId="5" pageNumber="549">70.1–72.6</td>
<td id="76EE5DE4CA2EB886FC53FA0AFBAD13AC" box="[981,1070,1445,1470]" gridcol="4" gridrow="18" pageId="5" pageNumber="549">66.7</td>
<td id="76EE5DE4CA2EB886FBC8FA0AFB3E13AC" box="[1102,1213,1445,1470]" gridcol="5" gridrow="18" pageId="5" pageNumber="549">71.8 (2.0)</td>
<td id="76EE5DE4CA2EB886FB7DFA0AFA1213AC" box="[1275,1425,1445,1470]" gridcol="6" gridrow="18" pageId="5" pageNumber="549">68.0–74.9</td>
</tr>
<tr id="353F3498CA2EB886FF25FA62FA1213F4" box="[163,1425,1485,1510]" gridrow="19" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FA62FE5D13F4" box="[163,478,1485,1510]" gridcol="0" gridrow="19" pageId="5" pageNumber="549">Pre-pectoral distance</th>
<td id="76EE5DE4CA2EB886FD87FA62FDDC13F4" box="[513,607,1485,1510]" gridcol="1" gridrow="19" pageId="5" pageNumber="549">28.2</td>
<td id="76EE5DE4CA2EB886FD0BFA62FD7613F4" box="[653,757,1485,1510]" gridcol="2" gridrow="19" pageId="5" pageNumber="549">33.2 (9.4)</td>
<td id="76EE5DE4CA2EB886FCA2FA62FC0913F4" box="[804,906,1485,1510]" gridcol="3" gridrow="19" pageId="5" pageNumber="549">26.5–49.3</td>
<td id="76EE5DE4CA2EB886FC53FA62FBAD13F4" box="[981,1070,1485,1510]" gridcol="4" gridrow="19" pageId="5" pageNumber="549">28.5</td>
<td id="76EE5DE4CA2EB886FBC8FA62FB3E13F4" box="[1102,1213,1485,1510]" gridcol="5" gridrow="19" pageId="5" pageNumber="549">27.9 (2.4)</td>
<td id="76EE5DE4CA2EB886FB7DFA62FA1213F4" box="[1275,1425,1485,1510]" gridcol="6" gridrow="19" pageId="5" pageNumber="549">23.5–31.0</td>
</tr>
<tr id="353F3498CA2EB886FF25FA5AFA12101C" box="[163,1425,1525,1550]" gridrow="20" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25FA5AFE5D101C" box="[163,478,1525,1550]" gridcol="0" gridrow="20" pageId="5" pageNumber="549">Dorsal fin length</th>
<td id="76EE5DE4CA2EB886FD87FA5AFDDC101C" box="[513,607,1525,1550]" gridcol="1" gridrow="20" pageId="5" pageNumber="549">31.5</td>
<td id="76EE5DE4CA2EB886FD0BFA5AFD76101C" box="[653,757,1525,1550]" gridcol="2" gridrow="20" pageId="5" pageNumber="549">31.3 (1.3)</td>
<td id="76EE5DE4CA2EB886FCA2FA5AFC09101C" box="[804,906,1525,1550]" gridcol="3" gridrow="20" pageId="5" pageNumber="549">30.3–33.4</td>
<td id="76EE5DE4CA2EB886FC53FA5AFBAD101C" box="[981,1070,1525,1550]" gridcol="4" gridrow="20" pageId="5" pageNumber="549">27.1</td>
<td id="76EE5DE4CA2EB886FBC8FA5AFB3E101C" box="[1102,1213,1525,1550]" gridcol="5" gridrow="20" pageId="5" pageNumber="549">30.9 (1.5)</td>
<td id="76EE5DE4CA2EB886FB7DFA5AFA12101C" box="[1275,1425,1525,1550]" gridcol="6" gridrow="20" pageId="5" pageNumber="549">27.7–33.6</td>
</tr>
<tr id="353F3498CA2EB886FF25F9B2FA121046" box="[163,1425,1565,1620]" gridrow="21" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F9B2FE5D1046" box="[163,478,1565,1620]" gridcol="0" gridrow="21" pageId="5" pageNumber="549">Length of stiff portion of last unbranched dorsal-fin ray</th>
<td id="76EE5DE4CA2EB886FD87F9B2FDDC1046" box="[513,607,1565,1620]" gridcol="1" gridrow="21" pageId="5" pageNumber="549">19.8</td>
<td id="76EE5DE4CA2EB886FD0BF9B2FD761046" box="[653,757,1565,1620]" gridcol="2" gridrow="21" pageId="5" pageNumber="549">18.2 (1.2)</td>
<td id="76EE5DE4CA2EB886FCA2F9B2FC091046" box="[804,906,1565,1620]" gridcol="3" gridrow="21" pageId="5" pageNumber="549">16.6–19.7</td>
<td id="76EE5DE4CA2EB886FC53F9B2FBAD1046" box="[981,1070,1565,1620]" gridcol="4" gridrow="21" pageId="5" pageNumber="549">17.9</td>
<td id="76EE5DE4CA2EB886FBC8F9B2FB3E1046" box="[1102,1213,1565,1620]" gridcol="5" gridrow="21" pageId="5" pageNumber="549">18.3 (1.5)</td>
<td id="76EE5DE4CA2EB886FB7DF9B2FA121046" box="[1275,1425,1565,1620]" gridcol="6" gridrow="21" pageId="5" pageNumber="549">15.8–20.6</td>
</tr>
<tr id="353F3498CA2EB886FF25F9CCFA12106E" box="[163,1425,1635,1660]" gridrow="22" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F9CCFE5D106E" box="[163,478,1635,1660]" gridcol="0" gridrow="22" pageId="5" pageNumber="549">Length of dorsal-fin base</th>
<td id="76EE5DE4CA2EB886FD87F9CCFDDC106E" box="[513,607,1635,1660]" gridcol="1" gridrow="22" pageId="5" pageNumber="549">18.4</td>
<td id="76EE5DE4CA2EB886FD0BF9CCFD76106E" box="[653,757,1635,1660]" gridcol="2" gridrow="22" pageId="5" pageNumber="549">18.5 (1.2)</td>
<td id="76EE5DE4CA2EB886FCA2F9CCFC09106E" box="[804,906,1635,1660]" gridcol="3" gridrow="22" pageId="5" pageNumber="549">16.6–19.8</td>
<td id="76EE5DE4CA2EB886FC53F9CCFBAD106E" box="[981,1070,1635,1660]" gridcol="4" gridrow="22" pageId="5" pageNumber="549">15.0</td>
<td id="76EE5DE4CA2EB886FBC8F9CCFB3E106E" box="[1102,1213,1635,1660]" gridcol="5" gridrow="22" pageId="5" pageNumber="549">17.7 (1.3)</td>
<td id="76EE5DE4CA2EB886FB7DF9CCFA12106E" box="[1275,1425,1635,1660]" gridcol="6" gridrow="22" pageId="5" pageNumber="549">14.3–19.3</td>
</tr>
<tr id="353F3498CA2EB886FF25F924FA1210B6" box="[163,1425,1675,1700]" gridrow="23" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F924FE5D10B6" box="[163,478,1675,1700]" gridcol="0" gridrow="23" pageId="5" pageNumber="549">Pectoral-fin length</th>
<td id="76EE5DE4CA2EB886FD87F924FDDC10B6" box="[513,607,1675,1700]" gridcol="1" gridrow="23" pageId="5" pageNumber="549">22.4</td>
<td id="76EE5DE4CA2EB886FD0BF924FD7610B6" box="[653,757,1675,1700]" gridcol="2" gridrow="23" pageId="5" pageNumber="549">23.0 (0.9)</td>
<td id="76EE5DE4CA2EB886FCA2F924FC0910B6" box="[804,906,1675,1700]" gridcol="3" gridrow="23" pageId="5" pageNumber="549">21.8–24.0</td>
<td id="76EE5DE4CA2EB886FC53F924FBAD10B6" box="[981,1070,1675,1700]" gridcol="4" gridrow="23" pageId="5" pageNumber="549">16.9</td>
<td id="76EE5DE4CA2EB886FBC8F924FB3E10B6" box="[1102,1213,1675,1700]" gridcol="5" gridrow="23" pageId="5" pageNumber="549">22.6 (1.3)</td>
<td id="76EE5DE4CA2EB886FB7DF924FA1210B6" box="[1275,1425,1675,1700]" gridcol="6" gridrow="23" pageId="5" pageNumber="549">20.6–24.4</td>
</tr>
<tr id="353F3498CA2EB886FF25F91CFA1210DE" box="[163,1425,1715,1740]" gridrow="24" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F91CFE5D10DE" box="[163,478,1715,1740]" gridcol="0" gridrow="24" pageId="5" pageNumber="549">Anal-fin depth</th>
<td id="76EE5DE4CA2EB886FD87F91CFDDC10DE" box="[513,607,1715,1740]" gridcol="1" gridrow="24" pageId="5" pageNumber="549">20.7</td>
<td id="76EE5DE4CA2EB886FD0BF91CFD7610DE" box="[653,757,1715,1740]" gridcol="2" gridrow="24" pageId="5" pageNumber="549">19.9 (0.8)</td>
<td id="76EE5DE4CA2EB886FCA2F91CFC0910DE" box="[804,906,1715,1740]" gridcol="3" gridrow="24" pageId="5" pageNumber="549">19.2–21.1</td>
<td id="76EE5DE4CA2EB886FC53F91CFBAD10DE" box="[981,1070,1715,1740]" gridcol="4" gridrow="24" pageId="5" pageNumber="549">17.4</td>
<td id="76EE5DE4CA2EB886FBC8F91CFB3E10DE" box="[1102,1213,1715,1740]" gridcol="5" gridrow="24" pageId="5" pageNumber="549">20.3 (1.0)</td>
<td id="76EE5DE4CA2EB886FB7DF91CFA1210DE" box="[1275,1425,1715,1740]" gridcol="6" gridrow="24" pageId="5" pageNumber="549">18.6–22.4</td>
</tr>
<tr id="353F3498CA2EB886FF25F973FA1210E7" box="[163,1425,1756,1781]" gridrow="25" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F973FE5D10E7" box="[163,478,1756,1781]" gridcol="0" gridrow="25" pageId="5" pageNumber="549">Caudal-peduncle length</th>
<td id="76EE5DE4CA2EB886FD87F973FDDC10E7" box="[513,607,1756,1781]" gridcol="1" gridrow="25" pageId="5" pageNumber="549">19.3</td>
<td id="76EE5DE4CA2EB886FD0BF973FD7610E7" box="[653,757,1756,1781]" gridcol="2" gridrow="25" pageId="5" pageNumber="549">20.4 (0.6)</td>
<td id="76EE5DE4CA2EB886FCA2F973FC0910E7" box="[804,906,1756,1781]" gridcol="3" gridrow="25" pageId="5" pageNumber="549">19.6–21.1</td>
<td id="76EE5DE4CA2EB886FC53F973FBAD10E7" box="[981,1070,1756,1781]" gridcol="4" gridrow="25" pageId="5" pageNumber="549">17.4</td>
<td id="76EE5DE4CA2EB886FBC8F973FB3E10E7" box="[1102,1213,1756,1781]" gridcol="5" gridrow="25" pageId="5" pageNumber="549">19.8 (1.8)</td>
<td id="76EE5DE4CA2EB886FB7DF973FA1210E7" box="[1275,1425,1756,1781]" gridcol="6" gridrow="25" pageId="5" pageNumber="549">16.2–22.6</td>
</tr>
<tr id="353F3498CA2EB886FF25F8ABFA12110F" box="[163,1425,1796,1821]" gridrow="26" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F8ABFE5D110F" box="[163,478,1796,1821]" gridcol="0" gridrow="26" pageId="5" pageNumber="549">Caudal-peduncle depth</th>
<td id="76EE5DE4CA2EB886FD87F8ABFDDC110F" box="[513,607,1796,1821]" gridcol="1" gridrow="26" pageId="5" pageNumber="549">13.8</td>
<td id="76EE5DE4CA2EB886FD0BF8ABFD76110F" box="[653,757,1796,1821]" gridcol="2" gridrow="26" pageId="5" pageNumber="549">13.9 (0.7)</td>
<td id="76EE5DE4CA2EB886FCA2F8ABFC09110F" box="[804,906,1796,1821]" gridcol="3" gridrow="26" pageId="5" pageNumber="549">13.1–14.6</td>
<td id="76EE5DE4CA2EB886FC53F8ABFBAD110F" box="[981,1070,1796,1821]" gridcol="4" gridrow="26" pageId="5" pageNumber="549">12.1</td>
<td id="76EE5DE4CA2EB886FBC8F8ABFB3E110F" box="[1102,1213,1796,1821]" gridcol="5" gridrow="26" pageId="5" pageNumber="549">14.1 (0.4)</td>
<td id="76EE5DE4CA2EB886FB7DF8ABFA12110F" box="[1275,1425,1796,1821]" gridcol="6" gridrow="26" pageId="5" pageNumber="549">13.2–14.9</td>
</tr>
<tr id="353F3498CA2EB886FF25F883FA121157" box="[163,1425,1836,1861]" gridrow="27" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F883FA121157" box="[163,1425,1836,1861]" colspan="7" colspanRight="6" gridcol="0" gridrow="27" pageId="5" pageNumber="549">
<emphasis id="B97BEAC8CA2E477CFF25F883FF641157" bold="true" box="[163,231,1836,1861]" pageId="5" pageNumber="549">% HL</emphasis>
</th>
</tr>
<tr id="353F3498CA2EB886FF25F8FBFA12117F" box="[163,1425,1876,1901]" gridrow="28" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F8FBFE5D117F" box="[163,478,1876,1901]" gridcol="0" gridrow="28" pageId="5" pageNumber="549">Post-orbital head length</th>
<td id="76EE5DE4CA2EB886FD87F8FBFDDC117F" box="[513,607,1876,1901]" gridcol="1" gridrow="28" pageId="5" pageNumber="549">38.7</td>
<td id="76EE5DE4CA2EB886FD0BF8FBFD76117F" box="[653,757,1876,1901]" gridcol="2" gridrow="28" pageId="5" pageNumber="549">39.7 (1.4)</td>
<td id="76EE5DE4CA2EB886FCA2F8FBFC09117F" box="[804,906,1876,1901]" gridcol="3" gridrow="28" pageId="5" pageNumber="549">38.3–42.0</td>
<td id="76EE5DE4CA2EB886FC53F8FBFBAD117F" box="[981,1070,1876,1901]" gridcol="4" gridrow="28" pageId="5" pageNumber="549">37.1</td>
<td id="76EE5DE4CA2EB886FBC8F8FBFB3E117F" box="[1102,1213,1876,1901]" gridcol="5" gridrow="28" pageId="5" pageNumber="549">38.7 (4.2)</td>
<td id="76EE5DE4CA2EB886FB7DF8FBFA12117F" box="[1275,1425,1876,1901]" gridcol="6" gridrow="28" pageId="5" pageNumber="549">29.8–45.3</td>
</tr>
<tr id="353F3498CA2EB886FF25F8D2FA121184" box="[163,1425,1917,1942]" gridrow="29" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F8D2FE5D1184" box="[163,478,1917,1942]" gridcol="0" gridrow="29" pageId="5" pageNumber="549">Head depth</th>
<td id="76EE5DE4CA2EB886FD87F8D2FDDC1184" box="[513,607,1917,1942]" gridcol="1" gridrow="29" pageId="5" pageNumber="549">73.4</td>
<td id="76EE5DE4CA2EB886FD0BF8D2FD761184" box="[653,757,1917,1942]" gridcol="2" gridrow="29" pageId="5" pageNumber="549">80.8 (2.6)</td>
<td id="76EE5DE4CA2EB886FCA2F8D2FC091184" box="[804,906,1917,1942]" gridcol="3" gridrow="29" pageId="5" pageNumber="549">77.7–85.0</td>
<td id="76EE5DE4CA2EB886FC53F8D2FBAD1184" box="[981,1070,1917,1942]" gridcol="4" gridrow="29" pageId="5" pageNumber="549">71.0</td>
<td id="76EE5DE4CA2EB886FBC8F8D2FB3E1184" box="[1102,1213,1917,1942]" gridcol="5" gridrow="29" pageId="5" pageNumber="549">84.0 (5.3)</td>
<td id="76EE5DE4CA2EB886FB7DF8D2FA121184" box="[1275,1425,1917,1942]" gridcol="6" gridrow="29" pageId="5" pageNumber="549">73.2–93.2</td>
</tr>
<tr id="353F3498CA2EB886FF25F80AFA1211AC" box="[163,1425,1957,1982]" gridrow="30" pageId="5" pageNumber="549">
<th id="76EE5DE4CA2EB886FF25F80AFE5D11AC" box="[163,478,1957,1982]" gridcol="0" gridrow="30" pageId="5" pageNumber="549">Head width</th>
<td id="76EE5DE4CA2EB886FD87F80AFDDC11AC" box="[513,607,1957,1982]" gridcol="1" gridrow="30" pageId="5" pageNumber="549">54.8</td>
<td id="76EE5DE4CA2EB886FD0BF80AFD7611AC" box="[653,757,1957,1982]" gridcol="2" gridrow="30" pageId="5" pageNumber="549">59.8 (2.2)</td>
<td id="76EE5DE4CA2EB886FCA2F80AFC0911AC" box="[804,906,1957,1982]" gridcol="3" gridrow="30" pageId="5" pageNumber="549">56.2–61.7</td>
<td id="76EE5DE4CA2EB886FC53F80AFBAD11AC" box="[981,1070,1957,1982]" gridcol="4" gridrow="30" pageId="5" pageNumber="549">50.0</td>
<td id="76EE5DE4CA2EB886FBC8F80AFB3E11AC" box="[1102,1213,1957,1982]" gridcol="5" gridrow="30" pageId="5" pageNumber="549">60.5 (3.4)</td>
<td id="76EE5DE4CA2EB886FB7DF80AFA1211AC" box="[1275,1425,1957,1982]" gridcol="6" gridrow="30" pageId="5" pageNumber="549">54.6–65.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB036DACA2E477CFBE1F860FA1211F4" blockId="5.[1127,1425,1999,2023]" box="[1127,1425,1999,2023]" pageId="5" pageNumber="549">
<tableNote id="76E93754CA2E477CFBE1F860FA1211F4" box="[1127,1425,1999,2023]" pageId="5" pageNumber="549" targetBox="[163,1425,662,1982]" targetPageId="5">...Continued on the next page</tableNote>
</paragraph>
<caption id="DF706652CA2D477FFF25FF0BFE1216AF" box="[163,401,164,189]" pageId="6" pageNumber="550">
<paragraph id="8BB036DACA2D477FFF25FF0BFE1216AF" blockId="6.[163,401,164,189]" box="[163,401,164,189]" pageId="6" pageNumber="550">
<emphasis id="B97BEAC8CA2D477FFF25FF0BFE9616AE" bold="true" box="[163,277,164,188]" pageId="6" pageNumber="550">TABLE 2.</emphasis>
(continued)
</paragraph>
</caption>
<paragraph id="8BB036DACA2D477FFF25FF63FAA0127F" pageId="6" pageNumber="550">
<table id="F90FC47ACA2DB886FF25FF63FA12127F" box="[163,1425,204,1133]" colsContinueFrom="5.[163,1425,662,1982]" gridcols="7" gridrows="22" pageId="6" pageNumber="550">
<tr id="353F3498CA2DB886FF25FF63FA1216F7" box="[163,1425,204,229]" gridrow="0" pageId="6" pageNumber="550" rowspan-3="1" rowspan-4="1">
<th id="76EE5DE4CA2DB886FF25FF63FE5D16F7" box="[163,478,204,229]" gridcol="0" gridrow="0" pageId="6" pageNumber="550">Characters</th>
<th id="76EE5DE4CA2DB886FD87FF63FD7616F7" box="[513,757,204,229]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="550">
<taxonomicName id="4C0F4D59CA2D477FFD87FF62FD2116F7" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[513,674,205,229]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="550" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2D477FFD87FF62FD2116F7" box="[513,674,205,229]" italics="true" pageId="6" pageNumber="550">Waikhomia hira</emphasis>
</taxonomicName>
</th>
<th id="76EE5DE4CA2DB886FBC8FF63FA1216F7" box="[1102,1425,204,229]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="6" pageNumber="550">
<taxonomicName id="4C0F4D59CA2D477FFBC8FF62FAD316F7" box="[1102,1360,205,229]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="550" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA2D477FFBC8FF62FAD316F7" box="[1102,1360,205,229]" italics="true" pageId="6" pageNumber="550">Waikhomia sahyadriensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F3498CA2DB886FF25FF5BFA12175F" box="[163,1425,244,333]" gridrow="1" pageId="6" pageNumber="550" rowspan-0="1">
<td id="76EE5DE4CA2DB886FD87FF5BFDDC175F" box="[513,607,244,333]" gridcol="1" gridrow="1" pageId="6" pageNumber="550">Holotype (602)</td>
<td id="76EE5DE4CA2DB886FD0BFF5BFC1B175F" box="[653,920,244,333]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="6" pageNumber="550">Paratypes (n= 5) (603–605, 607, 1002)</td>
<td id="76EE5DE4CA2DB886FC53FF5BFBAD175F" box="[981,1070,244,333]" gridcol="4" gridrow="1" pageId="6" pageNumber="550">Paratype (997)</td>
<td id="76EE5DE4CA2DB886FBC8FF5BFA12175F" box="[1102,1425,244,333]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="6" pageNumber="550">Topotypes + additional material (n=19) (582–597, 599–601)</td>
</tr>
<tr id="353F3498CA2DB886FF25FEF8FA121762" box="[163,1425,343,368]" gridrow="2" pageId="6" pageNumber="550" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="76EE5DE4CA2DB886FD0BFEF8FD761762" box="[653,757,343,368]" gridcol="2" gridrow="2" pageId="6" pageNumber="550">Mean (sd)</td>
<td id="76EE5DE4CA2DB886FCA2FEF8FC1B1762" box="[804,920,343,368]" gridcol="3" gridrow="2" pageId="6" pageNumber="550">Range</td>
<td id="76EE5DE4CA2DB886FBC8FEF8FB351762" box="[1102,1206,343,368]" gridcol="5" gridrow="2" pageId="6" pageNumber="550">Mean (sd)</td>
<td id="76EE5DE4CA2DB886FB7DFEF8FA121762" box="[1275,1425,343,368]" gridcol="6" gridrow="2" pageId="6" pageNumber="550">Range</td>
</tr>
<tr id="353F3498CA2DB886FF25FED0FA12178A" box="[163,1425,383,408]" gridrow="3" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FED0FE5D178A" box="[163,478,383,408]" gridcol="0" gridrow="3" pageId="6" pageNumber="550">Body width at dorsal-fin origin</th>
<td id="76EE5DE4CA2DB886FD87FED0FDDC178A" box="[513,607,383,408]" gridcol="1" gridrow="3" pageId="6" pageNumber="550">54.8</td>
<td id="76EE5DE4CA2DB886FD0BFED0FD76178A" box="[653,757,383,408]" gridcol="2" gridrow="3" pageId="6" pageNumber="550">52.7 (2.7)</td>
<td id="76EE5DE4CA2DB886FCA2FED0FC1B178A" box="[804,920,383,408]" gridcol="3" gridrow="3" pageId="6" pageNumber="550">50.0–56.1</td>
<td id="76EE5DE4CA2DB886FC53FED0FBAD178A" box="[981,1070,383,408]" gridcol="4" gridrow="3" pageId="6" pageNumber="550">38.7</td>
<td id="76EE5DE4CA2DB886FBC8FED0FB35178A" box="[1102,1206,383,408]" gridcol="5" gridrow="3" pageId="6" pageNumber="550">54.6 (6.2)</td>
<td id="76EE5DE4CA2DB886FB7DFED0FA12178A" box="[1275,1425,383,408]" gridcol="6" gridrow="3" pageId="6" pageNumber="550">40.2–63.4</td>
</tr>
<tr id="353F3498CA2DB886FF25FE07FA1217D3" box="[163,1425,424,449]" gridrow="4" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FE07FE5D17D3" box="[163,478,424,449]" gridcol="0" gridrow="4" pageId="6" pageNumber="550">Body width at anal-fin origin</th>
<td id="76EE5DE4CA2DB886FD87FE07FDDC17D3" box="[513,607,424,449]" gridcol="1" gridrow="4" pageId="6" pageNumber="550">43.5</td>
<td id="76EE5DE4CA2DB886FD0BFE07FD7617D3" box="[653,757,424,449]" gridcol="2" gridrow="4" pageId="6" pageNumber="550">35.0 (5.2)</td>
<td id="76EE5DE4CA2DB886FCA2FE07FC1B17D3" box="[804,920,424,449]" gridcol="3" gridrow="4" pageId="6" pageNumber="550">30.2–41.1</td>
<td id="76EE5DE4CA2DB886FC53FE07FBAD17D3" box="[981,1070,424,449]" gridcol="4" gridrow="4" pageId="6" pageNumber="550">25.8</td>
<td id="76EE5DE4CA2DB886FBC8FE07FB3517D3" box="[1102,1206,424,449]" gridcol="5" gridrow="4" pageId="6" pageNumber="550">37.8 (8.7)</td>
<td id="76EE5DE4CA2DB886FB7DFE07FA1217D3" box="[1275,1425,424,449]" gridcol="6" gridrow="4" pageId="6" pageNumber="550">21.6–51.5</td>
</tr>
<tr id="353F3498CA2DB886FF25FE7FFA1217FB" box="[163,1425,464,489]" gridrow="5" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FE7FFE5D17FB" box="[163,478,464,489]" gridcol="0" gridrow="5" pageId="6" pageNumber="550">Snout length</th>
<td id="76EE5DE4CA2DB886FD87FE7FFDDC17FB" box="[513,607,464,489]" gridcol="1" gridrow="5" pageId="6" pageNumber="550">28.2</td>
<td id="76EE5DE4CA2DB886FD0BFE7FFD7617FB" box="[653,757,464,489]" gridcol="2" gridrow="5" pageId="6" pageNumber="550">28.7 (1.7)</td>
<td id="76EE5DE4CA2DB886FCA2FE7FFC1B17FB" box="[804,920,464,489]" gridcol="3" gridrow="5" pageId="6" pageNumber="550">26.2–31.0</td>
<td id="76EE5DE4CA2DB886FC53FE7FFBAD17FB" box="[981,1070,464,489]" gridcol="4" gridrow="5" pageId="6" pageNumber="550">24.2</td>
<td id="76EE5DE4CA2DB886FBC8FE7FFB3517FB" box="[1102,1206,464,489]" gridcol="5" gridrow="5" pageId="6" pageNumber="550">32.2 (3.6)</td>
<td id="76EE5DE4CA2DB886FB7DFE7FFA1217FB" box="[1275,1425,464,489]" gridcol="6" gridrow="5" pageId="6" pageNumber="550">27.3–38.1</td>
</tr>
<tr id="353F3498CA2DB886FF25FE57FA121403" box="[163,1425,504,529]" gridrow="6" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FE57FE5D1403" box="[163,478,504,529]" gridcol="0" gridrow="6" pageId="6" pageNumber="550">Eye diameter</th>
<td id="76EE5DE4CA2DB886FD87FE57FDDC1403" box="[513,607,504,529]" gridcol="1" gridrow="6" pageId="6" pageNumber="550">31.5</td>
<td id="76EE5DE4CA2DB886FD0BFE57FD761403" box="[653,757,504,529]" gridcol="2" gridrow="6" pageId="6" pageNumber="550">34.6 (1.8)</td>
<td id="76EE5DE4CA2DB886FCA2FE57FC1B1403" box="[804,920,504,529]" gridcol="3" gridrow="6" pageId="6" pageNumber="550">33.0–37.4</td>
<td id="76EE5DE4CA2DB886FC53FE57FBAD1403" box="[981,1070,504,529]" gridcol="4" gridrow="6" pageId="6" pageNumber="550">35.5</td>
<td id="76EE5DE4CA2DB886FBC8FE57FB351403" box="[1102,1206,504,529]" gridcol="5" gridrow="6" pageId="6" pageNumber="550">31.8 (2.6)</td>
<td id="76EE5DE4CA2DB886FB7DFE57FA121403" box="[1275,1425,504,529]" gridcol="6" gridrow="6" pageId="6" pageNumber="550">25.5–36.8</td>
</tr>
<tr id="353F3498CA2DB886FF25FD8FFA12142B" box="[163,1425,544,569]" gridrow="7" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FD8FFE5D142B" box="[163,478,544,569]" gridcol="0" gridrow="7" pageId="6" pageNumber="550">Internarial width</th>
<td id="76EE5DE4CA2DB886FD87FD8FFDDC142B" box="[513,607,544,569]" gridcol="1" gridrow="7" pageId="6" pageNumber="550">22.6</td>
<td id="76EE5DE4CA2DB886FD0BFD8FFD76142B" box="[653,757,544,569]" gridcol="2" gridrow="7" pageId="6" pageNumber="550">21.4 (1.8)</td>
<td id="76EE5DE4CA2DB886FCA2FD8FFC1B142B" box="[804,920,544,569]" gridcol="3" gridrow="7" pageId="6" pageNumber="550">19.8–24.0</td>
<td id="76EE5DE4CA2DB886FC53FD8FFBAD142B" box="[981,1070,544,569]" gridcol="4" gridrow="7" pageId="6" pageNumber="550">24.2</td>
<td id="76EE5DE4CA2DB886FBC8FD8FFB35142B" box="[1102,1206,544,569]" gridcol="5" gridrow="7" pageId="6" pageNumber="550">21.9 (3.5)</td>
<td id="76EE5DE4CA2DB886FB7DFD8FFA12142B" box="[1275,1425,544,569]" gridcol="6" gridrow="7" pageId="6" pageNumber="550">14.4–28.3</td>
</tr>
<tr id="353F3498CA2DB886FF25FDE6FA121470" box="[163,1425,585,610]" gridrow="8" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FDE6FE5D1470" box="[163,478,585,610]" gridcol="0" gridrow="8" pageId="6" pageNumber="550">Inter orbital width</th>
<td id="76EE5DE4CA2DB886FD87FDE6FDDC1470" box="[513,607,585,610]" gridcol="1" gridrow="8" pageId="6" pageNumber="550">34.7</td>
<td id="76EE5DE4CA2DB886FD0BFDE6FD761470" box="[653,757,585,610]" gridcol="2" gridrow="8" pageId="6" pageNumber="550">35.6 (0.9)</td>
<td id="76EE5DE4CA2DB886FCA2FDE6FC1B1470" box="[804,920,585,610]" gridcol="3" gridrow="8" pageId="6" pageNumber="550">34.5–36.7</td>
<td id="76EE5DE4CA2DB886FC53FDE6FBAD1470" box="[981,1070,585,610]" gridcol="4" gridrow="8" pageId="6" pageNumber="550">30.6</td>
<td id="76EE5DE4CA2DB886FBC8FDE6FB351470" box="[1102,1206,585,610]" gridcol="5" gridrow="8" pageId="6" pageNumber="550">36.4 (3.6)</td>
<td id="76EE5DE4CA2DB886FB7DFDE6FA121470" box="[1275,1425,585,610]" gridcol="6" gridrow="8" pageId="6" pageNumber="550">28.9–41.7</td>
</tr>
<tr id="353F3498CA2DB886FF25FDDEFA12149B" box="[163,1425,625,649]" gridrow="9" pageId="6" pageNumber="550">
<th id="76EE5DE4CA2DB886FF25FDDEFA12149B" box="[163,1425,625,649]" colspan="7" colspanRight="6" gridcol="0" gridrow="9" pageId="6" pageNumber="550">
<emphasis id="B97BEAC8CA2D477FFF25FDDEFE80149B" bold="true" box="[163,259,625,649]" italics="true" pageId="6" pageNumber="550">Meristics</emphasis>
</th>
</tr>
<tr id="353F3498CA2DB886FF25FD36FA1214A0" box="[163,1425,665,690]" gridrow="10" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FD36FE5D14A0" box="[163,478,665,690]" gridcol="0" gridrow="10" pageId="6" pageNumber="550">Lateral line series scales</th>
<td id="76EE5DE4CA2DB886FD87FD36FDDC14A0" box="[513,607,665,690]" gridcol="1" gridrow="10" pageId="6" pageNumber="550">24</td>
<td id="76EE5DE4CA2DB886FCA2FD36FC1B14A0" box="[804,920,665,690]" gridcol="3" gridrow="10" pageId="6" pageNumber="550">23–25</td>
<td id="76EE5DE4CA2DB886FC53FD36FBAD14A0" box="[981,1070,665,690]" gridcol="4" gridrow="10" pageId="6" pageNumber="550">24</td>
<td id="76EE5DE4CA2DB886FB7DFD36FA1214A0" box="[1275,1425,665,690]" gridcol="6" gridrow="10" pageId="6" pageNumber="550">23–24</td>
</tr>
<tr id="353F3498CA2DB886FF25FD6EFA1214C8" box="[163,1425,705,730]" gridrow="11" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FD6EFE5D14C8" box="[163,478,705,730]" gridcol="0" gridrow="11" pageId="6" pageNumber="550">Transverse row scales</th>
<td id="76EE5DE4CA2DB886FD87FD6EFDDC14C8" box="[513,607,705,730]" gridcol="1" gridrow="11" pageId="6" pageNumber="550">4/1/3½</td>
<td id="76EE5DE4CA2DB886FCA2FD6EFC1B14C8" box="[804,920,705,730]" gridcol="3" gridrow="11" pageId="6" pageNumber="550">4–½4/1/3½</td>
<td id="76EE5DE4CA2DB886FC53FD6EFBAD14C8" box="[981,1070,705,730]" gridcol="4" gridrow="11" pageId="6" pageNumber="550">4/1/3½</td>
<td id="76EE5DE4CA2DB886FB7DFD6EFA1214C8" box="[1275,1425,705,730]" gridcol="6" gridrow="11" pageId="6" pageNumber="550">½3/1/3½</td>
</tr>
<tr id="353F3498CA2DB886FF25FD45FA121511" box="[163,1425,746,771]" gridrow="12" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FD45FE5D1511" box="[163,478,746,771]" gridcol="0" gridrow="12" pageId="6" pageNumber="550">Pre-dorsal scales</th>
<td id="76EE5DE4CA2DB886FD87FD45FDDC1511" box="[513,607,746,771]" gridcol="1" gridrow="12" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FCA2FD45FC1B1511" box="[804,920,746,771]" gridcol="3" gridrow="12" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FC53FD45FBAD1511" box="[981,1070,746,771]" gridcol="4" gridrow="12" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FB7DFD45FA121511" box="[1275,1425,746,771]" gridcol="6" gridrow="12" pageId="6" pageNumber="550">7–8</td>
</tr>
<tr id="353F3498CA2DB886FF25FCBDFA121539" box="[163,1425,786,811]" gridrow="13" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FCBDFE5D1539" box="[163,478,786,811]" gridcol="0" gridrow="13" pageId="6" pageNumber="550">Pre-pelvic scales</th>
<td id="76EE5DE4CA2DB886FD87FCBDFDDC1539" box="[513,607,786,811]" gridcol="1" gridrow="13" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FCA2FCBDFC1B1539" box="[804,920,786,811]" gridcol="3" gridrow="13" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FC53FCBDFBAD1539" box="[981,1070,786,811]" gridcol="4" gridrow="13" pageId="6" pageNumber="550">10</td>
<td id="76EE5DE4CA2DB886FB7DFCBDFA121539" box="[1275,1425,786,811]" gridcol="6" gridrow="13" pageId="6" pageNumber="550">10–11</td>
</tr>
<tr id="353F3498CA2DB886FF25FC95FA121541" box="[163,1425,826,851]" gridrow="14" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FC95FE5D1541" box="[163,478,826,851]" gridcol="0" gridrow="14" pageId="6" pageNumber="550">Pre-anal scales</th>
<td id="76EE5DE4CA2DB886FD87FC95FDDC1541" box="[513,607,826,851]" gridcol="1" gridrow="14" pageId="6" pageNumber="550">18</td>
<td id="76EE5DE4CA2DB886FCA2FC95FC1B1541" box="[804,920,826,851]" gridcol="3" gridrow="14" pageId="6" pageNumber="550">17–18</td>
<td id="76EE5DE4CA2DB886FC53FC95FBAD1541" box="[981,1070,826,851]" gridcol="4" gridrow="14" pageId="6" pageNumber="550">18</td>
<td id="76EE5DE4CA2DB886FB7DFC95FA121541" box="[1275,1425,826,851]" gridcol="6" gridrow="14" pageId="6" pageNumber="550">17–20</td>
</tr>
<tr id="353F3498CA2DB886FF25FCCDFA121569" box="[163,1425,866,891]" gridrow="15" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FCCDFE5D1569" box="[163,478,866,891]" gridcol="0" gridrow="15" pageId="6" pageNumber="550">Circumpeduncular scales</th>
<td id="76EE5DE4CA2DB886FD87FCCDFDDC1569" box="[513,607,866,891]" gridcol="1" gridrow="15" pageId="6" pageNumber="550">12</td>
<td id="76EE5DE4CA2DB886FCA2FCCDFC1B1569" box="[804,920,866,891]" gridcol="3" gridrow="15" pageId="6" pageNumber="550">12</td>
<td id="76EE5DE4CA2DB886FC53FCCDFBAD1569" box="[981,1070,866,891]" gridcol="4" gridrow="15" pageId="6" pageNumber="550">12</td>
<td id="76EE5DE4CA2DB886FB7DFCCDFA121569" box="[1275,1425,866,891]" gridcol="6" gridrow="15" pageId="6" pageNumber="550">12</td>
</tr>
<tr id="353F3498CA2DB886FF25FC24FA1215B6" box="[163,1425,907,932]" gridrow="16" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FC24FE5D15B6" box="[163,478,907,932]" gridcol="0" gridrow="16" pageId="6" pageNumber="550">Dorsal-fin rays</th>
<td id="76EE5DE4CA2DB886FD87FC24FDDC15B6" box="[513,607,907,932]" gridcol="1" gridrow="16" pageId="6" pageNumber="550">ii-i-8</td>
<td id="76EE5DE4CA2DB886FCA2FC24FC1B15B6" box="[804,920,907,932]" gridcol="3" gridrow="16" pageId="6" pageNumber="550">ii-i-8</td>
<td id="76EE5DE4CA2DB886FC53FC24FBAD15B6" box="[981,1070,907,932]" gridcol="4" gridrow="16" pageId="6" pageNumber="550">ii-i-8</td>
<td id="76EE5DE4CA2DB886FB7DFC24FA1215B6" box="[1275,1425,907,932]" gridcol="6" gridrow="16" pageId="6" pageNumber="550">ii-i-8</td>
</tr>
<tr id="353F3498CA2DB886FF25FC1CFA1215DE" box="[163,1425,947,972]" gridrow="17" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FC1CFE5D15DE" box="[163,478,947,972]" gridcol="0" gridrow="17" pageId="6" pageNumber="550">Pectoral-fin rays</th>
<td id="76EE5DE4CA2DB886FD87FC1CFDDC15DE" box="[513,607,947,972]" gridcol="1" gridrow="17" pageId="6" pageNumber="550">i-12</td>
<td id="76EE5DE4CA2DB886FCA2FC1CFC1B15DE" box="[804,920,947,972]" gridcol="3" gridrow="17" pageId="6" pageNumber="550">i-11–12</td>
<td id="76EE5DE4CA2DB886FC53FC1CFBAD15DE" box="[981,1070,947,972]" gridcol="4" gridrow="17" pageId="6" pageNumber="550">i-12</td>
<td id="76EE5DE4CA2DB886FB7DFC1CFA1215DE" box="[1275,1425,947,972]" gridcol="6" gridrow="17" pageId="6" pageNumber="550">i-12–13</td>
</tr>
<tr id="353F3498CA2DB886FF25FC74FA1215E6" box="[163,1425,987,1012]" gridrow="18" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FC74FE5D15E6" box="[163,478,987,1012]" gridcol="0" gridrow="18" pageId="6" pageNumber="550">Pelvic-fin rays</th>
<td id="76EE5DE4CA2DB886FD87FC74FDDC15E6" box="[513,607,987,1012]" gridcol="1" gridrow="18" pageId="6" pageNumber="550">i-8</td>
<td id="76EE5DE4CA2DB886FCA2FC74FC1B15E6" box="[804,920,987,1012]" gridcol="3" gridrow="18" pageId="6" pageNumber="550">i-8</td>
<td id="76EE5DE4CA2DB886FC53FC74FBAD15E6" box="[981,1070,987,1012]" gridcol="4" gridrow="18" pageId="6" pageNumber="550">i-8</td>
<td id="76EE5DE4CA2DB886FB7DFC74FA1215E6" box="[1275,1425,987,1012]" gridcol="6" gridrow="18" pageId="6" pageNumber="550">i-8</td>
</tr>
<tr id="353F3498CA2DB886FF25FBACFA12120E" box="[163,1425,1027,1052]" gridrow="19" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FBACFE5D120E" box="[163,478,1027,1052]" gridcol="0" gridrow="19" pageId="6" pageNumber="550">Anal-fin rays</th>
<td id="76EE5DE4CA2DB886FD87FBACFDDC120E" box="[513,607,1027,1052]" gridcol="1" gridrow="19" pageId="6" pageNumber="550">ii-i-5</td>
<td id="76EE5DE4CA2DB886FCA2FBACFC1B120E" box="[804,920,1027,1052]" gridcol="3" gridrow="19" pageId="6" pageNumber="550">ii-i-5</td>
<td id="76EE5DE4CA2DB886FC53FBACFBAD120E" box="[981,1070,1027,1052]" gridcol="4" gridrow="19" pageId="6" pageNumber="550">ii-i-5</td>
<td id="76EE5DE4CA2DB886FB7DFBACFA12120E" box="[1275,1425,1027,1052]" gridcol="6" gridrow="19" pageId="6" pageNumber="550">ii-i-5</td>
</tr>
<tr id="353F3498CA2DB886FF25FB83FA121257" box="[163,1425,1068,1093]" gridrow="20" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FB83FE5D1257" box="[163,478,1068,1093]" gridcol="0" gridrow="20" pageId="6" pageNumber="550">Caudal-fin rays (procurrent)</th>
<td id="76EE5DE4CA2DB886FD87FB83FDDC1257" box="[513,607,1068,1093]" gridcol="1" gridrow="20" pageId="6" pageNumber="550">8+7</td>
<td id="76EE5DE4CA2DB886FCA2FB83FC1B1257" box="[804,920,1068,1093]" gridcol="3" gridrow="20" pageId="6" pageNumber="550">7–8+6–7</td>
<td id="76EE5DE4CA2DB886FC53FB83FBAD1257" box="[981,1070,1068,1093]" gridcol="4" gridrow="20" pageId="6" pageNumber="550">8+7</td>
<td id="76EE5DE4CA2DB886FB7DFB83FA121257" box="[1275,1425,1068,1093]" gridcol="6" gridrow="20" pageId="6" pageNumber="550">7–8+6</td>
</tr>
<tr id="353F3498CA2DB886FF25FBFBFA12127F" box="[163,1425,1108,1133]" gridrow="21" pageId="6" pageNumber="550" rowspan-2="1" rowspan-5="1">
<th id="76EE5DE4CA2DB886FF25FBFBFE5D127F" box="[163,478,1108,1133]" gridcol="0" gridrow="21" pageId="6" pageNumber="550">Caudal-fin rays (principal)</th>
<td id="76EE5DE4CA2DB886FD87FBFBFDDC127F" box="[513,607,1108,1133]" gridcol="1" gridrow="21" pageId="6" pageNumber="550">9+8</td>
<td id="76EE5DE4CA2DB886FCA2FBFBFC1B127F" box="[804,920,1108,1133]" gridcol="3" gridrow="21" pageId="6" pageNumber="550">9+8</td>
<td id="76EE5DE4CA2DB886FC53FBFBFBAD127F" box="[981,1070,1108,1133]" gridcol="4" gridrow="21" pageId="6" pageNumber="550">9+8</td>
<td id="76EE5DE4CA2DB886FB7DFBFBFA12127F" box="[1275,1425,1108,1133]" gridcol="6" gridrow="21" pageId="6" pageNumber="550">9+8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB036DACA2D477FFF41FB33FCC1100C" blockId="6.[151,1437,1180,2035]" pageId="6" pageNumber="550">Body elongate, deep, its standard length 2.6–3.3 times of maximum body depth, compressed laterally; predorsal contour almost straight, steadily rising to dorsal fin origin, thereafter, gradually sloping ventral towards caudal-fin base. Ventral profile convex up to pelvic-fin base, abruptly sloping dorsal along anal-fin base, thereafter gradually sloping dorsal to caudal-fin base. Caudal peduncle slender, elongate, its length 1.4–1.6 times its depth. Head small, stout, compressed, its length 1.2–1.4 times its depth. Snout long (3.2–4.1 times in head length), pointed, lateral fold present, its length marginally shorter than eye diameter. Eyes large, mid-laterally positioned, closer to snout tip than posterior margin of operculum, diameter almost equal to (0.8–1.2 times) interorbital width. Mouth small, terminal, ventrally U-shaped, angle of gape not reaching vertical from anterior margin of eye. Lips thin, not interrupted, upper lip relatively fleshier than lower lip. Barbels absent. Nuptial tubercles present in mature males, scattered across snout, lower sides of head and ventral surface as far back as pelvic-fin base; also scattered over outer margins of unbranched rays of pectoral and pelvic fins.</paragraph>
<paragraph id="8BB036DACA2D477FFF41F987FF6811B9" blockId="6.[151,1437,1180,2035]" pageId="6" pageNumber="550">Dorsal fin high, length 0.9–1.2 times head length, originating approximately above pelvic-fin origin, its origin situated closer to tip of snout than to base of caudal fin; posterior margin deeply concave, extending beyond vertical through middle of anal fin, tip pointed. Dorsal fin with three unbranched rays (two supernumerary and one serially associated unbranched ray) and eight branched rays, the single unbranched serially associated ray feeble, length of segmented apex half that of entire fin ray. Pectoral fin with one simple and 11(1) or 12(6) branched rays, its tip rounded, when adpressed overlapping pelvic-fin base in mature males (4), but falling well short of pelvic-fin base in females (2). Pelvic fin with one simple and eight branched rays, its tip rounded, reaching vent when adpressed. Anal fin with three unbranched (two supernumerary and one serially associated unbranched ray) and five branched rays, its distal margin concave, with pointed tip. Caudal fin deeply forked, lobes measuring almost two-thirds of caudal-fin length, tips pointed. Principal caudal-fin rays 9+8; procurrent rays dorsally 7(2) or 8(5) and ventrally 6(1) or 7(6).</paragraph>
<paragraph id="8BB036DACA2D477EFF41F81BFE8F16C5" blockId="6.[151,1437,1180,2035]" lastBlockId="7.[151,1436,153,215]" lastPageId="7" lastPageNumber="551" pageId="6" pageNumber="550">Lateral line complete, with 23(1), 24(3) or 25(3) perforated scales, originating at cleithral spot, proceeding horizontally to caudal-fin base. Scales in transverse row 4(5) or ½4(2)/1/3½, pre-dorsal and pre-pelvic scales 10, pre-anal scales 17(1) or 18(6), circumpeduncular scales 12. Pelvic axillary scale present, one-fourth the length of pelvic-fin.</paragraph>
<caption id="DF706652CA2C477EFF11F9F6FB501060" ID-DOI="http://doi.org/10.5281/zenodo.3920623" ID-Zenodo-Dep="3920623" box="[151,1235,1625,1650]" httpUri="https://zenodo.org/record/3920623/files/figure.png" pageId="7" pageNumber="551" startId="7.[151,250,1625,1650]" targetBox="[151,1435,237,1600]" targetPageId="7">
<paragraph id="8BB036DACA2C477EFF11F9F6FB501060" blockId="7.[151,1235,1625,1650]" box="[151,1235,1625,1650]" pageId="7" pageNumber="551">
<emphasis id="B97BEAC8CA2C477EFF11F9F6FE901060" bold="true" box="[151,275,1625,1650]" pageId="7" pageNumber="551">FIGURE 3.</emphasis>
<taxonomicName id="4C0F4D59CA2C477EFE9FF9F5FE381060" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[281,443,1626,1650]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="551" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2C477EFE9FF9F5FE381060" box="[281,443,1626,1650]" italics="true" pageId="7" pageNumber="551">Waikhomia hira</emphasis>
</taxonomicName>
holotype (male, BNHS FWF 602, 42.9 mm SL) in (A) life and (B) preserved.
</paragraph>
</caption>
<paragraph id="8BB036DACA2C4770FF41F935FF491751" blockId="7.[151,1437,1690,1969]" lastBlockId="9.[151,1437,153,1728]" lastPageId="9" lastPageNumber="553" pageId="7" pageNumber="551">
<emphasis id="B97BEAC8CA2C477EFF41F935FEC010A7" bold="true" box="[199,323,1690,1717]" pageId="7" pageNumber="551">Osteology.</emphasis>
The skeletal anatomy of cleared and double stained 
<taxonomicName id="4C0F4D59CA2C477EFC09F933FC6210A7" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[911,993,1691,1717]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="551" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA2C477EFC09F933FC6210A7" box="[911,993,1691,1717]" italics="true" pageId="7" pageNumber="551">W. hira</emphasis>
</taxonomicName>
<typeStatus id="54B48878CA2C477EFC6EF934FBCB10A7" box="[1000,1096,1691,1717]" pageId="7" pageNumber="551" type="paratype">paratype</typeStatus>
BNHS FWF 604 is illustrated in 
<figureCitation id="13342A5FCA2C477EFF30F910FE9A10CB" box="[182,281,1727,1753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="7" pageNumber="551">Figure 4</figureCitation>
. Infraorbital 2 narrow, slender, less wide than infraorbital 4; Infraorbital 3 largest among infraorbital series, deep, broadest at the middle; infraorbital 4 rectangular, narrower than infraorbitals 1 and 3; infraorbital 5 with well-developed anterior and posterior laminae (
<figureCitation id="13342A5FCA2C477EFD28F8A8FCA61133" box="[686,805,1799,1825]" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="7" pageNumber="551">Figure 4B</figureCitation>
). Proximal process of maxilla rounded; premaxilla with vertically extended anterior and blunt posterior tip; dentary with large, expanded posterodorsal coronoid process (
<figureCitation id="13342A5FCA2C477EFF19F8E0FE82117B" box="[159,257,1871,1897]" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="7" pageNumber="551">Figure 4</figureCitation>
C–E). Wing shaped, large epibranchial 1 with posterior lamina plain, anterior lamina on medial margin protruding; gill rakers simple, sparsely placed, 2–3 on lateral and 3–4 on medial margin of epibranchial 1; basihyal large, 2 gill rakers on lateral and 9–10 on medial margin of ceratobranchial 1 (
<figureCitation id="13342A5FCA2C477EFB9DF838FB3111A3" box="[1051,1202,1943,1969]" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="7" pageNumber="551">Figure 4F, G</figureCitation>
). Total vertebrae 30, including 17 abdominal and 13 caudal vertebrae, inclusive of last caudal centrum; supraneural 3 broad, not extending over anterior plane of proximal-middle radial of dorsal-fin, leaving prominent gap between posterior most supraneural and proximal-middle radial of dorsal fin; lateral process of second vertebral centrum large, elongate, extending over base of outer arm of os suspensorium and up to medial base of neural arch 4. Epural curved (
<figureCitation id="13342A5FCA224770FAD7FEAAFF3D1751" captionStart="FIGURE 4" captionStartId="8.[151,250,1647,1672]" captionTargetBox="[258,1321,199,1615]" captionTargetId="figure@8.[238,1349,181,1622]" captionTargetPageId="8" captionText="FIGURE 4. Comparison of the skeleton of Waikhomia hira (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for W. sahyadriensis left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="9" pageNumber="553">Figure 4H</figureCitation>
).
</paragraph>
<caption id="DF706652CA234771FF11F9C0FDCA11FD" ID-DOI="http://doi.org/10.5281/zenodo.3920625" ID-Zenodo-Dep="3920625" httpUri="https://zenodo.org/record/3920625/files/figure.png" pageId="8" pageNumber="552" startId="8.[151,250,1647,1672]" targetBox="[258,1321,199,1615]" targetPageId="8">
<paragraph id="8BB036DACA234771FF11F9C0FDCA11FD" blockId="8.[151,1437,1647,2032]" pageId="8" pageNumber="552">
<emphasis id="B97BEAC8CA234771FF11F9C0FE97109A" bold="true" box="[151,276,1647,1672]" pageId="8" pageNumber="552">FIGURE 4.</emphasis>
Comparison of the skeleton of 
<taxonomicName id="4C0F4D59CA234771FDDAF9DFFD7C109A" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[604,767,1648,1672]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="552" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA234771FDDAF9DFFD7C109A" box="[604,767,1648,1672]" italics="true" pageId="8" pageNumber="552">Waikhomia hira</emphasis>
</taxonomicName>
(paratype, BNHS FWF 604, 41.2 mm SL) and 
<taxonomicName id="4C0F4D59CA234771FB6BF9DFFF7710BE" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="552" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA234771FB6BF9DFFF7710BE" italics="true" pageId="8" pageNumber="552">Waikhomia sahyadriensis</emphasis>
</taxonomicName>
(male, BNHS FWF 588, 49.1 mm SL). (A) Lateral view of cleared and double-stained specimens. (B) Infraorbital series, lateral view. (C) Left maxilla, lateral view. (D) Left premaxilla, lateral view. (E) Left dentary, lateral view. (F) Dorsal gill arch, ventral view. (G) Ventral gill arch, dorsal view. (H) Hypural plate, lateral view. Abbreviations: IO1–5, infraorbital 1–5; So, supraorbital; PP, palatine process; Ana, anguloarticular; Cm, coronomeckelian; De, dentary; MC, Meckel’s cartilage; Ra, retroarticular; Bb1–3, basibranchial 1–3; Bb4C, basibranchial 4 cartilage; Bh, basihyal; Cb1–5, ceratobranchial 1–5; CC, compound centrum; CIHPU3–4, inter-hemal spine cartilage of preural centrum 3–4; Eb1–4, epibranchial 1–4; Eb5C, epibranchial 5 cartilage; Ep, epural; Gr, gill rakers; H1–6, hypurals 1–6; Hb1–3, hypobranchial 1–3; HS, hemal spine; NS, neural spine; Pb2–3, pharyngobranchial 2–3; Pb4C, pharyngobranchial 4 cartilage; Ph, parhypural; Pls, pleurostyle; PU2–3, preural centra 2–3. Asterisk (*) in red for 
<taxonomicName id="4C0F4D59CA234771FE29F81AFDD911DE" box="[431,602,1972,1996]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="552" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA234771FE29F81AFDD911DE" box="[431,602,1972,1996]" italics="true" pageId="8" pageNumber="552">W. sahyadriensis</emphasis>
</taxonomicName>
left premaxilla indicates mechanical damage occurred while dissecting the specimen or anomaly detected. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BB036DACA224770FF41FEE3FCFD14B9" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FEE3FE1F1775" bold="true" box="[199,412,332,359]" pageId="9" pageNumber="553">Coloration in life.</emphasis>
Body bright iridescent silver, becoming reddish in mature males (
<figureCitation id="13342A5FCA224770FB01FEE2FB6B1775" box="[1159,1256,333,359]" captionStart="FIGURE 1" captionStartId="1.[151,250,1517,1542]" captionTargetBox="[152,1434,490,1492]" captionTargetId="figure@1.[151,1435,489,1492]" captionTargetPageId="1" captionText="FIGURE 1. Type locality and collection locations of Waikhomia hira in yellow (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (BNHS FWF 591, 48.8 mm SL) in red in the Western Ghats of India. Star indicates type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.3920613" httpUri="https://zenodo.org/record/3920613/files/figure.png" pageId="9" pageNumber="553">Figure 1</figureCitation>
, 
<figureCitation id="13342A5FCA224770FB71FEE2FA9F1775" box="[1271,1308,333,359]" captionStart="FIGURE 3" captionStartId="7.[151,250,1625,1650]" captionTargetBox="[151,1435,237,1600]" captionTargetId="figure@7.[151,1435,237,1600]" captionTargetPageId="7" captionText="FIGURE 3. Waikhomia hira holotype (male, BNHS FWF 602, 42.9 mm SL) in (A) life and (B) preserved." figureDoi="http://doi.org/10.5281/zenodo.3920623" httpUri="https://zenodo.org/record/3920623/files/figure.png" pageId="9" pageNumber="553">3A</figureCitation>
). Head and dorsum olive green. Lower head, chest, and abdomen silvery. Iris iridescent silver, a dark bar above and below pupil, running vertically across iris. Numerous (7 (3) or 8 (4)) small dark rhomboidal spots on side of body, restricted to lateral-line scale row: first (cleithral) spot small, mostly confined to first scale of lateral-line row, rarely extending onto scale above; post-pectoral spot small, between pectoral fin and pelvic-fin origin, covering 7 
<superScript id="7C7A9B92CA224770FB67FE73FB6D17F8" attach="left" box="[1249,1262,476,490]" fontSize="6" pageId="9" pageNumber="553">th</superScript>
scale of lateral line, marginally extending over ventral half of scale above; lateral spot small, above pelvic fin base, on or between 8 
<superScript id="7C7A9B92CA224770FF23FD8BFF42142D" attach="none" box="[165,193,548,575]" fontSize="6" pageId="9" pageNumber="553">th–</superScript>
9 
<superScript id="7C7A9B92CA224770FF48FD8BFF581420" attach="left" box="[206,219,548,562]" fontSize="6" pageId="9" pageNumber="553">th</superScript>
lateral-line scales; 1 (2) or 2 (5) post-pelvic spots between base of pelvic fin and anal-fin origin; small superanal spot above anal-fin base; two small caudal spots (
<figureCitation id="13342A5FCA224770FD74FDE6FCEA1471" box="[754,873,585,611]" captionStart="FIGURE 2" captionStartId="2.[151,250,1466,1491]" captionTargetBox="[158,1429,279,1431]" captionTargetId="figure@2.[151,1436,273,1442]" captionTargetPageId="2" captionText="FIGURE 2. Schematic drawing of colour pattern in (A) Waikhomia hira and (B) Waikhomia sahyadriensis. Abbreviations: Cs, caudal spot; Cls, cleithral spot; Dbs, dorsal-base spot; Ls, lateral spot; Sans, Super-anal spot; Ppcs, post-pectoral spot; Ppls, post-pelvic spot; Nps, Nape spot." figureDoi="http://doi.org/10.5281/zenodo.3920617" httpUri="https://zenodo.org/record/3920617/files/figure.png" pageId="9" pageNumber="553">Figure 2A</figureCitation>
). Dorsal-fin red in both sexes, however its anterior margin and tip black in mature males; pelvic fin jet black with white tip; pectoral, anal and caudal fins hyaline, with no markings. Each body scale bordered with melanophore at base.
</paragraph>
<paragraph id="8BB036DACA224770FF41FD1BFD241529" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FD1BFD8514DD" bold="true" box="[199,518,692,719]" pageId="9" pageNumber="553">Coloration in preservative.</emphasis>
Body pale cream; snout, head, dorsum, lower lip, cheek and gill cover dark brown; lower head, chest and abdominal region uniformly pale cream (
<figureCitation id="13342A5FCA224770FCD7FD76FC4614E1" box="[849,965,729,755]" captionStart="FIGURE 3" captionStartId="7.[151,250,1625,1650]" captionTargetBox="[151,1435,237,1600]" captionTargetId="figure@7.[151,1435,237,1600]" captionTargetPageId="7" captionText="FIGURE 3. Waikhomia hira holotype (male, BNHS FWF 602, 42.9 mm SL) in (A) life and (B) preserved." figureDoi="http://doi.org/10.5281/zenodo.3920623" httpUri="https://zenodo.org/record/3920623/files/figure.png" pageId="9" pageNumber="553">Figure 3B</figureCitation>
). Iris whitish. Body markings as described for live specimens. Dorsal-fin hyaline in both sexes with tip of fin black in mature males; pelvic, pectoral, anal and caudal fins are as described for live specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3156551CA224770FF41FCEAFF7B15FD" pageId="9" pageNumber="553" type="etymology">
<paragraph id="8BB036DACA224770FF41FCEAFCA21591" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FCEAFECD154D" bold="true" box="[199,334,837,863]" pageId="9" pageNumber="553">Etymology.</emphasis>
The species epithet hira (phonetic hira means diamond and refers to the small symmetrical, rhomboidal spots on the side of the body. A noun in apposition.
</paragraph>
<paragraph id="8BB036DACA224770FF41FC23FF7B15FD" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FC23FE0715B5" bold="true" box="[199,388,908,935]" pageId="9" pageNumber="553">Common name.</emphasis>
Kohinoor Barb, after the Indian Koh-i-Noor diamond, which is now a part of the crown jewels of the 
<collectingCountry id="F318764ACA224770FF66FC1EFE2215D9" box="[224,417,945,971]" name="United Kingdom" pageId="9" pageNumber="553">United Kingdom</collectingCountry>
, in reference to the rhomboidal, ‘diamond-shaped’ spots along the lateral-line scale row of this fish.
</paragraph>
</subSubSection>
<subSubSection id="C3156551CA224770FF41FC57FB4910D2" pageId="9" pageNumber="553" type="distribution">
<paragraph id="8BB036DACA224770FF41FC57FDED1249" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FC57FEE21201" bold="true" box="[199,353,1016,1043]" pageId="9" pageNumber="553">Distribution.</emphasis>
<taxonomicName id="4C0F4D59CA224770FEEEFC56FD9F1201" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[360,540,1017,1043]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FEEEFC56FD9F1201" box="[360,540,1017,1043]" italics="true" pageId="9" pageNumber="553">Waikhomia hira</emphasis>
</taxonomicName>
is currently known only from the upstream catchment of the west-flowing Kali River system in the northern Western Ghats, with the 
<typeStatus id="54B48878CA224770FD7CFBB2FCA91225" box="[762,810,1053,1079]" pageId="9" pageNumber="553">type</typeStatus>
series collected near Chandewadi, Kamra, Uttara Kannada District, 
<collectingRegion id="49CBF838CA224770FEB3FBEEFE281249" box="[309,427,1089,1115]" country="India" name="Karnataka" pageId="9" pageNumber="553">Karnataka</collectingRegion>
, 
<collectingCountry id="F318764ACA224770FE30FBEEFE731249" box="[438,496,1089,1115]" name="India" pageId="9" pageNumber="553">India</collectingCountry>
(
<figureCitation id="13342A5FCA224770FE78FBEEFDDE1249" box="[510,605,1089,1115]" captionStart="FIGURE 1" captionStartId="1.[151,250,1517,1542]" captionTargetBox="[152,1434,490,1492]" captionTargetId="figure@1.[151,1435,489,1492]" captionTargetPageId="1" captionText="FIGURE 1. Type locality and collection locations of Waikhomia hira in yellow (paratype, BNHS FWF 604, 41.2 mm SL) and Waikhomia sahyadriensis (BNHS FWF 591, 48.8 mm SL) in red in the Western Ghats of India. Star indicates type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.3920613" httpUri="https://zenodo.org/record/3920613/files/figure.png" pageId="9" pageNumber="553">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="8BB036DACA224770FF41FBCBFF4C136E" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FBCBFE3D126D" bold="true" box="[199,446,1124,1151]" pageId="9" pageNumber="553">Habitat and ecology.</emphasis>
<taxonomicName id="4C0F4D59CA224770FE40FBCAFDF9126D" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[454,634,1125,1151]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FE40FBCAFDF9126D" box="[454,634,1125,1151]" italics="true" pageId="9" pageNumber="553">Waikhomia hira</emphasis>
</taxonomicName>
occurs in large, deep (~ 
<quantity id="4CF79B3FCA224770FC16FBCAFC39126D" box="[912,954,1125,1151]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="553" unit="m" value="1.0">1 m</quantity>
) pools in the main river channel, in which the flow is sluggish, with fallen branches and other detritus, sand, large boulders and gravel as substratum. Co-occurring fishes in the 
<typeStatus id="54B48878CA224770FEFAFB02FE2F12D5" box="[380,428,1197,1223]" pageId="9" pageNumber="553">type</typeStatus>
locality of 
<taxonomicName id="4C0F4D59CA224770FDAAFB01FDFD12D5" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[556,638,1197,1223]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FDAAFB01FDFD12D5" box="[556,638,1197,1223]" italics="true" pageId="9" pageNumber="553">W. hira</emphasis>
</taxonomicName>
included 
<taxonomicName id="4C0F4D59CA224770FD6DFB02FCE312D5" baseAuthorityName="Pethiyagoda" baseAuthorityYear="2013" box="[747,864,1197,1223]" class="Actinopterygii" family="Cyprinidae" genus="Haludaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FD6DFB02FCE312D5" box="[747,864,1197,1223]" italics="true" pageId="9" pageNumber="553">Haludaria</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C0F4D59CA224770FC12FB02FAC512D5" authority="(Day, 1870)" baseAuthorityName="Day" baseAuthorityYear="1870" box="[916,1350,1197,1223]" class="Actinopterygii" family="Cyprinidae" genus="Hypselobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="B97BEAC8CA224770FC12FB02FB3B12D5" box="[916,1208,1197,1223]" italics="true" pageId="9" pageNumber="553">Hypselobarbus pulchellus</emphasis>
(Day, 1870)
</taxonomicName>
, 
<taxonomicName id="4C0F4D59CA224770FAD4FB02FE9312F9" authorityName="Hora" authorityYear="1942" class="Actinopterygii" family="Cyprinidae" genus="Osteochilichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FAD4FB02FE9312F9" italics="true" pageId="9" pageNumber="553">Osteochilichthys</emphasis>
</taxonomicName>
cf. 
<emphasis id="B97BEAC8CA224770FEBDFB7EFEFC12F9" box="[315,383,1233,1259]" italics="true" pageId="9" pageNumber="553">nashii</emphasis>
(Day, 1869), 
<taxonomicName id="4C0F4D59CA224770FD9CFB7EFAB612F9" authority="Katwate, Paingankar, Raghavan &amp; Dahanukar 2014" authorityName="Katwate, Paingankar, Raghavan &amp; Dahanukar" authorityYear="2014" box="[538,1333,1233,1259]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="B97BEAC8CA224770FD9CFB7EFD6B12F9" box="[538,744,1233,1259]" italics="true" pageId="9" pageNumber="553">Pethia longicauda</emphasis>
<bibRefCitation id="EF9E4B2BCA224770FD69FB7EFAB612F9" author="Katwate, U. &amp; Paingankar, M. S. &amp; Raghavan, R. &amp; Dahanukar, N." box="[751,1333,1233,1259]" pageId="9" pageNumber="553" pagination="235 - 248" refId="ref9188" refString="Katwate, U., Paingankar, M. S., Raghavan, R. &amp; Dahanukar, N. (2014 b) Pethia longicauda, a new species of barb (Teleostei: Cyprinidae) from the northern Western Ghats, India. Zootaxa, 3846 (2), 235 - 248. https: // doi. org / 10.11646 / zootaxa. 3846.2.4" type="journal article" year="2014">Katwate, Paingankar, Raghavan &amp; Dahanukar 2014</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0F4D59CA224770FAC6FB7DFDE91302" authority="(Jerdon, 1849)" baseAuthorityName="Jerdon" baseAuthorityYear="1849" class="Actinopterygii" family="Cyprinidae" genus="Devario" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="malabaricus">
<emphasis id="B97BEAC8CA224770FAC6FB7DFEA0131D" italics="true" pageId="9" pageNumber="553">Devario malabaricus</emphasis>
(Jerdon, 1849) (Cyprinidae)
</taxonomicName>
; 
<taxonomicName id="4C0F4D59CA224770FDF3FB5AFF721326" authority="Kumar, Katwate, Raghavan &amp; Dahanukar 2016" authorityName="Kumar, Katwate, Raghavan &amp; Dahanukar" authorityYear="2016" class="Actinopterygii" family="Balitoridae" genus="Balitora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="chipkali">
<emphasis id="B97BEAC8CA224770FDF3FB5AFCB7131D" box="[629,820,1269,1295]" italics="true" pageId="9" pageNumber="553">Balitora chipkali</emphasis>
<bibRefCitation id="EF9E4B2BCA224770FCBAFB5AFAD0131D" author="Kumar, S. &amp; Stecher, G. &amp; Tamura, K." box="[828,1363,1269,1295]" pageId="9" pageNumber="553" pagination="1870 - 1874" refId="ref9411" refString="Kumar, S., Stecher, G. &amp; Tamura, K. (2016) MEGA 7: Molecular Evolutionary Genetics Analysis version 7.0 for bigger datasets. Molecular Biology and Evolution, 33, 1870 - 1874. https: // doi. org / 10.1093 / molbev / msw 054" type="journal article" year="2016">Kumar, Katwate, Raghavan &amp; Dahanukar 2016</bibRefCitation>
(Balitoridae)
</taxonomicName>
, 
<taxonomicName id="4C0F4D59CA224770FF7BFAB6FE021321" authorityName="McClelland" authorityYear="1839" box="[253,385,1305,1331]" class="Actinopterygii" family="Aplocheilidae" genus="Aplocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FF7BFAB6FE021321" box="[253,385,1305,1331]" italics="true" pageId="9" pageNumber="553">Aplocheilus</emphasis>
</taxonomicName>
cf. 
<emphasis id="B97BEAC8CA224770FE2BFAB6FD851321" box="[429,518,1305,1331]" italics="true" pageId="9" pageNumber="553">lineatus</emphasis>
(Valenciennes, 1846) (
<taxonomicName id="4C0F4D59CA224770FC8EFAB6FC281326" box="[776,939,1305,1332]" class="Actinopterygii" family="Aplocheilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="553" phylum="Chordata" rank="family">Aplocheilidae</taxonomicName>
), 
<taxonomicName id="4C0F4D59CA224770FC39FAB6FEF2134A" authority="(Lacepede, 1800)" baseAuthorityName="Lacepede" baseAuthorityYear="1800" class="Actinopterygii" family="Mastacembelidae" genus="Mastacembelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Synbranchiformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="armatus">
<emphasis id="B97BEAC8CA224770FC39FAB6FB521321" box="[959,1233,1305,1331]" italics="true" pageId="9" pageNumber="553">Mastacembelus armatus</emphasis>
(Lacepède, 1800) (Mastacembelidae)
</taxonomicName>
, 
<taxonomicName id="4C0F4D59CA224770FEFDFA92FD931345" authorityName="Blyth" authorityYear="1860" box="[379,528,1341,1367]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FEFDFA92FD931345" box="[379,528,1341,1367]" italics="true" pageId="9" pageNumber="553">Glyptothorax</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C0F4D59CA224770FDC3FA92FD30134A" authorityName="Bleeker" authorityYear="1858" box="[581,691,1341,1368]" class="Actinopterygii" family="Sisoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="553" phylum="Chordata" rank="family">Sisoridae</taxonomicName>
), and 
<taxonomicName id="4C0F4D59CA224770FD73FA92FCF61345" authorityName="Swainson" authorityYear="1839" box="[757,885,1341,1367]" class="Actinopterygii" family="Synbranchidae" genus="Ophichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Synbranchiformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FD73FA92FCF61345" box="[757,885,1341,1367]" italics="true" pageId="9" pageNumber="553">Ophichthys</emphasis>
</taxonomicName>
cf. 
<emphasis id="B97BEAC8CA224770FC18FA92FC6C1345" box="[926,1007,1341,1367]" italics="true" pageId="9" pageNumber="553">indicus</emphasis>
(Silas &amp; Dawson, 1961) (
<taxonomicName id="4C0F4D59CA224770FA94FA92FF41136E" class="Actinopterygii" family="Synbranchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Synbranchiformes" pageId="9" pageNumber="553" phylum="Chordata" rank="family">Synbranchidae</taxonomicName>
).
</paragraph>
<paragraph id="8BB036DACA224770FF41FA2AFB4910D2" blockId="9.[151,1437,153,1728]" pageId="9" pageNumber="553">
<emphasis id="B97BEAC8CA224770FF41FA2AFE35138D" bold="true" box="[199,438,1412,1439]" pageId="9" pageNumber="553">Molecular analysis.</emphasis>
The best partition scheme and nucleotide substitution model for the partition was TPM3u+I+G4 (BIC=63891.691, lnL=-30030.611, df=513) for the combined partition of the first two codon positions of 
<emphasis id="B97BEAC8CA224770FF74FA62FEA413F5" box="[242,295,1485,1511]" italics="true" pageId="9" pageNumber="553">cox1</emphasis>
and 
<emphasis id="B97BEAC8CA224770FEDBFA62FE0F13F5" box="[349,396,1485,1511]" italics="true" pageId="9" pageNumber="553">cytb</emphasis>
, and TIM2+R3 (BIC=63201.904, lnL=-30215.861, df=371) for the combined partition the of third codon positions of 
<emphasis id="B97BEAC8CA224770FE28FA5EFE601019" box="[430,483,1521,1547]" italics="true" pageId="9" pageNumber="553">cox1</emphasis>
and 
<emphasis id="B97BEAC8CA224770FD9AFA5EFDC81019" box="[540,587,1521,1547]" italics="true" pageId="9" pageNumber="553">cytb</emphasis>
. Maximum likelihood analysis revealed 
<taxonomicName id="4C0F4D59CA224770FB9CFA5DFBEE1019" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[1050,1133,1521,1547]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FB9CFA5DFBEE1019" box="[1050,1133,1521,1547]" italics="true" pageId="9" pageNumber="553">W. hira</emphasis>
</taxonomicName>
and the widely distributed 
<taxonomicName id="4C0F4D59CA224770FF11F9B9FED7103D" box="[151,340,1557,1583]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA224770FF11F9B9FED7103D" box="[151,340,1557,1583]" italics="true" pageId="9" pageNumber="553">W. sahyadriensis</emphasis>
</taxonomicName>
to have a sister-group relationship (
<figureCitation id="13342A5FCA224770FD6CF9BAFCCF103D" box="[746,844,1556,1583]" captionStart="FIGURE 5" captionStartId="10.[151,250,1946,1971]" captionTargetBox="[298,1292,187,1915]" captionTargetId="figure@10.[291,1297,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylogenetic position of Waikhomia hira based on maximum likelihood analysis using best partition scheme. Species of Garra are used as the out-group. Values along the nodes are percent bootstraps for 1000 iterations. Bootstrap values less than 50% are not shown." figureDoi="http://doi.org/10.5281/zenodo.3920629" httpUri="https://zenodo.org/record/3920629/files/figure.png" pageId="9" pageNumber="553">Figure 5</figureCitation>
). These two species form a well-supported (ultrafast bootstrap =100) monophyletic clade, which is recovered as the sister group to 
<taxonomicName id="4C0F4D59CA224770FB81F995FBDE1041" authorityName="Hamilton" authorityYear="1822" box="[1031,1117,1594,1619]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="genus">
<emphasis id="B97BEAC8CA224770FB81F995FBDE1041" box="[1031,1117,1594,1619]" italics="true" pageId="9" pageNumber="553">Puntius</emphasis>
</taxonomicName>
but with weak node support (ultrafast bootstrap = 42). The genetic distances between 
<taxonomicName id="4C0F4D59CA224770FCA3F9F1FCFA1065" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[805,889,1629,1655]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FCA3F9F1FCFA1065" box="[805,889,1629,1655]" italics="true" pageId="9" pageNumber="553">W. hira</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F4D59CA224770FC34F9F1FBF31065" box="[946,1136,1629,1655]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA224770FC34F9F1FBF31065" box="[946,1136,1629,1655]" italics="true" pageId="9" pageNumber="553">W. sahyadriensis</emphasis>
</taxonomicName>
are 4.3–4.9% in 
<emphasis id="B97BEAC8CA224770FAB0F9F2FAEF1065" box="[1334,1388,1629,1655]" italics="true" pageId="9" pageNumber="553">cox1</emphasis>
and 7.0–8.2% in 
<emphasis id="B97BEAC8CA224770FEA1F92EFED51089" box="[295,342,1665,1691]" italics="true" pageId="9" pageNumber="553">cytb</emphasis>
. The genetic gap between 
<taxonomicName id="4C0F4D59CA224770FD07F92DFD501089" authorityName="Katwate &amp; Kumkar &amp; Raghavan &amp; Dahanukar" authorityYear="2020" box="[641,723,1665,1691]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="hira">
<emphasis id="B97BEAC8CA224770FD07F92DFD501089" box="[641,723,1665,1691]" italics="true" pageId="9" pageNumber="553">W. hira</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0F4D59CA224770FC8CF92DFC441089" box="[778,967,1665,1691]" class="Actinopterygii" family="Cyprinidae" genus="Waikhomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="553" phylum="Chordata" rank="species" species="sahyadriensis">
<emphasis id="B97BEAC8CA224770FC8CF92DFC441089" box="[778,967,1665,1691]" italics="true" pageId="9" pageNumber="553">W. sahyadriensis</emphasis>
</taxonomicName>
, based on the maximum intraspecific distances and minimum interspecies distances, was 3.4% in 
<emphasis id="B97BEAC8CA224770FC9EF90AFCCE10AD" box="[792,845,1701,1727]" italics="true" pageId="9" pageNumber="553">cox1</emphasis>
and 4.3% in 
<emphasis id="B97BEAC8CA224770FC67F90AFB9310AD" box="[993,1040,1701,1727]" italics="true" pageId="9" pageNumber="553">cytb</emphasis>
gene (
<figureCitation id="13342A5FCA224770FBDCF90AFB3F10D2" box="[1114,1212,1701,1728]" captionStart="FIGURE 6" captionStartId="11.[151,250,833,858]" captionTargetBox="[162,1422,187,798]" captionTargetId="figure@11.[157,1430,181,807]" captionTargetPageId="11" captionText="FIGURE 6. Genetic gap analysis for Waikhomia hira and Waikhomia sahyadriensis based on intra and inter species genetic distances for (A) cytochrome oxidase subunit 1 (cox1) and (B) cytochrome b (cytb). There is a genetic gap of 3.4%, which separates the two species based on cox1, and 4.3%, which separates them based on cytb." figureDoi="http://doi.org/10.5281/zenodo.3920633" httpUri="https://zenodo.org/record/3920633/files/figure.png" pageId="9" pageNumber="553">Figure 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>