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<document id="99238B428B633D1B64AB130C467E906A" ID-DOI="10.11646/zootaxa.5027.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448029" ID-ZooBank="CCF10500-01A3-476C-A4BE-57161E164EBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464259914" checkinUser="plazi" docAuthor="Kompier, Tom, Karube, Haruki, Futahashi, Ryo &amp; Phan, Quoc Toan" docDate="2021" docId="03A687CBFFCCFFEEE0E77509FC4DFCF3" docLanguage="en" docName="zootaxa.5027.1.1.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Planaeschna celia : Wilson &amp; Reels 2001" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="FF9FFFB3FFC0FFFEE0707207FFC3FFCA" masterDocTitle="The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="13" updateTime="1699280300883" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1608207E8D107FB14ED36A39389D49F2">The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)</mods:title>
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<mods:namePart id="3A589E5998F546DFFE4DD6CDC188F1F1">Kompier, Tom</mods:namePart>
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<mods:namePart id="772B99467E0EE81AD9F1B682CC6277F8">Karube, Haruki</mods:namePart>
<mods:affiliation id="5E13FA6DB3FF1D48CC070673A4540E0C">Kanagawa Prefectural Museum of Natural History, Odawara, Japan.</mods:affiliation>
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<mods:namePart id="5504E8E6BB1CD0CA7A7B2114B9769F62">Futahashi, Ryo</mods:namePart>
<mods:affiliation id="DE1CD73142BFC84AC0CF7ACA5F922CCA">National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan.</mods:affiliation>
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<mods:namePart id="13E8F6914A03FE713B0C404546FDB87B">Phan, Quoc Toan</mods:namePart>
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<mods:affiliation id="6906F6FBAAB93E4F0669C9ADA6835B60">The Center for Entomology &amp; Parasitology Research, College of Medicine &amp; Pharmacy, Duy Tan University, 550000, Da Nang, Vietnam. pqtoan 84 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3154 - 6546</mods:affiliation>
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<paragraph id="8BB036DDFFCCFFF2E0E77509FD67F8E2" blockId="12.[151,676,1805,1868]" box="[151,676,1805,1832]" pageId="12" pageNumber="13">
<heading id="D0F881B1FFCCFFF2E0E77509FD67F8E2" bold="true" box="[151,676,1805,1832]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B97BEACFFFCCFFF2E0E77509FD67F8E2" bold="true" box="[151,676,1805,1832]" pageId="12" pageNumber="13">
6.
<taxonomicName id="4C0F4D5EFFCCFFF2E0C5750AFD67F8E2" ID-CoL="4JJ3M" authority="Wilson &amp; Reels, 2001" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[181,676,1805,1832]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFCCFFF2E0C5750AFE52F8E2" bold="true" box="[181,401,1805,1832]" italics="true" pageId="12" pageNumber="13">Planaeschna celia</emphasis>
<bibRefCitation id="EF9E4B2CFFCCFFF2E1E9750AFD67F8E2" author="Wilson, K. D. P. &amp; Reels, G. T." box="[409,676,1805,1832]" pageId="12" pageNumber="13" pagination="145 - 208" refId="ref17637" refString="Wilson, K. D. P. &amp; Reels, G. T. (2001) Odonata of Hainan, China. Odonatologica, 30, 145 - 208." type="journal article" year="2001">Wilson &amp; Reels, 2001</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BB036DDFFCCFFF2E0E77535FE64F886" blockId="12.[151,676,1805,1868]" box="[151,423,1842,1868]" pageId="12" pageNumber="13">
(
<figureCitation id="13342A58FFCCFFF2E0EF7535FF24F886" box="[159,231,1842,1868]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[201,1386,181,1920]" captionTargetId="figure-22@14.[201,1386,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Planaeschna celia males [A] scan of dorsal habitus, [B] face, [CD] lateral habitus alive. (A) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (BD) same location, 26-X-2014 (two individuals)." figureDoi="http://doi.org/10.5281/zenodo.5448047" httpUri="https://zenodo.org/record/5448047/files/figure.png" pageId="12" pageNumber="13">Figs 8</figureCitation>
,
<figureCitation id="13342A58FFCCFFF2E0847535FE63F886" box="[244,416,1842,1868]" captionStart="FIGURE 9" captionStartId="15.[151,250,1874,1898]" captionTargetBox="[187,1401,181,1850]" captionTargetId="figure-17@15.[187,1401,181,1850]" captionTargetPageId="15" captionText="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="12" pageNumber="13">9B, 9D, 9F, 9H</figureCitation>
)
</paragraph>
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<subSubSection id="C3156556FFCCFFF2E0E77571FB55F844" box="[151,1174,1910,1934]" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8BB036DDFFCCFFF2E0E77571FB55F844" blockId="12.[151,1174,1910,1934]" box="[151,1174,1910,1934]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB1F11F3FFCCFFF2E0E77571FB55F844" box="[151,1174,1910,1934]" pageId="12" pageNumber="13">
<taxonomicName id="4C0F4D5EFFCCFFF2E0E77571FD8AF844" ID-CoL="4JJ3M" authority=": Wilson &amp; Reels (2001)" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[151,585,1910,1934]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFCCFFF2E0E77571FE92F844" box="[151,337,1910,1934]" italics="true" pageId="12" pageNumber="13">Planaeschna celia</emphasis>
:
<bibRefCitation id="EF9E4B2CFFCCFFF2E12F7571FD8AF844" author="Wilson, K. D. P. &amp; Reels, G. T." box="[351,585,1910,1934]" pageId="12" pageNumber="13" pagination="145 - 208" refId="ref17637" refString="Wilson, K. D. P. &amp; Reels, G. T. (2001) Odonata of Hainan, China. Odonatologica, 30, 145 - 208." type="journal article" year="2001">Wilson &amp; Reels (2001)</bibRefCitation>
</taxonomicName>
pp. 178180, Figs 8891;
<treatmentCitation id="0AAE10CCFFCCFFF2E32A7571FBDEF844" author="Zhang, H." box="[858,1053,1910,1934]" page="77" pageId="12" pageNumber="13" year="2019">
<bibRefCitation id="EF9E4B2CFFCCFFF2E32A7571FC27F844" author="Zhang, H." box="[858,996,1910,1934]" pageId="12" pageNumber="13" refId="ref17812" refString="Zhang, H. (2019) Dragonflies and damselflies of China. Vol. 1. Chongqing University Press, Chongqing, 736 pp." type="book" year="2019">Zhang (2019)</bibRefCitation>
p. 77
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(checklist).
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</paragraph>
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<subSubSection id="C3156556FFCCFFEEE0E775BBFD46FCDF" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BB036DDFFCCFFF2E0E775BBFEE0F830" blockId="12.[151,1436,1980,2042]" pageId="12" pageNumber="13">
<emphasis id="B97BEACFFFCCFFF2E0E775BBFE4FF81C" bold="true" box="[151,396,1980,2006]" pageId="12" pageNumber="13">Materials examined.</emphasis>
<materialsCitation id="3B673C80FFCCFFF2E1E475BAFC74F81C" box="[404,951,1980,2007]" collectingDate="2014-10-26" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="12" pageNumber="13" specimenCount="2" specimenCount-male="2" stateProvince="Phu Tho">
<specimenCount id="9D09FD54FFCCFFF2E1E475BAFE16F81D" box="[404,469,1981,2007]" pageId="12" pageNumber="13" type="male">2 ♂♂</specimenCount>
,
<collectorName id="26FA530BFFCCFFF2E19275BAFDBDF81D" box="[482,638,1981,2007]" pageId="12" pageNumber="13">Xuan Son NP</collectorName>
,
<collectingRegion id="49CBF83FFFCCFFF2E2F975BBFCEDF81D" box="[649,814,1980,2007]" country="Vietnam" name="Phu Tho" pageId="12" pageNumber="13">Phu Tho Prov.</collectingRegion>
,
<date id="FFB1101DFFCCFFF2E34975BBFC74F81C" box="[825,951,1980,2006]" pageId="12" pageNumber="13" value="2014-10-26">
<collectingDate id="EFF5E9F5FFCCFFF2E34975BBFC74F81C" box="[825,951,1980,2006]" pageId="12" pageNumber="13" value="2014-10-26">26-X-2014</collectingDate>
</date>
</materialsCitation>
, TK leg.;
<materialsCitation id="3B673C80FFCCFFF2E45875BAFEDCF830" collectingDate="2016-11-20" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho">
<specimenCount id="9D09FD54FFCCFFF2E45875BAFB96F81D" box="[1064,1109,1981,2007]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
, same location and collector,
<date id="FFB1101DFFCCFFF2E0E775E7FEDCF830" box="[151,287,2016,2042]" pageId="12" pageNumber="13" value="2016-11-20">
<collectingDate id="EFF5E9F5FFCCFFF2E0E775E7FEDCF830" box="[151,287,2016,2042]" pageId="12" pageNumber="13" value="2016-11-20">20-XI-2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF706655FFCDFFF3E0E77590FC59F819" ID-DOI="http://doi.org/10.5281/zenodo.5448045" ID-Zenodo-Dep="5448045" httpUri="https://zenodo.org/record/5448045/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1943,1967]" targetBox="[225,1362,181,1920]" targetPageId="13">
<paragraph id="8BB036DDFFCDFFF3E0E77590FC59F819" blockId="13.[151,1437,1943,2004]" pageId="13" pageNumber="14">
<emphasis id="B97BEACFFFCDFFF3E0E77590FED0F87A" bold="true" box="[151,275,1943,1968]" pageId="13" pageNumber="14">FIGURE 7.</emphasis>
<taxonomicName id="4C0F4D5EFFCDFFF3E169759FFE39F87A" authorityName=": Karube" authorityYear="2011" box="[281,506,1944,1968]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="B97BEACFFFCDFFF3E169759FFE39F87A" box="[281,506,1944,1968]" italics="true" pageId="13" pageNumber="14">Planaeschna asahinai</emphasis>
</taxonomicName>
alive in hand [A, B] habitus, [C, D] face. (A, C) male, 7-IX-2016, Sao La NR, Thua Thien- Hue Prov., (B, D) female, 6-VIII-2016, Bach Ma NP, Thua Thien-Hue Prov.
</paragraph>
</caption>
<caption id="DF706655FFCEFFF0E0E77590FC55F81E" ID-DOI="http://doi.org/10.5281/zenodo.5448047" ID-Zenodo-Dep="5448047" httpUri="https://zenodo.org/record/5448047/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1943,1967]" targetBox="[201,1386,181,1920]" targetPageId="14">
<paragraph id="8BB036DDFFCEFFF0E0E77590FC55F81E" blockId="14.[151,1436,1943,2004]" pageId="14" pageNumber="15">
<emphasis id="B97BEACFFFCEFFF0E0E77590FED7F865" bold="true" box="[151,276,1943,1967]" pageId="14" pageNumber="15">FIGURE 8.</emphasis>
<taxonomicName id="4C0F4D5EFFCEFFF0E16B759FFE15F87A" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[283,470,1944,1968]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFCEFFF0E16B759FFE15F87A" box="[283,470,1944,1968]" italics="true" pageId="14" pageNumber="15">Planaeschna celia</emphasis>
</taxonomicName>
males [A] scan of dorsal habitus, [B] face, [CD] lateral habitus alive. (A) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (BD) same location, 26-X-2014 (two individuals).
</paragraph>
</caption>
<caption id="DF706655FFCFFFF1E0E77555FE50F830" ID-DOI="http://doi.org/10.5281/zenodo.5448049" ID-Zenodo-Dep="5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1874,1898]" targetBox="[187,1401,181,1850]" targetPageId="15">
<paragraph id="8BB036DDFFCFFFF1E0E77555FE50F830" blockId="15.[151,1436,1874,2043]" pageId="15" pageNumber="16">
<emphasis id="B97BEACFFFCFFFF1E0E77555FED5F8A0" bold="true" box="[151,278,1874,1898]" pageId="15" pageNumber="16">FIGURE 9.</emphasis>
Comparison of details of male
<taxonomicName id="4C0F4D5EFFCFFFF1E21A7555FCB6F8A0" authorityName=": Asahina" authorityYear="1996" box="[618,885,1874,1898]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFCFFFF1E21A7555FCB6F8A0" box="[618,885,1874,1898]" italics="true" pageId="15" pageNumber="16">Planaeschna tamdaoensis</emphasis>
</taxonomicName>
and male
<taxonomicName id="4C0F4D5EFFCFFFF1E3977554FBF6F8A0" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[999,1077,1874,1898]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFCFFFF1E3977554FBF6F8A0" box="[999,1077,1874,1898]" italics="true" pageId="15" pageNumber="16">P. celia</emphasis>
</taxonomicName>
[A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G]
<taxonomicName id="4C0F4D5EFFCFFFF1E45D7570FB07F844" authorityName=": Asahina" authorityYear="1996" box="[1069,1220,1910,1934]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFCFFFF1E45D7570FB07F844" box="[1069,1220,1910,1934]" italics="true" pageId="15" pageNumber="16">P. tamdaoensis</emphasis>
</taxonomicName>
, [B, D, F, H]
<taxonomicName id="4C0F4D5EFFCFFFF1E53C7570FA55F844" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[1356,1430,1910,1934]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFCFFFF1E53C7570FA55F844" box="[1356,1430,1910,1934]" italics="true" pageId="15" pageNumber="16">P. celia</emphasis>
</taxonomicName>
. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015.
</paragraph>
</caption>
<paragraph id="8BB036DDFFD0FFEEE0B77291FBEDFF7B" blockId="16.[151,1437,150,826]" box="[199,1070,150,177]" pageId="16" pageNumber="17">
<emphasis id="B97BEACFFFD0FFEEE0B77291FE2FFF7A" bold="true" box="[199,492,150,176]" pageId="16" pageNumber="17">Additional observations.</emphasis>
<materialsCitation id="3B673C80FFD0FFEEE1837290FBE8FF7B" box="[499,1067,151,177]" collectingDate="2014-11-09" collectorName="Xuan Son NP" location="Phu Tho Prov." pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho">
<specimenCount id="9D09FD54FFD0FFEEE1837290FDDCFF7B" box="[499,543,151,177]" pageId="16" pageNumber="17" type="male">1 ♂</specimenCount>
,
<collectorName id="26FA530BFFD0FFEEE25B7290FD06FF7B" box="[555,709,151,177]" pageId="16" pageNumber="17">Xuan Son NP</collectorName>
,
<collectingRegion id="49CBF83FFFD0FFEEE2A07290FCB0FF7B" box="[720,883,151,177]" country="Vietnam" name="Phu Tho" pageId="16" pageNumber="17">Phu Tho Prov.</collectingRegion>
,
<date id="FFB1101DFFD0FFEEE30E7290FC34FF7B" box="[894,1015,151,177]" pageId="16" pageNumber="17" value="2014-11-09">
<collectingDate id="EFF5E9F5FFD0FFEEE30E7290FC34FF7B" box="[894,1015,151,177]" pageId="16" pageNumber="17" value="2014-11-09">9-XI-2014</collectingDate>
</date>
, TK
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFD0FFEEE0B772BCFBCEFE67" blockId="16.[151,1437,150,826]" pageId="16" pageNumber="17">
<emphasis id="B97BEACFFFD0FFEEE0B772BCFED2FF1F" bold="true" box="[199,273,187,213]" pageId="16" pageNumber="17">Notes.</emphasis>
<materialsCitation id="3B673C80FFD0FFEEE16B72BCFBCAFE67" collectorName="At Xuan Son NP &amp; Xuan Son NP" country="China" location="Hainan" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Hainan">
This species was assumed endemic to
<collectingRegion id="49CBF83FFFD0FFEEE2AD72BCFCF2FF1F" box="[733,817,187,213]" country="China" name="Hainan" pageId="16" pageNumber="17">Hainan</collectingRegion>
,
<collectingCountry id="F318764DFFD0FFEEE34F72BCFC46FF1F" box="[831,901,187,213]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
, where it is known from only a few locations (
<bibRefCitation id="EF9E4B2CFFD0FFEEE0EF72D8FEF0FF33" author="Wilson, K. D. P." box="[159,307,223,249]" pageId="16" pageNumber="17" pagination="168049" refId="ref17581" refString="Wilson, K. D. P. (2009) Planaeschna celia. The IUCN Red List of Threatened Species, 2009, e. T 168049 A 6454669. https: // doi. org / 10.2305 / IUCN. UK. 2009 - 2. RLTS. T 168049 A 6454669. en" type="journal article" year="2009">Wilson 2009</bibRefCitation>
) and considered vulnerable.
<collectorName id="26FA530BFFD0FFEEE20C72D8FCFCFF33" box="[636,831,223,249]" pageId="16" pageNumber="17">At Xuan Son NP</collectorName>
the species was found at only one location, where it inhabits a densely vegetated shallow stream of 67 meters wide under heavy forest cover. The stream springs forth from the karst rocks and is only several hundred meters long before joining a larger stream. Males were observed hovering low over small open areas of water. Although the area was visited many times, this species was only encountered three times. In
<collectingRegion id="49CBF83FFFD0FFEEE1AB7368FDEFFE43" box="[475,556,367,393]" country="China" name="Hainan" pageId="16" pageNumber="17">Hainan</collectingRegion>
it was recorded in June, but at
<collectorName id="26FA530BFFD0FFEEE3D27368FB82FE43" box="[930,1089,367,393]" pageId="16" pageNumber="17">Xuan Son NP</collectorName>
it was found during October- November. It occurred with five other
<taxonomicName id="4C0F4D5EFFD0FFEEE2377394FCECFE67" box="[583,815,403,429]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BEACFFFD0FFEEE2377394FD1BFE67" box="[583,728,403,429]" italics="true" pageId="16" pageNumber="17">Planaeschna</emphasis>
species
</taxonomicName>
at the same stream
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFD0FFEEE0B773B0FE45FD4F" blockId="16.[151,1437,150,826]" pageId="16" pageNumber="17">
<materialsCitation id="3B673C80FFD0FFEEE0B773B0FE42FD4F" location="Hainan" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
Vietnamese specimens of
<taxonomicName id="4C0F4D5EFFD0FFEEE19F73B0FD81FE1B" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[495,578,439,465]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFD0FFEEE19F73B0FD81FE1B" box="[495,578,439,465]" italics="true" pageId="16" pageNumber="17">P. celia</emphasis>
</taxonomicName>
are almost identical to the
<collectingRegion id="49CBF83FFFD0FFEEE30A73B0FC08FE1B" box="[890,971,439,465]" country="China" name="Hainan" pageId="16" pageNumber="17">Hainan</collectingRegion>
specimens from the original description, displaying the characteristic basal ventral hump adorned with small spines of the cerci and the strongly inwardcurved anterior hamule with a strongly pointed tip. Meanwhile, the Xuan Son specimens have two squarish yellow spots on the labrum (
<figureCitation id="13342A58FFD0FFEEE1F47024FE23FDF7" box="[388,480,547,573]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[201,1386,181,1920]" captionTargetId="figure-22@14.[201,1386,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Planaeschna celia males [A] scan of dorsal habitus, [B] face, [CD] lateral habitus alive. (A) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (BD) same location, 26-X-2014 (two individuals)." figureDoi="http://doi.org/10.5281/zenodo.5448047" httpUri="https://zenodo.org/record/5448047/files/figure.png" pageId="16" pageNumber="17">Figs 8B</figureCitation>
,
<figureCitation id="13342A58FFD0FFEEE19D7024FDD3FDF7" box="[493,528,547,573]" captionStart="FIGURE 9" captionStartId="15.[151,250,1874,1898]" captionTargetBox="[187,1401,181,1850]" captionTargetId="figure-17@15.[187,1401,181,1850]" captionTargetPageId="15" captionText="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="16" pageNumber="17">9B</figureCitation>
), which are absent in the
<typeStatus id="54B4887FFFD0FFEEE3417024FC54FDF7" box="[817,919,547,573]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
. Given only a few specimens are known from
<collectingRegion id="49CBF83FFFD0FFEEE0E77040FF28FDAB" box="[151,235,583,609]" country="China" name="Hainan" pageId="16" pageNumber="17">Hainan</collectingRegion>
, it is difficult to say whether this is within normal variation, or whether this may imply some difference between populations
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFD0FFEEE0B77088FD46FCDF" blockId="16.[151,1437,150,826]" pageId="16" pageNumber="17">
<taxonomicName id="4C0F4D5EFFD0FFEEE0B77088FE5BFD63" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[199,408,655,681]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFD0FFEEE0B77088FE5BFD63" box="[199,408,655,681]" italics="true" pageId="16" pageNumber="17">Planaeschna celia</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="4C0F4D5EFFD0FFEEE2BC7088FCB4FD63" authorityName=": Asahina" authorityYear="1996" box="[716,887,655,681]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFD0FFEEE2BC7088FCB4FD63" box="[716,887,655,681]" italics="true" pageId="16" pageNumber="17">P. tamdaoensis</emphasis>
</taxonomicName>
, but differs by the pattern on S2, facial pattern and anal appendages.
<taxonomicName id="4C0F4D5EFFD0FFEEE1E070B4FD9DFD07" authorityName=": Wilson &amp; Reels" authorityYear="2001" box="[400,606,691,717]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="celia">
<emphasis id="B97BEACFFFD0FFEEE1E070B4FD9DFD07" box="[400,606,691,717]" italics="true" pageId="16" pageNumber="17">Planaeschna celia</emphasis>
</taxonomicName>
lacks the yellow lower border of the antefrons (
<figureCitation id="13342A58FFD0FFEEE4F170B4FB19FD07" box="[1153,1242,691,717]" captionStart="FIGURE 9" captionStartId="15.[151,250,1874,1898]" captionTargetBox="[187,1401,181,1850]" captionTargetId="figure-17@15.[187,1401,181,1850]" captionTargetPageId="15" captionText="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
) seen in many
<taxonomicName id="4C0F4D5EFFD0FFEEE5F870B4FEE0FD3B" authorityName=": Asahina" authorityYear="1996" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFD0FFEEE5F870B4FEE0FD3B" italics="true" pageId="16" pageNumber="17">P. tamdaoensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0F4D5EFFD0FFEEE14170D0FD97FD3B" authorityName=": Asahina" authorityYear="1996" box="[305,596,727,753]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="B97BEACFFFD0FFEEE14170D0FD97FD3B" box="[305,596,727,753]" italics="true" pageId="16" pageNumber="17">Planaeschna tamdaoensis</emphasis>
</taxonomicName>
also lacks the characteristic ventral basal hump (
<figureCitation id="13342A58FFD0FFEEE40570D0FB0EFD3B" box="[1141,1229,727,753]" captionStart="FIGURE 9" captionStartId="15.[151,250,1874,1898]" captionTargetBox="[187,1401,181,1850]" captionTargetId="figure-17@15.[187,1401,181,1850]" captionTargetPageId="15" captionText="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="16" pageNumber="17">Fig. 9D</figureCitation>
) and has one large central yellow area on the labrum (
<figureCitation id="13342A58FFD0FFEEE26E70FCFDBBFCDF" box="[542,632,763,789]" captionStart="FIGURE 9" captionStartId="15.[151,250,1874,1898]" captionTargetBox="[187,1401,181,1850]" captionTargetId="figure-17@15.[187,1401,181,1850]" captionTargetPageId="15" captionText="FIGURE 9. Comparison of details of male Planaeschna tamdaoensis and male P. celia [A, B] facial pattern, [C, D] anal appendages lateral, [E, F] anal appendages dorsal, [G, H] dorsal pattern of S2. [A, C, E, G] P. tamdaoensis, [B, D, F, H] P. celia. (A) Pia Oac NP, Cao Bang Prov., 12-IX-2015, (B) Xuan Son NP, Phu Tho Prov., 26-X-2014 (different specimen than Fig. 2B), (C, E) Tam Dao NP, Vinh Phuc Prov., 27-VIII-2016, (D, F, H) Xuan Son NP, Phu Tho Prov., 20-XI-2016, (G) Pia Oac NP, Cao Bang Prov., 27-VI-2015." figureDoi="http://doi.org/10.5281/zenodo.5448049" httpUri="https://zenodo.org/record/5448049/files/figure.png" pageId="16" pageNumber="17">Fig. 9A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFD0FFEEE0B77119FC4DFCF3" box="[199,910,798,826]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BB036DDFFD0FFEEE0B77119FC4DFCF3" blockId="16.[151,1437,150,826]" box="[199,910,798,826]" pageId="16" pageNumber="17">
<emphasis id="B97BEACFFFD0FFEEE0B77119FEA2FCF2" bold="true" box="[199,353,798,824]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="F318764DFFD0FFEEE1177118FE69FCF3" box="[359,426,799,825]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
(
<collectingRegion id="49CBF83FFFD0FFEEE1C97118FDCFFCF3" box="[441,524,799,825]" country="China" name="Hainan" pageId="16" pageNumber="17">Hainan</collectingRegion>
Island),
<collectingCountry id="F318764DFFD0FFEEE21E7118FD0EFCF3" box="[622,717,799,825]" name="Vietnam" pageId="16" pageNumber="17">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF83FFFD0FFEEE2AB7118FC41FCF3" box="[731,898,799,825]" country="Vietnam" name="Phu Tho" pageId="16" pageNumber="17">Phu Tho Prov.</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>