treatments-xml/data/03/A6/87/03A687A0FD7EE318D6F96DA8FC1BF96B.xml
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<document id="3029041129EA4A9F9BE6440B1689096E" ID-CLB-Dataset="7605" ID-DOI="10.11646/zootaxa.4942.2.3" ID-GBIF-Dataset="cebb6e47-b2e0-45bb-85d2-e73b7fae63a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4600445" ID-ZooBank="0F533688-AB41-47D1-95A3-AF0B501A9D26" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615535301405" checkinUser="plazi" docAuthor="Chamorro, William, Lopera-Toro, Alejandro &amp; Rossini, Michele" docDate="2021" docId="03A687A0FD7EE318D6F96DA8FC1BF96B" docLanguage="en" docName="zootaxa.4942.2.3.pdf" docOrigin="Zootaxa 4942 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Dichotomius) quadrilobatus Chamorro, Lopera, &amp; Rossini 2021, new species" docType="treatment" docUuid="337EA216-4014-4EA1-8992-DA5D5B2DF143" docUuidSource="ZooBank" docVersion="7" lastPageNumber="202" masterDocId="FF9FFFD8FD79E311D66E6F2CFFFBFFF2" masterDocTitle="A new species and distribution records of Dichotomius Hope, 1838 (Coleoptera Scarabaeidae: Scarabaeinae) in Colombia" masterLastPageNumber="206" masterPageNumber="193" pageNumber="200" updateTime="1698933328055" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3E283D102D2213E3AF1483868FB2D917">A new species and distribution records of Dichotomius Hope, 1838 (Coleoptera Scarabaeidae: Scarabaeinae) in Colombia</mods:title>
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<mods:namePart id="4932C020A8113C3381318408688887CE">Chamorro, William</mods:namePart>
<mods:affiliation id="E1BE7B2F38C6F0A08C6B22C28979AB9C">Carrera 3 no. 25 - 32. Villa Recreo, San Juan de Pasto, Nariño, Colombia.</mods:affiliation>
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<mods:namePart id="CD19B295BFED81B2A54D5640F8968602">Lopera-Toro, Alejandro</mods:namePart>
<mods:nameIdentifier id="38F306B84548643B946E2839F8BFCD93" type="ORCID">0000-0001-5097-1655</mods:nameIdentifier>
<mods:affiliation id="2B605388F83616063411FE2B41652C6C">Investigador Asociado Fundación Ecotrópico Colombia y Grupo de Investigación Sistemática Biológica, Universidad Pedagógica y Tecnológica de Colombia, Av. Central del Norte 39 - 115, Tunja, Boyacá, Colombia. https: // orcid. org / 0000 - 0001 - 5097 - 1655 Finnish Museum of Natural History “ Luonnontieteellinen keskusmuseo (LUOMUS) ”, University of Helsinki, Pohjoinen Rautatiekatu 13, Helsinki, 00014, Finland. micros. naturae @ gmail. com; https: // orcid. org / 0000 - 0002 - 1938 - 6105</mods:affiliation>
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<mods:namePart id="0CBA4D1C6F28ACBE5BCCC660B0206026">Rossini, Michele</mods:namePart>
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<mods:date id="1F88AF3C8F7A961493CB3F993492059C">2021</mods:date>
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<heading id="D0F881DAFD7EE316D6F96DA8FB4FFD6C" bold="true" box="[151,1204,644,671]" fontSize="11" level="1" pageId="7" pageNumber="200" reason="1">
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<taxonomicName id="4C0F4D35FD7EE316D6F96DA8FBECFD6C" ID-CoL="7ZN53" authority="Chamorro, Lopera, &amp; Rossini" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[151,1047,644,671]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="200" phylum="Arthropoda" rank="species" species="quadrilobatus" status="sp. nov." subGenus="Dichotomius">
<emphasis id="B97BEAA4FD7EE316D6F96DA8FEC9FD6D" bold="true" box="[151,306,644,671]" italics="true" pageId="7" pageNumber="200">Dichotomius</emphasis>
(
<emphasis id="B97BEAA4FD7EE316D72D6DA8FE25FD6D" bold="true" box="[323,478,644,671]" italics="true" pageId="7" pageNumber="200">Dichotomius</emphasis>
)
<emphasis id="B97BEAA4FD7EE316D7816DA8FD6CFD6D" bold="true" box="[495,663,644,671]" italics="true" pageId="7" pageNumber="200">quadrilobatus</emphasis>
Chamorro, Lopera, &amp; Rossini
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,
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(
<figureCitation id="13342A33FD7EE316D6F16D84FF13FD31" box="[159,232,680,707]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="7" pageNumber="200">Figs. 1</figureCitation>
AF, 5)
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD7EE319D6F96DDDFC29FBE3" lastPageId="8" lastPageNumber="201" pageId="7" pageNumber="200" type="materials_examined">
<paragraph id="8BB036B6FD7EE316D6F96DDDFB24FCDC" blockId="7.[151,1437,752,2039]" pageId="7" pageNumber="200">
<emphasis id="B97BEAA4FD7EE316D6F96DDDFED8FCF8" bold="true" box="[151,291,752,779]" pageId="7" pageNumber="200">Type series.</emphasis>
<materialsCitation id="3B673CEBFD7EE316D7456DDCFB24FCDC" ID-GBIF-Occurrence="3061003330" collectingDate="1997-04-03" collectionCode="IAvH" collectorName="A. Lopera &amp; S. Amezquita" country="Colombia" elevation="200" latitude="3.0455556" location="Finca El Esfuerzo" longLatPrecision="21" longitude="-73.595" municipality="Pinalito" pageId="7" pageNumber="200" specimenCount="1" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="54B48814FD7EE316D7456DDCFE68FCF8" box="[299,403,752,778]" pageId="7" pageNumber="200" type="holotype">Holotype</typeStatus>
male (deposited in
<collectionCode id="ED1EAE73FD7EE316D41C6DDDFD4FFCF9" box="[626,692,753,779]" pageId="7" pageNumber="200">IAvH</collectionCode>
), labeled: “
<collectingCountry id="F3187626FD7EE316D55F6DDDFC35FCF9" box="[817,974,753,779]" name="Colombia" pageId="7" pageNumber="200">COLOMBIA</collectingCountry>
:
<collectingRegion id="49CBF854FD7EE316D5B66DDDFBD1FCF9" box="[984,1066,753,779]" country="Colombia" name="Meta" pageId="7" pageNumber="200">META</collectingRegion>
:
<collectingMunicipality id="6BD4ACCCFD7EE316D25B6DDCFB74FCF8" box="[1077,1167,752,778]" pageId="7" pageNumber="200">Piñalito</collectingMunicipality>
,
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,
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<elevation id="0022D185FD7EE316D31C6DDDFF56FCDD" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="200" unit="m" value="200.0">200 m</elevation>
</quantity>
(
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,
<geoCoordinate id="EE3B5071FD7EE316D7246C39FE0BFCDD" box="[330,496,788,815]" degrees="73" direction="west" minutes="35" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="42" value="-73.595">73° 35 42&quot; W</geoCoordinate>
).
<date id="FFB11076FD7EE316D46C6C38FD8FFCDC" box="[514,628,788,814]" pageId="7" pageNumber="200" value="1997-04-03">
<collectingDate id="EFF5E99EFD7EE316D46C6C38FD8FFCDC" box="[514,628,788,814]" pageId="7" pageNumber="200" value="1997-04-03">3-iv-1997</collectingDate>
</date>
.
<collectingMethod id="524E4EA1FD7EE316D4116C38FD38FCDC" box="[639,707,788,814]" pageId="7" pageNumber="200">Pitfall</collectingMethod>
human excrement.
<collectorName id="26FA5360FD7EE316D5F16C39FBEBFCDD" box="[927,1040,789,815]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D25A6C39FB2AFCDC" box="[1076,1233,788,815]" pageId="7" pageNumber="200">S. Amezquita</collectorName>
”.
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</paragraph>
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<materialsCitation id="3B673CEBFD7EE316D6A96C15FEBBFC85" ID-GBIF-Occurrence="3061003308" collectingDate="1958-02" collectionCode="CMNC" collectorName="Oliveira" country="Brazil" latitude="-4.227778" location="Tabatinga" longLatPrecision="21" longitude="-69.90667" pageId="7" pageNumber="200" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54B48814FD7EE316D6A96C15FEC3FCA1" box="[199,312,825,851]" pageId="7" pageNumber="200" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B97BEAA4FD7EE316D7296C14FE49FCA0" bold="true" box="[327,434,824,850]" pageId="7" pageNumber="200">
<collectingCountry id="F3187626FD7EE316D7296C14FE49FCA0" box="[327,434,824,850]" name="Brazil" pageId="7" pageNumber="200">BRAZIL</collectingCountry>
</emphasis>
: AMAZONAS:
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(
<geoCoordinate id="EE3B5071FD7EE316D4946C15FC8DFCA1" box="[762,886,825,851]" degrees="4" direction="south" minutes="13" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="40" value="-4.227778">4°1340S</geoCoordinate>
,
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),
<date id="FFB11076FD7EE316D2566C14FB73FCA1" box="[1080,1160,824,851]" pageId="7" pageNumber="200" value="1958-02">
<collectingDate id="EFF5E99EFD7EE316D2566C14FB73FCA1" box="[1080,1160,824,851]" pageId="7" pageNumber="200" value="1958-02">ii.1958</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D2F76C14FB0EFCA0" box="[1177,1269,824,850]" pageId="7" pageNumber="200">Oliveira</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D3676C15FA85FCA0" box="[1289,1406,824,851]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D3E06C15FF23FC84" pageId="7" pageNumber="200" type="male">2 males</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D68F6C71FEC2FC85" box="[225,313,861,887]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D7256C70FCEAFC69" ID-GBIF-Occurrence="3061003339" collectingDate="1997-03" collectionCode="CALT-ECC" collectorName="C. Duran" country="Colombia" elevation="100" location="Buenos Aires" longitude="-69.983116" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B97BEAA4FD7EE316D7256C70FE11FC84" bold="true" box="[331,490,860,886]" pageId="7" pageNumber="200">
<collectingCountry id="F3187626FD7EE316D7256C70FE11FC84" box="[331,490,860,886]" name="Colombia" pageId="7" pageNumber="200">COLOMBIA</collectingCountry>
</emphasis>
: AMAZONAS:
<location id="8ED0606DFD7EE316D4C76C71FCB9FC84" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D4C76C71FCB9FC84" box="[681,834,860,887]" country="Colombia" longitude="-69.983116" name="Buenos Aires" pageId="7" pageNumber="200">Buenos Aires</location>
,
<quantity id="4CF79B53FD7EE316D5216C71FC6CFC85" box="[847,919,861,887]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="200" unit="m" value="100.0">
<elevation id="0022D185FD7EE316D5216C71FC6CFC85" box="[847,919,861,887]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="200" unit="m" value="100.0">100 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5C66C70FBB9FC85" box="[936,1090,860,887]" pageId="7" pageNumber="200">3°958.19S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D2206C70FAF3FC85" box="[1102,1288,860,887]" degrees="69" direction="west" minutes="58" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="59.22" value="-69.983116">69°5859.22W</geoCoordinate>
),
<date id="FFB11076FD7EE316D3736C70FA8DFC84" box="[1309,1398,860,886]" pageId="7" pageNumber="200" value="1997-03">
<collectingDate id="EFF5E99EFD7EE316D3736C70FA8DFC84" box="[1309,1398,860,886]" pageId="7" pageNumber="200" value="1997-03">iii.1997</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D3ED6C71FF1BFC69" pageId="7" pageNumber="200">C. Durán</collectorName>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D7BA6CACFDECFC68" box="[468,535,896,922]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
<specimenCount id="9D09FD3FFD7EE316D4456CADFD8CFC68" box="[555,631,896,923]" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D4EE6CADFCF1FC69" box="[640,778,897,923]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D5706CACFC32FC4D" ID-GBIF-Occurrence="3061003336" collectingDate="1972-02-20" collectingDateMax="1972-02-25" collectingDateMin="1972-02-20" collectionCode="CMNC" collectorName="Peck &amp; Howden" country="Colombia" elevation="215" location="Leticia" longitude="-69.93584" pageId="7" pageNumber="200" specimenCount="7" specimenCount-female="2" specimenCount-male="5" typeStatus="paratype">
<location id="8ED0606DFD7EE316D5706CACFC94FC68" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D5706CACFC94FC68" box="[798,879,896,922]" country="Colombia" longitude="-69.93584" name="Leticia" pageId="7" pageNumber="200">Leticia</location>
,
<quantity id="4CF79B53FD7EE316D5126CACFC3EFC69" box="[892,965,896,923]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="200" unit="m" value="215.0">
<elevation id="0022D185FD7EE316D5126CACFC3EFC69" box="[892,965,896,923]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="200" unit="m" value="215.0">215 m</elevation>
</quantity>
(
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,
<geoCoordinate id="EE3B5071FD7EE316D2346CACFB18FC69" box="[1114,1251,896,923]" degrees="69" direction="west" minutes="56" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="9" value="-69.93584">69°569W</geoCoordinate>
),
<date id="FFB11076FD7EE316D2946CACFA6CFC69" box="[1274,1431,896,923]" pageId="7" pageNumber="200" value="1972-02-20" valueMax="1972-02-25" valueMin="1972-02-20">
<collectingDate id="EFF5E99EFD7EE316D2946CACFA6CFC69" box="[1274,1431,896,923]" pageId="7" pageNumber="200" value="1972-02-20" valueMax="1972-02-25" valueMin="1972-02-20">2025.ii.1972</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D6F96C88FF36FC4C" box="[151,205,932,958]" pageId="7" pageNumber="200">Peck</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D69C6C88FEAFFC4C" box="[242,340,932,958]" pageId="7" pageNumber="200">Howden</collectorName>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D4506C88FD7AFC4C" box="[574,641,932,958]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
<specimenCount id="9D09FD3FFD7EE316D4FE6C89FCFBFC4C" box="[656,768,932,959]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D5656C88FC99FC4C" box="[779,866,932,959]" pageId="7" pageNumber="200" type="male">5 males</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D5076C89FC3AFC4D" box="[873,961,933,959]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D5BD6C88FBB4FC11" ID-GBIF-Occurrence="3061003347" collectingDate="1974-02-23" collectingDateMax="1974-03-02" collectingDateMin="1974-02-23" collectionCode="CMNC, BDGC" collectorName="Howden &amp; Nealis" country="Colombia" elevation="215" location="Leticia" longitude="-69.93584" pageId="7" pageNumber="200" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<location id="8ED0606DFD7EE316D5BD6C88FBDFFC4C" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D5BD6C88FBDFFC4C" box="[979,1060,932,958]" country="Colombia" longitude="-69.93584" name="Leticia" pageId="7" pageNumber="200">Leticia</location>
,
<quantity id="4CF79B53FD7EE316D2416C88FB8DFC4D" box="[1071,1142,932,959]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="200" unit="m" value="215.0">
<elevation id="0022D185FD7EE316D2416C88FB8DFC4D" box="[1071,1142,932,959]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="200" unit="m" value="215.0">215 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D2E86C89FB01FC4D" box="[1158,1274,933,959]" pageId="7" pageNumber="200">4°1120S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D3686C88FA74FC4D" box="[1286,1423,932,959]" degrees="69" direction="west" minutes="56" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="9" value="-69.93584">69°569W</geoCoordinate>
),
<date id="FFB11076FD7EE316D6F96CE4FEBEFC10" box="[151,325,968,994]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">
<collectingDate id="EFF5E99EFD7EE316D6F96CE4FEBEFC10" box="[151,325,968,994]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">23.ii2.iii.1974</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D7366CE4FE4CFC10" box="[344,439,968,994]" pageId="7" pageNumber="200">Howden</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D7866CE4FDCBFC10" box="[488,560,968,994]" pageId="7" pageNumber="200">Nealis</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D4266CE5FD63FC10" box="[584,664,968,995]" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D4C86CE5FD07FC11" box="[678,764,969,995]" pageId="7" pageNumber="200">CMNC</collectionCode>
;
<specimenCount id="9D09FD3FFD7EE316D5606CE5FC96FC10" box="[782,877,968,995]" pageId="7" pageNumber="200" type="male">2 males</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D5116CE5FC1CFC10" box="[895,999,968,995]" pageId="7" pageNumber="200" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D59B6CE5FBB3FC11" box="[1013,1096,969,995]" pageId="7" pageNumber="200">BDGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D20E6CE5FADAFBF5" ID-GBIF-Occurrence="3061003349" collectingDate="1974-02-23" collectingDateMax="1974-03-02" collectingDateMin="1974-02-23" collectionCode="CMNC" collectorName="S. Peck" country="Colombia" elevation="213" latitude="-4.0736113" location="Leticia" longLatPrecision="21" longitude="-70.00139" municipality="Los Alpes" pageId="7" pageNumber="200" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D20E6CE5FB21FC10" box="[1120,1242,968,995]" pageId="7" pageNumber="200">Los Alpes</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D2826CE4FAC6FC10" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D2826CE4FAC6FC10" box="[1260,1341,968,994]" country="Colombia" latitude="-4.0736113" longLatPrecision="21" longitude="-70.00139" municipality="Los Alpes" name="Leticia" pageId="7" pageNumber="200">Leticia</location>
,
<quantity id="4CF79B53FD7EE316D3216CE5FA67FC11" box="[1359,1436,969,995]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="200" unit="m" value="213.0">
<elevation id="0022D185FD7EE316D3216CE5FA67FC11" box="[1359,1436,969,995]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="200" unit="m" value="213.0">213 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D6CE6CC0FEF4FBF5" box="[160,271,1004,1031]" degrees="4" direction="south" minutes="4" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="25" value="-4.0736113">4°425S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D7756CC0FE6EFBF5" box="[283,405,1004,1031]" degrees="70" direction="west" minutes="0" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="5" value="-70.00139">70°05W</geoCoordinate>
),
<date id="FFB11076FD7EE316D7C46CC0FDA3FBF4" box="[426,600,1004,1030]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">
<collectingDate id="EFF5E99EFD7EE316D7C46CC0FDA3FBF4" box="[426,600,1004,1030]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">23.ii2.iii.1974</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D40D6CC1FD42FBF4" box="[611,697,1004,1031]" pageId="7" pageNumber="200">S. Peck</collectorName>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D5CD6CC0FC1DFBF4" box="[931,998,1004,1030]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
<specimenCount id="9D09FD3FFD7EE316D59B6CC1FB9EFBF4" box="[1013,1125,1004,1031]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D21E6CC1FB41FBF4" box="[1136,1210,1004,1031]" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D2AF6CC1FAE2FBF5" box="[1217,1305,1005,1031]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D3456CC0FA62FBD9" ID-GBIF-Occurrence="3061003306" collectingDate="1974-02-23" collectingDateMax="1974-03-02" collectingDateMin="1974-02-23" collectionCode="CMNC" collectorName="S. Peck" country="Colombia" elevation="213" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" municipality="Road" pageId="7" pageNumber="200" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D3456CC0FA9DFBF4" box="[1323,1382,1004,1030]" pageId="7" pageNumber="200">Road</collectingMunicipality>
end,
<location id="8ED0606DFD7EE316D6F96B3CFF13FBD8" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D6F96B3CFF13FBD8" box="[151,232,1040,1066]" country="Colombia" latitude="-4.0691667" longLatPrecision="21" longitude="-69.998886" municipality="Road" name="Leticia" pageId="7" pageNumber="200">Leticia</location>
,
<quantity id="4CF79B53FD7EE316D69B6B3DFEC6FBD9" box="[245,317,1041,1067]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="200" unit="m" value="213.0">
<elevation id="0022D185FD7EE316D69B6B3DFEC6FBD9" box="[245,317,1041,1067]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="200" unit="m" value="213.0">213 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D7216B3DFE54FBD9" box="[335,431,1041,1067]" degrees="4" direction="south" minutes="4" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="9" value="-4.0691667">4°49S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D7D36B3CFDAFFBD9" box="[445,596,1040,1067]" degrees="69" direction="west" minutes="59" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="56" value="-69.998886">69°5956W</geoCoordinate>
),
<date id="FFB11076FD7EE316D4046B3CFCE3FBD8" box="[618,792,1040,1066]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">
<collectingDate id="EFF5E99EFD7EE316D4046B3CFCE3FBD8" box="[618,792,1040,1066]" pageId="7" pageNumber="200" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">23.ii2.iii.1974</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D54B6B3DFC86FBD8" box="[805,893,1040,1067]" pageId="7" pageNumber="200">S. Peck</collectorName>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D2006B3CFB4AFBD8" box="[1134,1201,1040,1066]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
<specimenCount id="9D09FD3FFD7EE316D2AC6B3DFACAFBD8" box="[1218,1329,1040,1067]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D3576B3DFA6AFBD9" box="[1337,1425,1041,1067]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D6F96B1AFE1CFB81" ID-GBIF-Occurrence="3061003326" collectingDate="2017-10-03" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" county="Florencia" elevation="365" latitude="1.6522222" location="Vereda Brasil" longLatPrecision="21" longitude="-75.59361" municipality="Carano" pageId="7" pageNumber="200" specimenCount="1" stateProvince="Caqueta" typeStatus="paratype">
<collectingRegion id="49CBF854FD7EE316D6F96B1AFEDAFBA3" box="[151,289,1078,1105]" country="Colombia" name="Caqueta" pageId="7" pageNumber="200">CAQUETÁ</collectingRegion>
:
<collectingCounty id="62D14E3AFD7EE316D7436B18FE61FBBC" box="[301,410,1076,1102]" pageId="7" pageNumber="200">Florencia</collectingCounty>
, C.
<collectingMunicipality id="6BD4ACCCFD7EE316D7A66B19FDE7FBBD" box="[456,540,1077,1103]" pageId="7" pageNumber="200">Caraño</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D4466B18FD3FFBBC" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D4466B18FD3FFBBC" box="[552,708,1076,1102]" country="Colombia" county="Florencia" latitude="1.6522222" longLatPrecision="21" longitude="-75.59361" municipality="Carano" name="Vereda Brasil" pageId="7" pageNumber="200" stateProvince="Caqueta">Vereda Brasil</location>
,
<quantity id="4CF79B53FD7EE316D4BE6B18FCE3FBBD" box="[720,792,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="7" pageNumber="200" unit="m" value="365.0">
<elevation id="0022D185FD7EE316D4BE6B18FCE3FBBD" box="[720,792,1076,1103]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="7" pageNumber="200" unit="m" value="365.0">365 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5446B19FC67FBBD" box="[810,924,1077,1103]" degrees="1" direction="north" minutes="39" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="8" value="1.6522222">1°398N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D5C46B18FBBAFBBD" box="[938,1089,1076,1103]" degrees="75" direction="west" minutes="35" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="37" value="-75.59361">75°3537W</geoCoordinate>
),
<date id="FFB11076FD7EE316D2386B19FB41FBBD" box="[1110,1210,1077,1103]" pageId="7" pageNumber="200" value="2017-10-03">
<collectingDate id="EFF5E99EFD7EE316D2386B19FB41FBBD" box="[1110,1210,1077,1103]" pageId="7" pageNumber="200" value="2017-10-03">3.x.2017</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D2A86B19FA67FBBD" box="[1222,1436,1077,1103]" pageId="7" pageNumber="200">Y. Ramos-Pastrana</collectorName>
(15 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D7FE6B75FE1BFB81" box="[400,480,1113,1139]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D79F6B74FCBCFB65" ID-GBIF-Occurrence="3061003318" collectingDate="2015-09-30" collectionCode="LEUA" collectorName="Vereda El Limon &amp; Y. Ramos-Pastrana" country="Colombia" elevation="644" latitude="1.3734167" location="Finca El Limon" longLatPrecision="1" longitude="-75.61017" municipality="Florencia" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D79F6B74FDA5FB80" box="[497,606,1112,1138]" pageId="7" pageNumber="200">Florencia</collectingMunicipality>
,
<collectorName id="26FA5360FD7EE316D4066B74FCD1FB80" box="[616,810,1112,1138]" pageId="7" pageNumber="200">Vereda El Limon</collectorName>
,
<location id="8ED0606DFD7EE316D55A6B74FC1DFB80" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D55A6B74FC1DFB80" box="[820,998,1112,1138]" country="Colombia" latitude="1.3734167" longLatPrecision="1" longitude="-75.61017" municipality="Florencia" name="Finca El Limon" pageId="7" pageNumber="200">Finca El Limon</location>
,
<quantity id="4CF79B53FD7EE316D59E6B74FBCCFB81" box="[1008,1079,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="6.44" pageId="7" pageNumber="200" unit="m" value="644.0">
<elevation id="0022D185FD7EE316D59E6B74FBCCFB81" box="[1008,1079,1112,1139]" metricMagnitude="2" metricUnit="m" metricValue="6.44" pageId="7" pageNumber="200" unit="m" value="644.0">644 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D2286B75FB20FB81" box="[1094,1243,1113,1139]" degrees="1" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="24.3" value="1.3734167">1º2224.3N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D2886B74FA74FB81" box="[1254,1423,1112,1139]" degrees="75" direction="west" minutes="36" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="36.6" value="-75.61017">75º3636.6W</geoCoordinate>
),
<date id="FFB11076FD7EE316D6F96B50FEEAFB65" box="[151,273,1148,1175]" pageId="7" pageNumber="200" value="2015-09-30">
<collectingDate id="EFF5E99EFD7EE316D6F96B50FEEAFB65" box="[151,273,1148,1175]" pageId="7" pageNumber="200" value="2015-09-30">30.ix.2015</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D7706B51FE0FFB65" box="[286,500,1149,1175]" pageId="7" pageNumber="200">Y. Ramos-Pastrana</collectorName>
(4 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D4816B51FCC4FB65" box="[751,831,1149,1175]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D53D6B50FB35FB49" ID-GBIF-Occurrence="3061003315" collectingDate="2010-09-04" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" elevation="249" latitude="1.4988416" location="Vereda La Viciosa. Finca Macagual" longLatPrecision="1" longitude="-75.65698" municipality="Florencia" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D53D6B50FC3BFB64" box="[851,960,1148,1174]" pageId="7" pageNumber="200">Florencia</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D5A26B50FA9EFB64" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D5A26B50FA9EFB64" box="[972,1381,1148,1175]" country="Colombia" latitude="1.4988416" longLatPrecision="1" longitude="-75.65698" municipality="Florencia" name="Vereda La Viciosa. Finca Macagual" pageId="7" pageNumber="200">Vereda La Viciosa. Finca Macagual</location>
,
<quantity id="4CF79B53FD7EE316D31C6B51FF56FB49" metricMagnitude="2" metricUnit="m" metricValue="2.4899999999999998" pageId="7" pageNumber="200" unit="m" value="249.0">
<elevation id="0022D185FD7EE316D31C6B51FF56FB49" metricMagnitude="2" metricUnit="m" metricValue="2.4899999999999998" pageId="7" pageNumber="200" unit="m" value="249.0">249 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D6D26B8CFE99FB49" box="[188,354,1184,1211]" degrees="1" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="55.83" value="1.4988416">1°2955.83N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D7036B8CFDDCFB49" box="[365,551,1184,1211]" degrees="75" direction="west" minutes="39" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="25.12" value="-75.65698">75°3925.12W</geoCoordinate>
),
<date id="FFB11076FD7EE316D4546B8CFD5CFB48" box="[570,679,1184,1210]" pageId="7" pageNumber="200" value="2010-09-04">
<collectingDate id="EFF5E99EFD7EE316D4546B8CFD5CFB48" box="[570,679,1184,1210]" pageId="7" pageNumber="200" value="2010-09-04">4.ix.2010</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D4DF6B8DFC7EFB49" box="[689,901,1185,1211]" pageId="7" pageNumber="200">Y. Ramos-Pastrana</collectorName>
(3 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D2186B8DFB3DFB49" box="[1142,1222,1185,1211]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D2B66B8CFA62FB2D" ID-GBIF-Occurrence="3061003303" collectingDate="2015-05-15" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" elevation="348" latitude="1.4902222" location="Vereda Sabastopol" longLatPrecision="1" longitude="-75.40267" municipality="Florencia" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D2B66B8CFABEFB48" box="[1240,1349,1184,1210]" pageId="7" pageNumber="200">Florencia</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D3216B8CFEEEFB2C" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D3216B8CFEEEFB2C" country="Colombia" latitude="1.4902222" longLatPrecision="1" longitude="-75.40267" municipality="Florencia" name="Vereda Sabastopol" pageId="7" pageNumber="200">Vereda Sabastopol</location>
,
<quantity id="4CF79B53FD7EE316D74B6BE9FE8BFB2D" box="[293,368,1221,1247]" metricMagnitude="2" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="200" unit="m" value="348.0">
<elevation id="0022D185FD7EE316D74B6BE9FE8BFB2D" box="[293,368,1221,1247]" metricMagnitude="2" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="200" unit="m" value="348.0">348 m</elevation>
</quantity>
(
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,
<geoCoordinate id="EE3B5071FD7EE316D4426BE8FD31FB2D" box="[556,714,1220,1247]" degrees="75" direction="west" minutes="24" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="9.6" value="-75.40267">75°249.6W</geoCoordinate>
),
<date id="FFB11076FD7EE316D48D6BE8FCA8FB2D" box="[739,851,1220,1247]" pageId="7" pageNumber="200" value="2015-05-15">
<collectingDate id="EFF5E99EFD7EE316D48D6BE8FCA8FB2D" box="[739,851,1220,1247]" pageId="7" pageNumber="200" value="2015-05-15">15.v.2015</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D50D6BE9FBC7FB2D" box="[867,1084,1221,1247]" pageId="7" pageNumber="200">Y. Ramos-Pastrana</collectorName>
(2 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D32F6BE9FA6AFB2D" box="[1345,1425,1221,1247]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D6F96BC4FDA8FAD5" ID-GBIF-Occurrence="3061003324" collectingDate="2015-04-17" collectionCode="LEUA" collectorName="Pastrana" country="Colombia" county="Montanita" elevation="348" latitude="1.4902222" location="Vereda Las Dalias" longLatPrecision="1" longitude="-75.40267" municipality="Vereda. Morros" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingCounty id="62D14E3AFD7EE316D6F96BC4FEF6FAF0" box="[151,269,1256,1282]" pageId="7" pageNumber="200">Montañita</collectingCounty>
,
<collectingMunicipality id="6BD4ACCCFD7EE316D7706BC4FE2EFAF1" box="[286,469,1256,1283]" pageId="7" pageNumber="200">Vereda. Morros</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D7886BC4FD3AFAF0" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D7886BC4FD3AFAF0" box="[486,705,1256,1283]" country="Colombia" county="Montanita" latitude="1.4902222" longLatPrecision="1" longitude="-75.40267" municipality="Vereda. Morros" name="Vereda Las Dalias" pageId="7" pageNumber="200">Vereda Las Dalias</location>
,
<quantity id="4CF79B53FD7EE316D4BD6BC5FCDBFAF1" box="[723,800,1257,1283]" metricMagnitude="2" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="200" unit="m" value="348.0">
<elevation id="0022D185FD7EE316D4BD6BC5FCDBFAF1" box="[723,800,1257,1283]" metricMagnitude="2" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="200" unit="m" value="348.0">348 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5586BC5FC36FAF1" box="[822,973,1257,1283]" degrees="1" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="24.8" value="1.4902222">1°2924.8N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D5B16BC4FB86FAF1" box="[991,1149,1256,1283]" degrees="75" direction="west" minutes="24" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="9.6" value="-75.40267">75°249.6W</geoCoordinate>
),
<date id="FFB11076FD7EE316D2F96BC4FAF4FAF1" box="[1175,1295,1256,1283]" pageId="7" pageNumber="200" value="2015-04-17">
<collectingDate id="EFF5E99EFD7EE316D2F96BC4FAF4FAF1" box="[1175,1295,1256,1283]" pageId="7" pageNumber="200" value="2015-04-17">17.iv.2015</collectingDate>
</date>
, Y.
<collectorName id="26FA5360FD7EE316D32B6BC5FF0DFAD5" pageId="7" pageNumber="200">Ramos-Pastrana</collectorName>
(28 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D7956A21FDB0FAD5" box="[507,587,1293,1319]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D4306A21FB6FFAB9" ID-GBIF-Occurrence="3061003319" collectingDate="2017-04-03" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" county="San Vicente del Caguan" latitude="2.2988334" location="Finca Rancho Veracruz" longLatPrecision="1" longitude="-74.74147" municipality="Vereda Alto Quebradon" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingCounty id="62D14E3AFD7EE316D4306A21FC94FAD5" box="[606,879,1292,1319]" pageId="7" pageNumber="200">San Vicente del Caguán</collectingCounty>
,
<collectingMunicipality id="6BD4ACCCFD7EE316D5126A20FB72FAD4" box="[892,1161,1292,1318]" pageId="7" pageNumber="200">Vereda Alto Quebradón</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D2FB6A20FA67FAD5" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D2FB6A20FA67FAD5" box="[1173,1436,1292,1319]" country="Colombia" county="San Vicente del Caguan" latitude="2.2988334" longLatPrecision="1" longitude="-74.74147" municipality="Vereda Alto Quebradon" name="Finca Rancho Veracruz" pageId="7" pageNumber="200">Finca Rancho Veracruz</location>
(
<geoCoordinate id="EE3B5071FD7EE316D6CE6A1CFECCFAB9" box="[160,311,1328,1355]" degrees="2" direction="north" minutes="17" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="55.8" value="2.2988334">2°1755.8N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D72D6A1DFE15FAB9" box="[323,494,1329,1355]" degrees="74" direction="west" minutes="44" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="29.3" value="-74.74147">74°4429.3W</geoCoordinate>
),
<date id="FFB11076FD7EE316D46C6A1CFD96FAB8" box="[514,621,1328,1354]" pageId="7" pageNumber="200" value="2017-04-03">
<collectingDate id="EFF5E99EFD7EE316D46C6A1CFD96FAB8" box="[514,621,1328,1354]" pageId="7" pageNumber="200" value="2017-04-03">3.iv.2017</collectingDate>
</date>
,
<collectorName id="26FA5360FD7EE316D4196A1DFCB0FAB9" box="[631,843,1329,1355]" pageId="7" pageNumber="200">Y. Ramos-Pastrana</collectorName>
(1 sex undetermined
<collectionCode id="ED1EAE73FD7EE316D2536A1DFB76FAB9" box="[1085,1165,1329,1355]" pageId="7" pageNumber="200">LEUA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D2F06A1DFE73FA61" ID-GBIF-Occurrence="3061003331" collectingDate="2008-01" collectionCode="CALT-ECC" collectorName="M. C. Santos" country="Colombia" elevation="200" latitude="2.3628333" location="Finca Eli" longLatPrecision="1" longitude="-72.64164" municipality="San Jose" pageId="7" pageNumber="200" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Guaviare" typeStatus="paratype">
<collectingRegion id="49CBF854FD7EE316D2F06A1DFACAFAB9" box="[1182,1329,1329,1355]" country="Colombia" name="Guaviare" pageId="7" pageNumber="200">GUAVIARE</collectingRegion>
:
<collectingMunicipality id="6BD4ACCCFD7EE316D3556A1DFA67FAB9" box="[1339,1436,1329,1355]" pageId="7" pageNumber="200">San José</collectingMunicipality>
del
<collectingRegion id="49CBF854FD7EE316D6AA6A78FED6FA9C" box="[196,301,1364,1390]" country="Colombia" name="Guaviare" pageId="7" pageNumber="200">Guaviare</collectingRegion>
,
<location id="8ED0606DFD7EE316D7526A78FE53FA9C" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D7526A78FE53FA9C" box="[316,424,1364,1390]" country="Colombia" latitude="2.3628333" longLatPrecision="1" longitude="-72.64164" municipality="San Jose" name="Finca Eli" pageId="7" pageNumber="200" stateProvince="Guaviare">Finca Eli</location>
,
<quantity id="4CF79B53FD7EE316D7D96A79FDFAFA9D" box="[439,513,1365,1391]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="200" unit="m" value="200.0">
<elevation id="0022D185FD7EE316D7D96A79FDFAFA9D" box="[439,513,1365,1391]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="200" unit="m" value="200.0">200 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D47A6A78FD52FA9C" box="[532,681,1364,1390]" degrees="2" direction="north" minutes="21" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="46.2" value="2.3628333">2º2146.2N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D4D66A79FC9AFA9D" box="[696,865,1365,1391]" degrees="72" direction="west" minutes="38" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="29.9" value="-72.64164">72º3829.9W</geoCoordinate>
),
<date id="FFB11076FD7EE316D5176A78FC39FA9C" box="[889,962,1364,1390]" pageId="7" pageNumber="200" value="2008-01">
<collectingDate id="EFF5E99EFD7EE316D5176A78FC39FA9C" box="[889,962,1364,1390]" pageId="7" pageNumber="200" value="2008-01">i.2008</collectingDate>
</date>
, monkey feces,
<collectorName id="26FA5360FD7EE316D2EF6A79FAEBFA9D" box="[1153,1296,1365,1391]" pageId="7" pageNumber="200">M.C. Santos</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D34D6A79FA84FA9C" box="[1315,1407,1364,1391]" pageId="7" pageNumber="200" type="male">3 males</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D3E06A79FF15FA60" pageId="7" pageNumber="200" type="female">3 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D6966A55FE79FA61" box="[248,386,1401,1427]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D7FB6A55FBB0FA45" ID-GBIF-Occurrence="3061003352" collectingDate="2017-11-30" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; W. Chamorro" country="Colombia" elevation="754" latitude="3.8282917" location="Finca El Porvenir" longLatPrecision="1" longitude="-73.84244" municipality="Cubarral" pageId="7" pageNumber="200" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Meta" typeStatus="paratype">
<collectingRegion id="49CBF854FD7EE316D7FB6A55FE1CFA61" box="[405,487,1401,1427]" country="Colombia" name="Meta" pageId="7" pageNumber="200">META</collectingRegion>
:
<collectingMunicipality id="6BD4ACCCFD7EE316D79D6A54FDA3FA60" box="[499,600,1400,1426]" pageId="7" pageNumber="200">Cubarral</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D40B6A54FCCAFA60" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D40B6A54FCCAFA60" box="[613,817,1400,1426]" country="Colombia" latitude="3.8282917" longLatPrecision="1" longitude="-73.84244" municipality="Cubarral" name="Finca El Porvenir" pageId="7" pageNumber="200" stateProvince="Meta">Finca El Porvenir</location>
,
<quantity id="4CF79B53FD7EE316D5516A54FC73FA61" box="[831,904,1400,1427]" metricMagnitude="2" metricUnit="m" metricValue="7.54" pageId="7" pageNumber="200" unit="m" value="754.0">
<elevation id="0022D185FD7EE316D5516A54FC73FA61" box="[831,904,1400,1427]" metricMagnitude="2" metricUnit="m" metricValue="7.54" pageId="7" pageNumber="200" unit="m" value="754.0">754 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5F46A54FBC4FA61" box="[922,1087,1400,1427]" degrees="3" direction="north" minutes="49" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="41.85" value="3.8282917">3°4941.85N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D2236A54FAFCFA61" box="[1101,1287,1400,1427]" degrees="73" direction="west" minutes="50" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="32.78" value="-73.84244">73°5032.78W</geoCoordinate>
),
<date id="FFB11076FD7EE316D3736A54FA63FA60" box="[1309,1432,1400,1426]" pageId="7" pageNumber="200" value="2017-11-30">
<collectingDate id="EFF5E99EFD7EE316D3736A54FA63FA60" box="[1309,1432,1400,1426]" pageId="7" pageNumber="200" value="2017-11-30">30.xi.2017</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D71C6AB0FE43FA44" box="[370,440,1436,1462]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D7D16AB1FDD4FA45" box="[447,559,1437,1463]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D4216AB1FD1DFA44" box="[591,742,1436,1463]" pageId="7" pageNumber="200">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D4986AB1FCBBFA44" box="[758,832,1436,1463]" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D5246AB1FC4FFA44" box="[842,948,1436,1463]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D5D46AB1FBBFFA45" box="[954,1092,1437,1463]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D23D6AB0FE12FA0D" ID-GBIF-Occurrence="3061003327" collectingDate="2017-11-30" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; W. Chamorro" country="Colombia" elevation="661" latitude="3.8204973" location="Finca La Cristalina" longLatPrecision="1" longitude="-73.84047" municipality="Cubarral" pageId="7" pageNumber="200" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D23D6AB0FB43FA44" box="[1107,1208,1436,1462]" pageId="7" pageNumber="200">Cubarral</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D2AC6AB0FA63FA44" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D2AC6AB0FA63FA44" box="[1218,1432,1436,1463]" country="Colombia" latitude="3.8204973" longLatPrecision="1" longitude="-73.84047" municipality="Cubarral" name="Finca La Cristalina" pageId="7" pageNumber="200">Finca La Cristalina</location>
,
<quantity id="4CF79B53FD7EE316D6F96AECFF19FA29" box="[151,226,1472,1499]" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" pageId="7" pageNumber="200" unit="m" value="661.0">
<elevation id="0022D185FD7EE316D6F96AECFF19FA29" box="[151,226,1472,1499]" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" pageId="7" pageNumber="200" unit="m" value="661.0">661 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D69B6AEDFE62FA29" box="[245,409,1473,1499]" degrees="3" direction="north" minutes="49" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="13.79" value="3.8204973">3°4913.79N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D7C76AECFD98FA29" box="[425,611,1472,1499]" degrees="73" direction="west" minutes="50" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="25.68" value="-73.84047">73°5025.68W</geoCoordinate>
),
<date id="FFB11076FD7EE316D4146AECFD0EFA28" box="[634,757,1472,1498]" pageId="7" pageNumber="200" value="2017-11-30">
<collectingDate id="EFF5E99EFD7EE316D4146AECFD0EFA28" box="[634,757,1472,1498]" pageId="7" pageNumber="200" value="2017-11-30">30.xi.2017</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D5836AECFBC8FA28" box="[1005,1075,1472,1498]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D22E6AEDFB4EFA29" box="[1088,1205,1473,1499]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D2B16AEDFA80FA28" box="[1247,1403,1472,1499]" pageId="7" pageNumber="200">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D3E36AEDFF20FA0C" pageId="7" pageNumber="200" type="male">2 males</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D6886AC9FEA9FA0C" box="[230,338,1508,1535]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D7366AC9FE19FA0D" box="[344,482,1509,1535]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D79D6AC8FC8BF9D1" ID-GBIF-Occurrence="3061003357" collectingDate="2013-05" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; D. Martinez" country="Colombia" elevation="576" latitude="3.8255556" location="Finca La Pradera" longLatPrecision="21" longitude="-73.82111" municipality="Cubarral" pageId="7" pageNumber="200" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D79D6AC8FDA3FA0C" box="[499,600,1508,1534]" pageId="7" pageNumber="200">Cubarral</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D40D6AC8FCD8FA0C" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D40D6AC8FCD8FA0C" box="[611,803,1508,1535]" country="Colombia" latitude="3.8255556" longLatPrecision="21" longitude="-73.82111" municipality="Cubarral" name="Finca La Pradera" pageId="7" pageNumber="200">Finca La Pradera</location>
,
<quantity id="4CF79B53FD7EE316D5416AC8FC8EFA0D" box="[815,885,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="5.76" pageId="7" pageNumber="200" unit="m" value="576.0">
<elevation id="0022D185FD7EE316D5416AC8FC8EFA0D" box="[815,885,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="5.76" pageId="7" pageNumber="200" unit="m" value="576.0">576 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5EA6AC9FBFFFA0D" box="[900,1028,1509,1535]" degrees="3" direction="north" minutes="49" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="32" value="3.8255556">3º4932N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D2616AC8FB58FA0D" box="[1039,1187,1508,1535]" degrees="73" direction="west" minutes="49" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="16" value="-73.82111">73º4916W</geoCoordinate>
),
<date id="FFB11076FD7EE316D2D96AC9FAFEFA0D" box="[1207,1285,1509,1535]" pageId="7" pageNumber="200" value="2013-05">
<collectingDate id="EFF5E99EFD7EE316D2D96AC9FAFEFA0D" box="[1207,1285,1509,1535]" pageId="7" pageNumber="200" value="2013-05">v.2013</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D68A6924FED1F9D0" box="[228,298,1544,1570]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D75B6925FE5CF9D1" box="[309,423,1545,1571]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D7A06925FDACF9D1" box="[462,599,1545,1571]" pageId="7" pageNumber="200">D. Martínez</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D4066925FD2DF9D0" box="[616,726,1544,1571]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D4B16925FC92F9D1" box="[735,873,1545,1571]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D5156924FC05F9B5" ID-GBIF-Occurrence="3061003338" collectingDate="2017-12-06" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; W. Chamorro." country="Colombia" elevation="576" latitude="3.8255556" location="Finca La Pradera" longLatPrecision="21" longitude="-73.82111" municipality="Cubarral" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D5156924FC1BF9D0" box="[891,992,1544,1570]" pageId="7" pageNumber="200">Cubarral</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D5836924FB4AF9D0" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D5836924FB4AF9D0" box="[1005,1201,1544,1571]" country="Colombia" latitude="3.8255556" longLatPrecision="21" longitude="-73.82111" municipality="Cubarral" name="Finca La Pradera" pageId="7" pageNumber="200">Finca La Pradera</location>
,
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(
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),
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<collectingDate id="EFF5E99EFD7EE316D72C6900FE4DF9B4" box="[322,438,1580,1606]" pageId="7" pageNumber="200" value="2017-12-06">6.xii.2017</collectingDate>
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,
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&amp;
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(
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<collectionCode id="ED1EAE73FD7EE316D5006901FC03F9B5" box="[878,1016,1581,1607]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D2626901FE9EF97D" ID-GBIF-Occurrence="3061003341" collectingDate="2016-10-25" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; W. Chamorro." country="Colombia" elevation="331" latitude="3.363847" location="Finca Buena Vista" longLatPrecision="1" longitude="-73.72247" municipality="Fuente de Oro" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D2626901FB4FF9B5" box="[1036,1204,1580,1607]" pageId="7" pageNumber="200">Fuente de Oro</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D2AD6900FA63F9B4" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D2AD6900FA63F9B4" box="[1219,1432,1580,1607]" country="Colombia" latitude="3.363847" longLatPrecision="1" longitude="-73.72247" municipality="Fuente de Oro" name="Finca Buena Vista" pageId="7" pageNumber="200">Finca Buena Vista</location>
,
<quantity id="4CF79B53FD7EE316D6F9697DFF19F999" box="[151,226,1617,1643]" metricMagnitude="2" metricUnit="m" metricValue="3.31" pageId="7" pageNumber="200" unit="m" value="331.0">
<elevation id="0022D185FD7EE316D6F9697DFF19F999" box="[151,226,1617,1643]" metricMagnitude="2" metricUnit="m" metricValue="3.31" pageId="7" pageNumber="200" unit="m" value="331.0">331 m</elevation>
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(
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,
<geoCoordinate id="EE3B5071FD7EE316D7C7697DFD98F999" box="[425,611,1617,1643]" degrees="73" direction="west" minutes="43" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="20.89" value="-73.72247">73°4320.89W</geoCoordinate>
),
<date id="FFB11076FD7EE316D415697CFD16F999" box="[635,749,1616,1643]" pageId="7" pageNumber="200" value="2016-10-25">
<collectingDate id="EFF5E99EFD7EE316D415697CFD16F999" box="[635,749,1616,1643]" pageId="7" pageNumber="200" value="2016-10-25">25.x.2016</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D588697CFBD7F998" box="[998,1068,1616,1642]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D257697DFB55F999" box="[1081,1198,1617,1643]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D2B6697DFA80F998" box="[1240,1403,1616,1643]" pageId="7" pageNumber="200">W. Chamorro.</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D3FE697DFF36F97C" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D6BA6959FEA5F97D" box="[212,350,1653,1679]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D7016959FC4AF941" ID-GBIF-Occurrence="3061003346" collectingDate="2017-03-25" collectionCode="CALT-ECC" collectorName="A. Lopera &amp; W. Chamorro." country="Colombia" elevation="331" latitude="3.363847" location="Finca Buena Vista" longLatPrecision="1" longitude="-73.72247" municipality="Fuente de Oro" pageId="7" pageNumber="200" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D7016959FDEAF97D" box="[367,529,1652,1679]" pageId="7" pageNumber="200">Fuente de Oro</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D4726958FD11F97C" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D4726958FD11F97C" box="[540,746,1652,1679]" country="Colombia" latitude="3.363847" longLatPrecision="1" longitude="-73.72247" municipality="Fuente de Oro" name="Finca Buena Vista" pageId="7" pageNumber="200">Finca Buena Vista</location>
,
<quantity id="4CF79B53FD7EE316D49B6959FCC7F97D" box="[757,828,1653,1679]" metricMagnitude="2" metricUnit="m" metricValue="3.31" pageId="7" pageNumber="200" unit="m" value="331.0">
<elevation id="0022D185FD7EE316D49B6959FCC7F97D" box="[757,828,1653,1679]" metricMagnitude="2" metricUnit="m" metricValue="3.31" pageId="7" pageNumber="200" unit="m" value="331.0">331 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D5256958FC0AF97D" box="[843,1009,1652,1679]" degrees="3" direction="north" minutes="21" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="49.85" value="3.363847">3°2149.85N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D5926959FB4EF97D" box="[1020,1205,1653,1679]" degrees="73" direction="west" minutes="43" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="20.89" value="-73.72247">73°4320.89W</geoCoordinate>
),
<date id="FFB11076FD7EE316D2A76958FABEF97D" box="[1225,1349,1652,1679]" pageId="7" pageNumber="200" value="2017-03-25">
<collectingDate id="EFF5E99EFD7EE316D2A76958FABEF97D" box="[1225,1349,1652,1679]" pageId="7" pageNumber="200" value="2017-03-25">25.iii.2017</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D74A69B4FE91F940" box="[292,362,1688,1714]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D71D69B5FE1EF941" box="[371,485,1689,1715]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D46769B5FD53F940" box="[521,680,1688,1715]" pageId="7" pageNumber="200">W. Chamorro.</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D4D469B5FCE2F940" box="[698,793,1688,1715]" pageId="7" pageNumber="200" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D54F69B5FC50F941" box="[801,939,1689,1715]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D5D269B5FF20F909" ID-GBIF-Occurrence="3061003353" collectingDate="2016-10-17" collectorName="A. Lopera &amp; W. Chamorro." country="Colombia" elevation="296" location="Finca La Herradura" longitude="-73.609215" municipality="Fuente de Oro" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D5D269B5FBA4F941" box="[956,1119,1688,1715]" pageId="7" pageNumber="200">Fuente de Oro</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D20569B4FAB2F940" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D20569B4FAB2F940" box="[1131,1353,1688,1715]" country="Colombia" longitude="-73.609215" municipality="Fuente de Oro" name="Finca La Herradura" pageId="7" pageNumber="200">Finca La Herradura</location>
,
<quantity id="4CF79B53FD7EE316D33B69B4FA67F941" box="[1365,1436,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="7" pageNumber="200" unit="m" value="296.0">
<elevation id="0022D185FD7EE316D33B69B4FA67F941" box="[1365,1436,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="2.96" pageId="7" pageNumber="200" unit="m" value="296.0">296 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D6CE6990FEC5F925" box="[160,318,1724,1751]" pageId="7" pageNumber="200">3°1740.75N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D7276990FDF9F925" box="[329,514,1724,1751]" degrees="73" direction="west" minutes="36" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="33.17" value="-73.609215">73°3633.17W</geoCoordinate>
),
<date id="FFB11076FD7EE316D4786990FD73F924" box="[534,648,1724,1750]" pageId="7" pageNumber="200" value="2016-10-17">
<collectingDate id="EFF5E99EFD7EE316D4786990FD73F924" box="[534,648,1724,1750]" pageId="7" pageNumber="200" value="2016-10-17">17.x.2016</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D51E6990FC4DF924" box="[880,950,1724,1750]" pageId="7" pageNumber="200">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD7EE316D5D06991FBD4F925" box="[958,1071,1725,1751]" pageId="7" pageNumber="200">A. Lopera</collectorName>
&amp;
<collectorName id="26FA5360FD7EE316D23F6991FB14F924" box="[1105,1263,1724,1751]" pageId="7" pageNumber="200">W. Chamorro.</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D36E6991FAB3F924" box="[1280,1352,1724,1751]" pageId="7" pageNumber="200" type="male">1 male</specimenCount>
CALTECC)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D68969CCFC25F909" ID-GBIF-Occurrence="3061003343" box="[231,990,1760,1787]" country="Colombia" elevation="334" latitude="3.3821" location="Los Cambulos" longLatPrecision="1" longitude="-73.742775" municipality="Granada" pageId="7" pageNumber="200" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D68969CCFEB2F908" box="[231,329,1760,1786]" pageId="7" pageNumber="200">Granada</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D73969CDFE05F908" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D73969CDFE05F908" box="[343,510,1760,1787]" country="Colombia" latitude="3.3821" longLatPrecision="1" longitude="-73.742775" municipality="Granada" name="Los Cambulos" pageId="7" pageNumber="200">Los Cambulos</location>
,
<quantity id="4CF79B53FD7EE316D46269CDFDADF909" box="[524,598,1761,1787]" metricMagnitude="2" metricUnit="m" metricValue="3.34" pageId="7" pageNumber="200" unit="m" value="334.0">
<elevation id="0022D185FD7EE316D46269CDFDADF909" box="[524,598,1761,1787]" metricMagnitude="2" metricUnit="m" metricValue="3.34" pageId="7" pageNumber="200" unit="m" value="334.0">334 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D40969CCFCF6F909" box="[615,781,1760,1787]" degrees="3" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="55.56" value="3.3821">3°2255.56N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D57569CDFC2DF909" box="[795,982,1761,1787]" degrees="73" direction="west" minutes="44" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="34.00" value="-73.742775">73°4434.00W</geoCoordinate>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D58569CCFD2FF8ED" ID-GBIF-Occurrence="3061003334" collectingDate="2016-10-28" collectionCode="CALT-ECC" collectorName="Lopera &amp; Chamorro" country="Colombia" pageId="7" pageNumber="200" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="FFB11076FD7EE316D58569CCFBA5F908" box="[1003,1118,1760,1786]" pageId="7" pageNumber="200" value="2016-10-28">
<collectingDate id="EFF5E99EFD7EE316D58569CCFBA5F908" box="[1003,1118,1760,1786]" pageId="7" pageNumber="200" value="2016-10-28">28.x.2016</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D33C69CCFA63F908" box="[1362,1432,1760,1786]" pageId="7" pageNumber="200">pitfall</collectingMethod>
, A.
<collectorName id="26FA5360FD7EE316D6D46829FEF2F8ED" box="[186,265,1797,1823]" pageId="7" pageNumber="200">Lopera</collectorName>
&amp; W.
<collectorName id="26FA5360FD7EE316D7386828FE33F8EC" box="[342,456,1796,1822]" pageId="7" pageNumber="200">Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D7B56829FDC0F8EC" box="[475,571,1796,1823]" pageId="7" pageNumber="200" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D42D6829FD36F8ED" box="[579,717,1797,1823]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D4B16828FB7BF8B1" ID-GBIF-Occurrence="3061003360" collectingDate="1997-04-03" collectionCode="CALT-ECC, ICN" collectorName="Finca El Esfuerzo" country="Colombia" elevation="200" latitude="3.0455556" location="Pinalito" longLatPrecision="21" longitude="-73.595" pageId="7" pageNumber="200" specimenCount="95" specimenCount-female="46" specimenCount-male="49" typeStatus="paratype">
<location id="8ED0606DFD7EE316D4B16828FCC2F8EC" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D4B16828FCC2F8EC" box="[735,825,1796,1822]" country="Colombia" latitude="3.0455556" longLatPrecision="21" longitude="-73.595" name="Pinalito" pageId="7" pageNumber="200">Piñalito</location>
,
<collectorName id="26FA5360FD7EE316D52B6828FBE9F8EC" box="[837,1042,1796,1822]" pageId="7" pageNumber="200">Finca El Esfuerzo</collectorName>
,
<quantity id="4CF79B53FD7EE316D2716829FB9CF8ED" box="[1055,1127,1797,1823]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="200" unit="m" value="200.0">
<elevation id="0022D185FD7EE316D2716829FB9CF8ED" box="[1055,1127,1797,1823]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="200" unit="m" value="200.0">200 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D2166829FB17F8ED" box="[1144,1260,1797,1823]" degrees="3" direction="north" minutes="2" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="44" value="3.0455556">3°244N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D2966828FA74F8ED" box="[1272,1423,1796,1823]" degrees="73" direction="west" minutes="35" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="42" value="-73.595">73°3542W</geoCoordinate>
),
<date id="FFB11076FD7EE316D6F96804FEF9F8B0" box="[151,258,1832,1858]" pageId="7" pageNumber="200" value="1997-04-03">
<collectingDate id="EFF5E99EFD7EE316D6F96804FEF9F8B0" box="[151,258,1832,1858]" pageId="7" pageNumber="200" value="1997-04-03">3.iv.1997</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD7EE316D7996804FDC1F8B0" box="[503,570,1832,1858]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
<specimenCount id="9D09FD3FFD7EE316D4226805FD4CF8B0" box="[588,695,1832,1859]" pageId="7" pageNumber="200" type="male">47 males</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D4AB6804FCB9F8B0" box="[709,834,1832,1858]" pageId="7" pageNumber="200" type="female">46 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D5256805FC28F8B1" box="[843,979,1833,1859]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
;
<specimenCount id="9D09FD3FFD7EE316D58F6805FBC2F8B0" box="[993,1081,1832,1859]" pageId="7" pageNumber="200" type="male">2 males</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D22D6805FB83F8B1" box="[1091,1144,1833,1859]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="7" pageNumber="200">ICN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD7EE316D2E06805FA9CF895" ID-GBIF-Occurrence="3061003351" collectionCode="CALT-ECC" collectorName="J. L. Montes" country="Colombia" elevation="214" latitude="3.748625" location="Estacion Rubiales" longLatPrecision="1" longitude="-71.463036" municipality="Puerto Gaitan" pageId="7" pageNumber="200" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD7EE316D2E06805FAD6F8B0" box="[1166,1325,1832,1859]" pageId="7" pageNumber="200">Puerto Gaitán</collectingMunicipality>
,
<location id="8ED0606DFD7EE316D3526804FF07F894" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D3526804FF07F894" country="Colombia" latitude="3.748625" longLatPrecision="1" longitude="-71.463036" municipality="Puerto Gaitan" name="Estacion Rubiales" pageId="7" pageNumber="200">Estación Rubiales</location>
,
<quantity id="4CF79B53FD7EE316D7666861FEAAF895" box="[264,337,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="7" pageNumber="200" unit="m" value="214.0">
<elevation id="0022D185FD7EE316D7666861FEAAF895" box="[264,337,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="7" pageNumber="200" unit="m" value="214.0">214 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD7EE316D70F6860FDFDF895" box="[353,518,1868,1895]" degrees="3" direction="north" minutes="44" orientation="latitude" pageId="7" pageNumber="200" precision="1" seconds="55.05" value="3.748625">3°4455.05N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD7EE316D47D6860FD35F895" box="[531,718,1868,1895]" degrees="71" direction="west" minutes="27" orientation="longitude" pageId="7" pageNumber="200" precision="1" seconds="46.93" value="-71.463036">71°2746.93W</geoCoordinate>
), ixx.2015,
<collectorName id="26FA5360FD7EE316D50C6861FC13F895" box="[866,1000,1869,1895]" pageId="7" pageNumber="200">J.L. Montes</collectorName>
(
<specimenCount id="9D09FD3FFD7EE316D5976861FBAFF894" box="[1017,1108,1868,1895]" pageId="7" pageNumber="200" type="male">2 males</specimenCount>
,
<specimenCount id="9D09FD3FFD7EE316D20F6861FB35F894" box="[1121,1230,1868,1895]" pageId="7" pageNumber="200" type="female">2 females</specimenCount>
<collectionCode id="ED1EAE73FD7EE316D2B86861FA9BF895" box="[1238,1376,1869,1895]" pageId="7" pageNumber="200">CALT-ECC</collectionCode>
)
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(
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<collectingMethod id="524E4EA1FD7EE316D74C6894FE9EF820" box="[290,357,1976,2002]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
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<collectionCode id="ED1EAE73FD7EE316D7B26895FDCFF821" box="[476,564,1977,2003]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
</materialsCitation>
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<location id="8ED0606DFD7EE316D4286895FCC4F820" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD7EE316D4286895FCC4F820" box="[582,831,1976,2003]" country="Colombia" latitude="4.078611" longLatPrecision="21" longitude="-73.373055" name="30 km E Villavicencio" pageId="7" pageNumber="200">
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E Villavicencio
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(
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, S. &amp; J.
<collectorName id="26FA5360FD7EE316D3316894FA63F820" box="[1375,1432,1976,2002]" pageId="7" pageNumber="200">Peck</collectorName>
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<collectingMethod id="524E4EA1FD7EE316D7A868F0FDF2F804" box="[454,521,2012,2038]" pageId="7" pageNumber="200">pitfall</collectingMethod>
(
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<collectionCode id="ED1EAE73FD7EE316D56F68F1FCA2F805" box="[769,857,2013,2039]" pageId="7" pageNumber="200">CMNC</collectionCode>
)
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E Villavicencio
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(
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, human feces baited
<collectingMethod id="524E4EA1FD71E319D4606FBBFDAFFF43" box="[526,596,151,177]" pageId="8" pageNumber="201">pitfall</collectingMethod>
, S. &amp; J.
<collectorName id="26FA5360FD71E319D4DB6FBBFD08FF43" box="[693,755,151,177]" pageId="8" pageNumber="201">Peck.</collectorName>
(
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,
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<collectionCode id="ED1EAE73FD71E319D5A96FBBFBE4FF43" box="[967,1055,151,177]" pageId="8" pageNumber="201">CMNC</collectionCode>
)
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<materialsCitation id="3B673CEBFD71E319D25C6FBBFD8BFF27" ID-GBIF-Occurrence="3061003332" collectingDate="1980-01" collectionCode="CMNC" collectorName="Bolle" country="Colombia" latitude="1.15" location="Santiago" longLatPrecision="1308" longitude="-77.0" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" stateProvince="Putumayo" typeStatus="paratype">
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:
<location id="8ED0606DFD71E319D2B36FBBFAC4FF43" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D2B36FBBFAC4FF43" box="[1245,1343,151,177]" country="Colombia" latitude="1.15" longLatPrecision="1308" longitude="-77.0" name="Santiago" pageId="8" pageNumber="201" stateProvince="Putumayo">Santiago</location>
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,
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),
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,
<collectorName id="26FA5360FD71E319D70B6F97FE59FF27" box="[357,418,187,213]" pageId="8" pageNumber="201">Bolle</collectorName>
(
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<collectionCode id="ED1EAE73FD71E319D47F6F97FD92FF27" box="[529,617,187,213]" pageId="8" pageNumber="201">CMNC</collectionCode>
)
</materialsCitation>
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<materialsCitation id="3B673CEBFD71E319D4106F90FDE9FF0B" ID-GBIF-Occurrence="3061003322" collectingDate="1995-09-30" collectionCode="CALT-ECC" collectorName="Amezquita" country="Colombia" elevation="100" location="Caparu" pageId="8" pageNumber="201" specimenCount="101" specimenCount-male="1" stateProvince="Vaupes" typeStatus="paratype">
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:
<location id="8ED0606DFD71E319D4956F97FCB4FF27" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D4956F97FCB4FF27" box="[763,847,187,213]" country="Colombia" name="Caparu" pageId="8" pageNumber="201" stateProvince="Vaupes">Caparú</location>
,
<specimenCount id="9D09FD3FFD71E319D5306F97FC4BFF27" box="[862,944,187,213]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D5306F97FC57FF27" box="[862,940,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="201" unit="m" value="100.0">
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.
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<date id="FFB11076FD71E319D5D56F96FBCEFF27" box="[955,1077,186,213]" pageId="8" pageNumber="201" value="1995-09-30">
<collectingDate id="EFF5E99EFD71E319D5D56F96FBCEFF27" box="[955,1077,186,213]" pageId="8" pageNumber="201" value="1995-09-30">30.ix.1995</collectingDate>
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<collectingMethod id="524E4EA1FD71E319D35F6F97FA8CFF27" box="[1329,1399,187,213]" pageId="8" pageNumber="201">pitfall</collectingMethod>
, S.
<collectorName id="26FA5360FD71E319D6F96FF3FEE9FF0B" box="[151,274,223,249]" pageId="8" pageNumber="201">Amézquita</collectorName>
(
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<collectionCode id="ED1EAE73FD71E319D7EF6FF3FDF0FF0B" box="[385,523,223,249]" pageId="8" pageNumber="201">CALT-ECC</collectionCode>
)
</materialsCitation>
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<materialsCitation id="3B673CEBFD71E319D44F6FF3FE22FEEF" ID-GBIF-Occurrence="3061003356" collectingDate="1995-02-01" collectionCode="BDGC" collectorName="B. D. Gill" country="Colombia" county="Rio Apaporis" elevation="200" latitude="-1.0166667" location="Biological Station" longLatPrecision="1308" longitude="-69.083336" municipality="Caparu" pageId="8" pageNumber="201" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<collectingMunicipality id="6BD4ACCCFD71E319D4A26FF3FCDBFF0B" box="[716,800,223,249]" pageId="8" pageNumber="201">Caparú</collectingMunicipality>
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(
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)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D78B6E2FFE7EFEB3" ID-GBIF-Occurrence="3061003317" collectingDate="1995-11-27" collectionCode="BDGC" collectorName="B. D. Gill" country="Colombia" county="Rio Apaporis" elevation="200" latitude="-1.0166667" location="Biological Station" longLatPrecision="1308" longitude="-69.083336" municipality="Caparu" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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(
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),
<date id="FFB11076FD71E319D28E6E2EFAA1FEEF" box="[1248,1370,258,285]" pageId="8" pageNumber="201" value="1995-11-27">
<collectingDate id="EFF5E99EFD71E319D28E6E2EFAA1FEEF" box="[1248,1370,258,285]" pageId="8" pageNumber="201" value="1995-11-27">27.xi.1995</collectingDate>
</date>
,
<collectorName id="26FA5360FD71E319D3096E2FFF38FEB3" pageId="8" pageNumber="201">B.D. Gill</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D6B86E0BFEDAFEB3" box="[214,289,295,321]" pageId="8" pageNumber="201" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD71E319D7446E0BFE86FEB3" box="[298,381,295,321]" pageId="8" pageNumber="201">BDGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D7FE6E0AFB73FE97" ID-GBIF-Occurrence="3061003302" collectingDate="2007-08-25" collectionCode="CALT-ECC" collectorName="D. Bunay" country="Ecuador" county="Aucayacu" elevation="291" latitude="-0.055833332" location="Lago Agrio" longLatPrecision="21" longitude="-76.5925" municipality="Rio Eno" pageId="8" pageNumber="201" specimenCount="1" specimenCount-female="1" stateProvince="Sucumbios" typeStatus="paratype">
<emphasis id="B97BEAA4FD71E319D7FE6E0AFDE5FEB2" bold="true" box="[400,542,294,320]" pageId="8" pageNumber="201">
<collectingCountry id="F3187626FD71E319D7FE6E0AFDE5FEB2" box="[400,542,294,320]" name="Ecuador" pageId="8" pageNumber="201">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49CBF854FD71E319D4406E04FD2DFEB1" box="[558,726,296,323]" country="Ecuador" name="Sucumbios" pageId="8" pageNumber="201">SUCUMBÍOS</collectingRegion>
:
<collectingCounty id="62D14E3AFD71E319D48E6E0BFCA8FEB3" box="[736,851,295,321]" pageId="8" pageNumber="201">Aucayacu</collectingCounty>
,
<collectingMunicipality id="6BD4ACCCFD71E319D5316E0BFC3BFEB3" box="[863,960,295,321]" pageId="8" pageNumber="201">Río Eno</collectingMunicipality>
,
<quantity id="4CF79B53FD71E319D5A36E0AFBEEFEB3" box="[973,1045,294,321]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="201" unit="km" value="15.0">15 km</quantity>
de
<location id="8ED0606DFD71E319D22E6E0BFB3FFEB3" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D22E6E0BFB3FFEB3" box="[1088,1220,295,321]" country="Ecuador" county="Aucayacu" latitude="-0.055833332" longLatPrecision="21" longitude="-76.5925" municipality="Rio Eno" name="Lago Agrio" pageId="8" pageNumber="201" stateProvince="Sucumbios">Lago Agrio</location>
,
<quantity id="4CF79B53FD71E319D2BF6E0BFAE1FEB3" box="[1233,1306,295,321]" metricMagnitude="2" metricUnit="m" metricValue="2.91" pageId="8" pageNumber="201" unit="m" value="291.0">
<elevation id="0022D185FD71E319D2BF6E0BFAE1FEB3" box="[1233,1306,295,321]" metricMagnitude="2" metricUnit="m" metricValue="2.91" pageId="8" pageNumber="201" unit="m" value="291.0">291 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D3456E0BFA63FEB3" box="[1323,1432,295,321]" degrees="0" direction="south" minutes="3" orientation="latitude" pageId="8" pageNumber="201" precision="15" seconds="21" value="-0.055833332">0º321S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D6F96E66FED0FE97" box="[151,299,330,357]" degrees="76" direction="west" minutes="35" orientation="longitude" pageId="8" pageNumber="201" precision="15" seconds="33" value="-76.5925">76º3533W</geoCoordinate>
),
<date id="FFB11076FD71E319D72F6E66FE30FE97" box="[321,459,330,357]" pageId="8" pageNumber="201" value="2007-08-25">
<collectingDate id="EFF5E99EFD71E319D72F6E66FE30FE97" box="[321,459,330,357]" pageId="8" pageNumber="201" value="2007-08-25">25.viii.2007</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D4D56E67FCFAFE97" box="[699,769,331,357]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D5606E67FC81FE97" box="[782,890,331,357]" pageId="8" pageNumber="201">D. Buñay</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D5E06E67FC14FE97" box="[910,1007,331,357]" pageId="8" pageNumber="201" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD71E319D5996E67FB7AFE97" box="[1015,1153,331,357]" pageId="8" pageNumber="201">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D2FA6E67FEA2FE5F" ID-GBIF-Occurrence="3061003310" collectingDate="2019-09-30" collectionCode="CONRAZ" collectorName="W. Chamorro" country="Ecuador" elevation="265" latitude="-0.090622224" location="Rio Aguarico" longLatPrecision="1" longitude="-76.511024" municipality="Chiritza" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD71E319D2FA6E67FB0AFE97" box="[1172,1265,331,357]" pageId="8" pageNumber="201">Chiritza</collectingMunicipality>
,
<location id="8ED0606DFD71E319D2906E67FA63FE97" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D2906E67FA63FE97" box="[1278,1432,331,357]" country="Ecuador" latitude="-0.090622224" longLatPrecision="1" longitude="-76.511024" municipality="Chiritza" name="Rio Aguarico" pageId="8" pageNumber="201">Río Aguarico</location>
, várzea forest,
<quantity id="4CF79B53FD71E319D7516E42FE71FE7B" box="[319,394,366,393]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="201" unit="m" value="265.0">
<elevation id="0022D185FD71E319D7516E42FE71FE7B" box="[319,394,366,393]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="201" unit="m" value="265.0">265 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D7F06E42FDC1FE7B" box="[414,570,366,393]" degrees="0" direction="south" minutes="5" orientation="latitude" pageId="8" pageNumber="201" precision="1" seconds="26.24" value="-0.090622224">0°526.24 S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D4256E43FD03FE78" box="[587,760,367,394]" degrees="76" direction="west" minutes="30" orientation="longitude" pageId="8" pageNumber="201" precision="1" seconds="39.7" value="-76.511024">76°3039.7W</geoCoordinate>
),
<date id="FFB11076FD71E319D57E6E43FC71FE7B" box="[784,906,367,393]" pageId="8" pageNumber="201" value="2019-09-30">
<collectingDate id="EFF5E99EFD71E319D57E6E43FC71FE7B" box="[784,906,367,393]" pageId="8" pageNumber="201" value="2019-09-30">30.ix.2019</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D2E66E43FB35FE7B" box="[1160,1230,367,393]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D2B36E43FA81FE7B" box="[1245,1402,367,393]" pageId="8" pageNumber="201">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D3FE6E43FF36FE5F" pageId="8" pageNumber="201" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD71E319D6B76EBFFEA9FE5F" box="[217,338,403,429]" pageId="8" pageNumber="201">CONRAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D7096EBFFC3FFE23" ID-GBIF-Occurrence="3061003316" collectingDate="2019-10-11" collectionCode="CALT-ECC" collectorName="W. Chamorro" country="Ecuador" elevation="260" latitude="-0.215625" location="Rio Shushufindi" longLatPrecision="1" longitude="-76.53652" municipality="Comunidad Siona" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD71E319D7096EBFFDC3FE5F" box="[359,568,403,429]" pageId="8" pageNumber="201">Comunidad Siona</collectingMunicipality>
,
<location id="8ED0606DFD71E319D4296EBFFCFEFE5F" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D4296EBFFCFEFE5F" box="[583,773,403,429]" country="Ecuador" latitude="-0.215625" longLatPrecision="1" longitude="-76.53652" municipality="Comunidad Siona" name="Rio Shushufindi" pageId="8" pageNumber="201">Río Shushufindi</location>
,
<quantity id="4CF79B53FD71E319D57A6EBFFC99FE5F" box="[788,866,403,429]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="201" unit="m" value="260.0">
<elevation id="0022D185FD71E319D57A6EBFFC99FE5F" box="[788,866,403,429]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="201" unit="m" value="260.0">260 m</elevation>
</quantity>
, moretal forest (
<geoCoordinate id="EE3B5071FD71E319D2486EBEFB3DFE5F" box="[1062,1222,402,429]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="8" pageNumber="201" precision="1" seconds="56.25" value="-0.215625">0°1256.25S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D2B86EBFFA74FE5C" box="[1238,1423,403,430]" degrees="76" direction="west" minutes="32" orientation="longitude" pageId="8" pageNumber="201" precision="1" seconds="11.48" value="-76.53652">76°3211.48W</geoCoordinate>
),
<date id="FFB11076FD71E319D6F96E9BFEF3FE23" box="[151,264,439,465]" pageId="8" pageNumber="201" value="2019-10-11">
<collectingDate id="EFF5E99EFD71E319D6F96E9BFEF3FE23" box="[151,264,439,465]" pageId="8" pageNumber="201" value="2019-10-11">11.x.2019</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D7806E9BFDCFFE23" box="[494,564,439,465]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D4506E9BFD2FFE23" box="[574,724,439,465]" pageId="8" pageNumber="201">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D48B6E9BFCD6FE23" box="[741,813,439,465]" pageId="8" pageNumber="201" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD71E319D55D6E9BFC46FE23" box="[819,957,439,465]" pageId="8" pageNumber="201">CALT-ECC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D5A36E9BFCE9FE07" ID-GBIF-Occurrence="3061003354" collectingDate="1977-09-23" collectingDateMax="1977-09-28" collectingDateMin="1977-09-23" collectionCode="CMNC" collectorName="L. Pena" country="Ecuador" elevation="150" latitude="0.044444446" location="Rio Aguarico" longLatPrecision="21" longitude="-76.69722" municipality="Dureno" pageId="8" pageNumber="201" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingMunicipality id="6BD4ACCCFD71E319D5A36E9BFBDFFE23" box="[973,1060,439,465]" pageId="8" pageNumber="201">Dureno</collectingMunicipality>
,
<location id="8ED0606DFD71E319D2406E9BFB3EFE23" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D2406E9BFB3EFE23" box="[1070,1221,439,465]" country="Ecuador" latitude="0.044444446" longLatPrecision="21" longitude="-76.69722" municipality="Dureno" name="Rio Aguarico" pageId="8" pageNumber="201">Río Aguarico</location>
,
<quantity id="4CF79B53FD71E319D2A16E9AFAEEFE23" box="[1231,1301,438,465]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="201" unit="m" value="150.0">
<elevation id="0022D185FD71E319D2A16E9AFAEEFE23" box="[1231,1301,438,465]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="201" unit="m" value="150.0">150 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D34A6E9BFA6CFE23" box="[1316,1431,439,465]" degrees="0" direction="north" minutes="2" orientation="latitude" pageId="8" pageNumber="201" precision="15" seconds="40" value="0.044444446">0°240N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D6F96EF6FED6FE07" box="[151,301,474,501]" degrees="76" direction="west" minutes="41" orientation="longitude" pageId="8" pageNumber="201" precision="15" seconds="50" value="-76.69722">76°4150W</geoCoordinate>
),
<date id="FFB11076FD71E319D7516EF7FE1DFE07" box="[319,486,475,501]" pageId="8" pageNumber="201" value="1977-09-23" valueMax="1977-09-28" valueMin="1977-09-23">
<collectingDate id="EFF5E99EFD71E319D7516EF7FE1DFE07" box="[319,486,475,501]" pageId="8" pageNumber="201" value="1977-09-23" valueMax="1977-09-28" valueMin="1977-09-23">2328-ix.1977</collectingDate>
</date>
,
<collectorName id="26FA5360FD71E319D7816EF7FDBAFE07" box="[495,577,475,501]" pageId="8" pageNumber="201">L. Peña</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D43F6EF7FD55FE07" box="[593,686,475,501]" pageId="8" pageNumber="201" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD71E319D4DC6EF7FCF1FE07" box="[690,778,475,501]" pageId="8" pageNumber="201">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D5776EF7FB80FDEB" ID-GBIF-Occurrence="3061003345" collectingDate="2019-10-09" collectionCode="CONRAZ" collectorName="W. Chamorro" country="Ecuador" county="Shushufindi" elevation="260" latitude="-0.16352223" location="Rio Eno" longLatPrecision="1" longitude="-76.566864" municipality="Miss Ecuador" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCounty id="62D14E3AFD71E319D5776EF7FC58FE07" box="[793,931,475,501]" pageId="8" pageNumber="201">Shushufindi</collectingCounty>
,
<collectingMunicipality id="6BD4ACCCFD71E319D5C26EF7FBBDFE07" box="[940,1094,475,501]" pageId="8" pageNumber="201">Miss Ecuador</collectingMunicipality>
,
<location id="8ED0606DFD71E319D2206EF7FB50FE07" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D2206EF7FB50FE07" box="[1102,1195,475,501]" country="Ecuador" county="Shushufindi" latitude="-0.16352223" longLatPrecision="1" longitude="-76.566864" municipality="Miss Ecuador" name="Rio Eno" pageId="8" pageNumber="201">Río Eno</location>
,
<quantity id="4CF79B53FD71E319D2DA6EF7FB03FE07" box="[1204,1272,475,501]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="201" unit="m" value="260.0">
<elevation id="0022D185FD71E319D2DA6EF7FB03FE07" box="[1204,1272,475,501]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="201" unit="m" value="260.0">260 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D36B6EF7FA6CFE07" box="[1285,1431,475,501]" degrees="0" direction="south" minutes="9" orientation="latitude" pageId="8" pageNumber="201" precision="1" seconds="48.68" value="-0.16352223">0°948.68S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D6F96ED3FEB8FDE8" box="[151,323,511,538]" degrees="76" direction="west" minutes="34" orientation="longitude" pageId="8" pageNumber="201" precision="1" seconds="0.71" value="-76.566864">76°340.71W</geoCoordinate>
),
<date id="FFB11076FD71E319D7366ED3FE46FDEB" box="[344,445,511,537]" pageId="8" pageNumber="201" value="2019-10-09">
<collectingDate id="EFF5E99EFD71E319D7366ED3FE46FDEB" box="[344,445,511,537]" pageId="8" pageNumber="201" value="2019-10-09">9.x.2019</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D4C56ED3FD0AFDEB" box="[683,753,511,537]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D4936ED3FC6EFDEB" box="[765,917,511,537]" pageId="8" pageNumber="201">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D5C66ED3FC09FDEB" box="[936,1010,511,537]" pageId="8" pageNumber="201" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD71E319D5956ED3FB8FFDEB" box="[1019,1140,511,537]" pageId="8" pageNumber="201">CONRAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D2E86ED3FDA8FD93" ID-GBIF-Occurrence="3061003314" collectingDate="2020-11-15" collectionCode="CONRAZ" collectorName="P. Araujo." country="Ecuador" county="Tivacuno" elevation="234" latitude="-0.6483472" location="Parque Nacional Yasuni" longLatPrecision="1" longitude="-76.35732" municipality="Rio Tiputini" pageId="8" pageNumber="201" specimenCount="1" specimenCount-female="1" stateProvince="Orellana" typeStatus="paratype">
<collectingRegion id="49CBF854FD71E319D2E86ED3FADAFDEB" box="[1158,1313,511,537]" country="Ecuador" name="Orellana" pageId="8" pageNumber="201">ORELLANA</collectingRegion>
:
<collectingCounty id="62D14E3AFD71E319D3426ED3FA63FDEB" box="[1324,1432,511,537]" pageId="8" pageNumber="201">Tivacuno</collectingCounty>
,
<collectingMunicipality id="6BD4ACCCFD71E319D6F96D0FFEDEFDCF" box="[151,293,547,573]" pageId="8" pageNumber="201">Río Tiputini</collectingMunicipality>
,
<location id="8ED0606DFD71E319D75A6D0FFDB2FDCF" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D75A6D0FFDB2FDCF" box="[308,585,547,573]" country="Ecuador" county="Tivacuno" latitude="-0.6483472" longLatPrecision="1" longitude="-76.35732" municipality="Rio Tiputini" name="Parque Nacional Yasuni" pageId="8" pageNumber="201" stateProvince="Orellana">Parque Nacional Yasuní</location>
,
<quantity id="4CF79B53FD71E319D4396D0FFD59FDCF" box="[599,674,547,573]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="8" pageNumber="201" unit="m" value="234.0">
<elevation id="0022D185FD71E319D4396D0FFD59FDCF" box="[599,674,547,573]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="8" pageNumber="201" unit="m" value="234.0">234 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D4DA6D0EFCAEFDCF" box="[692,853,546,573]" degrees="0" direction="south" minutes="38" orientation="latitude" pageId="8" pageNumber="201" precision="1" seconds="54.05" value="-0.6483472">0°3854.05S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D50A6D0FFBE6FDCC" box="[868,1053,547,574]" degrees="76" direction="west" minutes="21" orientation="longitude" pageId="8" pageNumber="201" precision="1" seconds="26.37" value="-76.35732">76°2126.37W</geoCoordinate>
),
<date id="FFB11076FD71E319D25B6D0EFB54FDCF" box="[1077,1199,546,573]" pageId="8" pageNumber="201" value="2020-11-15">
<collectingDate id="EFF5E99EFD71E319D25B6D0EFB54FDCF" box="[1077,1199,546,573]" pageId="8" pageNumber="201" value="2020-11-15">15.xi.2020</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D6F96D6BFF26FD93" box="[151,221,583,609]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D6876D6BFEA3FD93" box="[233,344,583,609]" pageId="8" pageNumber="201">P. Araujo.</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D7046D6BFE31FD93" box="[362,458,583,609]" pageId="8" pageNumber="201" type="female">1 female</specimenCount>
<collectionCode id="ED1EAE73FD71E319D7BC6D6BFDB0FD93" box="[466,587,583,609]" pageId="8" pageNumber="201">CONRAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B673CEBFD71E319D4306D6BFB63FD77" ID-GBIF-Occurrence="3061003323" collectingDate="2008-04-20" collectionCode="CONRAZ" collectorName="W. Chamorro" country="Peru" elevation="12200" latitude="-2.156303" location="Helipuerto" longLatPrecision="1" longitude="-75.716415" municipality="Rio Pucacuro" pageId="8" pageNumber="201" specimenCount="1" specimenCount-male="1" stateProvince="Loreto" typeStatus="paratype">
<emphasis id="B97BEAA4FD71E319D4306D6BFD51FD93" bold="true" box="[606,682,583,609]" pageId="8" pageNumber="201">
<collectingCountry id="F3187626FD71E319D4306D6BFD51FD93" box="[606,682,583,609]" name="Peru" pageId="8" pageNumber="201">PERU</collectingCountry>
</emphasis>
:
<collectingRegion id="49CBF854FD71E319D4D46D6BFCD7FD93" box="[698,812,583,609]" country="Peru" name="Loreto" pageId="8" pageNumber="201">LORETO</collectingRegion>
:
<collectingMunicipality id="6BD4ACCCFD71E319D5596D6BFC29FD93" box="[823,978,583,609]" pageId="8" pageNumber="201">Río Pucacuro</collectingMunicipality>
,
<location id="8ED0606DFD71E319D5B06D6BFBADFD93" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D5B06D6BFBADFD93" box="[990,1110,583,609]" country="Peru" latitude="-2.156303" longLatPrecision="1" longitude="-75.716415" municipality="Rio Pucacuro" name="Helipuerto" pageId="8" pageNumber="201" stateProvince="Loreto">Helipuerto</location>
HP
<quantity id="4CF79B53FD71E319D2EB6D6BFB0EFD93" box="[1157,1269,583,609]" metricMagnitude="4" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="201" unit="m" value="12200.0">
<elevation id="0022D185FD71E319D2EB6D6BFB0EFD93" box="[1157,1269,583,609]" metricMagnitude="4" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="201" unit="m" value="12200.0">12, 200 m</elevation>
</quantity>
(
<geoCoordinate id="EE3B5071FD71E319D36B6D6BFA6CFD93" box="[1285,1431,583,609]" degrees="2" direction="south" minutes="9" orientation="latitude" pageId="8" pageNumber="201" precision="1" seconds="22.69" value="-2.156303">2°922.69S</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD71E319D6F96D46FEAAFD77" box="[151,337,618,645]" degrees="75" direction="west" minutes="42" orientation="longitude" pageId="8" pageNumber="201" precision="1" seconds="59.12" value="-75.716415">75°4259.12W</geoCoordinate>
),
<date id="FFB11076FD71E319D70B6D47FE25FD77" box="[357,478,619,645]" pageId="8" pageNumber="201" value="2008-04-20">
<collectingDate id="EFF5E99EFD71E319D70B6D47FE25FD77" box="[357,478,619,645]" pageId="8" pageNumber="201" value="2008-04-20">20.iv.2008</collectingDate>
</date>
, human feces baited
<collectingMethod id="524E4EA1FD71E319D4A66D47FCF5FD77" box="[712,782,619,645]" pageId="8" pageNumber="201">pitfall</collectingMethod>
,
<collectorName id="26FA5360FD71E319D5766D47FC4BFD77" box="[792,944,619,645]" pageId="8" pageNumber="201">W. Chamorro</collectorName>
(
<specimenCount id="9D09FD3FFD71E319D5AC6D47FBF0FD77" box="[962,1035,619,645]" pageId="8" pageNumber="201" type="male">1 male</specimenCount>
<collectionCode id="ED1EAE73FD71E319D27D6D47FB6BFD77" box="[1043,1168,619,645]" pageId="8" pageNumber="201">CONRAZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036B6FD71E319D6A96DA2FCCDFC73" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="201">
<emphasis id="B97BEAA4FD71E319D6A96DA2FE4FFD5B" bold="true" box="[199,436,654,681]" pageId="8" pageNumber="201">Literature records.</emphasis>
As “
<taxonomicName id="4C0F4D35FD71E319D7936DA3FD06FD5B" authority="" box="[509,765,655,681]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="subGenus" subGenus="Boreus">
<emphasis id="B97BEAA4FD71E319D7936DA3FD77FD5B" box="[509,652,655,681]" italics="true" pageId="8" pageNumber="201">Dichotomius</emphasis>
(
<emphasis id="B97BEAA4FD71E319D4C86DA3FD0FFD5A" box="[678,756,655,680]" italics="true" pageId="8" pageNumber="201">Boreus</emphasis>
)
</taxonomicName>
sp.” (in
<bibRefCitation id="EF9E4B47FD71E319D5166DA3FA70FD5B" author="Sarmiento-Garces, R. &amp; Amat-Garcia, G." box="[888,1419,655,681]" pageId="8" pageNumber="201" pagination="1 - 132" refId="ref9846" refString="Sarmiento-Garces, R. &amp; Amat-Garcia, G. (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia Monografia, 4, 1 - 132." type="journal article" year="2014">Sarmiento-Garcés &amp; Amat-García 2014: 44</bibRefCitation>
).:
<emphasis id="B97BEAA4FD71E319D6F96D9EFECDFD3E" bold="true" box="[151,310,690,716]" pageId="8" pageNumber="201">
<collectingCountry id="F3187626FD71E319D6F96D9EFECDFD3E" box="[151,310,690,716]" name="Colombia" pageId="8" pageNumber="201">COLOMBIA</collectingCountry>
</emphasis>
:
<collectingRegion id="49CBF854FD71E319D72C6D9FFE1EFD3F" box="[322,485,691,717]" country="Colombia" name="AMAZONAS" pageId="8" pageNumber="201">AMAZONAS</collectingRegion>
: Leticia vía a Tarapacá km
<quantity id="4CF79B53FD71E319D57F6D9FFC8EFD3F" box="[785,885,691,717]" metricMagnitude="3" metricUnit="m" metricValue="7.12" pageId="8" pageNumber="201" unit="m" value="7120.0">
7,
<specimenCount id="9D09FD3FFD71E319D5426D9FFC8EFD3F" box="[812,885,691,717]" pageId="8" pageNumber="201" type="generic">120 m</specimenCount>
</quantity>
. Leticia vía a Tarapacá km 11,
<specimenCount id="9D09FD3FFD71E319D2A46D9FFAF2FD3F" box="[1226,1289,691,717]" pageId="8" pageNumber="201" type="generic">90 m.</specimenCount>
<collectingRegion id="49CBF854FD71E319D3616D98FA62FD3D" box="[1295,1433,692,719]" country="Colombia" name="Caqueta" pageId="8" pageNumber="201">CAQUETÁ</collectingRegion>
: Puerto Solano, Parque Nacional Natural Chiribiquete, Río Saramano,
<specimenCount id="9D09FD3FFD71E319D5F46DFAFC1DFD03" box="[922,998,726,753]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D5F46DFAFC19FD03" box="[922,994,726,753]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="201" unit="m" value="250.0">250 m</quantity>
.
</specimenCount>
Chiribiquete, Río Cuñare-Amu,
<specimenCount id="9D09FD3FFD71E319D33E6DFAFA67FD03" box="[1360,1436,726,753]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D33E6DFAFA63FD03" box="[1360,1432,726,753]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="201" unit="m" value="250.0">250 m</quantity>
.
</specimenCount>
<collectingRegion id="49CBF854FD71E319D6F96DD7FED1FCE7" box="[151,298,763,789]" country="Colombia" name="Guaviare" pageId="8" pageNumber="201">GUAVIARE</collectingRegion>
: San José del Guaviare. Reserva Nukak, Santa Marta,
<specimenCount id="9D09FD3FFD71E319D5EA6DD6FC2AFCE7" box="[900,977,762,789]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D5EA6DD6FC36FCE7" box="[900,973,762,789]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="201" unit="m" value="250.0">250 m</quantity>
.
</specimenCount>
Nukak, Cerro. Tomachipan, caño Cocuy, Cerro Moyano,
<specimenCount id="9D09FD3FFD71E319D7276C33FE6CFCCB" box="[329,407,799,825]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D7276C33FE68FCCB" box="[329,403,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="201" unit="m" value="200.0">200 m</quantity>
.
</specimenCount>
<collectingRegion id="49CBF854FD71E319D7F06C33FE0BFCCB" box="[414,496,799,825]" country="Colombia" name="Meta" pageId="8" pageNumber="201">META</collectingRegion>
: Parque Nacional Natural Tinigua, Río Duda,
<specimenCount id="9D09FD3FFD71E319D59D6C32FBBAFCCB" box="[1011,1089,798,825]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D59D6C32FBC7FCCB" box="[1011,1084,798,825]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="201" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
Puerto López El Naranjal,
<specimenCount id="9D09FD3FFD71E319D31C6C33FF4FFCAF" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D31C6C33FF4BFCAF" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="201" unit="m" value="220.0">220 m</quantity>
.
</specimenCount>
<collectingRegion id="49CBF854FD71E319D6D36C68FED5FCAD" box="[189,302,836,863]" country="Colombia" name="Narino" pageId="8" pageNumber="201">NARIÑO</collectingRegion>
: Ipiales, Territorio Kofán, Cuenca alta de los ríos RumiyacoRanchería,
<specimenCount id="9D09FD3FFD71E319D21F6C6FFB39FCAF" box="[1137,1218,835,861]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D21F6C6FFB45FCAF" box="[1137,1214,835,861]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="201" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
<collectingRegion id="49CBF854FD71E319D2A46C68FAC3FCAD" box="[1226,1336,836,863]" country="Colombia" name="Vaupes" pageId="8" pageNumber="201">VAUPÉS</collectingRegion>
: Caparú. Igapó,
<specimenCount id="9D09FD3FFD71E319D68B6C4BFEC8FC73" box="[229,307,871,897]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D68B6C4BFED4FC73" box="[229,303,871,897]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="201" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
Terraza. R. N. Mosiro-Itajura, Colina,
<specimenCount id="9D09FD3FFD71E319D4866C4BFCCDFC73" box="[744,822,871,897]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D4866C4BFCC9FC73" box="[744,818,871,897]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="201" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BB036B6FD71E319D6A96CA7FF1EFC3B" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="201">
<materialsCitation id="3B673CEBFD71E319D6A96CA7FF1EFC3B" ID-GBIF-Occurrence="3061003328" country="Ecuador" elevation="570" location="Bosque Protector Oglan Alto" municipality="As" pageId="8" pageNumber="201" specimenCount="570" stateProvince="Pastaza">
<collectingMunicipality id="6BD4ACCCFD71E319D6A96CA7FF1DFC57" box="[199,230,907,933]" pageId="8" pageNumber="201">As</collectingMunicipality>
<taxonomicName id="4C0F4D35FD71E319D6956CA7FE71FC57" authorityName="Hope" authorityYear="1838" box="[251,394,907,933]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAA4FD71E319D6956CA7FE71FC57" box="[251,394,907,933]" italics="true" pageId="8" pageNumber="201">Dichotomius</emphasis>
</taxonomicName>
sp.” (in
<bibRefCitation id="EF9E4B47FD71E319D7806CA7FCF4FC57" author="Chamorro, W. R. &amp; Gallo, F. O. &amp; Delgado, S. &amp; Enriquez, S. I. &amp; Guasumba, V. &amp; Lopez-Iborra, G." box="[494,783,907,933]" pageId="8" pageNumber="201" pagination="34 - 49" refId="ref8115" refString="Chamorro, W. R., Gallo, F. O., Delgado, S., Enriquez, S. I., Guasumba, V. &amp; Lopez-Iborra, G. (2019 b) Los escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) del Bosque Protector Oglan Alto, Pastaza, Ecuador. Biota Colombiana, 20, 34 - 49. https: // doi. org / 10.21068 / c 2019. v 20 n 01 a 03." type="journal article" year="2019">
Chamorro
<emphasis id="B97BEAA4FD71E319D4096CA7FD61FC57" box="[615,666,907,933]" italics="true" pageId="8" pageNumber="201">et al</emphasis>
. 2019b: 9
</bibRefCitation>
):
<emphasis id="B97BEAA4FD71E319D5486CA6FC46FC56" bold="true" box="[806,957,906,932]" pageId="8" pageNumber="201">
<collectingCountry id="F3187626FD71E319D5486CA6FC43FC56" box="[806,952,906,932]" name="Ecuador" pageId="8" pageNumber="201">ECUADOR</collectingCountry>
:
</emphasis>
<collectingRegion id="49CBF854FD71E319D5AB6CA7FBB8FC57" box="[965,1091,907,933]" country="Ecuador" name="Pastaza" pageId="8" pageNumber="201">PASTAZA</collectingRegion>
:
<location id="8ED0606DFD71E319D2206CA7FA63FC57" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D2206CA7FA63FC57" box="[1102,1432,907,933]" country="Ecuador" municipality="As" name="Bosque Protector Oglan Alto" pageId="8" pageNumber="201" stateProvince="Pastaza">Bosque Protector Oglán Alto</location>
,
<specimenCount id="9D09FD3FFD71E319D6F96C82FF1EFC3B" box="[151,229,942,969]" pageId="8" pageNumber="201" type="generic">
<quantity id="4CF79B53FD71E319D6F96C82FF1AFC3B" box="[151,225,942,969]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="201" unit="m" value="570.0">
<elevation id="0022D185FD71E319D6F96C82FF1AFC3B" box="[151,225,942,969]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="201" unit="m" value="570.0">570 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="8BB036B6FD71E319D6A96CFFFC29FBE3" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="201">
<materialsCitation id="3B673CEBFD71E319D6A96CFFFC29FBE3" ID-GBIF-Occurrence="3061003329" country="Peru" county="As" elevation="148" location="Iquitos-Nauta" municipality="Reserva Nacional Allpahuayo Mishana" pageId="8" pageNumber="201" specimenCount="1" stateProvince="Loreto">
<collectingCounty id="62D14E3AFD71E319D6A96CFFFF1DFC1F" box="[199,230,979,1005]" pageId="8" pageNumber="201">As</collectingCounty>
<taxonomicName id="4C0F4D35FD71E319D6936CFFFE19FC1F" box="[253,482,979,1005]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D6936CFFFE19FC1F" box="[253,482,979,1005]" italics="true" pageId="8" pageNumber="201">Dichotomius boreus</emphasis>
</taxonomicName>
” (see
<bibRefCitation id="EF9E4B47FD71E319D4416CFFFCAAFC1C" author="Ampudia, G. C. &amp; Estrella, R. V. &amp; Perez, P. E." box="[559,849,979,1006]" pageId="8" pageNumber="201" pagination="53 - 70" refId="ref7922" refString="Ampudia, G. C., Estrella, R. V. &amp; Perez, P. E. (2020) Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) en bosques sobre arena blanca de la Reserva Nacional Allpahuayo Mishana, en la Amazonia peruana. Ciencia Amazonica (Iquitos), 8, 53 - 70. https: // doi. org / 10.22386 / ca. v 8 i 1.280" type="journal article" year="2020">
Ampudia
<emphasis id="B97BEAA4FD71E319D4CF6CFFFD2DFC1F" box="[673,726,979,1005]" italics="true" pageId="8" pageNumber="201">et al</emphasis>
. 2020: 61
</bibRefCitation>
):
<emphasis id="B97BEAA4FD71E319D5076CFFFC4EFC1F" bold="true" box="[873,949,979,1005]" pageId="8" pageNumber="201">
<collectingCountry id="F3187626FD71E319D5076CFFFC4EFC1F" box="[873,949,979,1005]" name="Peru" pageId="8" pageNumber="201">PERU</collectingCountry>
</emphasis>
:
<collectingRegion id="49CBF854FD71E319D5A96CFFFBC2FC1F" box="[967,1081,979,1005]" country="Peru" name="Loreto" pageId="8" pageNumber="201">LORETO</collectingRegion>
:
<collectingMunicipality id="6BD4ACCCFD71E319D2296CFFFF01FBE3" pageId="8" pageNumber="201">Reserva Nacional Allpahuayo Mishana</collectingMunicipality>
, near km 24.6 &amp; km 28.0 on the
<location id="8ED0606DFD71E319D4056CDBFCF1FBE3" LSID="urn:lsid:plazi:treatment:03A687A0FD7EE318D6F96DA8FC1BF96B:8ED0606DFD71E319D4056CDBFCF1FBE3" box="[619,778,1015,1041]" country="Peru" county="As" municipality="Reserva Nacional Allpahuayo Mishana" name="Iquitos-Nauta" pageId="8" pageNumber="201" stateProvince="Loreto">IquitosNauta</location>
road,
<quantity id="4CF79B53FD71E319D53F6CDAFC35FBE3" box="[849,974,1014,1041]" metricMagnitude="2" metricUnit="m" metricValue="1.48" metricValueMax="1.51" metricValueMin="1.45" pageId="8" pageNumber="201" unit="m" value="148.0" valueMax="151.0" valueMin="145.0">
<elevation id="0022D185FD71E319D53F6CDAFC35FBE3" box="[849,974,1014,1041]" metricMagnitude="2" metricUnit="m" metricValue="1.48" metricValueMax="1.51" metricValueMin="1.45" pageId="8" pageNumber="201" unit="m" value="148.0" valueMax="151.0" valueMin="145.0">145-151 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD71E319D6A96B37FEF7FBAB" pageId="8" pageNumber="201" type="etymology">
<paragraph id="8BB036B6FD71E319D6A96B37FEF7FBAB" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="201">
<emphasis id="B97BEAA4FD71E319D6A96B37FEB3FBC7" bold="true" box="[199,328,1051,1077]" pageId="8" pageNumber="201">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C0F4D35FD71E319D4296B37FD1FFBC7" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[583,740,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B97BEAA4FD71E319D4296B37FD1FFBC7" box="[583,740,1051,1077]" italics="true" pageId="8" pageNumber="201">quadrilobatus</emphasis>
</taxonomicName>
means “with four lobes” and refers to the structures on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD71E319D6A96B4EFC3BFA17" pageId="8" pageNumber="201" type="diagnosis">
<paragraph id="8BB036B6FD71E319D6A96B4EFC3BFA17" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="201">
<emphasis id="B97BEAA4FD71E319D6A96B4EFEB9FB8E" bold="true" box="[199,322,1122,1148]" pageId="8" pageNumber="201">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D35FD71E319D7286B4FFD8CFB8F" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[326,631,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B97BEAA4FD71E319D7286B4FFD8CFB8F" box="[326,631,1123,1149]" italics="true" pageId="8" pageNumber="201">Dichotomius quadrilobatus</emphasis>
</taxonomicName>
differs from other species of the
<taxonomicName id="4C0F4D35FD71E319D5B26B4FFBBCFB8F" box="[988,1095,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D5B26B4FFC0BFB8E" box="[988,1008,1123,1148]" italics="true" pageId="8" pageNumber="201">D</emphasis>
.
<emphasis id="B97BEAA4FD71E319D5926B4FFBBCFB8F" box="[1020,1095,1123,1149]" italics="true" pageId="8" pageNumber="201">boreus</emphasis>
</taxonomicName>
species group by the following combination of characters: (1) clypeus with symmetrically transverse carina, upper margin of clypeal carina concave medially (
<figureCitation id="13342A33FD71E319D7666B87FEABFB37" box="[264,336,1195,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="8" pageNumber="201">Figs. 1</figureCitation>
AB, D), (2) anterior pronotal edge with margin beaded and distinctly widened medially; superior pronotal lobes with anterior edge obtusely acuminate (forked to transversally truncate in
<taxonomicName id="4C0F4D35FD71E319D2EA6BE3FB09FB1B" box="[1156,1266,1231,1257]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D2EA6BE3FB09FB1B" box="[1156,1266,1231,1257]" italics="true" pageId="8" pageNumber="201">D. boreus</emphasis>
</taxonomicName>
) and divergent (closer and not divergent in the other members of the
<taxonomicName id="4C0F4D35FD71E319D4806BDFFCA1FAFF" box="[750,858,1267,1293]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D4806BDFFCF9FAFE" box="[750,770,1267,1292]" italics="true" pageId="8" pageNumber="201">D</emphasis>
.
<emphasis id="B97BEAA4FD71E319D5616BDFFCA1FAFF" box="[783,858,1267,1293]" italics="true" pageId="8" pageNumber="201">boreus</emphasis>
</taxonomicName>
species group); external apices of the lobes marked and pointing forward. In addition, remarkable differences are found in the morphology of the male genitalia: apex of parameres pincer shaped (normal in
<taxonomicName id="4C0F4D35FD71E319D4306A17FD36FAA7" box="[606,717,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D4306A17FD36FAA7" box="[606,717,1339,1365]" italics="true" pageId="8" pageNumber="201">D. boreus</emphasis>
</taxonomicName>
); dorsal edges of the parameres parallel and close proximally, clearly divergent near the apex, and convergent apically (dorsal edges close throughout and divergent at the apex in
<taxonomicName id="4C0F4D35FD71E319D6F96AAFFEFDFA6F" box="[151,262,1411,1437]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D6F96AAFFEFDFA6F" box="[151,262,1411,1437]" italics="true" pageId="8" pageNumber="201">D. boreus</emphasis>
</taxonomicName>
); lamella copulatrix differs in shape of the left lobe, whose margin of inferior branch of the bifurcation is broadly rounded (clearly acuminate in
<taxonomicName id="4C0F4D35FD71E319D4086A8BFD2FFA33" box="[614,724,1447,1473]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D4086A8BFD2FFA33" box="[614,724,1447,1473]" italics="true" pageId="8" pageNumber="201">D. boreus</emphasis>
</taxonomicName>
). To date,
<taxonomicName id="4C0F4D35FD71E319D5236A8BFBF7FA33" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[845,1036,1447,1473]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B97BEAA4FD71E319D5236A8BFBF7FA33" box="[845,1036,1447,1473]" italics="true" pageId="8" pageNumber="201">D. quadrilobatus</emphasis>
</taxonomicName>
, is the largest species within the
<taxonomicName id="4C0F4D35FD71E319D3EF6A8BFF19FA17" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD71E319D3EF6A8BFA6EFA32" box="[1409,1429,1447,1472]" italics="true" pageId="8" pageNumber="201">D</emphasis>
.
<emphasis id="B97BEAA4FD71E319D6F96AE7FF19FA17" box="[151,226,1483,1509]" italics="true" pageId="8" pageNumber="201">boreus</emphasis>
</taxonomicName>
species group, with some specimens exceeding
<quantity id="4CF79B53FD71E319D56E6AE7FCB4FA17" box="[768,847,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="201" unit="mm" value="31.0">31 mm</quantity>
in length.
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD71E318D6A96AC2FBB6FBAB" lastPageId="9" lastPageNumber="202" pageId="8" pageNumber="201" type="description">
<paragraph id="8BB036B6FD71E318D6A96AC2FEF7FC57" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,150,1689]" lastPageId="9" lastPageNumber="202" pageId="8" pageNumber="201">
<emphasis id="B97BEAA4FD71E319D6A96AC2FDADF9FB" bold="true" box="[199,598,1518,1545]" pageId="8" pageNumber="201">Description. Body measurements.</emphasis>
Male (
<figureCitation id="13342A33FD71E319D4CA6AC3FCFDF9FB" box="[676,774,1519,1545]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="8" pageNumber="201">Figs. 1A</figureCitation>
, CD): length
<quantity id="4CF79B53FD71E319D5C76AC3FBDAF9FB" box="[937,1057,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="3.2" metricValueMin="1.9" pageId="8" pageNumber="201" unit="mm" value="25.5" valueMax="32.0" valueMin="19.0">1932 mm</quantity>
(
<typeStatus id="54B48814FD71E319D2406AC3FB69F9FB" box="[1070,1170,1519,1545]" pageId="8" pageNumber="201" type="holotype">holotype</typeStatus>
<quantity id="4CF79B53FD71E319D2F66AC3FB04F9FB" box="[1176,1279,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="8" pageNumber="201" unit="mm" value="31.2">31.2 mm</quantity>
); width
<quantity id="4CF79B53FD71E319D3396AC3FF38F9DF" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="8" pageNumber="201" unit="mm" value="15.0" valueMax="19.0" valueMin="11.0">1119 mm</quantity>
(
<typeStatus id="54B48814FD71E319D6BD693FFECCF9DF" box="[211,311,1555,1581]" pageId="8" pageNumber="201" type="holotype">holotype</typeStatus>
<quantity id="4CF79B53FD71E319D72E693FFE52F9DF" box="[320,425,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="8" pageNumber="201" unit="mm" value="18.6">18.6 mm</quantity>
). Female (
<figureCitation id="13342A33FD71E319D44C693FFD81F9DF" box="[546,634,1555,1581]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="8" pageNumber="201">Fig. 1B</figureCitation>
): length 22.127.0 mm in length; width
<quantity id="4CF79B53FD71E319D222693FFB0FF9DF" box="[1100,1268,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.7899999999999998" metricValueMin="1.28" pageId="8" pageNumber="201" unit="mm" value="15.35" valueMax="17.9" valueMin="12.8">12.817.9 mm</quantity>
.
<emphasis id="B97BEAA4FD71E319D36E693EFAB2F9DE" bold="true" box="[1280,1353,1554,1580]" pageId="8" pageNumber="201">Color.</emphasis>
Dorsal side of body black to reddish brown (
<typeStatus id="54B48814FD71E319D452691BFD5BF9A3" box="[572,672,1591,1617]" pageId="8" pageNumber="201" type="holotype">holotype</typeStatus>
matt black), antennomeres and mouthparts reddish brown, antennal club dark yellow and dull.
<emphasis id="B97BEAA4FD71E319D7AC6977FDF9F987" bold="true" box="[450,514,1627,1653]" pageId="8" pageNumber="201">Head</emphasis>
. (
<figureCitation id="13342A33FD71E319D4766977FD95F987" box="[536,622,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="8" pageNumber="201">Fig. 1D</figureCitation>
) Semicircular, wider than long. Clypeal margin feebly sinuate at middle, clypeal surface strongly wrinkled. Clypeal carina high and trapezoidal, located behind clypeal edge. Clypeogenal suture marked, slightly swollen medially. External edge of genae rounded, with small tubercle in proximity of the clypeogenal junction. Frons with trituberculate carina; central tubercle higher than the lateral tubercles, triangular shaped, and with acute apex; lateral tubercles low and wide.
<emphasis id="B97BEAA4FD71E319D52E69C7FC42F8F7" bold="true" box="[832,953,1771,1797]" pageId="8" pageNumber="201">Pronotum</emphasis>
. Wider than long; anterior pronotal margin beaded, posterior margin wider at the middle, posterior margin of bead obtusely pointed backwards, lateral edges of the pronotum with brown setae. Pronotal surface smooth and bright, with simple, irregular punctures; anterior region declivous. Medial suture deep, dividing the lobes longitudinally forming a “T” with lobes anterior margins (
<figureCitation id="13342A33FD71E319D30A687BFF5AF867" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="8" pageNumber="201">Figs. 1</figureCitation>
AB). Lobes square, outer edges normally with rounded projection at apex. Mediobasal region of pronotum with punctures simple and rounded. Lateral fovea of pronotum located on pronotal anterior ridge that connects superior lobes, ridge above foveae forming carina-like projection. Deeply excavated fovea located between anteromedial and posterolateral edge of pronotum.
<emphasis id="B97BEAA4FD70E318D4656FBAFD34FF43" bold="true" box="[523,719,150,177]" pageId="9" pageNumber="202">Thoracic sterna.</emphasis>
Proepisternum with small spiniform process between procoxae, procoxae with rounded apex, proepisternal surface heavily punctate anteriorly, punctures ocellate with long setae. Propleural surface pubescent on anterior and lateral regions, glabrous in posterior region. Prosternal surface rough, smooth in the middle, sparsely pubescent, with carina at middle. Mesosternal surface rough, anterior edge feebly rounded, central and posterior regions smooth, anterolateral regions with strong and ocellate punctures associated with long, dense setae. Mesepisternum densely punctate. Metasternal disc finely punctate, with feeble groove at middle.
<emphasis id="B97BEAA4FD70E318D69B6E43FEB2FE7B" bold="true" box="[245,329,367,393]" pageId="9" pageNumber="202">Elytra.</emphasis>
Surface smooth, without microsculpture. Striae shallow, more impressed on the elytral disc and apex, striae punctures rounded, separated by at least 1.5 times the size of the puncture. Interstriae feebly convex and with fine punctures.
<emphasis id="B97BEAA4FD70E318D7146E9BFE42FE23" bold="true" box="[378,441,439,465]" pageId="9" pageNumber="202">Legs.</emphasis>
Profemora, mesofemora, and metafemora completely smooth, with long setae on anterior and posterior edges, Protibial surface smooth dorsally and rugose ventrally, protibial spur acute, directed inwards. Mesotibial and metatibial surface smooth, external edge strongly serrate and internal edge finely serrate, both edges with long setae. Metatibial spur feebly emarginate at apex.
<emphasis id="B97BEAA4FD70E318D5176D0EFC0EFDCE" bold="true" box="[889,1013,546,572]" pageId="9" pageNumber="202">Abdomen.</emphasis>
Sternites 26 without microsculpture and with small, rounded punctures along anterior and posterior edges; posterolateral regions pubescent and with ocellate punctures.
<emphasis id="B97BEAA4FD70E318D71D6D46FE16FD76" bold="true" box="[371,493,618,644]" pageId="9" pageNumber="202">Pygidium.</emphasis>
Wider than long, surface smooth, with small, dense punctation, margins complete.
<emphasis id="B97BEAA4FD70E318D6F96DA3FEB9FD5A" bold="true" box="[151,322,654,681]" pageId="9" pageNumber="202">Male genitalia</emphasis>
(
<figureCitation id="13342A33FD70E318D73C6DA3FE53FD5B" box="[338,424,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="9" pageNumber="202">Fig. 1E</figureCitation>
). In dorsal view, parameres narrower apically, sides rather straight and not sinuate, apices rounded and curved inward, assuming the shape of pincers. Dorsal edges of parameres simple, parallel and close proximally and medially, divergent near apex and convergent apically. In lateral view, ventral side of parameres straight, excavated near the phallobase, dorsal side of parameres evenly curved toward apex.
<emphasis id="B97BEAA4FD70E318D29E6DD6FA6EFCE6" bold="true" box="[1264,1429,762,788]" pageId="9" pageNumber="202">Endophallites</emphasis>
. Lamella copulatrix (
<figureCitation id="13342A33FD70E318D7136C33FE29FCCB" box="[381,466,799,825]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="9" pageNumber="202">Fig. 1F</figureCitation>
): horseshoe shaped, with left inferior lobe elongate and developed upward, right lobe distinctly wider, bent upward at the bottom; apex of right lobe bifurcated, superior branch acuminate and directed upward, while inferior branch is widely rounded and directed internally. See
<figureCitation id="13342A33FD70E318D5996C4BFBB3FC73" box="[1015,1096,871,897]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="9" pageNumber="202">Fig. 1F</figureCitation>
for the remaining endophallic structures.
</paragraph>
<paragraph id="8BB036B6FD70E318D6A96C82FBB6FBAB" blockId="9.[151,1437,150,1689]" pageId="9" pageNumber="202">
<emphasis id="B97BEAA4FD70E318D6A96C82FE20FC3A" bold="true" box="[199,475,942,968]" pageId="9" pageNumber="202">Intraspecific variation.</emphasis>
Males and females of
<taxonomicName id="4C0F4D35FD70E318D48F6C83FC59FC3B" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[737,930,943,969]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B97BEAA4FD70E318D48F6C83FC59FC3B" box="[737,930,943,969]" italics="true" pageId="9" pageNumber="202">D. quadrilobatus</emphasis>
</taxonomicName>
are morphologically very similar, but males can be recognized by the shape of the posterior margin of the anterior pronotal bead, which is pointed backward (it is simply curved in females) (
<figureCitation id="13342A33FD70E318D7866CDBFDC4FBE3" box="[488,575,1015,1041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1917,1941]" captionTargetBox="[172,1421,193,1883]" captionTargetId="figure-17@3.[159,1429,181,1894]" captionTargetPageId="3" captionText="FIGURE 1. Dichotomius species. A, Dichotomius quadrilobatus (dorsal view), male; B, Dichotomius quadrilobatus (dorsal view), female; C, Dichotomius quadrilobatus (lateral view), male; D, Dichotomius quadrilobatus, head male; E, Dichotomius quadrilobatus, aedeagus; F, Dichotomius quadrilobatus, endophallites." figureDoi="http://doi.org/10.5281/zenodo.4600449" httpUri="https://zenodo.org/record/4600449/files/figure.png" pageId="9" pageNumber="202">Fig. 1B</figureCitation>
), clypeal carina is narrower in males (clearly wider in females), and the last abdominal sternite is narrower at middle in males (evenly wide in females). Small specimens (about
<quantity id="4CF79B53FD70E318D3656B37FAA7FBC7" box="[1291,1372,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="202" unit="mm" value="21.0">21 mm</quantity>
) may lack of clypeal carina and have reduced and somewhat rounded pronotal projections.
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD70E318D6A96B4FFB97FA6F" pageId="9" pageNumber="202" type="discussion">
<paragraph id="8BB036B6FD70E318D6A96B4FFB97FA6F" blockId="9.[151,1437,150,1689]" pageId="9" pageNumber="202">
<emphasis id="B97BEAA4FD70E318D6A96B4FFEC0FB8F" bold="true" box="[199,315,1123,1149]" pageId="9" pageNumber="202">Remarks.</emphasis>
<bibRefCitation id="EF9E4B47FD70E318D7276B4FFCDEFB8F" author="Sarmiento-Garces, R. &amp; Amat-Garcia, G." box="[329,805,1123,1149]" pageId="9" pageNumber="202" pagination="285 - 296" refId="ref9798" refString="Sarmiento-Garces, R. &amp; Amat-Garcia, G. (2009) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en la amazonia colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 33, 285 - 296." type="journal article" year="2009">Sarmiento-Garcés &amp; Amat-García (2009</bibRefCitation>
,
<bibRefCitation id="EF9E4B47FD70E318D5596B4FFC8AFB8F" author="Sarmiento-Garces, R. &amp; Amat-Garcia, G." box="[823,881,1123,1149]" pageId="9" pageNumber="202" pagination="1 - 132" refId="ref9846" refString="Sarmiento-Garces, R. &amp; Amat-Garcia, G. (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia Monografia, 4, 1 - 132." type="journal article" year="2014">2014</bibRefCitation>
) provided a short description, along with two illustrations of the head and pronotum of a
<taxonomicName id="4C0F4D35FD70E318D4EF6BABFCEBFB53" authorityName="Hope" authorityYear="1838" box="[641,784,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAA4FD70E318D4EF6BABFCEBFB53" box="[641,784,1159,1185]" italics="true" pageId="9" pageNumber="202">Dichotomius</emphasis>
</taxonomicName>
species, which was considered by the authors themselves to be a close relative of
<emphasis id="B97BEAA4FD70E318D7DD6B87FDD0FB37" box="[435,555,1195,1221]" italics="true" pageId="9" pageNumber="202">
<taxonomicName id="4C0F4D35FD70E318D7DD6B87FDDCFB37" box="[435,551,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="species" species="boreus">D. boreus</taxonomicName>
.
</emphasis>
Although apparently aware of describing a new species, the authors did not provide any new name for this
<taxonomicName id="4C0F4D35FD70E318D79E6BE3FD84FB1B" authorityName="Hope" authorityYear="1838" box="[496,639,1231,1257]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAA4FD70E318D79E6BE3FD84FB1B" box="[496,639,1231,1257]" italics="true" pageId="9" pageNumber="202">Dichotomius</emphasis>
</taxonomicName>
, nor selected any name-bearing
<typeStatus id="54B48814FD70E318D5876BE3FBE2FB1B" box="[1001,1049,1231,1257]" pageId="9" pageNumber="202">type</typeStatus>
specimen to fix the identity of this species, which was initially called “
<taxonomicName id="4C0F4D35FD70E318D45C6BDFFD3AFAFF" authorityName="Hope" authorityYear="1838" box="[562,705,1267,1293]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAA4FD70E318D45C6BDFFD3AFAFF" box="[562,705,1267,1293]" italics="true" pageId="9" pageNumber="202">Dichotomius</emphasis>
</taxonomicName>
af.
<taxonomicName id="4C0F4D35FD70E318D49E6BDFFCC0FAFF" box="[752,827,1267,1293]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="species" species="boreus">
<emphasis id="B97BEAA4FD70E318D49E6BDFFCC0FAFF" box="[752,827,1267,1293]" italics="true" pageId="9" pageNumber="202">boreus</emphasis>
</taxonomicName>
” (
<bibRefCitation id="EF9E4B47FD70E318D5366BDFFACBFAFF" author="Sarmiento-Garces, R. &amp; Amat-Garcia, G." box="[856,1328,1267,1293]" pageId="9" pageNumber="202" pagination="285 - 296" refId="ref9798" refString="Sarmiento-Garces, R. &amp; Amat-Garcia, G. (2009) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en la amazonia colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 33, 285 - 296." type="journal article" year="2009">Sarmiento-Garcés &amp; Amat-García (2009)</bibRefCitation>
and later “
<taxonomicName id="4C0F4D35FD70E318D6CA6A3BFE66FAC3" authority="" box="[164,413,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="subGenus" subGenus="Boreus">
<emphasis id="B97BEAA4FD70E318D6CA6A3BFEC8FAC3" box="[164,307,1303,1329]" italics="true" pageId="9" pageNumber="202">Dichotomius</emphasis>
(
<emphasis id="B97BEAA4FD70E318D7286A3BFE6FFAC2" box="[326,404,1303,1328]" italics="true" pageId="9" pageNumber="202">Boreus</emphasis>
)
</taxonomicName>
sp.” (
<bibRefCitation id="EF9E4B47FD70E318D78B6A3BFC38FAC3" author="Sarmiento-Garces, R. &amp; Amat-Garcia, G." box="[485,963,1303,1329]" pageId="9" pageNumber="202" pagination="1 - 132" refId="ref9846" refString="Sarmiento-Garces, R. &amp; Amat-Garcia, G. (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia Monografia, 4, 1 - 132." type="journal article" year="2014">Sarmiento-Garcés &amp; Amat-García (2014)</bibRefCitation>
. In both papers, the authors included the species in their identification keys, which would lead one to suppose that they considered the series of specimens examined to belong to a distinct species. It is unknown why these authors did not validate a name for this species under the articles of the
<bibRefCitation id="EF9E4B47FD70E318D7C96AAFFB93FA6F" author="International Commission on Zoological Nomenclature" box="[423,1128,1411,1437]" pageId="9" pageNumber="202" refId="ref8931" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315653DFD70E318D6A96A8AFC1BF96B" pageId="9" pageNumber="202" type="distribution">
<paragraph id="8BB036B6FD70E318D6A96A8AFC11F987" blockId="9.[151,1437,150,1689]" pageId="9" pageNumber="202">
<emphasis id="B97BEAA4FD70E318D6A96A8AFE9AFA32" bold="true" box="[199,353,1446,1472]" pageId="9" pageNumber="202">Distribution.</emphasis>
This species is distributed in
<collectingCountry id="F3187626FD70E318D4DA6A8BFD00FA33" box="[692,763,1447,1473]" name="Brazil" pageId="9" pageNumber="202">Brazil</collectingCountry>
,
<collectingCountry id="F3187626FD70E318D5666A8BFC82FA33" box="[776,889,1447,1473]" name="Colombia" pageId="9" pageNumber="202">Colombia</collectingCountry>
,
<collectingCountry id="F3187626FD70E318D5E86A8BFC1EFA33" box="[902,997,1447,1473]" name="Ecuador" pageId="9" pageNumber="202">Ecuador</collectingCountry>
, and
<collectingCountry id="F3187626FD70E318D24C6A8BFBAEFA33" box="[1058,1109,1447,1473]" name="Peru" pageId="9" pageNumber="202">Peru</collectingCountry>
and inhabits lowland forests (terrafirme forests), white sand forests (in
<collectingCountry id="F3187626FD70E318D4006AE7FD5FFA17" box="[622,676,1483,1509]" name="Peru" pageId="9" pageNumber="202">Peru</collectingCountry>
, locally known as “varillal”), and foothill forests in the Amazon and Orinoco regions (
<figureCitation id="13342A33FD70E318D7326AC3FE5AF9FB" box="[348,417,1518,1545]" captionStart="FIGURE5" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[196,1394,189,1909]" captionTargetId="figure-17@11.[184,1404,181,1919]" captionTargetPageId="11" captionText="FIGURE5. Distributionof Dichotomius (Dichotomius) quadrilobatusnewspecies(greencircle) andDichotomius(Selenocopris) fortepunctatus (Luederwaldt, 1923) (blue square). (?) Doubtful data." figureDoi="http://doi.org/10.5281/zenodo.4600459" httpUri="https://zenodo.org/record/4600459/files/figure.png" pageId="9" pageNumber="202">Fig. 5</figureCitation>
). According to the biogeographical analysis proposed by
<bibRefCitation id="EF9E4B47FD70E318D2466AC3FB25F9FB" author="Morrone, J. J." box="[1064,1246,1519,1545]" pageId="9" pageNumber="202" pagination="1 - 110" refId="ref9569" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782, 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone (2014)</bibRefCitation>
, and considering our current knowledge on its distribution,
<taxonomicName id="4C0F4D35FD70E318D40D693FFCDBF9DF" authorityName="Chamorro, Lopera, &amp; Rossini" authorityYear="2021" box="[611,800,1555,1581]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="202" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B97BEAA4FD70E318D40D693FFCDBF9DF" box="[611,800,1555,1581]" italics="true" pageId="9" pageNumber="202">D. quadrilobatus</emphasis>
</taxonomicName>
occurs in the biogeographical provinces of Sabana, Napo, and Imeri. This new species has been recorded from
<quantity id="4CF79B53FD70E318D483691AFC99F9A3" box="[749,866,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="4.175" metricValueMax="7.5" metricValueMin="0.85" pageId="9" pageNumber="202" unit="m" value="417.5" valueMax="750.0" valueMin="85.0">
85
<specimenCount id="9D09FD3FFD70E318D578691AFC99F9A3" box="[790,866,1590,1617]" pageId="9" pageNumber="202" type="generic">750 m</specimenCount>
</quantity>
. The record from Santiago,
<collectingRegion id="49CBF854FD70E318D2F1691BFA67F9A3" box="[1183,1436,1591,1617]" country="Colombia" name="Putumayo" pageId="9" pageNumber="202">Putumayo Department</collectingRegion>
(
<geoCoordinate id="EE3B5071FD70E318D6CF6977FF1CF987" box="[161,231,1627,1653]" degrees="1" direction="north" minutes="9" orientation="latitude" pageId="9" pageNumber="202" precision="925" value="1.15">1°9N</geoCoordinate>
,
<geoCoordinate id="EE3B5071FD70E318D69D6977FEB4F987" box="[243,335,1627,1653]" degrees="77" direction="west" minutes="0" orientation="longitude" pageId="9" pageNumber="202" precision="925" value="-77.0">77°0W</geoCoordinate>
),
<collectingCountry id="F3187626FD70E318D70A6977FE2EF987" box="[356,469,1627,1653]" name="Colombia" pageId="9" pageNumber="202">Colombia</collectingCountry>
, which is located at
<quantity id="4CF79B53FD70E318D4D66977FCF4F987" box="[696,783,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="202" unit="m" value="2000.0">2000 m</quantity>
, may be erroneous.
</paragraph>
<paragraph id="8BB036B6FD70E318D6A96953FC1BF96B" blockId="9.[151,1437,150,1689]" box="[199,992,1663,1689]" pageId="9" pageNumber="202">
<emphasis id="B97BEAA4FD70E318D6A96953FE85F96B" bold="true" box="[199,382,1663,1689]" pageId="9" pageNumber="202">Temporal data.</emphasis>
Collected all months of the year except June and July.
</paragraph>
</subSubSection>
</treatment>
</document>