treatments-xml/data/03/A0/87/03A087FA231EFF8AFF3D615AFDF0B82B.xml
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<document id="D172FF6992BBF9D46CBB574CAE4BE9E9" ID-CLB-Dataset="9218" ID-DOI="10.11646/zootaxa.4896.2.7" ID-GBIF-Dataset="7fbdd6a3-8010-4616-965d-241515af8423" ID-ISSN="1175-5326" ID-Zenodo-Dep="4381842" ID-ZooBank="71E90DA6-7595-4010-A849-0EBA20D15921" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608543618368" checkinUser="plazi" docAuthor="Bota-Sierra, Cornelio Andrés &amp; Novelo-Gutiérrez, Rodolfo" docDate="2020" docId="03A087FA231EFF8AFF3D615AFDF0B82B" docLanguage="en" docName="zootaxa.4896.2.7.pdf" docOrigin="Zootaxa 4896 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigomphus rufus Bota-Sierra &amp; Novelo-Gutiérrez 2020, sp.nov." docType="treatment" docVersion="10" lastPageNumber="271" masterDocId="FF99FF82231FFF8CFFAA6245FF94BE1D" masterDocTitle="Two new species of Colombian Epigomphus (Odonata: Gomphidae)" masterLastPageNumber="276" masterPageNumber="265" pageNumber="266" updateTime="1698847844051" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6D604B280C6DE533ECDCBAF4FB8D56E9">Two new species of Colombian Epigomphus (Odonata: Gomphidae)</mods:title>
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<mods:name id="43C316EF2F27CD9DC1D177B26731A527" type="personal">
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<mods:roleTerm id="328DC79E2A92DF32C0951B59E20796A6">Author</mods:roleTerm>
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<mods:namePart id="0B9B812ED96B9D1FC76408C5ED1010F8">Bota-Sierra, Cornelio Andrés</mods:namePart>
<mods:affiliation id="70EC8AE36AF586C97D5FA317FEAAE0AF">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera Antigua a Coatepec 351, El Haya, 91070 Xalapa, Veracruz, MEXICO. &amp; Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellín, Antioquia, COLOMBIA.</mods:affiliation>
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<mods:namePart id="38F0BDB24519B44F2C47DDD7E88028E2">Novelo-Gutiérrez, Rodolfo</mods:namePart>
<mods:affiliation id="96195D7564A942E15AF33ACE93EFA7F8">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera Antigua a Coatepec 351, El Haya, 91070 Xalapa, Veracruz, MEXICO. &amp; rodolfo. novelo @ inecol. mx; http: // orcid. org / 0000 - 0003 - 3163 - 2820</mods:affiliation>
<mods:nameIdentifier id="FD6370C5CECE9BE3064090FEE2E309D1" type="email">rodolfo.novelo@inecol.mx</mods:nameIdentifier>
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<mods:title id="F9485EBB5347673A219AD9CC96C6A42B">Zootaxa</mods:title>
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<mods:date id="81C705FC3DAFDE7ACC03F95192381619">2020</mods:date>
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<mods:number id="9F4A3F0F4800D49858B4E8047CAEF797">2020-12-21</mods:number>
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<treatment id="03A087FA231EFF8AFF3D615AFDF0B82B" ID-DOI="http://doi.org/10.5281/zenodo.4382787" ID-GBIF-Taxon="176234987" ID-Zenodo-Dep="4382787" LSID="urn:lsid:plazi:treatment:03A087FA231EFF8AFF3D615AFDF0B82B" httpUri="http://treatment.plazi.org/id/03A087FA231EFF8AFF3D615AFDF0B82B" lastPageId="6" lastPageNumber="271" pageId="1" pageNumber="266">
<subSubSection id="C3136567231EFF8DFF3D615AFE42BD24" box="[151,470,799,826]" pageId="1" pageNumber="266" type="nomenclature">
<paragraph id="8BB636EC231EFF8DFF3D615AFE42BD24" blockId="1.[151,470,799,861]" box="[151,470,799,826]" pageId="1" pageNumber="266">
<heading id="D0FE8180231EFF8DFF3D615AFE42BD24" bold="true" box="[151,470,799,826]" fontSize="11" level="1" pageId="1" pageNumber="266" reason="1">
<emphasis id="B97DEAFE231EFF8DFF3D615AFE42BD24" bold="true" box="[151,470,799,826]" pageId="1" pageNumber="266">
<taxonomicName id="4C094D6F231EFF8DFF3D615AFEEDBD24" ID-CoL="9X2MN" authority="Bota-Sierra &amp; Novelo-Gutiérrez, 2020" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[151,377,799,826]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
<emphasis id="B97DEAFE231EFF8DFF3D615AFEEDBD24" bold="true" box="[151,377,799,826]" italics="true" pageId="1" pageNumber="266">Epigomphus rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E5785231EFF8DFE2A615AFE42BD24" box="[384,470,799,825]" pageId="1" pageNumber="266" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3136567231EFF8DFF3D6106FE86BD40" box="[151,274,835,861]" pageId="1" pageNumber="266" type="description">
<paragraph id="8BB636EC231EFF8DFF3D6106FE86BD40" blockId="1.[151,470,799,861]" box="[151,274,835,861]" pageId="1" pageNumber="266">
(
<figureCitation id="13322A69231EFF8DFF356106FE9EBD40" box="[159,266,835,861]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1760,1784]" captionStartId-1="2.[151,250,1700,1724]" captionStartId-2="3.[151,250,1887,1911]" captionStartId-3="4.[151,250,1872,1896]" captionTargetBox-0="[153,1432,889,1735]" captionTargetBox-1="[224,1364,534,1677]" captionTargetBox-2="[257,1331,561,1864]" captionTargetBox-3="[148,1439,525,1834]" captionTargetId-0="figure-209@1.[151,1436,885,1735]" captionTargetId-1="figure-173@2.[224,1364,525,1677]" captionTargetId-2="figure-226@3.[257,1331,561,1864]" captionTargetId-3="figure-190@4.[148,1439,525,1849]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." captionText-1="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." captionText-2="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." captionText-3="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi-0="http://doi.org/10.5281/zenodo.4381846" figureDoi-1="http://doi.org/10.5281/zenodo.4381850" figureDoi-2="http://doi.org/10.5281/zenodo.4381854" figureDoi-3="http://doi.org/10.5281/zenodo.4381858" httpUri-0="https://zenodo.org/record/4381846/files/figure.png" httpUri-1="https://zenodo.org/record/4381850/files/figure.png" httpUri-2="https://zenodo.org/record/4381854/files/figure.png" httpUri-3="https://zenodo.org/record/4381858/files/figure.png" pageId="1" pageNumber="266">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF766664231EFF8DFF3D64A5FBBEB901" ID-DOI="http://doi.org/10.5281/zenodo.4381846" ID-Zenodo-Dep="4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="1" pageNumber="266" startId="1.[151,250,1760,1784]" targetBox="[153,1432,889,1735]" targetPageId="1">
<paragraph id="8BB636EC231EFF8DFF3D64A5FBBEB901" blockId="1.[151,1436,1760,1820]" pageId="1" pageNumber="266">
<emphasis id="B97DEAFE231EFF8DFF3D64A5FE86B8E5" bold="true" box="[151,274,1760,1784]" pageId="1" pageNumber="266">FIGURE 1.</emphasis>
<taxonomicName id="4C094D6F231EFF8DFEBD64A5FE5BB8E5" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[279,463,1760,1784]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="266" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231EFF8DFEBD64A5FE5BB8E5" box="[279,463,1760,1784]" italics="true" pageId="1" pageNumber="266">Epigomphus rufus</emphasis>
</taxonomicName>
head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge.
</paragraph>
</caption>
<subSubSection id="C3136567231EFF8EFF3D650FFB91BF00" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266" type="materials_examined">
<paragraph id="8BB636EC231EFF8EFF3D650FFB91BF00" blockId="1.[151,1437,1866,2037]" lastBlockId="2.[151,1437,150,502]" lastPageId="2" lastPageNumber="267" pageId="1" pageNumber="266">
<emphasis id="B97DEAFE231EFF8DFF3D650FFE12B979" bold="true" box="[151,390,1866,1892]" pageId="1" pageNumber="266">Material examined:</emphasis>
Three specimens were collected (1♂,
<specimenCount id="9D0FFD65231EFF8DFCEE650EFCE9B978" box="[836,893,1867,1893]" pageId="1" pageNumber="266" type="female">2♀♀</specimenCount>
), we assigned them as conspecific due to their share of the same habitat in the same forest patch, their conspicuous coloration, and mainly the correspondence of male cerci and epiproct with the posterior portion of the female head. All three specimens are teneral, and the two females were reared from the last larval instar to the adult stage (one of them was kept alive and feed with small insects for five days, which allowed the coloration pattern to get very close to the mature pattern).
<collectionCode id="ED18AE29231EFF8DFB65659EFAB4B9E8" box="[1231,1312,2011,2037]" pageId="1" pageNumber="266">CEUA</collectionCode>
:
<materialsCitation id="3B613CB1231EFF8EFA82659EFA82BEC8" ID-GBIF-Occurrence="3003801303" collectingDate="2012-05-27" collectionCode="R" collectorName="Leg &amp; C. Bota." elevation="1644" lastPageId="2" lastPageNumber="267" latitude="-75.13079" location="Arrierito Antioqueno Reserve" pageId="1" pageNumber="266" specimenCode="CEUA 113490, CEUA 99305" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B97DEAFE231EFF8DFA82659EFA01B9E8" bold="true" box="[1320,1429,2011,2037]" pageId="1" pageNumber="266">
<typeStatus id="54B2884E231EFF8DFA82659EFA01B9E8" box="[1320,1429,2011,2037]" pageId="1" pageNumber="266" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D0FFD65231DFF8EFF3D62D2FF38BEAC" box="[151,172,151,177]" pageId="2" pageNumber="267" type="female"></specimenCount>
teneral,
<collectionCode id="ED18AE29231DFF8EFEA662D2FEC3BEAC" box="[268,343,151,177]" pageId="2" pageNumber="267">CEUA</collectionCode>
113500,
<date id="FFB7102C231DFF8EFE1462D2FD12BEAC" box="[446,646,151,177]" pageId="2" pageNumber="267" value="2019-02-19">19 February 2019</date>
,
<collectingCountry id="F31E767C231DFF8EFD3B62D2FCBDBEAC" box="[657,809,151,177]" name="Colombia" pageId="2" pageNumber="267">COLOMBIA</collectingCountry>
,
<collectingRegion id="49CDF80E231DFF8EFC9862D2FBBBBEAC" box="[818,1071,151,177]" country="Colombia" name="Antioquia" pageId="2" pageNumber="267">Antioquia Department</collectingRegion>
, La Forzoza Reserve, Anorí Municipality, El Roble Township,
<geoCoordinate id="EE3D502B231DFF8EFE5262FEFDF7BEC8" box="[504,611,187,213]" pageId="2" pageNumber="267">7.001010</geoCoordinate>
,
<geoCoordinate id="EE3D502B231DFF8EFDDA62FFFD60BEC8" box="[624,756,186,213]" degrees="75.130792" direction="south" orientation="latitude" pageId="2" pageNumber="267" precision="1" value="-75.13079">-75.130792</geoCoordinate>
;
<elevation id="0024D1DF231DFF8EFCAB62FEFC08BEC8" box="[769,924,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.644" pageId="2" pageNumber="267" unit="m" value="1644.0">
<quantity id="4CF19B09231DFF8EFCAB62FEFCCABEC8" box="[769,862,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.644" pageId="2" pageNumber="267" unit="m" value="1644.0">1,644 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="26FC533A231DFF8EFC0062FEFC4EBEC8" box="[938,986,187,213]" pageId="2" pageNumber="267">Leg</collectorName>
:
<collectionCode id="ED18AE29231DFF8EFC4F62FEFC6FBEC8" box="[997,1019,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="267">R</collectionCode>
. Novelo-G. and
<collectorName id="26FC533A231DFF8EFB1D62FEFA82BEC8" box="[1207,1302,187,213]" pageId="2" pageNumber="267">C. Bota.</collectorName>
</materialsCitation>
<materialsCitation id="3B613CB1231DFF8EFAB762FEFB91BF00" ID-GBIF-Occurrence="3003801305" collectingDate="2012-05-27" collectionCode="R" collectorName="Leg &amp; C. Bota." elevation="1644" latitude="-75.13079" location="Arrierito Antioqueno Reserve" pageId="2" pageNumber="267" specimenCode="CEUA 113490, CEUA 99305" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B97DEAFE231DFF8EFAB762FEFA01BEC8" bold="true" box="[1309,1429,187,213]" pageId="2" pageNumber="267">
<typeStatus id="54B2884E231DFF8EFAB762FEFA01BEC8" box="[1309,1429,187,213]" pageId="2" pageNumber="267" type="paratype">Paratypes</typeStatus>
</emphasis>
,
<specimenCount id="9D0FFD65231DFF8EFF3D629AFF2EBEE4" box="[151,186,223,249]" pageId="2" pageNumber="267" type="female">1♀</specimenCount>
, teneral, same as
<typeStatus id="54B2884E231DFF8EFE2F629AFE73BEE4" box="[389,487,223,249]" pageId="2" pageNumber="267" type="holotype">holotype</typeStatus>
but
<specimenCode id="DBAF9E97231DFF8EFDB0629AFD50BEE4" box="[538,708,223,249]" collectionCode="CEUA" pageId="2" pageNumber="267">CEUA 113490</specimenCode>
. 1 ♂, teneral,
<specimenCode id="DBAF9E97231DFF8EFCCE629AFC6BBEE4" box="[868,1023,222,249]" collectionCode="CEUA" pageId="2" pageNumber="267">CEUA 99305</specimenCode>
,
<date id="FFB7102C231DFF8EFBA1629AFB31BEE4" box="[1035,1189,223,249]" pageId="2" pageNumber="267" value="2012-05-27">
<collectingDate id="EFF3E9C4231DFF8EFBA1629AFB31BEE4" box="[1035,1189,223,249]" pageId="2" pageNumber="267" value="2012-05-27">27 May 2012</collectingDate>
</date>
,
<location id="8ED66037231DFF8EFB1A629AFF60BF00" LSID="urn:lsid:plazi:treatment:03A087FA231EFF8AFF3D615AFDF0B82B:8ED66037231DFF8EFB1A629AFF60BF00" latitude="-75.13079" name="Arrierito Antioqueno Reserve" pageId="2" pageNumber="267">Arrierito Antioqueño Reserve</location>
,
<geoCoordinate id="EE3D502B231DFF8EFF556346FEFFBF00" box="[255,363,259,285]" pageId="2" pageNumber="267">6.982318</geoCoordinate>
,
<geoCoordinate id="EE3D502B231DFF8EFEDC6347FE6DBF00" box="[374,505,258,285]" degrees="75.111666" direction="south" orientation="latitude" pageId="2" pageNumber="267" precision="1" value="-75.111664">-75.111666</geoCoordinate>
;
<quantity id="4CF19B09231DFF8EFDA96346FDCBBF00" box="[515,607,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.6869999999999998" pageId="2" pageNumber="267" unit="m" value="1687.0">1,687 m</quantity>
a.s.l., Leg: J.A. Cogollo and C. Bota.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3136567231DFF8EFF6D6362FF56BF94" pageId="2" pageNumber="267" type="etymology">
<paragraph id="8BB636EC231DFF8EFF6D6362FF56BF94" blockId="2.[151,1437,150,502]" pageId="2" pageNumber="267">
<emphasis id="B97DEAFE231DFF8EFF6D6362FEC6BF5C" bold="true" box="[199,338,295,321]" pageId="2" pageNumber="267">Etymology:</emphasis>
The color of this species is unique among the species in the genus, since its dark ground coloration is reddish-brown instead of the common dark brown or black. Thus we named it
<taxonomicName id="4C094D6F231DFF8EFB91630EFBE1BF78" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[1083,1141,331,357]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="267" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231DFF8EFB91630EFBE1BF78" box="[1083,1141,331,357]" italics="true" pageId="2" pageNumber="267">rufus</emphasis>
</taxonomicName>
from Latin, which means red.
</paragraph>
</subSubSection>
<subSubSection id="C3136567231DFF89FF6D63D7FCB5BCD0" lastPageId="5" lastPageNumber="270" pageId="2" pageNumber="267" type="description">
<paragraph id="8BB636EC231DFF8EFF6D63D7FB1CBFEB" blockId="2.[151,1437,150,502]" pageId="2" pageNumber="267">
<emphasis id="B97DEAFE231DFF8EFF6D63D7FE76BFB0" bold="true" box="[199,482,402,429]" pageId="2" pageNumber="267">
Description of
<typeStatus id="54B2884E231DFF8EFED163D6FE76BFB0" box="[379,482,403,429]" pageId="2" pageNumber="267" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="B97DEAFE231DFF8EFE5863D6FDBBBFB0" box="[498,559,403,429]" italics="true" pageId="2" pageNumber="267">Head</emphasis>
reddish-brown, eyes when alive greenish-gray. Posterior ocelli over a tumid area, bordered by a C-shaped thick rim (
<figureCitation id="13322A69231DFF8EFDF263F2FD59BFCC" box="[600,717,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1760,1784]" captionTargetBox="[153,1432,889,1735]" captionTargetId="figure-209@1.[151,1436,885,1735]" captionTargetPageId="1" captionText="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." figureDoi="http://doi.org/10.5281/zenodo.4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="2" pageNumber="267">Figs. 1a, c</figureCitation>
); occipital posterior border sinuous with a small medial concave incision (
<figureCitation id="13322A69231DFF8EFF55639EFEDBBFE8" box="[255,335,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1760,1784]" captionTargetBox="[153,1432,889,1735]" captionTargetId="figure-209@1.[151,1436,885,1735]" captionTargetPageId="1" captionText="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." figureDoi="http://doi.org/10.5281/zenodo.4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="2" pageNumber="267">Fig. 1a</figureCitation>
); posterior eye margin tumid, with a conic hole located under it (
<figureCitation id="13322A69231DFF8EFB82639EFBEEBFEB" box="[1064,1146,475,502]" captionStart="FIGURE 1" captionStartId="1.[151,250,1760,1784]" captionTargetBox="[153,1432,889,1735]" captionTargetId="figure-209@1.[151,1436,885,1735]" captionTargetPageId="1" captionText="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." figureDoi="http://doi.org/10.5281/zenodo.4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="2" pageNumber="267">Fig. 1d</figureCitation>
).
</paragraph>
<caption id="DF766664231DFF8EFF3D64E1FB6FB8A1" ID-DOI="http://doi.org/10.5281/zenodo.4381850" ID-Zenodo-Dep="4381850" box="[151,1275,1700,1725]" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="2" pageNumber="267" startId="2.[151,250,1700,1724]" targetBox="[224,1364,534,1677]" targetPageId="2">
<paragraph id="8BB636EC231DFF8EFF3D64E1FB6FB8A1" blockId="2.[151,1275,1700,1725]" box="[151,1275,1700,1725]" pageId="2" pageNumber="267">
<emphasis id="B97DEAFE231DFF8EFF3D64E1FE87B8A1" bold="true" box="[151,275,1700,1724]" pageId="2" pageNumber="267">FIGURE 2.</emphasis>
<taxonomicName id="4C094D6F231DFF8EFEB364E1FE46B8A1" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[281,466,1700,1724]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="267" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231DFF8EFEB364E1FE46B8A1" box="[281,466,1700,1724]" italics="true" pageId="2" pageNumber="267">Epigomphus rufus</emphasis>
</taxonomicName>
habitus: a, teneral female holotype; b, teneral male paratype (alive in the field).
</paragraph>
</caption>
<paragraph id="8BB636EC231DFF8FFF6D64AFFBD3BF78" blockId="2.[151,1437,1770,2013]" lastBlockId="3.[151,1436,151,538]" lastPageId="3" lastPageNumber="268" pageId="2" pageNumber="267">
<emphasis id="B97DEAFE231DFF8EFF6D64AFFE8CB919" box="[199,280,1770,1796]" italics="true" pageId="2" pageNumber="267">Thorax</emphasis>
. Reddish brown,the following greenish-gray areas were visible when the specimen died, but unfortunately dispel with time (
<figureCitation id="13322A69231DFF8EFE0C654BFE6CB935" box="[422,504,1806,1832]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="2" pageNumber="267">Fig. 2a</figureCitation>
): two medial spots and two lateroapical spots on posterior lobe of pronotum, mesepisternum with a longitudinal line, a transversal line on anterior border, the second pale antehumeral stripe reduced to a single small posterior spot, the rest of the pleura with three longitudinal lines: one on mesepimeron, one on metepisternum, and one on metepimeron. The antealar crest yellow. Legs brown except distal portion of femora, tibiae, and tarsi which are black. Metafemora approximately 1 ½ the length of pro- and mesofemora. All femora with an external row of spines. Tibiae with long, slender spines on external border, except protibiae with a row of eleven scale-like spines on the external side. Tarsal claws brown with a subapical tooth.
<emphasis id="B97DEAFE231CFF8FFBC762D2FB25BEAD" box="[1133,1201,151,176]" italics="true" pageId="3" pageNumber="268">Wings</emphasis>
. Hyaline with amber tinge on base, veins black and Pt brown (
<figureCitation id="13322A69231CFF8FFDCE62FEFD22BEC8" box="[612,694,187,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="3" pageNumber="268">Fig. 2a</figureCitation>
). Pt covering 7 cells. Left FW with 20 Ax, right FW with 21, FWs with 15 Px; left HW with 14 Ax, right HW with 15 Ax; HWs with 14 Px; supratriangles, triangles and subtriangles without crossevins. Second reinforced antenodal the 8th in left FW, the 7th in right FW, in both HW the 7th; (
<figureCitation id="13322A69231CFF8FFAC56346FF21BF5C" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="3" pageNumber="268">Fig. 4a</figureCitation>
). Cubito-anal crossveins in addition to inner side of subtriangle,
<quantity id="4CF19B09231CFF8FFC0A6362FC5ABF5C" box="[928,974,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="268" unit="in" value="3.0">3 in</quantity>
FWs,
<quantity id="4CF19B09231CFF8FFBB66362FBDDBF5C" box="[1052,1097,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="268" unit="in" value="2.0">2 in</quantity>
left, and
<quantity id="4CF19B09231CFF8FFB106362FB73BF5C" box="[1210,1255,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="268" unit="in" value="3.0">3 in</quantity>
right HW. Five crossveins in space between sectors of arculus and point of branching of RP on HW.
</paragraph>
<paragraph id="8BB636EC231CFF8FFF6D632AFBF1BC07" blockId="3.[151,1436,151,538]" pageId="3" pageNumber="268">
<emphasis id="B97DEAFE231CFF8FFF6D632AFEA5BF94" box="[199,305,367,393]" italics="true" pageId="3" pageNumber="268">Abdomen</emphasis>
. Club-like, narrowing slightly on S3 and widening again on S8, S9 widest. Brown, but S1S2 darker, the distal part of S3 to S7 light brown, S8S10 dark brown dorsal and light brown laterally, black distal rings on S2S7 (
<figureCitation id="13322A69231CFF8FFF5863F2FED2BFCC" box="[242,326,439,465]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="3" pageNumber="268">Fig. 2a</figureCitation>
). Auricles rounded and smooth. Epiproct smooth, long with convex apex, almost as long as S10 (
<figureCitation id="13322A69231CFF8FFF35639EFE8BBFE8" box="[159,287,475,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="3" pageNumber="268">Figs. 3 a, c</figureCitation>
). Cerci approximately ¾ of epiproct length, cylindrical with pointed tips slightly curved ventrad, their apices convergent (
<figureCitation id="13322A69231CFF8FFEC563BAFE28BC04" box="[367,444,511,537]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="3" pageNumber="268">Figs. 3</figureCitation>
ac). Paraprocts conical, straight, with pointed tips (
<figureCitation id="13322A69231CFF8FFBAE63BAFBCCBC07" box="[1028,1112,511,538]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="3" pageNumber="268">Fig. 3b</figureCitation>
).
</paragraph>
<caption id="DF766664231CFF8FFF3D651AFDADB9A2" ID-DOI="http://doi.org/10.5281/zenodo.4381854" ID-Zenodo-Dep="4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="3" pageNumber="268" startId="3.[151,250,1887,1911]" targetBox="[257,1331,561,1864]" targetPageId="3">
<paragraph id="8BB636EC231CFF8FFF3D651AFDADB9A2" blockId="3.[151,1436,1887,1983]" pageId="3" pageNumber="268">
<emphasis id="B97DEAFE231CFF8FFF3D651AFE86B96A" bold="true" box="[151,274,1887,1911]" pageId="3" pageNumber="268">FIGURE 3.</emphasis>
<taxonomicName id="4C094D6F231CFF8FFEB2651AFE45B96A" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[280,465,1887,1911]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231CFF8FFEB2651AFE45B96A" box="[280,465,1887,1911]" italics="true" pageId="3" pageNumber="268">Epigomphus rufus</emphasis>
</taxonomicName>
abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce.
</paragraph>
</caption>
<paragraph id="8BB636EC231BFF88FF6D62D2FCB7BEAC" blockId="4.[151,1437,150,502]" box="[199,803,150,177]" pageId="4" pageNumber="269">
<emphasis id="B97DEAFE231BFF88FF6D62D2FEE2BEAC" bold="true" box="[199,374,151,177]" pageId="4" pageNumber="269">Measurements</emphasis>
(mm): TL 58, Ab 44, FW 43, HW 41.
</paragraph>
<paragraph id="8BB636EC231BFF88FF6D62FEFAA8BF5C" blockId="4.[151,1437,150,502]" pageId="4" pageNumber="269">
<emphasis id="B97DEAFE231BFF88FF6D62FEFEEBBEC8" bold="true" box="[199,383,187,213]" pageId="4" pageNumber="269">
Male
<typeStatus id="54B2884E231BFF88FEA062FEFEEEBEC8" box="[266,378,187,213]" pageId="4" pageNumber="269" type="paratype">paratype</typeStatus>
:
</emphasis>
Teneral, probably do not bear the final coloration pattern. As female
<typeStatus id="54B2884E231BFF88FBD562FEFB75BEC8" box="[1151,1249,187,213]" pageId="4" pageNumber="269" type="holotype">holotype</typeStatus>
but with following differences:
<emphasis id="B97DEAFE231BFF88FEE4629AFE06BEE4" box="[334,402,223,249]" italics="true" pageId="4" pageNumber="269">Head.</emphasis>
With greenish-gray spots on antefrons and in the tumid area behind ocelli, the vertex lacks the C-shaped thick rim, instead it presents a complete transverse postocellar ridge, emarginate medially (
<figureCitation id="13322A69231BFF88FA926346FA19BF03" box="[1336,1421,259,286]" captionStart="FIGURE 1" captionStartId="1.[151,250,1760,1784]" captionTargetBox="[153,1432,889,1735]" captionTargetId="figure-209@1.[151,1436,885,1735]" captionTargetPageId="1" captionText="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." figureDoi="http://doi.org/10.5281/zenodo.4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="4" pageNumber="269">Fig. 1b</figureCitation>
). The posterior part of head behind eyes is not tumid and lacks the conic groove on the occipital region.
</paragraph>
<paragraph id="8BB636EC231BFF88FF6D630EFE00BFEB" blockId="4.[151,1437,150,502]" pageId="4" pageNumber="269">
<emphasis id="B97DEAFE231BFF88FF6D630EFEB7BF78" box="[199,291,331,357]" italics="true" pageId="4" pageNumber="269">Thorax.</emphasis>
Without greenish gray lines on mesepisternum. Metatibia and first two tarsomeres with a row of peglike spines on internal side.
<emphasis id="B97DEAFE231BFF88FE67632AFD85BF95" box="[461,529,367,392]" italics="true" pageId="4" pageNumber="269">Wings</emphasis>
. Pt covering 5 ½ cells. Left FW with 20 Ax, left FW with 14 Px; left HW with 16 Ax, left HW with 13 Px; supratriangles, triangles and subtriangles without crossevins. Second reinforced antenodal the 6th in left FW, in left HW the 7th. Cubito-anal crossveins in addition to inner side of subtriangle,
<quantity id="4CF19B09231BFF88FAA463F2FAAEBFCC" box="[1294,1338,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="269" unit="in" value="3.0">3 in</quantity>
left FW,
<quantity id="4CF19B09231BFF88FF3D639EFF56BFE8" box="[151,194,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="269" unit="in" value="2.0">2 in</quantity>
left HW (
<figureCitation id="13322A69231BFF88FE99639EFE13BFEB" box="[307,391,475,502]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="4" pageNumber="269">Fig. 2b</figureCitation>
).
</paragraph>
<caption id="DF766664231BFF88FF3D6515FDBCB9B2" ID-DOI="http://doi.org/10.5281/zenodo.4381858" ID-Zenodo-Dep="4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="4" pageNumber="269" startId="4.[151,250,1872,1896]" targetBox="[148,1439,525,1834]" targetPageId="4">
<paragraph id="8BB636EC231BFF88FF3D6515FDBCB9B2" blockId="4.[151,1436,1872,1968]" pageId="4" pageNumber="269">
<emphasis id="B97DEAFE231BFF88FF3D6515FE80B975" bold="true" box="[151,276,1872,1896]" pageId="4" pageNumber="269">FIGURE 4.</emphasis>
<taxonomicName id="4C094D6F231BFF88FEB16515FE40B975" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[283,468,1872,1896]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231BFF88FEB16515FE40B975" box="[283,468,1872,1896]" italics="true" pageId="4" pageNumber="269">Epigomphus rufus</emphasis>
</taxonomicName>
male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth.
</paragraph>
</caption>
<paragraph id="8BB636EC231AFF89FF6D62D2FADEBC7C" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF6D62D2FEADBEAC" box="[199,313,151,177]" italics="true" pageId="5" pageNumber="270">Abdomen.</emphasis>
The width of abdominal segments is more uniform, just slightly wider on S810 (
<figureCitation id="13322A69231AFF89FB7C62D2FABFBEAF" box="[1238,1323,151,178]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="5" pageNumber="270">Fig. 2b</figureCitation>
). Auricles subquadrangular, each with rows of 14 black denticles on posterior border (
<figureCitation id="13322A69231AFF89FB8C62FEFB3EBECB" box="[1062,1194,187,214]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4b, d</figureCitation>
). Anterior lamina of genital fossa swollen and rounded, densely covered by long golden setae (
<figureCitation id="13322A69231AFF89FBA4629AFB07BEE7" box="[1038,1171,223,250]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4b, d</figureCitation>
); anterior hamuli with pointed black apical margins (
<figureCitation id="13322A69231AFF89FE5E6346FDD6BF00" box="[500,578,259,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="5" pageNumber="270">Figs. 3</figureCitation>
de, 4b, d); posterior hamuli long, approximately twice the length of anterior hamuli, subtriangular in lateral view, their mesial side covered by black tubercles (
<figureCitation id="13322A69231AFF89FBD76362FB5FBF5C" box="[1149,1227,295,321]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="5" pageNumber="270">Figs. 3</figureCitation>
de, 4b, d).
<emphasis id="B97DEAFE231AFF89FAFE6362FE90BF78" italics="true" pageId="5" pageNumber="270">Vesica spermalis</emphasis>
reddish brown: segment 1 large, concave in ventral view, in lateral view with a ventroapical rounded process directed distally (
<figureCitation id="13322A69231AFF89FE60632AFD88BF94" box="[458,540,367,393]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="5" pageNumber="270">Figs. 3</figureCitation>
de, 4b, d); segment 3 widening towards its distal portion, two ventro apical pointed horns directed ventrally; segment 4, in lateral view, with prepuce subquadrangular, lateral lobes strongly bulging dorsally, in ventral view, forming a nozzle-like slender projection directed posteriorly and parallel to cornua; cornua short (
<figureCitation id="13322A69231AFF89FEC4639EFE28BFEB" box="[366,444,475,502]" captionStart="FIGURE 3" captionStartId="3.[151,250,1887,1911]" captionTargetBox="[257,1331,561,1864]" captionTargetId="figure-226@3.[257,1331,561,1864]" captionTargetPageId="3" captionText="FIGURE 3. Epigomphus rufus abdomen: last segments of female holotype: a, dorsal view; b ventral view; c, lateral view. Male paratype: d, lateral view of S2; e, ventral view of S2. ce: cercus; cor: cornua; ep: epiproct; hrn: horns; l. l. p.: lateral lobe slender projection; pa: paraprocts; prep: prepuce." figureDoi="http://doi.org/10.5281/zenodo.4381854" httpUri="https://zenodo.org/record/4381854/files/figure.png" pageId="5" pageNumber="270">Figs. 3</figureCitation>
de); Cercus and epiproct subequal in length, less than half the length of S10 (
<figureCitation id="13322A69231AFF89FAC8639EFF46BC04" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
). Cerci concave mesially, in lateral view strongly curved ventrally with a pointed dark tip (
<figureCitation id="13322A69231AFF89FAA563BAFA1ABC04" box="[1295,1422,511,537]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
). Epiproct divided in two lateroapical processes, in ventral view widely V-shaped forming an angle close to 90°, each lateroapical process with two black sharp teeth, one apical and another one subapical (
<figureCitation id="13322A69231AFF89FB6B6002FAA8BC7C" box="[1217,1340,583,609]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
).
</paragraph>
<paragraph id="8BB636EC231AFF89FF6D602EFCB5BC98" blockId="5.[151,1437,151,717]" box="[199,801,618,645]" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF6D602EFEE2BC98" bold="true" box="[199,374,619,645]" pageId="5" pageNumber="270">Measurements</emphasis>
(mm): TL 54, Ab 42, FW 40, HW 38.
</paragraph>
<paragraph id="8BB636EC231AFF89FF6D60CBFB25BCB4" blockId="5.[151,1437,151,717]" box="[199,1201,654,681]" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF6D60CBFDBCBCB4" bold="true" box="[199,552,654,681]" pageId="5" pageNumber="270">
Variation of female
<typeStatus id="54B2884E231AFF89FE1960CAFDB7BCB4" box="[435,547,655,681]" pageId="5" pageNumber="270" type="paratype">paratype</typeStatus>
:
</emphasis>
Ax in FW 19, in HW 16; Px in FW 1314, in HW 1214.
</paragraph>
<paragraph id="8BB636EC231AFF89FF6D60F6FCB5BCD0" blockId="5.[151,1437,151,717]" box="[199,801,690,717]" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF6D60F6FEE2BCD0" bold="true" box="[199,374,691,717]" pageId="5" pageNumber="270">Measurements</emphasis>
(mm): TL 59, Ab 44, FW 44, HW 42.
</paragraph>
</subSubSection>
<caption id="DF766664231AFF89FF3D6425FC44B865" ID-DOI="http://doi.org/10.5281/zenodo.4381862" ID-Zenodo-Dep="4381862" box="[151,976,1632,1657]" httpUri="https://zenodo.org/record/4381862/files/figure.png" pageId="5" pageNumber="270" startId="5.[151,250,1632,1656]" targetBox="[185,1406,746,1601]" targetPageId="5">
<paragraph id="8BB636EC231AFF89FF3D6425FC44B865" blockId="5.[151,976,1632,1657]" box="[151,976,1632,1657]" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF3D6425FE87B865" bold="true" box="[151,275,1632,1656]" pageId="5" pageNumber="270">FIGURE 5.</emphasis>
Map with the type localities of
<taxonomicName id="4C094D6F231AFF89FDFC6425FC9BB865" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[598,783,1632,1656]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231AFF89FDFC6425FC9BB865" box="[598,783,1632,1656]" italics="true" pageId="5" pageNumber="270">Epigomphus rufus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D6F231AFF89FCEA6424FC58B865" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[832,972,1632,1656]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="brillantina">E. brillantina</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3136567231AFF8AFF6D64E3FBD6BF94" lastPageId="6" lastPageNumber="271" pageId="5" pageNumber="270" type="diagnosis">
<paragraph id="8BB636EC231AFF8AFF6D64E3FBD6BF94" blockId="5.[151,1437,1702,2017]" lastBlockId="6.[151,1436,151,393]" lastPageId="6" lastPageNumber="271" pageId="5" pageNumber="270">
<emphasis id="B97DEAFE231AFF89FF6D64E3FED0B8DD" bold="true" box="[199,324,1702,1728]" pageId="5" pageNumber="270">Diagnosis:</emphasis>
The typical color pattern of species of this genus usually alternates black or dark brown with yellow, green and light blue; this is the only known species that has reddish brown coloration as dark background color making it easily recognizable (
<figureCitation id="13322A69231AFF89FE5264ABFDDCB914" box="[504,584,1774,1801]" captionStart="FIGURE 2" captionStartId="2.[151,250,1700,1724]" captionTargetBox="[224,1364,534,1677]" captionTargetId="figure-173@2.[224,1364,525,1677]" captionTargetPageId="2" captionText="FIGURE 2. Epigomphus rufus habitus: a, teneral female holotype; b, teneral male paratype (alive in the field)." figureDoi="http://doi.org/10.5281/zenodo.4381850" httpUri="https://zenodo.org/record/4381850/files/figure.png" pageId="5" pageNumber="270">Figs. 2</figureCitation>
ab).
<taxonomicName id="4C094D6F231AFF89FD2064ABFCCDB915" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[650,857,1774,1800]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231AFF89FD2064ABFCCDB915" box="[650,857,1774,1800]" italics="true" pageId="5" pageNumber="270">Epigomphus rufus</emphasis>
</taxonomicName>
shares with a group of 16 species, including most of the South American and some Central American species, its second pale antehumeral stripe reduced to a single small posterior spot (
<tableCitation id="C68B0357231AFF89FE2C6573FE4FB94D" box="[390,475,1846,1872]" captionStart="TABLE 1" captionStartId="6.[151,239,440,464]" captionTargetPageId="6" captionText="TABLE 1. Epigomphus species with second pale antehumeral stripe reduced to a single small posterior spot, their distribution, and male cercus length compared with S10 length." httpUri="http://table.plazi.org/id/DF7666642319FF8AFF3D63FDFC91BFE9" pageId="5" pageNumber="270" tableUuid="DF7666642319FF8AFF3D63FDFC91BFE9">Table 1</tableCitation>
). Among them, the males of
<emphasis id="B97DEAFE231AFF89FC806572FA08B94D" box="[810,1436,1846,1872]" italics="true" pageId="5" pageNumber="270">
<taxonomicName id="4C094D6F231AFF89FC806572FC3CB94D" authorityName="Calvert" authorityYear="1905" box="[810,936,1846,1872]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="camelus">E. camelus</taxonomicName>
,
<taxonomicName id="4C094D6F231AFF89FC1F6572FBC4B94D" authorityName="Belle" authorityYear="1994" box="[949,1104,1847,1872]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="compactus">E. compactus</taxonomicName>
,
<taxonomicName id="4C094D6F231AFF89FBF66572FB7BB94D" authorityName="Calvert" authorityYear="1929" box="[1116,1263,1846,1872]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="subsimilis">E. subsimilis</taxonomicName>
,
<taxonomicName id="4C094D6F231AFF89FB566572FA0CB94D" authorityName="Calvert" authorityYear="1903" box="[1276,1432,1846,1872]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="tumefactus">E. tumefactus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C094D6F231AFF89FF6C651EFEE6B969" authorityName="Calvert" authorityYear="1908" box="[198,370,1883,1908]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="verticicornis">
<emphasis id="B97DEAFE231AFF89FF6C651EFEE6B969" box="[198,370,1883,1908]" italics="true" pageId="5" pageNumber="270">E. verticicornis</emphasis>
</taxonomicName>
share with
<taxonomicName id="4C094D6F231AFF89FE5B651EFDDDB969" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[497,585,1882,1908]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231AFF89FE5B651EFDDDB969" box="[497,585,1882,1908]" italics="true" pageId="5" pageNumber="270">E. rufus</emphasis>
</taxonomicName>
cercus length shorter than S10 length (
<figureCitation id="13322A69231AFF89FC50651FFBE0B968" box="[1018,1140,1882,1909]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
), but males of
<taxonomicName id="4C094D6F231AFF89FABD651EFAE4B969" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[1303,1392,1882,1908]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE231AFF89FABD651EFAE4B969" box="[1303,1392,1882,1908]" italics="true" pageId="5" pageNumber="270">E. rufus</emphasis>
</taxonomicName>
can be easily distinguished from all these species by cercus strongly curved downwards and with a sharp tip which is unique among all 33 known
<taxonomicName id="4C094D6F231AFF89FE7D65E7FDF7B9A1" authorityName="Hagen in Selys" authorityYear="1854" box="[471,611,1954,1980]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAFE231AFF89FE7D65E7FDF7B9A1" box="[471,611,1954,1980]" italics="true" pageId="5" pageNumber="270">Epigomphus</emphasis>
</taxonomicName>
species (
<figureCitation id="13322A69231AFF89FD6165E7FCD3B9A0" box="[715,839,1954,1981]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
), and so is the shape of male epiproct by having two teeth, one apical and one subapical, on each lateroapical process (
<figureCitation id="13322A69231AFF89FC2C6583FB90B9FC" box="[902,1028,1990,2017]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="5" pageNumber="270">Figs. 4a, c</figureCitation>
). As pointed out by
<bibRefCitation id="EF984B1D231AFF89FB456583FA03B9FD" author="Calvert, P. P." box="[1263,1431,1990,2016]" pageId="5" pageNumber="270" pagination="323 - 354" refId="ref5110" refString="Calvert, P. P. (1920) The Costa Rican species of Epigomphus and their mutual mating adaptations (Odonata). Transactions of the American Entomological Society, 46, 323 - 354." type="journal article" year="1920">Calvert (1920)</bibRefCitation>
, these caudal structures of males perfectly match with structures on vertex, occiput, and posterior area of females head, including the C-shaped tubercles behind the posterior ocelli and the conical holes where cerci match on rear of head (
<figureCitation id="13322A692319FF8AFF70629AFEA0BEE4" box="[218,308,223,249]" captionStart="FIGURE 1" captionStartId="1.[151,250,1760,1784]" captionTargetBox="[153,1432,889,1735]" captionTargetId="figure-209@1.[151,1436,885,1735]" captionTargetPageId="1" captionText="FIGURE 1. Epigomphus rufus head. Female holotype: a, dorsal view; c, frontal view; d, posterior view. Male paratype: b, dorsal view. c-s. r.: C-shaped thick rim; h: posterior conic hole; p. r.: transverse postocellar ridge." figureDoi="http://doi.org/10.5281/zenodo.4381846" httpUri="https://zenodo.org/record/4381846/files/figure.png" pageId="6" pageNumber="271">Figs. 1a</figureCitation>
, cd), the females of only ten species in this group of
<taxonomicName id="4C094D6F2319FF8AFC2E629AFB84BEE4" authorityName="Hagen in Selys" authorityYear="1854" box="[900,1040,223,249]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAFE2319FF8AFC2E629AFB84BEE4" box="[900,1040,223,249]" italics="true" pageId="6" pageNumber="271">Epigomphus</emphasis>
</taxonomicName>
are known (
<tableCitation id="C68B03572319FF8AFB3D629AFB7DBEE4" box="[1175,1257,223,249]" captionStart="TABLE 1" captionStartId="6.[151,239,440,464]" captionTargetPageId="6" captionText="TABLE 1. Epigomphus species with second pale antehumeral stripe reduced to a single small posterior spot, their distribution, and male cercus length compared with S10 length." httpUri="http://table.plazi.org/id/DF7666642319FF8AFF3D63FDFC91BFE9" pageId="6" pageNumber="271" tableUuid="DF7666642319FF8AFF3D63FDFC91BFE9">Table 1</tableCitation>
), unfortunately, the descriptions of the posterior area of the head are not meticulous for all of them, but in all the cases where the structures are well described
<taxonomicName id="4C094D6F2319FF8AFE486362FDA9BF5C" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[482,573,295,321]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="B97DEAFE2319FF8AFE486362FDA9BF5C" box="[482,573,295,321]" italics="true" pageId="6" pageNumber="271">E. rufus</emphasis>
</taxonomicName>
females can be easily distinguishable since no other has C-shaped tubercles behind the posterior ocelli or conical holes in the rear area and this combination of characteristics is expected to be unique as the shape of the male cercus and epiproct among the species in the genus.
</paragraph>
</subSubSection>
<caption id="DF7666642319FF8AFF3D63FDFC91BFE9" ID-Table-UUID="DF7666642319FF8AFF3D63FDFC91BFE9" httpUri="http://table.plazi.org/id/DF7666642319FF8AFF3D63FDFC91BFE9" pageId="6" pageNumber="271" targetBox="[166,1406,516,1373]" targetIsTable="true" targetPageId="6">
<paragraph id="8BB636EC2319FF8AFF3D63FDFC91BFE9" blockId="6.[151,1437,440,501]" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF3D63FDFE9EBFCD" bold="true" box="[151,266,440,464]" pageId="6" pageNumber="271">TABLE 1.</emphasis>
<emphasis id="B97DEAFE2319FF8AFEBB63FDFE33BFCD" box="[273,423,440,464]" italics="true" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFEBB63FDFE02BFCD" authorityName="Hagen in Selys" authorityYear="1854" box="[273,406,440,464]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">Epigomphus</taxonomicName>
s
</emphasis>
pecies with second pale antehumeral stripe reduced to a single small posterior spot, their distribution, and male cercus length compared with S10 length.
</paragraph>
</caption>
<paragraph id="8BB636EC2319FF8AFF0C6041FAF0BB41" pageId="6" pageNumber="271">
<table id="F909C44C23190073FF0C6041FAEABB40" box="[166,1406,516,1373]" gridcols="5" gridrows="19" pageId="6" pageNumber="271">
<tr id="353934AE23190073FF0C6041FAEABC01" box="[166,1406,516,540]" gridrow="0" pageId="6" pageNumber="271">
<th id="76E85DD223190073FF0C6041FDC5BC01" box="[166,593,516,540]" gridcol="0" gridrow="0" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF0C6041FF61BC01" bold="true" box="[166,245,516,540]" pageId="6" pageNumber="271">Species</emphasis>
</th>
<th id="76E85DD223190073FDD16041FB80BC01" box="[635,1044,516,540]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFCAF6041FC1DBC01" bold="true" box="[773,905,516,540]" pageId="6" pageNumber="271">Distribution</emphasis>
</th>
<th id="76E85DD223190073FB966041FB64BC01" box="[1084,1264,516,540]" gridcol="3" gridrow="0" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFBE16041FB75BC01" bold="true" box="[1099,1249,516,540]" pageId="6" pageNumber="271">Cercus length</emphasis>
</th>
<th id="76E85DD223190073FA8F6041FAEABC01" box="[1317,1406,516,540]" gridcol="4" gridrow="0" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFA806041FAEDBC01" bold="true" box="[1322,1401,516,540]" pageId="6" pageNumber="271">Known</emphasis>
</th>
</tr>
<tr id="353934AE23190073FF0C606DFAEABC59" box="[166,1406,552,580]" gridrow="1" pageId="6" pageNumber="271" rowspan-0="1">
<td id="76E85DD223190073FDD1606DFCA0BC59" box="[635,820,552,580]" gridcol="1" gridrow="1" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFDD16069FCA0BC59" bold="true" box="[635,820,556,580]" italics="true" pageId="6" pageNumber="271">West of the Andes</emphasis>
</td>
<td id="76E85DD223190073FCF3606DFB80BC59" box="[857,1044,552,580]" gridcol="2" gridrow="1" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFCF16069FB86BC59" bold="true" box="[859,1042,556,580]" italics="true" pageId="6" pageNumber="271">East of the Andes</emphasis>
</td>
<td id="76E85DD223190073FB96606DFB64BC59" box="[1084,1264,552,580]" gridcol="3" gridrow="1" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFB96606DFB64BC5D" bold="true" box="[1084,1264,552,576]" pageId="6" pageNumber="271">shorter than S10</emphasis>
</td>
<td id="76E85DD223190073FA8F606DFAEABC59" box="[1317,1406,552,580]" gridcol="4" gridrow="1" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFA8F606DFAEABC5D" bold="true" box="[1317,1406,552,576]" pageId="6" pageNumber="271">Females</emphasis>
</td>
</tr>
<tr id="353934AE23190073FF0C6016FAEABC71" box="[166,1406,595,620]" gridrow="2" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6016FDC5BC71" box="[166,593,595,620]" gridcol="0" gridrow="2" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF0C6016FE62BC71" bold="true" box="[166,502,595,620]" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6016FE0BBC76" authority="Bota-Sierra &amp; Novelo-Gutiérrez, 2020" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[166,415,595,619]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="brillantina" status="sp. nov.">
<emphasis id="B97DEAFE2319FF8AFF0C6016FE0BBC76" bold="true" box="[166,415,595,619]" italics="true" pageId="6" pageNumber="271">Epigomphus brillantina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57852319FF8AFE0C6011FE62BC71" box="[422,502,596,620]" pageId="6" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76E85DD223190073FDD16016FCA0BC71" box="[635,820,595,620]" gridcol="1" gridrow="2" pageId="6" pageNumber="271">Col</td>
<td id="76E85DD223190073FB966016FB64BC71" box="[1084,1264,595,620]" gridcol="3" gridrow="2" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F6016FAEABC71" box="[1317,1406,595,620]" gridcol="4" gridrow="2" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C6039FAEABC88" box="[166,1406,636,661]" gridrow="3" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6039FDC5BC88" box="[166,593,636,661]" gridcol="0" gridrow="3" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6038FD9ABC89" authority="Calvert, 1905" authorityName="Calvert" authorityYear="1905" box="[166,526,636,661]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="camelus">
<emphasis id="B97DEAFE2319FF8AFF0C6038FEEABC88" box="[166,382,637,661]" italics="true" pageId="6" pageNumber="271">Epigomphus camelus</emphasis>
Calvert, 1905
</taxonomicName>
</th>
<td id="76E85DD223190073FDD16039FCA0BC88" box="[635,820,636,661]" gridcol="1" gridrow="3" pageId="6" pageNumber="271">Co R</td>
<td id="76E85DD223190073FB966039FB64BC88" box="[1084,1264,636,661]" gridcol="3" gridrow="3" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F6039FAEABC88" box="[1317,1406,636,661]" gridcol="4" gridrow="3" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C60E0FAEABCA0" box="[166,1406,677,701]" gridrow="4" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C60E0FDC5BCA0" box="[166,593,677,701]" gridcol="0" gridrow="4" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C60E0FD86BCA0" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[166,530,677,701]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B97DEAFE2319FF8AFF0C60E0FE03BCA0" box="[166,407,677,701]" italics="true" pageId="6" pageNumber="271">Epigomphus compactus</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE3760E0FD86BCA0" author="Belle, J." box="[413,530,677,701]" pageId="6" pageNumber="271" pagination="45" refId="ref4939" refString="Belle, J. (1994) Three new Neotropical Gomphidae from the genera Archaeogomphus Williamson, Cyanogomphus Selys and Epigomphus Hagen (Anisoptera). Odonatologica, 23, 45." type="journal article" year="1994">Belle, 1994</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD160E0FCA0BCA0" box="[635,820,677,701]" gridcol="1" gridrow="4" pageId="6" pageNumber="271">Pan</td>
<td id="76E85DD223190073FB9660E0FB64BCA0" box="[1084,1264,677,701]" gridcol="3" gridrow="4" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F60E0FAEABCA0" box="[1317,1406,677,701]" gridcol="4" gridrow="4" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C6088FAEABCF8" box="[166,1406,717,741]" gridrow="5" pageId="6" pageNumber="271" rowspan-1="1">
<th id="76E85DD223190073FF0C6088FDC5BCF8" box="[166,593,717,741]" gridcol="0" gridrow="5" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6088FD80BCF8" authority="Belle, 1988" authorityName="Belle" authorityYear="1988" box="[166,532,717,741]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="gibberosus">
<emphasis id="B97DEAFE2319FF8AFF0C6088FE0DBCF8" box="[166,409,717,741]" italics="true" pageId="6" pageNumber="271">Epigomphus gibberosus</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE356088FD80BCF8" author="Belle, J." box="[415,532,717,741]" pageId="6" pageNumber="271" pagination="135 - 140" refId="ref4896" refString="Belle, J. (1988) Epigomphus gibberosus, a new species from Peru, with lectotype designations for the eligible species of the genus Epigomphus (Odonata: Gomphidae). Tijdschrift voor entomologie, 131, 135 - 140." type="journal article" year="1988">Belle, 1988</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FCF36088FB80BCF8" box="[857,1044,717,741]" gridcol="2" gridrow="5" pageId="6" pageNumber="271">Per</td>
<td id="76E85DD223190073FB966088FB64BCF8" box="[1084,1264,717,741]" gridcol="3" gridrow="5" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F6088FAEABCF8" box="[1317,1406,717,741]" gridcol="4" gridrow="5" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C60B0FAEABD13" box="[166,1406,757,782]" gridrow="6" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C60B0FDC5BD13" box="[166,593,757,782]" gridcol="0" gridrow="6" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C60B0FDB5BD13" authority="Brooks, 1989" authorityName="Brooks" authorityYear="1989" box="[166,545,757,782]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="houghtoni">
<emphasis id="B97DEAFE2319FF8AFF0C60B0FE06BD10" box="[166,402,757,781]" italics="true" pageId="6" pageNumber="271">Epigomphus houghtoni</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE3260B0FDB5BD13" author="Brooks, S. J." box="[408,545,757,782]" pageId="6" pageNumber="271" pagination="163 - 176" refId="ref5041" refString="Brooks, S. J. (1989) New dragonflies (Odonata) from Costa Rica. Tijdschrift Voor Entomologie, 132, 163 - 176." type="journal article" year="1989">Brooks, 1989</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD160B0FCA0BD13" box="[635,820,757,782]" gridcol="1" gridrow="6" pageId="6" pageNumber="271">Co R</td>
<td id="76E85DD223190073FB9660B0FB64BD13" box="[1084,1264,757,782]" gridcol="3" gridrow="6" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F60B0FAEABD13" box="[1317,1406,757,782]" gridcol="4" gridrow="6" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C615BFAEABD47" box="[166,1406,798,858]" gridrow="7" pageId="6" pageNumber="271" rowspan-1="1">
<th id="76E85DD223190073FF0C615BFDC5BD47" box="[166,593,798,858]" gridcol="0" gridrow="7" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C615BFE74BD2B" authority="Ris, 1918" authorityName="Ris" authorityYear="1918" box="[166,480,798,822]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="hylaeus">
<emphasis id="B97DEAFE2319FF8AFF0C615BFEECBD2B" box="[166,376,798,822]" italics="true" pageId="6" pageNumber="271">Epigomphus hylaeus</emphasis>
Ris, 1918
</taxonomicName>
</th>
<td id="76E85DD223190073FCF3615BFB80BD47" box="[857,1044,798,858]" gridcol="2" gridrow="7" pageId="6" pageNumber="271">Bra, Per, Sur, and FrG</td>
<td id="76E85DD223190073FB96615BFB64BD47" box="[1084,1264,798,858]" gridcol="3" gridrow="7" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F615BFAEABD47" box="[1317,1406,798,858]" gridcol="4" gridrow="7" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C612FFAEABD9F" box="[166,1406,874,898]" gridrow="8" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C612FFDC5BD9F" box="[166,593,874,898]" gridcol="0" gridrow="8" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C612FFE62BD9F" authority="Belle, 1993" authorityName="Belle" authorityYear="1993" box="[166,502,874,898]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="jannyae">
<emphasis id="B97DEAFE2319FF8AFF0C612FFEEFBD9F" box="[166,379,874,898]" italics="true" pageId="6" pageNumber="271">Epigomphus jannyae</emphasis>
Belle, 1993
</taxonomicName>
</th>
<td id="76E85DD223190073FDD1612FFCA0BD9F" box="[635,820,874,898]" gridcol="1" gridrow="8" pageId="6" pageNumber="271">Pan</td>
<td id="76E85DD223190073FB96612FFB64BD9F" box="[1084,1264,874,898]" gridcol="3" gridrow="8" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F612FFAEABD9F" box="[1317,1406,874,898]" gridcol="4" gridrow="8" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C61D7FAEABDB7" box="[166,1406,914,938]" gridrow="9" pageId="6" pageNumber="271" rowspan-1="1">
<th id="76E85DD223190073FF0C61D7FDC5BDB7" box="[166,593,914,938]" gridcol="0" gridrow="9" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C61D7FE61BDB7" authority="Calvert, 1903" authorityName="Calvert" authorityYear="1903" box="[166,501,914,938]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="llama">
<emphasis id="B97DEAFE2319FF8AFF0C61D7FEF1BDB7" box="[166,357,914,938]" italics="true" pageId="6" pageNumber="271">Epigomphus llama</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFEC161D7FE61BDB7" author="Calvert, P. P." box="[363,501,914,938]" pageId="6" pageNumber="271" pagination="183 - 192" refId="ref5069" refString="Calvert, P. P. (1903) On some American Gomphinae (Odonata). Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia, 14, 183 - 192." type="journal article" year="1903">Calvert, 1903</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FCF361D7FB80BDB7" box="[857,1044,914,938]" gridcol="2" gridrow="9" pageId="6" pageNumber="271">Bol and Per</td>
<td id="76E85DD223190073FB9661D7FB64BDB7" box="[1084,1264,914,938]" gridcol="3" gridrow="9" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F61D7FAEABDB7" box="[1317,1406,914,938]" gridcol="4" gridrow="9" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C61FFFAEABDEB" box="[166,1406,954,1014]" gridrow="10" pageId="6" pageNumber="271" rowspan-1="1">
<th id="76E85DD223190073FF0C61FFFDC5BDEB" box="[166,593,954,1014]" gridcol="0" gridrow="10" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C61FFFE61BDCF" authority="Selys, 1869" authorityName="Selys" authorityYear="1869" box="[166,501,954,978]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis id="B97DEAFE2319FF8AFF0C61FFFEECBDCF" box="[166,376,954,978]" italics="true" pageId="6" pageNumber="271">Epigomphus obtusus</emphasis>
Selys, 1869
</taxonomicName>
</th>
<td id="76E85DD223190073FCF361FFFB80BDEB" box="[857,1044,954,1014]" gridcol="2" gridrow="10" pageId="6" pageNumber="271">Bra, Col, Ecu, and Per</td>
<td id="76E85DD223190073FB9661FFFB64BDEB" box="[1084,1264,954,1014]" gridcol="3" gridrow="10" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F61FFFAEABDEB" box="[1317,1406,954,1014]" gridcol="4" gridrow="10" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C6642FAEABA02" box="[166,1406,1031,1055]" gridrow="11" pageId="6" pageNumber="271" rowspan-1="1">
<th id="76E85DD223190073FF0C6642FDC5BA02" box="[166,593,1031,1055]" gridcol="0" gridrow="11" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6642FD8DBA02" authority="Hagen, 1854" authorityName="Hagen" authorityYear="1854" box="[166,537,1031,1055]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="paludosus">
<emphasis id="B97DEAFE2319FF8AFF0C6642FE06BA02" box="[166,402,1031,1055]" italics="true" pageId="6" pageNumber="271">Epigomphus paludosus</emphasis>
Hagen, 1854
</taxonomicName>
</th>
<td id="76E85DD223190073FCF36642FB80BA02" box="[857,1044,1031,1055]" gridcol="2" gridrow="11" pageId="6" pageNumber="271">Arg, Bra, and Par</td>
<td id="76E85DD223190073FB966642FB64BA02" box="[1084,1264,1031,1055]" gridcol="3" gridrow="11" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F6642FAEABA02" box="[1317,1406,1031,1055]" gridcol="4" gridrow="11" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C666AFAEABA5A" box="[166,1406,1071,1095]" gridrow="12" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C666AFDC5BA5A" box="[166,593,1071,1095]" gridcol="0" gridrow="12" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C666AFD81BA5A" authority="Belle, 1970" authorityName="Belle" authorityYear="1970" box="[166,533,1071,1095]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="pechumani">
<emphasis id="B97DEAFE2319FF8AFF0C666AFE0EBA5A" box="[166,410,1071,1095]" italics="true" pageId="6" pageNumber="271">Epigomphus pechumani</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE0A666AFD81BA5A" author="Belle, J." box="[416,533,1071,1095]" pageId="6" pageNumber="271" pagination="3 - 154" refId="ref4811" refString="Belle, J. (1970) Studies on South American Gomphidae (Odonata). Studies on the Fauna of Surinam and other Guyanas, 11, 3 - 154." type="journal article" year="1970">Belle, 1970</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD1666AFCA0BA5A" box="[635,820,1071,1095]" gridcol="1" gridrow="12" pageId="6" pageNumber="271">Col</td>
<td id="76E85DD223190073FB96666AFB64BA5A" box="[1084,1264,1071,1095]" gridcol="3" gridrow="12" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F666AFAEABA5A" box="[1317,1406,1071,1095]" gridcol="4" gridrow="12" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C6612FAEABA8E" box="[166,1406,1111,1171]" gridrow="13" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6612FDC5BA8E" box="[166,593,1111,1171]" gridcol="0" gridrow="13" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6612FDB8BA72" authority="Calvert, 1903" authorityName="Calvert" authorityYear="1903" box="[166,556,1111,1135]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2319FF8AFF0C6612FE08BA72" box="[166,412,1111,1135]" italics="true" pageId="6" pageNumber="271">Epigomphus quadracies</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE0B6612FDB8BA72" author="Calvert, P. P." box="[417,556,1111,1135]" pageId="6" pageNumber="271" pagination="183 - 192" refId="ref5069" refString="Calvert, P. P. (1903) On some American Gomphinae (Odonata). Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia, 14, 183 - 192." type="journal article" year="1903">Calvert, 1903</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD16612FCA0BA8E" box="[635,820,1111,1171]" gridcol="1" gridrow="13" pageId="6" pageNumber="271">Co R, Gua, and Pan</td>
<td id="76E85DD223190073FB966612FB64BA8E" box="[1084,1264,1111,1171]" gridcol="3" gridrow="13" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F6612FAEABA8E" box="[1317,1406,1111,1171]" gridcol="4" gridrow="13" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C66E6FAEABAA6" box="[166,1406,1187,1211]" gridrow="14" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C66E6FDC5BAA6" box="[166,593,1187,1211]" gridcol="0" gridrow="14" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF0C66E6FE2BBAA6" bold="true" box="[166,447,1187,1211]" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C66E6FEFCBAA6" authority="Bota-Sierra &amp; Novelo-Gutiérrez, 2020" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[166,360,1187,1211]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
<emphasis id="B97DEAFE2319FF8AFF0C66E6FEFCBAA6" bold="true" box="[166,360,1187,1211]" italics="true" pageId="6" pageNumber="271">Epigomphus rufus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57852319FF8AFEC566E6FE2BBAA6" box="[367,447,1187,1211]" pageId="6" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76E85DD223190073FDD166E6FCA0BAA6" box="[635,820,1187,1211]" gridcol="1" gridrow="14" pageId="6" pageNumber="271">Col</td>
<td id="76E85DD223190073FB9666E6FB64BAA6" box="[1084,1264,1187,1211]" gridcol="3" gridrow="14" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F66E6FAEABAA6" box="[1317,1406,1187,1211]" gridcol="4" gridrow="14" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C6689FAEABAF9" box="[166,1406,1228,1252]" gridrow="15" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6689FDC5BAF9" box="[166,593,1228,1252]" gridcol="0" gridrow="15" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6689FDC5BAF9" authority="Kennedy, 1946" authorityName="Kennedy" authorityYear="1946" box="[166,593,1228,1252]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="subquadrices">
<emphasis id="B97DEAFE2319FF8AFF0C6689FE26BAF9" box="[166,434,1228,1252]" italics="true" pageId="6" pageNumber="271">Epigomphus subquadrices</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE126689FDC5BAF9" author="Kennedy, C. H." box="[440,593,1228,1252]" pageId="6" pageNumber="271" pagination="662 - 666" refId="ref5390" refString="Kennedy, C. H. (1946) Epigomphus subquadrices, a New Dragonfly (Odonata: Gomphidae) from Panama, with notes on E. quadrices and Eugomphus N. Subgen. Annals of the Entomological Society of America, 39, 662 - 666. https: // doi. org / 10.1093 / aesa / 39.4.662" type="journal article" year="1946">Kennedy, 1946</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD16689FCA0BAF9" box="[635,820,1228,1252]" gridcol="1" gridrow="15" pageId="6" pageNumber="271">Pan</td>
<td id="76E85DD223190073FB966689FB64BAF9" box="[1084,1264,1228,1252]" gridcol="3" gridrow="15" pageId="6" pageNumber="271">No</td>
<td id="76E85DD223190073FA8F6689FAEABAF9" box="[1317,1406,1228,1252]" gridcol="4" gridrow="15" pageId="6" pageNumber="271">No</td>
</tr>
<tr id="353934AE23190073FF0C66B1FAEABB11" box="[166,1406,1268,1292]" gridrow="16" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C66B1FDC5BB11" box="[166,593,1268,1292]" gridcol="0" gridrow="16" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C66B1FDB5BB11" authority="Calvert, 1929" authorityName="Calvert" authorityYear="1929" box="[166,545,1268,1292]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B97DEAFE2319FF8AFF0C66B1FE04BB11" box="[166,400,1268,1292]" italics="true" pageId="6" pageNumber="271">Epigomphus subsimilis</emphasis>
Calvert, 1929
</taxonomicName>
</th>
<td id="76E85DD223190073FDD166B1FCA0BB11" box="[635,820,1268,1292]" gridcol="1" gridrow="16" pageId="6" pageNumber="271">Co R</td>
<td id="76E85DD223190073FB9666B1FB64BB11" box="[1084,1264,1268,1292]" gridcol="3" gridrow="16" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F66B1FAEABB11" box="[1317,1406,1268,1292]" gridcol="4" gridrow="16" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C6759FAEABB29" box="[166,1406,1308,1332]" gridrow="17" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6759FDC5BB29" box="[166,593,1308,1332]" gridcol="0" gridrow="17" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6759FDBDBB29" authority="Calvert, 1903" authorityName="Calvert" authorityYear="1903" box="[166,553,1308,1332]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="tumefactus">
<emphasis id="B97DEAFE2319FF8AFF0C6759FE0DBB29" box="[166,409,1308,1332]" italics="true" pageId="6" pageNumber="271">Epigomphus tumefactus</emphasis>
<bibRefCitation id="EF984B1D2319FF8AFE356759FDBDBB29" author="Calvert, P. P." box="[415,553,1308,1332]" pageId="6" pageNumber="271" pagination="183 - 192" refId="ref5069" refString="Calvert, P. P. (1903) On some American Gomphinae (Odonata). Entomological news, and proceedings of the Entomological Section of the Academy of Natural Sciences of Philadelphia, 14, 183 - 192." type="journal article" year="1903">Calvert, 1903</bibRefCitation>
</taxonomicName>
</th>
<td id="76E85DD223190073FDD16759FCA0BB29" box="[635,820,1308,1332]" gridcol="1" gridrow="17" pageId="6" pageNumber="271">Co R</td>
<td id="76E85DD223190073FB966759FB64BB29" box="[1084,1264,1308,1332]" gridcol="3" gridrow="17" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F6759FAEABB29" box="[1317,1406,1308,1332]" gridcol="4" gridrow="17" pageId="6" pageNumber="271">Yes</td>
</tr>
<tr id="353934AE23190073FF0C6701FAEABB40" box="[166,1406,1348,1373]" gridrow="18" pageId="6" pageNumber="271" rowspan-2="1">
<th id="76E85DD223190073FF0C6701FDC5BB40" box="[166,593,1348,1373]" gridcol="0" gridrow="18" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF0C6701FDAFBB40" authority="Calvert, 1908" authorityName="Calvert" authorityYear="1908" box="[166,571,1348,1373]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="verticicornis">
<emphasis id="B97DEAFE2319FF8AFF0C6701FE3FBB41" box="[166,427,1348,1372]" italics="true" pageId="6" pageNumber="271">Epigomphus verticicornis</emphasis>
Calvert, 1908
</taxonomicName>
</th>
<td id="76E85DD223190073FDD16701FCA0BB40" box="[635,820,1348,1373]" gridcol="1" gridrow="18" pageId="6" pageNumber="271">Co R</td>
<td id="76E85DD223190073FB966701FB64BB40" box="[1084,1264,1348,1373]" gridcol="3" gridrow="18" pageId="6" pageNumber="271">Yes</td>
<td id="76E85DD223190073FA8F6701FAEABB40" box="[1317,1406,1348,1373]" gridcol="4" gridrow="18" pageId="6" pageNumber="271">Yes</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB636EC2319FF8AFF3D672FFCF5BBBB" blockId="6.[151,1437,1386,1446]" pageId="6" pageNumber="271">
<tableNote id="76EF37622319FF8AFF3D672FFCF5BBBB" pageId="6" pageNumber="271" targetBox="[166,1406,516,1373]" targetPageId="6">Arg: Argentina, Bol: Bolivia, Bra: Brazil, Col: Colombia, Co R: Costa Rica, Ecu: Ecuador, FrG: French Guyana, Gua: Guatemala, Pan: Panama, Par: Paraguay, Per: Peru, Sur: Suriname.</tableNote>
</paragraph>
<subSubSection id="C31365672319FF8AFF6D6791FDF0B82B" pageId="6" pageNumber="271" type="distribution">
<paragraph id="8BB636EC2319FF8AFF6D6791FDF0B82B" blockId="6.[151,1437,1492,1591]" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF6D6791FE66BBF3" bold="true" box="[199,498,1492,1518]" pageId="6" pageNumber="271">Distribution and habitat:</emphasis>
This species is so far only known to occur in two close localities, which are actually part of the same forest patch at the northern portion of the Colombian Central Cordillera, close to first and second order streams in the interior of the forest.
</paragraph>
</subSubSection>
</treatment>
</document>