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<document ID-DOI="http://doi.org/10.5281/zenodo.6167528" ID-GBIF-Dataset="1fd9017d-6a56-470b-aa83-68944cd8ac46" ID-GBIF-Taxon="130509476" ID-Zenodo-Dep="6167528" checkinTime="1492608005255" checkinUser="jeremy" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2012" docId="03A6879FA85FFFBDE595FF7B0561613A" docLanguage="en" docName="DankittipakulTavano_Singtripop2012FINAL.imf" docOrigin="Journal of Natural History 46" docSource="https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae" docStyle="DocumentStyle{}" docTitle="Teutamus hirtellus Dankittipakul, Tavano &amp; Singtripop, 2012, sp. nov." docType="treatment" docVersion="18" lastPageId="30" lastPageNumber="1719" masterDocId="FF9FFFE7A842FFA3E422FFF90169654C" masterDocTitle="Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae)" masterLastPageNumber="1730" masterPageNumber="1689" pageId="29" pageNumber="1718" updateTime="1645288162421" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dankittipakul, Pakawin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tavano, Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Singtripop, Tippawan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1689</mods:start>
<mods:end>1730</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.110.1080/00222933.2012.681314</mods:identifier>
<mods:identifier type="GBIF-Dataset">1fd9017d-6a56-470b-aa83-68944cd8ac46</mods:identifier>
<mods:identifier type="Zenodo-Dep">574642</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6167528" ID-GBIF-Taxon="130509476" ID-Zenodo-Dep="6167528" LSID="urn:lsid:plazi:treatment:03A6879FA85FFFBDE595FF7B0561613A" httpUri="http://treatment.plazi.org/id/03A6879FA85FFFBDE595FF7B0561613A" lastPageId="30" lastPageNumber="1719" pageId="29" pageNumber="1718">
<subSubSection box="[439,732,129,154]" pageId="29" pageNumber="1718" type="nomenclature">
<paragraph blockId="29.[439,732,129,186]" box="[439,732,129,154]" pageId="29" pageNumber="1718">
<heading box="[439,732,129,154]" centered="true" fontSize="10" level="3" pageId="29" pageNumber="1718" reason="4">
<taxonomicName box="[439,641,129,153]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="hirtellus" status="sp. nov.">
<emphasis bold="true" box="[439,641,129,153]" italics="true" pageId="29" pageNumber="1718">Teutamus hirtellus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[648,732,131,154]" pageId="29" pageNumber="1718" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[483,688,161,186]" pageId="29" pageNumber="1718" type="description">
<paragraph blockId="29.[439,732,129,186]" box="[483,688,161,186]" pageId="29" pageNumber="1718">
<heading box="[483,688,161,186]" centered="true" fontSize="10" level="3" pageId="29" pageNumber="1718" reason="4">
(
<figureCitation box="[491,598,161,186]" captionStart="Figure 2" captionStartId="4.[173,240,1479,1501]" captionTargetId="figure@3.[161,1010,124,1660]" captionTargetPageId="3" captionText="Figure 2. Teutamus orthogonus sp. nov., male holotype (A). Teutamus jambiensis, male paratype (MHNG) (B). Teutamus andrewdavisi, male paratype (MHNG) (C). Teutamus politus, male (D, I) from southern Thailand, female holotype (MSNG) (E). Teutamus brachiatus sp. nov., male paratype (F). Teutamus lioneli sp. nov., male paratype (G). Teutamus hirtellus sp. nov., male holotype (H). (A G) Habitus, dorsal and (H) ventral; (I) opisthosoma, ventral. Abbreviations: ES, epigastric scutum; PS, post-epigastric scutum; SM, submarginal lobe." httpUri="https://zenodo.org/record/574644/files/figure.png" pageId="29" pageNumber="1718">Figures 2</figureCitation>
H,
<figureCitation box="[632,659,162,186]" captionStart="Figure 14" captionText="Figure 14. Teutamus hirtellus sp. nov., male holotype (A). Teutamus andrewdavisi, male paratype (MHNG) (B). Teutamus jambiensis, male paratype (MHNG) (C). (A C) Male palp, ventral. Scale bars 0.25 mm." pageId="29" pageNumber="1718">14</figureCitation>
A)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="1718" type="diagnosis">
<paragraph blockId="29.[105,1066,244,562]" box="[105,213,244,267]" pageId="29" pageNumber="1718">
<emphasis box="[105,213,244,267]" italics="true" pageId="29" pageNumber="1718">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="29.[105,1066,244,562]" pageId="29" pageNumber="1718">
The male holotype of
<taxonomicName box="[380,504,283,307]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="hirtellus" status="sp. nov.">
<emphasis box="[380,504,283,307]" italics="true" pageId="29" pageNumber="1718">T. hirtellus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[518,609,284,307]" pageId="29" pageNumber="1718" rank="species">sp. nov.</taxonomicNameLabel>
closely resembles that of
<taxonomicName box="[928,1065,283,307]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="rothorum">
<emphasis box="[928,1065,283,307]" italics="true" pageId="29" pageNumber="1718">T. rothorum</emphasis>
</taxonomicName>
in having a large conductor but can be easily distinguished by the more elongate conductor pointing anteriad instead of prolaterad (
<figureCitation box="[724,841,346,371]" captionStart="Figure 14" captionStartId="29.[105,172,1518,1540]" captionTargetBox="[105,1052,713,1485]" captionTargetId="figure@29.[105,1065,711,1485]" captionTargetPageId="29" captionText="Figure 14. Teutamus hirtellus sp. nov., male holotype (A). Teutamus andrewdavisi, male paratype (MHNG) (B). Teutamus jambiensis, male paratype (MHNG) (C). (A C) Male palp, ventral. Scale bars 0.25 mm." httpUri="https://zenodo.org/record/574656/files/figure.png" pageId="29" pageNumber="1718">Figure 14</figureCitation>
A). To our knowledge, only the male of this new species and those of
<taxonomicName box="[733,906,379,403]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="andrewdavisi">
<emphasis box="[733,906,379,403]" italics="true" pageId="29" pageNumber="1718">T. andrewdavisi</emphasis>
</taxonomicName>
(
<figureCitation box="[926,1039,378,403]" captionStart="Figure 14" captionStartId="29.[105,172,1518,1540]" captionTargetBox="[105,1052,713,1485]" captionTargetId="figure@29.[105,1065,711,1485]" captionTargetPageId="29" captionText="Figure 14. Teutamus hirtellus sp. nov., male holotype (A). Teutamus andrewdavisi, male paratype (MHNG) (B). Teutamus jambiensis, male paratype (MHNG) (C). (A C) Male palp, ventral. Scale bars 0.25 mm." httpUri="https://zenodo.org/record/574656/files/figure.png" pageId="29" pageNumber="1718">Figure 14</figureCitation>
B) and
<taxonomicName box="[159,275,412,435]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis box="[159,275,412,435]" italics="true" pageId="29" pageNumber="1718">T. vittatus</emphasis>
</taxonomicName>
possess a group of bristles on the RTA. In other species the postepigastric scuta slightly extend posteriorly (
<figureCitation box="[597,695,442,467]" captionStart="Figure 2" captionStartId="4.[173,240,1479,1501]" captionTargetId="figure@3.[161,1010,124,1660]" captionTargetPageId="3" captionText="Figure 2. Teutamus orthogonus sp. nov., male holotype (A). Teutamus jambiensis, male paratype (MHNG) (B). Teutamus andrewdavisi, male paratype (MHNG) (C). Teutamus politus, male (D, I) from southern Thailand, female holotype (MSNG) (E). Teutamus brachiatus sp. nov., male paratype (F). Teutamus lioneli sp. nov., male paratype (G). Teutamus hirtellus sp. nov., male holotype (H). (A G) Habitus, dorsal and (H) ventral; (I) opisthosoma, ventral. Abbreviations: ES, epigastric scutum; PS, post-epigastric scutum; SM, submarginal lobe." httpUri="https://zenodo.org/record/574644/files/figure.png" pageId="29" pageNumber="1718">
Figure
<date box="[680,705,443,467]" pageId="29" pageNumber="1718">2I</date>
</figureCitation>
), whereas in
<taxonomicName box="[855,973,442,466]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="hirtellus" status="sp. nov.">
<emphasis box="[855,973,442,466]" italics="true" pageId="29" pageNumber="1718">T. hirtellus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[981,1066,443,466]" pageId="29" pageNumber="1718" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName box="[159,311,474,498]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="orthogonus" status="sp. nov.">
<emphasis box="[159,311,474,498]" italics="true" pageId="29" pageNumber="1718">T. orthogonus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[322,411,475,498]" pageId="29" pageNumber="1718" rank="species">sp. nov.</taxonomicNameLabel>
they extend further and fuse with the dorsal scutum to form a complete tubular ring enclosing anterior half of the opisthosoma entirely (
<figureCitation box="[113,210,538,562]" captionStart="Figure 2" captionStartId="4.[173,240,1479,1501]" captionTargetId="figure@3.[161,1010,124,1660]" captionTargetPageId="3" captionText="Figure 2. Teutamus orthogonus sp. nov., male holotype (A). Teutamus jambiensis, male paratype (MHNG) (B). Teutamus andrewdavisi, male paratype (MHNG) (C). Teutamus politus, male (D, I) from southern Thailand, female holotype (MSNG) (E). Teutamus brachiatus sp. nov., male paratype (F). Teutamus lioneli sp. nov., male paratype (G). Teutamus hirtellus sp. nov., male holotype (H). (A G) Habitus, dorsal and (H) ventral; (I) opisthosoma, ventral. Abbreviations: ES, epigastric scutum; PS, post-epigastric scutum; SM, submarginal lobe." httpUri="https://zenodo.org/record/574644/files/figure.png" pageId="29" pageNumber="1718">Figure 2</figureCitation>
H).
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/574656/files/figure.png" pageId="29" pageNumber="1718" targetBox="[105,1052,713,1485]" targetPageId="29">
<paragraph blockId="29.[105,1066,1518,1598]" pageId="29" pageNumber="1718">
Figure 14.
<taxonomicName box="[220,400,1518,1540]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="hirtellus" status="sp. nov.">
<emphasis box="[220,400,1518,1540]" italics="true" pageId="29" pageNumber="1718">Teutamus hirtellus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[405,479,1519,1540]" pageId="29" pageNumber="1718" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (A).
<taxonomicName box="[684,911,1518,1540]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="andrewdavisi">
<emphasis box="[684,911,1518,1540]" italics="true" pageId="29" pageNumber="1718">Teutamus andrewdavisi</emphasis>
</taxonomicName>
, male paratype (MHNG) (B).
<taxonomicName box="[260,468,1547,1569]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="1718" phylum="Arthropoda" rank="species" species="jambiensis">
<emphasis box="[260,468,1547,1569]" italics="true" pageId="29" pageNumber="1718">Teutamus jambiensis</emphasis>
</taxonomicName>
, male paratype (MHNG) (C). (AC) Male palp, ventral. Scale bars 0.25 mm.
</paragraph>
</caption>
<subSubSection pageId="30" pageNumber="1719" type="materials_examined">
<paragraph blockId="30.[173,1134,130,258]" box="[173,326,130,154]" pageId="30" pageNumber="1719">
<emphasis box="[173,326,130,154]" italics="true" pageId="30" pageNumber="1719">Type material</emphasis>
</paragraph>
<paragraph blockId="30.[173,1134,130,258]" pageId="30" pageNumber="1719">
<materialsCitation ID-GBIF-Occurrence="1502458267" collectingDate="2009-12-06" collectingDateMax="2009-12-08" collectingDateMin="2009-12-06" collectionCode="MHNG" collection_SpecimenCode="PAL-09/07" collectorName="A. Schulz" country="Philippines" elevation="150" location="Daylight Hole and Lions Cave" municipality="Sabang Underground NP" pageId="30" pageNumber="1719" specimenCount="1" specimenCount-male="1" stateProvince="Palawan, Puerto Princesa Region" typeStatus="holotype">
<emphasis box="[173,283,170,194]" italics="true" pageId="30" pageNumber="1719">
<typeStatus box="[173,278,170,194]" pageId="30" pageNumber="1719">Holotype</typeStatus>
.
</emphasis>
<specimenCount box="[290,311,170,194]" pageId="30" pageNumber="1719" type="male"></specimenCount>
,
<collectingCountry box="[324,453,169,193]" name="Philippines" pageId="30" pageNumber="1719" value="Philippines">Philippines</collectingCountry>
:
<collectingRegion box="[462,835,169,193]" pageId="30" pageNumber="1719">Palawan, Puerto Princesa Region</collectingRegion>
:
<collectingMunicipality box="[844,1129,169,193]" pageId="30" pageNumber="1719">Sabang Underground NP</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03A6879FA85FFFBDE595FF7B0561613A:8ED06052A85CFFBDE48FFF30036465AE" box="[173,525,201,226]" country="Philippines" municipality="Sabang Underground NP" name="Daylight Hole and Lions Cave" pageId="30" pageNumber="1719" stateProvince="Palawan, Puerto Princesa Region">Daylight Hole and Lions Cave</location>
,
<quantity box="[537,664,202,226]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="30" pageNumber="1719" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation box="[537,664,202,226]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="30" pageNumber="1719" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</elevation>
</quantity>
,
<location LSID="urn:lsid:plazi:treatment:03A6879FA85FFFBDE595FF7B0561613A:8ED06052A85CFFBDE686FF30022065AD" box="[676,841,201,225]" country="Philippines" municipality="Sabang Underground NP" name="primary forest" pageId="30" pageNumber="1719" stateProvince="Palawan, Puerto Princesa Region">primary forest</location>
,
<collectingDate box="[854,1081,201,226]" pageId="30" pageNumber="1719" value="2009-12-06" valueMax="2009-12-08" valueMin="2009-12-06">
<date box="[854,1081,201,226]" pageId="30" pageNumber="1719" value="2009-12-06" valueMax="2009-12-08" valueMin="2009-12-06">68 December 2009</date>
</collectingDate>
, leg.
<collectorName box="[173,282,233,258]" pageId="30" pageNumber="1719">A. Schulz</collectorName>
(
<collectionCode box="[297,532,234,258]" pageId="30" pageNumber="1719">
MHNG, PAL-09
<emphasis box="[489,501,234,258]" italics="true" pageId="30" pageNumber="1719">/</emphasis>
07
</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="1719" type="etymology">
<paragraph blockId="30.[173,1134,315,411]" box="[173,293,315,339]" pageId="30" pageNumber="1719">
<emphasis box="[173,293,315,339]" italics="true" pageId="30" pageNumber="1719">Etymology</emphasis>
</paragraph>
<paragraph blockId="30.[173,1134,315,411]" pageId="30" pageNumber="1719">
The specific epithet is a nominative form of an adjective (Latin:
<taxonomicName box="[925,1013,355,379]" class="Arachnida" family="Liocranidae" genus="Teutamus" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="1719" phylum="Arthropoda" rank="species" species="hirtellus">
<emphasis box="[925,1013,355,379]" italics="true" pageId="30" pageNumber="1719">hirtellus</emphasis>
</taxonomicName>
, -
<emphasis box="[1040,1053,356,379]" italics="true" pageId="30" pageNumber="1719">a</emphasis>
,-
<emphasis box="[1069,1103,356,379]" italics="true" pageId="30" pageNumber="1719">um</emphasis>
= slightly hairy), and refers to the tuft of bristles on RTA of the male palp.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="1719" type="description">
<paragraph blockId="30.[173,1134,470,979]" box="[173,300,470,493]" pageId="30" pageNumber="1719">
<emphasis box="[173,300,470,493]" italics="true" pageId="30" pageNumber="1719">Description</emphasis>
</paragraph>
<paragraph blockId="30.[173,1134,470,979]" pageId="30" pageNumber="1719">
<emphasis box="[173,231,509,533]" italics="true" pageId="30" pageNumber="1719">Male</emphasis>
(holotype). Total length 4.74; prosoma 2.26 long, 1.44 wide; opisthosoma 2.48 long, 1.26 wide. Leg formula: 1423; I 10.48 (2.36, 0.92, 3.38, 2.32, 1.50); II 8.08 (1.86, 0.90, 2.54, 1.76, 1.02); III 6.84 (1.54, 0.88, 2.00, 1.44, 0.98); IV 10.16 (2.46, 0.92, 2.88, 2.56, 1.34). Spination: Leg I: Femora p1-1; tibiae p10 r10; metatarsi p6 r6. Leg II: Tibiae p10 r8; metatarsi p6 r5. Carapace dark reddish brown; submarginal lobes blunt, slightly elevated. Sternum dark reddish brown. Legs hairless, bi-coloured: Coxae, trochanters and femora dark brown, other segments slightly paler, their distal portion white. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Venter covered by posteriorly extended post-epigastric scuta (
<figureCitation box="[181,278,795,820]" captionStart="Figure 2" captionStartId="4.[173,240,1479,1501]" captionTargetId="figure@3.[161,1010,124,1660]" captionTargetPageId="3" captionText="Figure 2. Teutamus orthogonus sp. nov., male holotype (A). Teutamus jambiensis, male paratype (MHNG) (B). Teutamus andrewdavisi, male paratype (MHNG) (C). Teutamus politus, male (D, I) from southern Thailand, female holotype (MSNG) (E). Teutamus brachiatus sp. nov., male paratype (F). Teutamus lioneli sp. nov., male paratype (G). Teutamus hirtellus sp. nov., male holotype (H). (A G) Habitus, dorsal and (H) ventral; (I) opisthosoma, ventral. Abbreviations: ES, epigastric scutum; PS, post-epigastric scutum; SM, submarginal lobe." httpUri="https://zenodo.org/record/574644/files/figure.png" pageId="30" pageNumber="1719">Figure 2</figureCitation>
H).
</paragraph>
<paragraph blockId="30.[173,1134,470,979]" pageId="30" pageNumber="1719">
Palp (
<figureCitation box="[286,401,827,852]" captionStart="Figure 14" captionStartId="29.[105,172,1518,1540]" captionTargetBox="[105,1052,713,1485]" captionTargetId="figure@29.[105,1065,711,1485]" captionTargetPageId="29" captionText="Figure 14. Teutamus hirtellus sp. nov., male holotype (A). Teutamus andrewdavisi, male paratype (MHNG) (B). Teutamus jambiensis, male paratype (MHNG) (C). (A C) Male palp, ventral. Scale bars 0.25 mm." httpUri="https://zenodo.org/record/574656/files/figure.png" pageId="30" pageNumber="1719">Figure 14</figureCitation>
A): RTA digitiform in ventral view, elongated-slender, its apex bluntly pointed, lateral margin covered with bristles. DTA short, almost indistinct. Tegulum elongate-ovoid, slightly excavated on apico-prolateral side. Conductor elongated, broad at base, its apex sharply pointed, almost reaching apex of cymbium. Embolic base clearly visible.
</paragraph>
<paragraph blockId="30.[173,388,1013,1037]" box="[173,388,1013,1037]" pageId="30" pageNumber="1719">
<emphasis box="[173,260,1013,1037]" italics="true" pageId="30" pageNumber="1719">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection box="[173,1032,1079,1143]" pageId="30" pageNumber="1719" type="distribution">
<paragraph blockId="30.[173,1032,1079,1143]" box="[173,306,1079,1103]" pageId="30" pageNumber="1719">
<emphasis box="[173,306,1079,1103]" italics="true" pageId="30" pageNumber="1719">Distribution</emphasis>
</paragraph>
<paragraph blockId="30.[173,1032,1079,1143]" box="[173,1032,1118,1143]" pageId="30" pageNumber="1719">Known only from the type locality on the island of Palawan, the Philippines.</paragraph>
</subSubSection>
</treatment>
</document>