245 lines
31 KiB
XML
245 lines
31 KiB
XML
<document id="101A3C4BF6B4577706E156BD3CBFBD6A" ID-CLB-Dataset="272041" ID-DOI="10.1093/zoolinnean/zlad013" ID-GBIF-Dataset="effa6181-8326-4c7d-8c91-809f7fabb862" ID-ISSN="0024-4082" ID-Zenodo-Dep="8432389" ID-ZooBank="C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696939870042" checkinUser="plazi" docAuthor="Silva, Felipe De Vasconcelos, Pola, Marta & Cervera, Juan Lucas" docDate="2023" docId="03A6878CFF866500FFDAFC39EC88FB39" docLanguage="en" docName="zlad013.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 199 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad013" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Marioniinae Silva 2023, subfam. nov." docType="treatment" docUuid="CAFEB7CF-064B-41A9-A25E-A3C2D9B1961B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="468" masterDocId="FF9FFFF4FF906517FC52FF84EE67FF8B" masterDocTitle="A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)" masterLastPageNumber="476" masterPageNumber="445" pageNumber="467" updateTime="1699694776904" updateUser="plazi">
|
||
<mods:mods id="52D485F6FE2562AD5111E8A273290CFF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1B8806676F0CC9F678D7A7EE24D11EB7">
|
||
<mods:title id="A3DB493EE384B97F7DD45C88AE7F52E7">A stomach plate to divide them all: a phylogenetic reassessment of the family Tritoniidae (Nudibranchia: Cladobranchia)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4339705E669A9FBE7D6E80397DECCD79" type="personal">
|
||
<mods:role id="32B4C52C4B5758800DEFBD514B4F85BC">
|
||
<mods:roleTerm id="B1DD7E8C5ADC2577BADBE3A30EADD59E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="30218CC1248CE285AD28CDBD08036048">Silva, Felipe De Vasconcelos</mods:namePart>
|
||
<mods:affiliation id="3E7D7791F561B1436E585F524F78891C">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
|
||
<mods:nameIdentifier id="F5A5BE69BDBE8C79E1981D230925FD87" type="email">felipe.devasconcelos@uca.es</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="2CDFB559F77CD1800262DDFF86310525" type="personal">
|
||
<mods:role id="56308D1F6FF7D2CDB44DBB935343652C">
|
||
<mods:roleTerm id="14AF0FA1B456F41CF3FEB91A728CA527">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AA059D89C059A5F705708D3CC5F26C65">Pola, Marta</mods:namePart>
|
||
<mods:affiliation id="287324E77A68D389CFFF9B9FFC0A80AA">Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid; Campus de Excelencia Internacional UAM + CSIC, C / Darwin, 2, 28049 Madrid, Spain</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="EEF11C0379A9FB0D74ED71F92901A808" type="personal">
|
||
<mods:role id="553706D08D5A363651520BA7929650F7">
|
||
<mods:roleTerm id="95BBB09F7EBDED6808A4B61B1CC56D02">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F4070F079141CDA3E950AFA532C85BA0">Cervera, Juan Lucas</mods:namePart>
|
||
<mods:affiliation id="D5E1B62056CEDE68F9168161FE8D1C26">Departamento de Biología, Facultad de Ciencias del Mar y Ambientales; Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain & Instituto Universitario de Investigación Marina (INMAR), Universidad de Cádiz; Campus de Excelencia Internacional del Mar (CEI · MAR), Universidad de Cádiz, Av. República Saharaui, s / n, Ap. 40. 11510 Puerto Real (Cádiz), Spain</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="062DE18A8847420E3B0EA45B3EC93ECE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A08F58DDB4B440424D9D3927123C364B" type="host">
|
||
<mods:titleInfo id="3526E16EC3B1B7AB52B578849CDFBC6E">
|
||
<mods:title id="8BE482A179FCF8E831260ACA8203F5BF">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="0F6E9D16694F5A72A1E1A32FCC9DE12B">
|
||
<mods:date id="4F76B87DF6E68A00286E133905456A0C">2023</mods:date>
|
||
<mods:detail id="3B03F6DD6E4BBD8809FC89CEE3AF2442" type="series">
|
||
<mods:title id="AB184F8C68506AF692C3D8C73F945316">Zool. J. Linn. Soc.</mods:title>
|
||
</mods:detail>
|
||
<mods:detail id="96AD0F38285F1B327DA56AFCA3565D24" type="pubDate">
|
||
<mods:number id="D0EA9DAEF47442181E4A0C2328E03885">2023-07-11</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="83123549D66EC2650217C2F71780ABF0" type="volume">
|
||
<mods:number id="ABA50B34A797E9D318C8C58D0FC07A8F">199</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CC8462E4C1813EE11991FEC67084B1C1" type="issue">
|
||
<mods:number id="9A352A7F319B6EF7C5F3B33C06E25907">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="EE98090AD72808EADFCB93230887B5CB" unit="page">
|
||
<mods:start id="798F3A9A656C7517243CEDF9CE9F2DCD">445</mods:start>
|
||
<mods:end id="AA8531DEDE3FC98C6167CC05C5829456">476</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="15CCB9C2BB5CADFCC92D68EC8BB3578B">
|
||
<mods:url id="293C3D01A7178078F767E9F048E8947F">http://dx.doi.org/10.1093/zoolinnean/zlad013</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5B8CECE0575A5127BE3E319EDC1D91C9">journal article</mods:classification>
|
||
<mods:identifier id="2250BB6ED8041873AFAB61EE9DFB81DC" type="CLB-Dataset">272041</mods:identifier>
|
||
<mods:identifier id="E29980FF5416C06F8733F5311B16B752" type="DOI">10.1093/zoolinnean/zlad013</mods:identifier>
|
||
<mods:identifier id="C0992FC3374B375E0B6F554057B7A6E8" type="GBIF-Dataset">effa6181-8326-4c7d-8c91-809f7fabb862</mods:identifier>
|
||
<mods:identifier id="AAD1ABAC53FFC6ADBAFD7FCBE5402BEE" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="3DA1B802937D56C69CD295C24F6D854C" type="Zenodo-Dep">8432389</mods:identifier>
|
||
<mods:identifier id="24506F57992D1D7ED8232B2F2DE66A76" type="ZooBank">C3E2DFF9-A0A6-41EA-A149-0F73A2BEE5E6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A6878CFF866500FFDAFC39EC88FB39" ID-DOI="http://doi.org/10.5281/zenodo.8432423" ID-GBIF-Taxon="213768865" ID-Zenodo-Dep="8432423" LSID="urn:lsid:zoobank.org:act:CAFEB7CF-064B-41A9-A25E-A3C2D9B1961B" httpUri="http://treatment.plazi.org/id/03A6878CFF866500FFDAFC39EC88FB39" lastPageId="23" lastPageNumber="468" pageId="22" pageNumber="467">
|
||
<subSubSection id="C3156511FF866501FFDAFC39EAD7FB91" pageId="22" pageNumber="467" type="nomenclature">
|
||
<paragraph id="8BB0369AFF866501FFDAFC39EB15FC5C" blockId="22.[904,1394,956,983]" box="[904,1394,956,983]" pageId="22" pageNumber="467">
|
||
<heading id="D0F881F6FF866501FFDAFC39EB15FC5C" bold="true" box="[904,1394,956,983]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="467" reason="3">
|
||
<emphasis id="B97BEA88FF866501FFDAFC39EB15FC5C" bold="true" box="[904,1394,956,983]" pageId="22" pageNumber="467">
|
||
Subfamily
|
||
<taxonomicName id="4C0F4D19FF866501FFAAFC39EA81FC5C" authority="Silva et al." authorityName="Silva" authorityYear="2023" box="[1016,1254,956,983]" class="Gastropoda" family="Tritoniidae" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Marioniinae">
|
||
Marioniinae Silva
|
||
<emphasis id="B97BEA88FF866501F8E5FC39EA81FC5C" bold="true" box="[1207,1254,956,983]" italics="true" pageId="22" pageNumber="467">et al.</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24857F3FF866501F8BDFC39EB15FC5C" box="[1263,1394,957,983]" pageId="22" pageNumber="467" rank="subFamily">subfam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF866501FF6BFC67EAD7FB91" blockId="22.[825,1474,995,1050]" pageId="22" pageNumber="467">
|
||
<emphasis id="B97BEA88FF866501FF6BFC67EA79FC70" box="[825,1054,995,1019]" italics="true" pageId="22" pageNumber="467">Zoobank registration:</emphasis>
|
||
<uri id="FF9E3A98FF866501F865FC67EACBFB91" pageId="22" pageNumber="467">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFA90C4FFF866501F966FC67EACBFB91" pageId="22" pageNumber="467">CAFEB7CF-064B-41A9-A25E-A3C2D9B1961B</uuid>
|
||
</uri>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3156511FF866500FF6BFBC6EC88FB39" lastPageId="23" lastPageNumber="468" pageId="22" pageNumber="467" type="diagnosis">
|
||
<paragraph id="8BB0369AFF866501FF6BFBC6EAAFFBD2" blockId="22.[825,1224,1089,1113]" box="[825,1224,1089,1113]" pageId="22" pageNumber="467">
|
||
<emphasis id="B97BEA88FF866501FF6BFBC6EDF8FBD2" box="[825,927,1090,1113]" italics="true" pageId="22" pageNumber="467">Diagnosis:</emphasis>
|
||
True stomach plates present.
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF866501FF6BFBFCEB39FAC0" blockId="22.[825,1474,1144,1356]" pageId="22" pageNumber="467">
|
||
<emphasis id="B97BEA88FF866501FF6BFBFCEDDFFB1B" box="[825,952,1144,1168]" italics="true" pageId="22" pageNumber="467">Morphology:</emphasis>
|
||
Notum smooth or tuberculate. Rachidian tooth uni- or tricuspid, smooth or denticulate. Masticatory border of the jaws denticulate. First lateral tooth smooth, hooked and sharp. True stomach plates present. ODG complex cladohepatic or holohepatic. Ampulla and bursa copulatrix large. Gonopore at one-third of body length, furthest from rhinophores. Anus at middle of body length; nephroproct oħen above anus.
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF866501FF6BFAF2EDD1FA27" blockId="22.[825,1474,1397,1452]" pageId="22" pageNumber="467">
|
||
<emphasis id="B97BEA88FF866501FF6BFAF2EA9EFA06" box="[825,1273,1397,1421]" italics="true" pageId="22" pageNumber="467">
|
||
Genera included in this study:
|
||
<taxonomicName id="4C0F4D19FF866501F8CFFAF2EA9EFA06" authorityName="Vayssiere" authorityYear="1877" box="[1181,1273,1398,1421]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="genus">Marionia</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF866501F95CFAF2EBEAFA06" authorityName="Odhner" authorityYear="1934" box="[1294,1421,1398,1421]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF866501F95CFAF2EBEAFA06" box="[1294,1421,1398,1421]" italics="true" pageId="22" pageNumber="467">Marioniopsis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF866501FF6BFA11EDD7FA27" authorityName="Baba" authorityYear="1949" box="[825,944,1429,1452]" class="Gastropoda" family="Tritoniidae" genus="Paratritonia" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF866501FF6BFA11EDD7FA27" box="[825,944,1429,1452]" italics="true" pageId="22" pageNumber="467">Paratritonia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF866501FF6BFA4FEB0CF832" blockId="22.[825,1474,1483,1977]" pageId="22" pageNumber="467">
|
||
<emphasis id="B97BEA88FF866501FF6BFA4FEDF0FA68" box="[825,919,1483,1507]" italics="true" pageId="22" pageNumber="467">Remarks:</emphasis>
|
||
Based on our integrative approach, the plate-bearing tritoniids are a separate and morphologically distinct clade from the plateless species (
|
||
<figureCitation id="13342A1FFF866501F84DF98EEA3DF9A9" box="[1055,1114,1546,1570]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="22" pageNumber="467">Fig. 2</figureCitation>
|
||
). This group is raised to the subfamily level as the monophyletic
|
||
<taxonomicName id="4C0F4D19FF866501F8D8F9ADEB6FF9CA" box="[1162,1288,1577,1601]" class="Gastropoda" family="Tritoniidae" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="subFamily" subFamily="Marioniinae">Marioniinae</taxonomicName>
|
||
and characterized by the presence of true stomach plates. The true stomach plates of marioniids are thick, cuticular structures aưached to a belt or girdle and are oħen found near the openings between the digestive gland and the stomach. Gosliner (1994) suggested the presence of stomach plates in
|
||
<taxonomicName id="4C0F4D19FF866501F835F942EA99F955" authorityName="Willan and Morton" authorityYear="1984" box="[1127,1278,1734,1758]" class="Gastropoda" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="456" phylum="Mollusca" rank="subOrder" subOrder="Cladobranchia">Cladobranchia</taxonomicName>
|
||
as a plesiomorphic condition, as they are common in this group. Similar cuticular structures are also found, for example, in the family
|
||
<taxonomicName id="4C0F4D19FF866501F91DF880EBA5F897" authorityName="Rafinesque" authorityYear="1815" box="[1359,1474,1796,1820]" class="Gastropoda" family="Tethydidae" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="family">Tethydidae</taxonomicName>
|
||
and in some genera of
|
||
<taxonomicName id="4C0F4D19FF866501F860F8A0EAACF8B7" authorityName="Willan and Morton" authorityYear="1984" box="[1074,1227,1828,1852]" class="Gastropoda" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="456" phylum="Mollusca" rank="subOrder" subOrder="Cladobranchia">Cladobranchia</taxonomicName>
|
||
, e.g.
|
||
<taxonomicName id="4C0F4D19FF866501F956F8A0EBA5F8B7" authority="Rang 1829" authorityName="Rang" authorityYear="1829" box="[1284,1474,1828,1852]" class="Gastropoda" family="Tethydidae" genus="Melibe" kingdom="Animalia" order="Nudibranchia" pageId="22" pageNumber="467" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF866501F956F8A0EB22F8B7" box="[1284,1349,1828,1852]" italics="true" pageId="22" pageNumber="467">Melibe</emphasis>
|
||
Rang 1829
|
||
</taxonomicName>
|
||
(Rudman 1998,
|
||
<bibRefCitation id="EF9E4B6BFF866501FFB2F8C7EAAAF8D0" author="Gosliner T & Pola M" box="[992,1229,1859,1883]" pageId="22" pageNumber="467" pagination="333 - 349" refId="ref25265" refString="Gosliner T, Pola M. Diversification of filter-feeding nudibranchs: two remarkable new species of Melibe (Opisthobranchia: Tethyiidae) from the tropical western Pacific. Systematics and Biodiversity 2012; 10: 333 - 349." type="journal article" year="2012">Gosliner and Pola 2012</bibRefCitation>
|
||
). However, we consider the presence of true stomach plates in the marioniids to be an apomorphic condition, consistent with the basal position of the
|
||
<taxonomicName id="4C0F4D19FF866501FF6BF825ED89F832" baseAuthorityName="Korshunova and Martynov" baseAuthorityYear="2020" box="[825,1006,1953,1977]" class="Gastropoda" family="Pleurobranchidae" kingdom="Animalia" order="Pleurobranchida" pageId="22" pageNumber="467" phylum="Mollusca" rank="family">Pleurobranchidae</taxonomicName>
|
||
in our phylogenetic analyses (
|
||
<figureCitation id="13342A1FFF866501F971F825EB3CF832" box="[1315,1371,1953,1977]" captionStart="Figure 2" captionStartId="10.[129,194,1719,1743]" captionTargetBox="[325,1278,148,1689]" captionTargetId="figure-6@10.[321,1281,144,1692]" captionTargetPageId="10" captionText="Figure 2. Bayesian phylogenetic tree based on the concatenated molecular data (COI + 16S + H3) of Tritoniidae (sensu Odhner 1963). Outgroup taxa are depicted in grey.Tritoniidae genera are independently coloured, and branches with several replicates for each species hypothesis were collapsed for clarity.Names in bold are species which sequences were generated for this study with the number of sequenced specimens showed in brackets. PP are shown right/top and BS leħ/boưom.Unsupported branches are not labelled. Green circles indicate maximum support by both analyses. Solid vertical bars indicate specimens grouped as species entities based on COI dataset. Empty vertical bars indicate the specimens grouped as species based on the 16S dataset." figureDoi="http://doi.org/10.5281/zenodo.8432395" httpUri="https://zenodo.org/record/8432395/files/figure.png" pageId="22" pageNumber="467">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF876500FCDFFF14ECB0FE08" blockId="23.[113,763,144,1202]" pageId="23" pageNumber="468">
|
||
Marioniidsaredividedintothreegenera:
|
||
<taxonomicName id="4C0F4D19FF876500FE4EFF15EC11FF23" authorityName="Vayssiere" authorityYear="1877" box="[540,630,145,168]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FE4EFF15EC11FF23" box="[540,630,145,168]" italics="true" pageId="23" pageNumber="468">Marionia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F4D19FF876500FE2CFF15EC9DFF23" authorityName="Odhner" authorityYear="1934" box="[638,762,145,168]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FE2CFF15EC9DFF23" box="[638,762,145,168]" italics="true" pageId="23" pageNumber="468">Marioniopsis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F4D19FF876500FCCCFF34EF75FF4C" authorityName="Baba" authorityYear="1949" box="[158,274,176,199]" class="Gastropoda" family="Tritoniidae" genus="Paratritonia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FCCCFF34EF75FF4C" box="[158,274,176,199]" italics="true" pageId="23" pageNumber="468">Paratritonia</emphasis>
|
||
</taxonomicName>
|
||
. They may be distinguished from each other by the rachidian tooth morphology (tricuspid smooth in
|
||
<taxonomicName id="4C0F4D19FF876500FEC9FF4BEC92FF6D" authorityName="Vayssiere" authorityYear="1877" box="[667,757,207,230]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FEC9FF4BEC92FF6D" box="[667,757,207,230]" italics="true" pageId="23" pageNumber="468">Marionia</emphasis>
|
||
</taxonomicName>
|
||
, tricuspid denticulate in
|
||
<taxonomicName id="4C0F4D19FF876500FD0FFF6BEFBEFE8D" authorityName="Odhner" authorityYear="1934" box="[349,473,239,262]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FD0FFF6BEFBEFE8D" box="[349,473,239,262]" italics="true" pageId="23" pageNumber="468">Marioniopsis</emphasis>
|
||
</taxonomicName>
|
||
and unicuspid denticulate in
|
||
<taxonomicName id="4C0F4D19FF876500FC23FE8AEE82FEAE" authorityName="Baba" authorityYear="1949" box="[113,229,270,293]" class="Gastropoda" family="Tritoniidae" genus="Paratritonia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FC23FE8AEE82FEAE" box="[113,229,270,293]" italics="true" pageId="23" pageNumber="468">Paratritonia</emphasis>
|
||
</taxonomicName>
|
||
), masticatory borders of the jaw (denticles are sharp and triangular or rounded in
|
||
<taxonomicName id="4C0F4D19FF876500FDC3FEA9EF8CFECF" authorityName="Vayssiere" authorityYear="1877" box="[401,491,301,324]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FDC3FEA9EF8CFECF" box="[401,491,301,324]" italics="true" pageId="23" pageNumber="468">Marionia</emphasis>
|
||
</taxonomicName>
|
||
, blunt and rounded or rod-shaped in
|
||
<taxonomicName id="4C0F4D19FF876500FC89FEC9EF31FEEF" authorityName="Odhner" authorityYear="1934" box="[219,342,333,356]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FC89FEC9EF31FEEF" box="[219,342,333,356]" italics="true" pageId="23" pageNumber="468">Marioniopsis</emphasis>
|
||
</taxonomicName>
|
||
, and ‘faint scales’ in
|
||
<taxonomicName id="4C0F4D19FF876500FE79FEC9ECF8FEEF" authorityName="Baba" authorityYear="1949" box="[555,671,333,356]" class="Gastropoda" family="Tritoniidae" genus="Paratritonia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FE79FEC9ECF8FEEF" box="[555,671,333,356]" italics="true" pageId="23" pageNumber="468">Paratritonia</emphasis>
|
||
</taxonomicName>
|
||
) and digestive system arrangement (see remarks for
|
||
<taxonomicName id="4C0F4D19FF876500FE7EFEE8ECE1FE08" authorityName="Vayssiere" authorityYear="1877" box="[556,646,364,387]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BEA88FF876500FE7EFEE8ECE1FE08" box="[556,646,364,387]" italics="true" pageId="23" pageNumber="468">Marionia</emphasis>
|
||
</taxonomicName>
|
||
ahead).
|
||
</paragraph>
|
||
<paragraph id="8BB0369AFF876500FCDFFE0FEC88FB39" blockId="23.[113,763,144,1202]" pageId="23" pageNumber="468">
|
||
The taxonomic status of Indo-Pacific marioniids is in a remarkable state of taxonomic confusion resulting from (1) incomplete descriptions, (2) numerous sequenced unnamed species, (3) numerous specimens identified based on external morphology, (4) lack of morpho-anatomical studies that would help link names to living specimens and (5) pseudocryptic speciation that misleads taxonomy based on morphology. This has led to several controversies about the taxonomic validity of some species included in this study. For example, a literature search revealed that
|
||
<taxonomicName id="4C0F4D19FF876500FC23FD21EF3DFD37" authorityName="Bergh" authorityYear="1890" box="[113,346,676,700]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="species" species="arborescens">
|
||
<emphasis id="B97BEA88FF876500FC23FD21EF3DFD37" box="[113,346,676,700]" italics="true" pageId="23" pageNumber="468">Marioniopsis arborescens</emphasis>
|
||
</taxonomicName>
|
||
has been used in the last two decades for some marioniids identified solely on the basis of external morphology (
|
||
<bibRefCitation id="EF9E4B6BFF876500FCEBFD67EF1DFD70" author="Gosliner TM & Behrens DW & Valdes A" box="[185,378,739,763]" pageId="23" pageNumber="468" refId="ref25304" refString="Gosliner TM, Behrens DW, Valdes A. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. San Francisco: California Academy of Sciences / Sea Challengers Natural History Books, 2008." type="book" year="2008">
|
||
Gosliner
|
||
<emphasis id="B97BEA88FF876500FD46FD60EF5BFD70" box="[276,316,739,763]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
, 2015, Hullet
|
||
<emphasis id="B97BEA88FF876500FE5AFD60EC57FD70" box="[520,560,739,763]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2015,
|
||
<bibRefCitation id="EF9E4B6BFF876500FE2BFD67EEC4FC90" author="Tibirica Y & Pola M & Cervera JL" pageId="23" pageNumber="468" pagination="1 - 133" refId="ref27808" refString="Tibirica Y, Pola M, Cervera JL. Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique. Zootaxa 2017; 4359: 1 - 133." type="journal article" year="2017">
|
||
Tibiriçá
|
||
<emphasis id="B97BEA88FF876500FE9FFD60EC93FD70" box="[717,756,739,763]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
). However,
|
||
<taxonomicName id="4C0F4D19FF876500FD70FC87EC6BFC91" authorityName="Bergh" authorityYear="1890" box="[290,524,770,794]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="species" species="arborescens">
|
||
<emphasis id="B97BEA88FF876500FD70FC87EC6BFC91" box="[290,524,770,794]" italics="true" pageId="23" pageNumber="468">Marioniopsis arborescens</emphasis>
|
||
</taxonomicName>
|
||
was synonymized with
|
||
<taxonomicName id="4C0F4D19FF876500FC23FCA7EFE0FCB1" baseAuthorityName="Ruppell and Leuckart" baseAuthorityYear="1828" box="[113,391,802,826]" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="species" species="cyanobranchiata">
|
||
<emphasis id="B97BEA88FF876500FC23FCA7EFE0FCB1" box="[113,391,802,826]" italics="true" pageId="23" pageNumber="468">Marioniopsis cyanobranchiata</emphasis>
|
||
</taxonomicName>
|
||
based on internal and external morphology long ago (
|
||
<bibRefCitation id="EF9E4B6BFF876500FD5CFCC5EFF3FCD2" author="Odhner NH" box="[270,404,833,857]" pageId="23" pageNumber="468" refId="ref26584" refString="Odhner NH. Nudibranchia dendronotacea: a revision of the system. Brussels: Musee Royal d'Histoire Naturelle de Belgique, 1936." type="book" year="1936">Odhner 1936</bibRefCitation>
|
||
), and since then no internal anatomical evidence has been presented to support the resurrection of the species. Similarly, the name
|
||
<taxonomicName id="4C0F4D19FF876500FD2DFC04ECC7FC13" authority="Bergh 1905" authorityName="Bergh" authorityYear="1905" box="[383,672,896,920]" class="Gastropoda" family="Tritoniidae" genus="Marionia" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="species" species="distincta">
|
||
<emphasis id="B97BEA88FF876500FD2DFC04EC4DFC13" box="[383,554,896,920]" italics="true" pageId="23" pageNumber="468">Marionia distincta</emphasis>
|
||
Bergh 1905
|
||
</taxonomicName>
|
||
has been used to identify marioniids on the basis of external morphology (
|
||
<bibRefCitation id="EF9E4B6BFF876500FC29FC3AEF5EFC5C" author="Gosliner TM & Behrens DW & Valdes A" box="[123,313,958,983]" pageId="23" pageNumber="468" refId="ref25304" refString="Gosliner TM, Behrens DW, Valdes A. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. San Francisco: California Academy of Sciences / Sea Challengers Natural History Books, 2008." type="book" year="2008">
|
||
Gosliner
|
||
<emphasis id="B97BEA88FF876500FC87FC3BEE9BFC5D" box="[213,252,958,982]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
, 2015;
|
||
<bibRefCitation id="EF9E4B6BFF876500FDD0FC3AEC4DFC5D" author="Huleu RE & Mahguib J & Gosliner TM" box="[386,554,958,982]" pageId="23" pageNumber="468" pagination="215 - 222" refId="ref25435" refString="Huleu RE, Mahguib J, Gosliner TM, et al. Molecular evaluation of the phylogenetic position of the enigmatic species Trivettea papalotla (Bertsch) (Mollusca: Nudibranchia). Invertebrate Systematics 2015; 29: 215 - 222." type="journal article" year="2015">
|
||
Huleư
|
||
<emphasis id="B97BEA88FF876500FD94FC3BEF8AFC5D" box="[454,493,958,982]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9E4B6BFF876500FE67FC3AEC8DFC5C" author="Tibirica Y & Pola M & Cervera JL" box="[565,746,958,983]" pageId="23" pageNumber="468" pagination="1 - 133" refId="ref27808" refString="Tibirica Y, Pola M, Cervera JL. Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique. Zootaxa 2017; 4359: 1 - 133." type="journal article" year="2017">
|
||
Tibiriçá
|
||
<emphasis id="B97BEA88FF876500FED5FC3BECCAFC5D" box="[647,685,958,982]" italics="true" pageId="23" pageNumber="468">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
), despite the established synonymy with
|
||
<taxonomicName id="4C0F4D19FF876500FDA7FC5AEEC9FB9E" authority="(Odhner 1936)" baseAuthorityName="Odhner" baseAuthorityYear="1936" class="Gastropoda" family="Tritoniidae" genus="Marioniopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="23" pageNumber="468" phylum="Mollusca" rank="species" species="levis">
|
||
<emphasis id="B97BEA88FF876500FDA7FC5AECFCFC7D" box="[501,667,990,1014]" italics="true" pageId="23" pageNumber="468">Marioniopsis levis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9E4B6BFF876500FEF9FC5AEEC4FB9E" author="Odhner NH" pageId="23" pageNumber="468" refId="ref26584" refString="Odhner NH. Nudibranchia dendronotacea: a revision of the system. Brussels: Musee Royal d'Histoire Naturelle de Belgique, 1936." type="book" year="1936">Odhner 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. These inconsistencies cause great confusion in the identification of these species. Because a broad taxonomic reassessment of the Indo-Pacific marioniids is beyond the scope of the present study, we have retained the original identification of the included marioniids from the Indo-Pacific region (
|
||
<tableCitation id="C68D0321FF876500FE57FBFEEC2BFB19" box="[517,588,1146,1170]" captionStart="Table 1" captionStartId="3.[110,165,401,426]" captionTargetBox="[110,1937,615,1719]" captionText="Table 1. List of specimens used for phylogenetic and morphological analyses. Accession numbers of the new sequences generated for this study are in bold.Museums abbreviations:California Academy of Sciences, CASIZ (San Francisco, USA); Museo de Zoologia, Universidad de Costa Rica, MZUCR (San José, Costa Rica); Museu de Zoologia da Universidade de São Paulo, MZUSP (São Paulo, Brazil); Museu Nacional da Universidade do Rio de Janeiro, MNRJ (Rio de Janeiro, Brazil); Natural History Museum of Los Angeles County, LACM (Los Angeles, USA); Zoological Museum of Moscow University, ZMMU (Moscow, Russia); University Museum of Bergen, ZMBN (Bergen, Norway); Museo Nacional de Ciencias Naturales, MNCN (Madrid, Spain); Museu Nacional de História e da Ciência de Lisboa, MNHNC (Lisbon, Portugal); Museu de História Natural de Maputo, MHN (Maputo, Mozambique); California State Polytechnic University, CPIC (California; USA).Geographic distribution abbreviations: NAO: North Atlantic Ocean; SAO: South Atlantic Ocean; NPO: North Pacific Ocean; SPO: South Pacific Ocean; SO, Southern Ocean; IO: Indian Ocean; M: Mediterranean Sea" httpUri="http://table.plazi.org/id/DF706612FF936514FC3CFE15EC59FDDA" pageId="23" pageNumber="468" tableUuid="DF706612FF936514FC3CFE15EC59FDDA">Table 1</tableCitation>
|
||
) and will address the taxonomy of our sequenced material in detail in future studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |