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<document ID-DOI="http://doi.org/10.5281/zenodo.6458594" ID-GBIF-Dataset="adeeb71f-7f8d-4e00-bc9f-35089363f76e" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6458594" approvalRequired="120" approvalRequired_for_taxonomicNames="95" approvalRequired_for_treatments="25" checkinTime="1600878147105" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03A687BCFFA6FFA613BFFC42FBE9FFC8" docLanguage="en" docName="hbmw_9_Phyllostomidae_444.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Phyllostomus elongatus E. Geoffroy Saint-Hilaire 1810" docType="treatment" docVersion="8" lastPageNumber="507" masterDocId="FF9FFFC4FFB1FFB1133CFFBAFFE0F244" masterDocTitle="Phyllostomidae" masterLastPageNumber="583" masterPageNumber="444" pageNumber="507" updateTime="1656353518553" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phyllostomidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>444</mods:start>
<mods:end>583</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6458594</mods:identifier>
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<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6726944" ID-Zenodo-Dep="6726944" LSID="urn:lsid:plazi:treatment:03A687BCFFA6FFA613BFFC42FBE9FFC8" httpUri="http://treatment.plazi.org/id/03A687BCFFA6FFA613BFFC42FBE9FFC8" lastPageNumber="507" pageId="23" pageNumber="507">
<subSubSection box="[131,189,1016,1062]" pageId="23" pageNumber="507" type="multiple">
<paragraph blockId="23.[129,1169,1016,1102]" box="[131,189,1016,1062]" pageId="23" pageNumber="507">
<heading box="[131,189,1016,1062]" pageId="23" pageNumber="507">
<figureCitation box="[131,189,1016,1062]" captionStart="Plate 36: Phyllostomidae" captionStartId="17.[137,167,3288,3313]" captionTargetBox="[23,2765,17,3655]" captionTargetPageId="16" captionText="24. Long-legged Bat (Macrophyllum macrophyllum), 25. Fringe-lipped Bat (Trachops cirrhosus), 26. Striped Hairy-nosed Bat (Gardnerycteris crenulatum), 27. Keenan's Hairy-nosed Bat (Gardnerycteris keenani), 28. Koepckes Hairy-nosed Bat (Gardnerycteris koepckeae), 29. Kalkos Round-eared Bat (Lophostoma kalkoae), 30. Pygmy Round-eared Bat (Lophostoma brasiliense), 31. Carrikers Round-eared Bat (Lophostoma carrikeri), 32. Schulzs Round-eared Bat (Lophostoma schulzi), 33. Western Round-eared Bat (Lophostoma occidentale), 34. Davis's Round-eared Bat (Lophostoma evotis), 35. White-throated Round-eared Bat (Lophostoma silvicola), 36. Greater Round-eared Bat (Tonatia bidens), 37. Stripe-headed Round-eared Bat (Tonatia saurophila), 38. Pale-faced Bat (Phylloderma stenops), 39. Pale Spear-nosed Bat (Phyllostomus discolor), 40. Lesser Spear-nosed Bat (Phyllostomus elongatus), 41. Greater Spear-nosed Bat (Phyllostomus hastatus), 42. Guianan Spear-nosed Bat (Phyllostomus latifolius), 43. Woolly False Vampire Bat (Chrotopterus auritus), 44. Southern Golden Bat (Mimon bennettii), 45. Cozumelan Golden Bat (Mimon cozumelae), 46. Spectral Bat (Vampyrum spectrum)" figureDoi="http://doi.org/10.5281/zenodo.6458675" httpUri="https://zenodo.org/record/6458675/files/figure.png" pageId="23" pageNumber="507">40.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[205,686,1016,1062]" pageId="23" pageNumber="507" type="vernacular_names">
<paragraph blockId="23.[129,1169,1016,1102]" box="[205,686,1016,1062]" pageId="23" pageNumber="507">
<heading box="[205,686,1016,1062]" pageId="23" pageNumber="507">
<vernacularName box="[205,686,1016,1062]" pageId="23" pageNumber="507">Lesser Spear-nosed Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[735,1169,1016,1062]" pageId="23" pageNumber="507" type="nomenclature">
<paragraph blockId="23.[129,1169,1016,1102]" box="[735,1169,1016,1062]" pageId="23" pageNumber="507">
<heading box="[735,1169,1016,1062]" pageId="23" pageNumber="507">
<taxonomicName authority="E. Geoffroy Saint-Hilaire, 1810" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1810" box="[735,1169,1016,1062]" class="Mammalia" family="Phyllostomidae" genus="Phyllostomus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="elongatus">
<emphasis box="[735,1169,1016,1062]" italics="true" pageId="23" pageNumber="507">Phyllostomus elongatus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[131,1104,1080,1101]" pageId="23" pageNumber="507" type="vernacular_names">
<paragraph blockId="23.[129,1169,1016,1102]" box="[131,1104,1080,1101]" pageId="23" pageNumber="507">
<heading box="[131,1104,1080,1101]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[131,206,1080,1101]" pageId="23" pageNumber="507">French:</emphasis>
<vernacularName box="[216,483,1080,1101]" pageId="23" pageNumber="507">Phyllostome des sous-bois</vernacularName>
/
<emphasis bold="true" box="[504,595,1080,1101]" pageId="23" pageNumber="507">German:</emphasis>
<vernacularName box="[605,790,1080,1101]" pageId="23" pageNumber="507">Kleine Lanzennase</vernacularName>
/
<emphasis bold="true" box="[811,903,1080,1101]" pageId="23" pageNumber="507">Spanish:</emphasis>
<vernacularName box="[913,1008,1080,1101]" pageId="23" pageNumber="507">Filostoma</vernacularName>
pequeno
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="reference_group">
<paragraph blockId="23.[743,1335,1146,1574]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[743,899,1146,1179]" pageId="23" pageNumber="507">Taxonomy.</emphasis>
<taxonomicName authority="E. Geoffroy Saint-Hilaire, 1810" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1810" class="Mammalia" family="Phyllostomidae" genus="Phyllostoma" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="elongatum">Phyllostoma elongatum E. Geoffroy Saint-Hilaire, 1810</taxonomicName>
,
<materialsCitation pageId="23" pageNumber="507">
type locality not given. Restricted by A. Cabrera in 1958 to Rio Branco,
<collectingRegion box="[920,1099,1269,1298]" country="Brazil" name="Mato Grosso" pageId="23" pageNumber="507">Mato Grosso</collectingRegion>
,
<collectingCountry box="[1115,1197,1269,1298]" name="Brazil" pageId="23" pageNumber="507">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[744,1106,1305,1338]" pageId="23" pageNumber="507" type="discussion">
<paragraph blockId="23.[743,1335,1146,1574]" box="[744,1106,1305,1338]" pageId="23" pageNumber="507">This species is monotypic.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6458683" ID-Zenodo-Dep="6458683" httpUri="https://zenodo.org/record/6458683/files/figure.png" inLine="true" pageId="23" pageNumber="507" targetBox="[129,720,1157,1571]" targetPageId="23">
<paragraph blockId="23.[743,1335,1146,1574]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[743,920,1344,1377]" pageId="23" pageNumber="507">Distribution.</emphasis>
Lowlands E of Andes in
<collectingCountry name="Colombia" pageId="23" pageNumber="507">Colombia</collectingCountry>
,
<collectingCountry box="[863,1010,1387,1416]" name="Venezuela" pageId="23" pageNumber="507">Venezuela</collectingCountry>
, the Guianas, Amazonian Basin to SE
<collectingCountry box="[990,1056,1427,1456]" name="Peru" pageId="23" pageNumber="507">Peru</collectingCountry>
and N &amp; C
<collectingCountry box="[1232,1328,1427,1456]" name="Bolivia" pageId="23" pageNumber="507">Bolivia</collectingCountry>
, and E
<collectingCountry box="[843,924,1462,1495]" name="Brazil" pageId="23" pageNumber="507">Brazil</collectingCountry>
, also an isolated population in W
<collectingCountry box="[819,956,1502,1535]" name="Colombia" pageId="23" pageNumber="507">Colombia</collectingCountry>
and NW
<collectingCountry box="[1091,1210,1502,1535]" name="Ecuador" pageId="23" pageNumber="507">Ecuador</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="description">
<paragraph blockId="23.[743,1335,1146,1574]" lastBlockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[743,986,1541,1574]" pageId="23" pageNumber="507">Descriptive notes.</emphasis>
Head-body 65-100 mm, tail 15-24 mm, ear 25-32 mm, hindfoot 14-20 mm, forearm 58-71 mm; weight 30-57 g. The Lesser Spear-nosed Bat is large and robust. Its morphology is very similar to the Greater Spear-nosed Bat (
<taxonomicName box="[483,624,1660,1693]" class="Mammalia" family="Phyllostomidae" genus="Phyllostomus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="hastatus">P. hastatus</taxonomicName>
), but it is much smaller and less robust in all aspects. Fur is short, soft, velvety, and glossy; dorsal fur is brown to reddish brown; head, neck, and shoulders are often paler than back; venter is paler than back to fairly dark. Head is large; snout is broad and robust; face and noseleaf are black to dark brown; noseleaf is medium-sized, broad, and pear-shaped, with base horseshoe-shaped and entire free edge separated from upper lip; ears are medium-sized, broad, and triangular, with rounded tips; eyes are large; and chin has small tubercles forming a Vsshape. Wing membranes are blackish, but tips are sometimes white; caudal membrane is developed; calcar is longer than foot (more than
<quantity box="[449,551,1979,2008]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="507" unit="mm" value="14.0">14 mm</quantity>
); and tail is short, reaching to middle of membrane or less. Teeth are strong and pointed. Chromosomal complement has 2n = 32 and FN = 58.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="biology_ecology">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[129,241,2054,2087]" pageId="23" pageNumber="507">Habitat.</emphasis>
Mainly evergreen forest habitats but also humid tropical to low subtropical forests, Andean foothills mainly in Amazonia, and humid tropical forest in
<collectingRegion box="[1243,1334,2093,2126]" country="Panama" name="Embera-Wounaan" pageId="23" pageNumber="507">Choco</collectingRegion>
Ecoregion at elevations of 60-1100 m. The Lesser Spearnosed Bat is occasionally found in drier habitats near streams, semi-humid forests, and flooded savannas. It also occurs in terra firma forest; flooded forests; swamps; primary, secondary, selectively logged and gallery forests; forest edges; cultivated areas; and pastures. It usually flies low over streams and rivers.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="food_feeding">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[129,386,2330,2363]" pageId="23" pageNumber="507">Food and Feeding.</emphasis>
Diet of the Lesser Spear-nosed Bat includes small vertebrates, flower parts, nectar, fruits, and large insects. In
<collectingCountry box="[699,765,2374,2403]" name="Peru" pageId="23" pageNumber="507">Peru</collectingCountry>
, pollen has been found on it. In
<collectingCountry box="[1234,1327,2374,2403]" name="Bolivia" pageId="23" pageNumber="507">Bolivia</collectingCountry>
, several families of insects occur in its diet (
<taxonomicName box="[727,852,2417,2442]" class="Insecta" family="Blattidae" kingdom="Animalia" order="Blattodea" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Blattidae</taxonomicName>
,
<taxonomicName box="[870,997,2417,2442]" class="Insecta" family="Gryllidae" kingdom="Animalia" order="Orthoptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Gryllidae</taxonomicName>
,
<taxonomicName box="[1013,1168,2417,2442]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Reduviidae</taxonomicName>
,
<taxonomicName box="[1186,1327,2417,2442]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
,
<taxonomicName box="[131,330,2448,2481]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
, Hydrophylidae,
<taxonomicName box="[576,758,2448,2481]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
, and
<taxonomicName box="[843,1006,2448,2481]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="507" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
), along with orthopterans, coleopterans, and other undetermined arthropods. In
<collectingCountry box="[968,1048,2492,2521]" name="Brazil" pageId="23" pageNumber="507">Brazil</collectingCountry>
, fruit of Rollinia mucosa (
<taxonomicName box="[203,371,2531,2560]" class="Magnoliopsida" kingdom="Plantae" order="Magnoliales" pageId="23" pageNumber="507" phylum="Tracheophyta" rank="order">Magnoliales</taxonomicName>
,
<taxonomicName box="[386,561,2531,2560]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="23" pageNumber="507" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
) was eaten. It is occasionally carnivorous; in Amazonas,
<collectingCountry box="[130,210,2566,2599]" name="Brazil" pageId="23" pageNumber="507">Brazil</collectingCountry>
, it was reported eating ajuvenile of Sebas Short-tailed Bat (
<taxonomicName box="[1038,1297,2566,2599]" class="Mammalia" family="Phyllostomidae" genus="Carollia" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="perspicillata">Carollia perspicillata</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="breeding">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[130,264,2606,2639]" pageId="23" pageNumber="507">Breeding.</emphasis>
Reproductive female Lesser Spear-nosed Bats have been found in June in
<collectingCountry box="[131,268,2645,2678]" name="Colombia" pageId="23" pageNumber="507">Colombia</collectingCountry>
and July-August in
<collectingCountry box="[552,620,2645,2678]" name="Peru" pageId="23" pageNumber="507">Peru</collectingCountry>
. In western
<collectingCountry box="[791,909,2645,2678]" name="Ecuador" pageId="23" pageNumber="507">Ecuador</collectingCountry>
, mothers with young attached to their nipples have been recorded in November and February. In northern
<collectingCountry box="[1246,1328,2685,2718]" name="Brazil" pageId="23" pageNumber="507">Brazil</collectingCountry>
, several pregnant and lactating females were caught at beginning of rainy season.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="activity">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[128,363,2764,2797]" pageId="23" pageNumber="507">Activity patterns.</emphasis>
The Lesser Spear-nosed Bat is nocturnal and aerial. It emerges from its roosts at complete darkness and forage for c.2 hours before returning to roosts. In
<collectingCountry box="[173,268,2843,2876]" name="Bolivia" pageId="23" pageNumber="507">Bolivia</collectingCountry>
, it has been captured in mist net at 18:00-20:00 h. Echolocation calls are 51-4-78-2 kHz. It roosts in hollow trees, culverts, and tunnels and under bridges.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="biology_ecology">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[130,824,2921,2954]" pageId="23" pageNumber="507">Movements, Home range and Social organization.</emphasis>
Lesser Spear-nosed Bats roost in aggregations, divided up into smaller social units consisting of permanent harems of a single male and up to twelve females and their young and groups of non-breeding males. This social structure is very similar to that of the Greater Spear-nosed Bat. In
<collectingRegion box="[1170,1328,3040,3073]" country="Ecuador" name="Esmeraldas" pageId="23" pageNumber="507">Esmeraldas</collectingRegion>
,
<collectingCountry box="[131,246,3079,3112]" name="Ecuador" pageId="23" pageNumber="507">Ecuador</collectingCountry>
, a colony of ¢.200 individuals was found sharing space with Sebas Short-tailed Bats. In
<collectingRegion box="[244,309,3118,3151]" country="Bolivia" name="Beni" pageId="23" pageNumber="507">Beni</collectingRegion>
,
<collectingCountry box="[325,419,3118,3151]" name="Bolivia" pageId="23" pageNumber="507">Bolivia</collectingCountry>
, an individual was found roosting in a tree hollow of
<taxonomicName class="Magnoliopsida" family="Phytolaccaceae" genus="Gallesia" kingdom="Plantae" order="Caryophyllales" pageId="23" pageNumber="507" phylum="Tracheophyta" rank="species" species="integrifolia">Gallesia integrifolia</taxonomicName>
(
<taxonomicName box="[226,442,3158,3191]" class="Magnoliopsida" family="Phytolaccaceae" kingdom="Plantae" order="Caryophyllales" pageId="23" pageNumber="507" phylum="Tracheophyta" rank="family">Phytolaccaceae</taxonomicName>
), sharing space with Sebas Short-tailed Bats, Common Vampire Bats (
<taxonomicName box="[206,459,3198,3231]" class="Mammalia" family="Phyllostomidae" genus="Desmodus" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="rotundus">Desmodus rotundus</taxonomicName>
), and Pallass Long-tongued Bats (
<taxonomicName box="[936,1200,3198,3231]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="23" pageNumber="507" phylum="Chordata" rank="species" species="soricina">Glossophaga soricina</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="conservation">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[130,478,3237,3270]" pageId="23" pageNumber="507">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Lesser Spear-nosed Batis fairly common and widespread. Its conservation status is considered stable and is found at several protected areas.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="507" type="bibRefCitation_list">
<paragraph blockId="23.[128,1337,1585,3468]" pageId="23" pageNumber="507">
<emphasis bold="true" box="[131,283,3364,3389]" pageId="23" pageNumber="507">Bibliography.</emphasis>
Aguirre (2002), Aguirre &amp; Teran (2007), Aguirre, Lens &amp; Matthysen (2003), Albuja (1999), Cabrera (1958), Fischer et al. (1997), Gardner (1977b), Handley (1976), Hill (1965a), Nogueira, Peracchi &amp; Moratelli (2007),
<collectingRegion box="[215,280,3443,3468]" country="Uruguay" name="Rivera" pageId="23" pageNumber="507">Rivera</collectingRegion>
et al. (2015), Tirira (2017), Tuttle (1970), Williams &amp; Genoways (2008).
</paragraph>
</subSubSection>
</treatment>
</document>