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<document id="6E831CDB2A534C2D0C203DBB374BC2BD" ID="10.11646/zootaxa.4238.2.9SLASH10384" ID-CLB-Dataset="36351" ID-DOI="10.11646/zootaxa.4238.2.9" ID-GBIF-Dataset="0afb74a5-ce8b-4852-890b-0e2a14c25a42" ID-ISSN="1175-5326" ID-Zenodo-Dep="345273" ID-ZooBank="AACD836B-8B48-415D-BD59-B11B164EBB5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488532071421" checkinUser="plazi" docAuthor="Saldaitis, Aidas &amp; Benedek, Balázs" docDate="2017" docId="03A087EEFFBDFFD9FF73FAB9FC76FE7D" docLanguage="en" docName="zootaxa.4238.2.9SLASH10384.pdf" docOrigin="Zootaxa 4238 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Irene Saldaitis &amp; Benedek, 2017, gen. nov." docType="treatment" docUuid="A15866B1-3228-43F4-938A-FDC3AD07A901" docUuidSource="ZooBank" docVersion="8" lastPageNumber="275" masterDocId="FF99FF96FFBDFFD8FFE4FFFBFFFDFFDC" masterDocTitle="Irene litanga, a new genus and new species in the Polia generic-complex from China (Lepidoptera, Noctuidae, Hadeninae, Hadenini)" masterLastPageNumber="280" masterPageNumber="275" pageNumber="275" updateTime="1698437587889" updateUser="ExternalLinkService">
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<mods:title id="D90C4BE580B8F0944D3E04E09F8319F1">Irene litanga, a new genus and new species in the Polia generic-complex from China (Lepidoptera, Noctuidae, Hadeninae, Hadenini)</mods:title>
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<mods:namePart id="5742515003958791FFF82BC64A673D82">Saldaitis, Aidas</mods:namePart>
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<mods:name id="30BD9D23601EE60893846480BEBE470A" type="personal">
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<mods:namePart id="2F19A6364B063D4CD1CF2B687D5DACAC">Benedek, Balázs</mods:namePart>
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<mods:typeOfResource id="39B78B73A081EE13A8D986B63A621E14">text</mods:typeOfResource>
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<mods:title id="47280B6CAFB1B57B66102355BDB01B7E">Zootaxa</mods:title>
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<mods:part id="530C5762F1591BB8E32FA99DB880815D">
<mods:date id="E9A8436C7F7BDEBA358B590246A60049">2017</mods:date>
<mods:detail id="616E49593C34558DB2E5C79A67B59EDF" type="volume">
<mods:number id="F67FFBA7D3CD1039CA3096862609B504">4238</mods:number>
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<mods:classification id="45FC61D63C55ED168F6284723328E6D1">journal article</mods:classification>
<mods:identifier id="ED3A08B1995F551A5371CC0B5646F26F" type="CLB-Dataset">36351</mods:identifier>
<mods:identifier id="103C489527A7A351F00F62B4D58D0CCC" type="DOI">10.11646/zootaxa.4238.2.9</mods:identifier>
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<treatment id="03A087EEFFBDFFD9FF73FAB9FC76FE7D" ID-DOI="http://doi.org/10.5281/zenodo.6040120" ID-GBIF-Taxon="127712265" ID-Zenodo-Dep="6040120" LSID="urn:lsid:zoobank.org:act:A15866B1-3228-43F4-938A-FDC3AD07A901" httpUri="http://treatment.plazi.org/id/03A087EEFFBDFFD9FF73FAB9FC76FE7D" lastPageId="1" lastPageNumber="275" pageId="0" pageNumber="275">
<subSubSection id="C3136573FFBDFFD8FF73FAB9FE45FAA0" pageId="0" pageNumber="275" type="nomenclature">
<paragraph id="8BB636F8FFBDFFD8FF73FAB9FEB0FA81" blockId="0.[151,1436,1312,2028]" box="[151,333,1346,1373]" pageId="0" pageNumber="275">
<heading id="D0FE8194FFBDFFD8FF73FAB9FEB0FA81" bold="true" box="[151,333,1346,1373]" fontSize="11" level="1" pageId="0" pageNumber="275" reason="1">
<emphasis id="B97DEAEAFFBDFFD8FF73FAB9FEB0FA81" bold="true" box="[151,333,1346,1373]" pageId="0" pageNumber="275">
<taxonomicName id="4C094D7BFFBDFFD8FF73FAB9FF25FA80" ID-CoL="54YC" box="[151,216,1346,1372]" class="Sordariomycetes" family="Meliolaceae" genus="Irene" kingdom="Fungi" order="Meliolales" pageId="0" pageNumber="275" phylum="Ascomycota" rank="genus" status="gen. nov.">
<emphasis id="B97DEAEAFFBDFFD8FF73FAB9FF25FA80" bold="true" box="[151,216,1346,1372]" italics="true" pageId="0" pageNumber="275">Irene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E5791FFBDFFD8FF3BFAB8FEB0FA81" box="[223,333,1347,1373]" pageId="0" pageNumber="275" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB636F8FFBDFFD8FF73FA9EFE45FAA0" blockId="0.[151,1436,1312,2028]" box="[151,440,1381,1404]" pageId="0" pageNumber="275">
Type-species:
<taxonomicName id="4C094D7BFFBDFFD8FED4FA9EFE45FAA0" box="[304,440,1381,1404]" class="Sordariomycetes" family="Meliolaceae" genus="Irene" kingdom="Fungi" order="Meliolales" pageId="0" pageNumber="275" phylum="Ascomycota" rank="species" species="litanga">
<emphasis id="B97DEAEAFFBDFFD8FED4FA9EFE45FAA0" box="[304,440,1381,1404]" italics="true" pageId="0" pageNumber="275">Irene litanga</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3136573FFBDFFD9FF23FA73FC76FE7D" lastPageId="1" lastPageNumber="276" pageId="0" pageNumber="275" type="diagnosis">
<paragraph id="8BB636F8FFBDFFD9FF23FA73FC76FE7D" blockId="0.[151,1436,1312,2028]" lastBlockId="1.[151,1436,152,417]" lastPageId="1" lastPageNumber="276" pageId="0" pageNumber="275">
<emphasis id="B97DEAEAFFBDFFD8FF23FA73FEC1FA43" bold="true" box="[199,316,1416,1439]" pageId="0" pageNumber="275">Diagnosis.</emphasis>
The new and yet monotypical genus represents a separate phyletic-line within the Poliina, standing rather remote from all other known members of the subtribus. The external morphology of the new species resembles members of the genus
<taxonomicName id="4C094D7BFFBDFFD8FECEFA36FE62FA38" box="[298,415,1485,1508]" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FECEFA36FE62FA38" box="[298,415,1485,1508]" italics="true" pageId="0" pageNumber="275">Kollariana</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="4C094D7BFFBDFFD8FE07FA36FBBDFA38" authority="Kollar &amp; Redtenbacher, 1844" authorityName="Kollar &amp; Redtenbacher" authorityYear="1844" box="[483,1088,1485,1508]" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="scotochlora">
<emphasis id="B97DEAEAFFBDFFD8FE07FA36FD1DFA38" box="[483,736,1485,1508]" italics="true" pageId="0" pageNumber="275">Kollariana scotochlora</emphasis>
(
<bibRefCitation id="EF984B09FFBDFFD8FD15FA36FBC5FA38" author="Kollar" box="[753,1080,1485,1508]" pageId="0" pageNumber="275" refString="Kollar, V. &amp; Redtenbacher, L. (1844) Aufzahlung und Beschreibung der von Freiherrn Carl v. Hugel auf seiner Reise durch Kaschmir und des Himaleya gebirge gesammelten Insectin. In: Hugel, C. F. von (Ed.), Kaschmir und das Reich der Sick. Viertes Band, Stuttgart, pp. 393 - 564." type="book chapter" year="1844">Kollar &amp; Redtenbacher, 1844</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="13322A7DFFBDFFD8FBB5FA36FB4AFA38" box="[1105,1207,1485,1508]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,258,1594,1616]" captionTargetBox="[152,1433,474,1568]" captionTargetId="figure@1.[151,1436,471,1572]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Irene gen. n. &amp; Kollariana spp., adults. 1. I. litanga, male, holotype, China, Sichuan (coll. GBG / ZSM); 2. I. litanga, female, paratype, China, Sichuan (coll. AFM); 3. I. litanga, female, paratype, China, Yunnan (coll. HNHM); 4. I. litanga, female, paratype, China, Yunnan (coll. HNHM); 5. K. scotochlora, male, Pakistan, Kaghan valley (PGM); 6. K. scotochlora, female, Pakistan, Kaghan valley (PGM)." httpUri="https://zenodo.org/record/345274/files/figure.png" pageId="0" pageNumber="275">Figs 5, 6</figureCitation>
,
<figureCitation id="13322A7DFFBDFFD8FB23FA36FAEDFA38" box="[1223,1296,1485,1508]" captionStart="FIGURES 17 – 22" captionStartId="4.[151,258,1931,1953]" captionTargetBox="[202,1340,191,1882]" captionTargetId="figure@4.[200,1357,191,1892]" captionTargetPageId="4" captionText="FIGURES 17 – 22. Kollariana, Polia spp. &amp; Bryoxena spp. male and female genitalia. 17. K. scotochlora, male, gen. prep. GYP 4442 m; 18. P. nebulosa, male, gen. prep. GYP 4447 m; 19. B. centralasiae, male, gen. prep. GYP 4449 m; 20. K. scotochlora, female, gen. prep. GYP 4445 f; 21. P. nebulosa, female, gen. prep. GYP 4439 f; 22. B. centralasiae, female, gen. prep. GYP 4447 f." httpUri="https://zenodo.org/record/345277/files/figure.png" pageId="0" pageNumber="275">17, 20</figureCitation>
),
<taxonomicName id="4C094D7BFFBDFFD8FAC3FA36FE5BF9DB" authority="Draudt, 1950" authorityName="Draudt" authorityYear="1950" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="albomixta">
<emphasis id="B97DEAEAFFBDFFD8FAC3FA36FEFFF9DB" italics="true" pageId="0" pageNumber="275">Kollariana albomixta</emphasis>
(
<bibRefCitation id="EF984B09FFBDFFD8FEEBFA0BFE63F9DB" author="Draudt" box="[271,414,1520,1543]" pageId="0" pageNumber="275" refString="Draudt, M. (1950) Beitrage zur Kenntnis der Agrotiden-Fauna Chinas aus den Ausbeuten Dr. H. Hone's (Beitrag zur Fauna Sinica). Mitteilungen der Munchner Entomologischen Gesellschaft, 40 (1), 1 - 174." type="journal article" year="1950">Draudt, 1950</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C094D7BFFBDFFD8FE3CFA0BFC89F9DB" authority="Plante, 1982" authorityName="Plante" authorityYear="1982" box="[472,884,1520,1543]" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="similissima">
<emphasis id="B97DEAEAFFBDFFD8FE3CFA0BFD36F9DB" box="[472,715,1520,1543]" italics="true" pageId="0" pageNumber="275">Kollariana similissima</emphasis>
<bibRefCitation id="EF984B09FFBDFFD8FD36FA0BFC89F9DB" author="Plante" box="[722,884,1520,1543]" pageId="0" pageNumber="275" refString="Plante, J. (1982) Quatre especes nouvelles de Noctuidae Hadeninae de l'Himalaya. Bulletin de la Societe entomologique de France, 87, 286 - 292." type="journal article" year="1982">(Plante, 1982))</bibRefCitation>
</taxonomicName>
, but
<taxonomicName id="4C094D7BFFBDFFD8FC43FA0BFBFAF9DB" box="[935,1031,1520,1543]" class="Sordariomycetes" family="Meliolaceae" genus="Irene" kingdom="Fungi" order="Meliolales" pageId="0" pageNumber="275" phylum="Ascomycota" rank="species" species="litanga" status="sp. nov.">
<emphasis id="B97DEAEAFFBDFFD8FC43FA0BFBFAF9DB" box="[935,1031,1520,1543]" italics="true" pageId="0" pageNumber="275">I. litanga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E5791FFBDFFD8FBEAFA0BFBB7F9DB" box="[1038,1098,1520,1543]" pageId="0" pageNumber="275" rank="species">
<emphasis id="B97DEAEAFFBDFFD8FBEAFA0BFBB7F9DB" bold="true" box="[1038,1098,1520,1543]" pageId="0" pageNumber="275">sp. n.</emphasis>
</taxonomicNameLabel>
has a more smoothly granulous wing-pattern with more diffuse patterns and a more contrasted dark medial and light subbasal and subterminal area. The olive-greenish ground colour mixed with a yellowish tint occurs in certain members of the genus
<taxonomicName id="4C094D7BFFBDFFD8FB29F9CEFF32F9B3" authority="Boursin, 1954" authorityName="Boursin" authorityYear="1954" class="Insecta" family="Noctuidae" genus="Bryopolia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FB29F9CEFAC5F990" box="[1229,1336,1589,1612]" italics="true" pageId="0" pageNumber="275">Bryopolia</emphasis>
<bibRefCitation id="EF984B09FFBDFFD8FAA5F9CEFF32F9B3" author="Boursin" pageId="0" pageNumber="275" refString="Boursin, Ch. (1954 b) Contribution to the knowledge of the Agrotidae-Trifinae of Kashmir. Bulletin de la Societe Fouad Ier d'Entomologie, 38, 81 - 106." type="journal article" year="1954">Boursin, 1954</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C094D7BFFBDFFD8FF3EF9A3FD61F9B3" authority="Varga, Ronkay &amp; Hacker, 1990" authorityName="Varga, Ronkay &amp; Hacker" authorityYear="1990" box="[218,668,1624,1647]" class="Insecta" family="Noctuidae" genus="Bryoxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FF3EF9A3FEBDF9B3" box="[218,320,1624,1647]" italics="true" pageId="0" pageNumber="275">Bryoxena</emphasis>
Varga, Ronkay &amp; Hacker, 1990
</taxonomicName>
(
<taxonomicName id="4C094D7BFFBDFFD8FD48F9A3FB90F9B3" authority="Staudinger, 1882" authorityName="Staudinger" authorityYear="1882" box="[684,1133,1624,1647]" class="Insecta" family="Noctuidae" genus="Bryoxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="centralasiae">
<emphasis id="B97DEAEAFFBDFFD8FD48F9A3FC62F9B3" box="[684,927,1624,1647]" italics="true" pageId="0" pageNumber="275">Bryoxena centralasiae</emphasis>
(
<bibRefCitation id="EF984B09FFBDFFD8FC49F9A3FB9BF9B3" author="Staudinger" box="[941,1126,1624,1647]" pageId="0" pageNumber="275" refString="Staudinger, O. (1882 b) Beitrag zur Lepidopteren-fauna Central-Asiens. Entomologische Zeitung. Entomologischen Vereine zu Stettin, 43, 35 - 78." type="journal article" year="1882">Staudinger, 1882</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="13322A7DFFBDFFD8FB99F9A3FB3EF9B3" box="[1149,1219,1624,1647]" captionStart="FIGURES 7 – 12" captionStartId="2.[151,258,1954,1976]" captionTargetBox="[165,1417,842,1917]" captionTargetId="figure@2.[160,1420,840,1924]" captionTargetPageId="2" captionText="FIGURES 7 – 12. Polia, Bryoxena &amp; Hada spp., adults. 7. P. nebulosa, male, Armenia, David-Bek (coll. ASV); 8. P. nebulosa, female, Japan, Honshu (coll. ASV); 9. B. centralasiae, female, Tadjikistan, Gissar (coll. PGM); 10. H. fraterna, male, holotype, E. Tibet, Taba (coll. PGM / HNHM); 11. H. fraterna, male, China, Sichuan (coll. AFM); 12. H. fraterna, female, China, Sichuan (coll. AFM)." httpUri="https://zenodo.org/record/345275/files/figure.png" pageId="0" pageNumber="275">Figs 9</figureCitation>
,
<figureCitation id="13322A7DFFBDFFD8FB34F9A3FAEAF9B3" box="[1232,1303,1624,1647]" captionStart="FIGURES 17 – 22" captionStartId="4.[151,258,1931,1953]" captionTargetBox="[202,1340,191,1882]" captionTargetId="figure@4.[200,1357,191,1892]" captionTargetPageId="4" captionText="FIGURES 17 – 22. Kollariana, Polia spp. &amp; Bryoxena spp. male and female genitalia. 17. K. scotochlora, male, gen. prep. GYP 4442 m; 18. P. nebulosa, male, gen. prep. GYP 4447 m; 19. B. centralasiae, male, gen. prep. GYP 4449 m; 20. K. scotochlora, female, gen. prep. GYP 4445 f; 21. P. nebulosa, female, gen. prep. GYP 4439 f; 22. B. centralasiae, female, gen. prep. GYP 4447 f." httpUri="https://zenodo.org/record/345277/files/figure.png" pageId="0" pageNumber="275">19, 22</figureCitation>
), occasional similarity, following genus are members of the subfamily
<taxonomicName id="4C094D7BFFBDFFD8FCF1F980FC78F94E" box="[789,901,1659,1682]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="subFamily" subFamily="Xyleninae">Xyleninae</taxonomicName>
, tribe
<taxonomicName id="4C094D7BFFBDFFD8FC28F980FBC4F94E" box="[972,1081,1659,1682]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="tribe" tribe="Apameini">Apameini</taxonomicName>
), or
<taxonomicName id="4C094D7BFFBDFFD8FB8AF980FAB3F94E" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" box="[1134,1358,1659,1682]" class="Insecta" family="Noctuidae" genus="Hada" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FB8AF980FB54F94E" box="[1134,1193,1659,1682]" italics="true" pageId="0" pageNumber="275">Hada</emphasis>
<bibRefCitation id="EF984B09FFBDFFD8FB57F980FAB3F94E" author="Billberg" box="[1203,1358,1659,1682]" pageId="0" pageNumber="275" refString="Billberg G. J. (1820) Enumeratio Insectorum in Museo G. J. Billberg. Typis Gadelianis, Stockholm, 138 pp. https: // doi. org / 10.5962 / bhl. title. 49763" type="book" year="1820">Billberg, 1820</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C094D7BFFBDFFD8FA85F980FE0BF968" authority="Gyulai &amp; Ronkay, 1998" authorityName="Gyulai &amp; Ronkay" authorityYear="1998" class="Insecta" family="Noctuidae" genus="Hada" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B97DEAEAFFBDFFD8FA85F980FF13F968" italics="true" pageId="0" pageNumber="275">Hada fraterna</emphasis>
<bibRefCitation id="EF984B09FFBDFFD8FF11F966FE0BF968" author="Gyulai" box="[245,502,1693,1716]" pageId="0" pageNumber="275" refString="Gyulai, P. &amp; Ronkay, L. (1998) Seven new species of Noctuidae (Lepidoptera) from Asia. Acta Zoolologica Academia Scientiarum Hungarica, 44 (4), 311 - 327." type="journal article" year="1998">Gyulai &amp; Ronkay, 1998</bibRefCitation>
</taxonomicName>
) (
<figureCitation id="13322A7DFFBDFFD8FDEFF966FD7AF968" box="[523,647,1693,1716]" captionStart="FIGURES 7 – 12" captionStartId="2.[151,258,1954,1976]" captionTargetBox="[165,1417,842,1917]" captionTargetId="figure@2.[160,1420,840,1924]" captionTargetPageId="2" captionText="FIGURES 7 – 12. Polia, Bryoxena &amp; Hada spp., adults. 7. P. nebulosa, male, Armenia, David-Bek (coll. ASV); 8. P. nebulosa, female, Japan, Honshu (coll. ASV); 9. B. centralasiae, female, Tadjikistan, Gissar (coll. PGM); 10. H. fraterna, male, holotype, E. Tibet, Taba (coll. PGM / HNHM); 11. H. fraterna, male, China, Sichuan (coll. AFM); 12. H. fraterna, female, China, Sichuan (coll. AFM)." httpUri="https://zenodo.org/record/345275/files/figure.png" pageId="0" pageNumber="275">Figs 10–12</figureCitation>
,
<figureCitation id="13322A7DFFBDFFD8FD75F966FD2AF968" box="[657,727,1693,1716]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,258,1951,1973]" captionTargetBox="[209,1342,193,1898]" captionTargetId="figure@3.[192,1360,191,1908]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Irene gen. n. &amp; Hada spp., male and female genitalia. 13. I. litanga, male, holotype, gen. prep. OP 3364 m; 14. H. fraterna, male, gen. prep. GYP 1863 m; 15. I. litanga, female, paratype, gen. prep. OP 3365 f; 16. H. fraterna, female, gen. prep. GYP 4514 f." httpUri="https://zenodo.org/record/345276/files/figure.png" pageId="0" pageNumber="275">14, 16</figureCitation>
), but
<taxonomicName id="4C094D7BFFBDFFD8FCF0F966FC8BF968" box="[788,886,1693,1716]" class="Sordariomycetes" family="Meliolaceae" genus="Irene" kingdom="Fungi" order="Meliolales" pageId="0" pageNumber="275" phylum="Ascomycota" rank="species" species="litanga">
<emphasis id="B97DEAEAFFBDFFD8FCF0F966FC8BF968" box="[788,886,1693,1716]" italics="true" pageId="0" pageNumber="275">I. litanga</emphasis>
</taxonomicName>
is larger in size than all the aforementioned similar species (
<quantity id="4CF19B1DFFBDFFD8FF15F93BFE98F90B" box="[241,357,1728,1751]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="0" pageNumber="275" unit="mm" value="53.0" valueMax="54.0" valueMin="52.0">52–54 mm</quantity>
comparing to
<quantity id="4CF19B1DFFBDFFD8FE1AF93BFD92F90B" box="[510,623,1728,1751]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="0" pageNumber="275" unit="mm" value="47.5" valueMax="50.0" valueMin="45.0">45–50 mm</quantity>
of the largest
<taxonomicName id="4C094D7BFFBDFFD8FCE1F93BFC5FF90B" box="[773,930,1728,1751]" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="scotochlora">
<emphasis id="B97DEAEAFFBDFFD8FCE1F93BFC5FF90B" box="[773,930,1728,1751]" italics="true" pageId="0" pageNumber="275">K. scotochlora</emphasis>
</taxonomicName>
) and the genitalia of
<taxonomicName id="4C094D7BFFBDFFD8FB66F93BFB10F90B" box="[1154,1261,1728,1751]" class="Insecta" family="Noctuidae" genus="Bryopolia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FB66F93BFB10F90B" box="[1154,1261,1728,1751]" italics="true" pageId="0" pageNumber="275">Bryopolia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D7BFFBDFFD8FB1DF93BFAA2F90B" box="[1273,1375,1728,1751]" class="Insecta" family="Noctuidae" genus="Bryoxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FB1DF93BFAA2F90B" box="[1273,1375,1728,1751]" italics="true" pageId="0" pageNumber="275">Bryoxena</emphasis>
</taxonomicName>
(
<figureCitation id="13322A7DFFBDFFD8FA89F93BFF22F926" captionStart="FIGURES 17 – 22" captionStartId="4.[151,258,1931,1953]" captionTargetBox="[202,1340,191,1882]" captionTargetId="figure@4.[200,1357,191,1892]" captionTargetPageId="4" captionText="FIGURES 17 – 22. Kollariana, Polia spp. &amp; Bryoxena spp. male and female genitalia. 17. K. scotochlora, male, gen. prep. GYP 4442 m; 18. P. nebulosa, male, gen. prep. GYP 4447 m; 19. B. centralasiae, male, gen. prep. GYP 4449 m; 20. K. scotochlora, female, gen. prep. GYP 4445 f; 21. P. nebulosa, female, gen. prep. GYP 4439 f; 22. B. centralasiae, female, gen. prep. GYP 4447 f." httpUri="https://zenodo.org/record/345277/files/figure.png" pageId="0" pageNumber="275">Figs 19, 22</figureCitation>
) are significantly different. In the male genitalia of
<taxonomicName id="4C094D7BFFBDFFD8FCCAF918FC68F926" box="[814,917,1763,1786]" class="Sordariomycetes" family="Meliolaceae" genus="Irene" kingdom="Fungi" order="Meliolales" pageId="0" pageNumber="275" phylum="Ascomycota" rank="species" species="litanga">
<emphasis id="B97DEAEAFFBDFFD8FCCAF918FC68F926" box="[814,917,1763,1786]" italics="true" pageId="0" pageNumber="275">I. litanga</emphasis>
</taxonomicName>
, several specialized characters corroborate the separation on a generic level as no species from
<taxonomicName id="4C094D7BFFBDFFD8FD7BF8FEFD24F8C0" box="[671,729,1797,1820]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FD7BF8FEFD24F8C0" box="[671,729,1797,1820]" italics="true" pageId="0" pageNumber="275">Polia</emphasis>
</taxonomicName>
genera group from the Palearctic region (
<bibRefCitation id="EF984B09FFBDFFD8FB7DF8FEFA97F8C0" author="Hacker" box="[1177,1386,1797,1820]" pageId="0" pageNumber="275" refString="Hacker, H., Ronkay, L. &amp; Hreblay, M. (2002) Hadeninae I. Noctuidae Europaeae. Vol. 4. Entomological press, Soro, 419 pp." type="book" year="2002">
Hacker
<emphasis id="B97DEAEAFFBDFFD8FB0BF8FEFADBF8C0" box="[1263,1318,1797,1820]" italics="true" pageId="0" pageNumber="275">et al.</emphasis>
, 2002
</bibRefCitation>
) nor the Nearctic (
<bibRefCitation id="EF984B09FFBDFFD8FECEF8D3FE31F8E3" author="McCabe" box="[298,460,1832,1855]" pageId="0" pageNumber="275" refString="McCabe, T. L. (1980) A reclassification of the Polia complex for North America (Lepidoptera: Noctuidae). The University of the State of New York, The State Education Department, Albany, New York, 141 pp." type="book" year="1980">McCabe, 1980</bibRefCitation>
) are known to have such characters. While the male genitalia are similar in their general configuration to those of
<taxonomicName id="4C094D7BFFBDFFD8FE49F8B0FDD5F8BE" box="[429,552,1867,1890]" class="Insecta" family="Noctuidae" genus="Hada" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B97DEAEAFFBDFFD8FE49F8B0FDD5F8BE" box="[429,552,1867,1890]" italics="true" pageId="0" pageNumber="275">H. fraterna</emphasis>
</taxonomicName>
, the narrow, hook-like uncus, the well-developed corona and the strong saccular extension are instead common characters within
<taxonomicName id="4C094D7BFFBDFFD8FD79F896FD2AF858" box="[669,727,1901,1924]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FD79F896FD2AF858" box="[669,727,1901,1924]" italics="true" pageId="0" pageNumber="275">Polia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C094D7BFFBDFFD8FD03F896FBFEF858" authority="Hufnagel" authorityName="Hufnagel" box="[743,1027,1901,1924]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B97DEAEAFFBDFFD8FD03F896FC7AF858" box="[743,903,1901,1924]" italics="true" pageId="0" pageNumber="275">Polia nebulosa</emphasis>
(Hufnagel)
</taxonomicName>
(
<figureCitation id="13322A7DFFBDFFD8FBF5F896FB8DF858" box="[1041,1136,1901,1924]" captionStart="FIGURES 7 – 12" captionStartId="2.[151,258,1954,1976]" captionTargetBox="[165,1417,842,1917]" captionTargetId="figure@2.[160,1420,840,1924]" captionTargetPageId="2" captionText="FIGURES 7 – 12. Polia, Bryoxena &amp; Hada spp., adults. 7. P. nebulosa, male, Armenia, David-Bek (coll. ASV); 8. P. nebulosa, female, Japan, Honshu (coll. ASV); 9. B. centralasiae, female, Tadjikistan, Gissar (coll. PGM); 10. H. fraterna, male, holotype, E. Tibet, Taba (coll. PGM / HNHM); 11. H. fraterna, male, China, Sichuan (coll. AFM); 12. H. fraterna, female, China, Sichuan (coll. AFM)." httpUri="https://zenodo.org/record/345275/files/figure.png" pageId="0" pageNumber="275">Figs 7, 8</figureCitation>
,
<figureCitation id="13322A7DFFBDFFD8FB98F896FB3DF858" box="[1148,1216,1901,1924]" captionStart="FIGURES 17 – 22" captionStartId="4.[151,258,1931,1953]" captionTargetBox="[202,1340,191,1882]" captionTargetId="figure@4.[200,1357,191,1892]" captionTargetPageId="4" captionText="FIGURES 17 – 22. Kollariana, Polia spp. &amp; Bryoxena spp. male and female genitalia. 17. K. scotochlora, male, gen. prep. GYP 4442 m; 18. P. nebulosa, male, gen. prep. GYP 4447 m; 19. B. centralasiae, male, gen. prep. GYP 4449 m; 20. K. scotochlora, female, gen. prep. GYP 4445 f; 21. P. nebulosa, female, gen. prep. GYP 4439 f; 22. B. centralasiae, female, gen. prep. GYP 4447 f." httpUri="https://zenodo.org/record/345277/files/figure.png" pageId="0" pageNumber="275">18, 21</figureCitation>
), the type-species of
<taxonomicName id="4C094D7BFFBDFFD8FF73F86BFF2CF87B" box="[151,209,1936,1959]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FF73F86BFF2CF87B" box="[151,209,1936,1959]" italics="true" pageId="0" pageNumber="275">Polia</emphasis>
</taxonomicName>
). The shape of the cucullus is similar to that of
<taxonomicName id="4C094D7BFFBDFFD8FD35F86BFCBBF87B" box="[721,838,1936,1959]" class="Insecta" family="Noctuidae" genus="Kollariana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAEAFFBDFFD8FD35F86BFCBBF87B" box="[721,838,1936,1959]" italics="true" pageId="0" pageNumber="275">Kollariana</emphasis>
</taxonomicName>
, but the configuration of the tegumen, the valva (nearly evenly broad, without anal spine), the clasper and harpe, the strongly sclerotised and elongated fultura and the very long and narrow, branch-like and apically rounded saccular extension are specialized characters of the new genus. The specific characters of the aedeagus are the presence of the sclerotised carinal bar on both the ventral and dorsal side, the medially broader, terminally curved vesica, the presence of the single strong cornuti on the first medial diverticulum and the narrow, dagger-like shaped second diverticulum. The female genitalia resemble
<taxonomicName id="4C094D7BFFBCFFD9FBF7FF26FB70FF28" box="[1043,1165,221,244]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B97DEAEAFFBCFFD9FBF7FF26FB70FF28" box="[1043,1165,221,244]" italics="true" pageId="1" pageNumber="276">P. nebulosa</emphasis>
</taxonomicName>
(
<figureCitation id="13322A7DFFBCFFD9FB7FFF26FB15FF28" box="[1179,1256,221,244]" captionStart="FIGURES 17 – 22" captionStartId="4.[151,258,1931,1953]" captionTargetBox="[202,1340,191,1882]" captionTargetId="figure@4.[200,1357,191,1892]" captionTargetPageId="4" captionText="FIGURES 17 – 22. Kollariana, Polia spp. &amp; Bryoxena spp. male and female genitalia. 17. K. scotochlora, male, gen. prep. GYP 4442 m; 18. P. nebulosa, male, gen. prep. GYP 4447 m; 19. B. centralasiae, male, gen. prep. GYP 4449 m; 20. K. scotochlora, female, gen. prep. GYP 4445 f; 21. P. nebulosa, female, gen. prep. GYP 4439 f; 22. B. centralasiae, female, gen. prep. GYP 4447 f." httpUri="https://zenodo.org/record/345277/files/figure.png" pageId="1" pageNumber="276">Fig. 21</figureCitation>
) and
<taxonomicName id="4C094D7BFFBCFFD9FAC0FF26FA66FF28" box="[1316,1435,221,244]" class="Insecta" family="Noctuidae" genus="Hada" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B97DEAEAFFBCFFD9FAC0FF26FA66FF28" box="[1316,1435,221,244]" italics="true" pageId="1" pageNumber="276">H. fraterna</emphasis>
</taxonomicName>
(
<figureCitation id="13322A7DFFBCFFD9FF7BFEFBFF13FECB" box="[159,238,256,279]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,258,1951,1973]" captionTargetBox="[209,1342,193,1898]" captionTargetId="figure@3.[192,1360,191,1908]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Irene gen. n. &amp; Hada spp., male and female genitalia. 13. I. litanga, male, holotype, gen. prep. OP 3364 m; 14. H. fraterna, male, gen. prep. GYP 1863 m; 15. I. litanga, female, paratype, gen. prep. OP 3365 f; 16. H. fraterna, female, gen. prep. GYP 4514 f." httpUri="https://zenodo.org/record/345276/files/figure.png" pageId="1" pageNumber="276">Fig. 16</figureCitation>
) in the heavily sclerotised antrum and ductus bursae, but the widely conical, large, tongue-shaped ostial plate, the broader, rounded distal part of the antrum, the presence of the two lateral lamias on the proximal half of the ductus (a common character in the
<taxonomicName id="4C094D7BFFBCFFD9FE55FEBEFD03FE80" authority="Draudt, 1950" authorityName="Draudt" authorityYear="1950" box="[433,766,325,348]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="costirufa">
<emphasis id="B97DEAEAFFBCFFD9FE55FEBEFDAEFE80" box="[433,595,325,348]" italics="true" pageId="1" pageNumber="276">Polia costirufa</emphasis>
(
<bibRefCitation id="EF984B09FFBCFFD9FD87FEBEFD0BFE80" author="Draudt" box="[611,758,325,348]" pageId="1" pageNumber="276" refString="Draudt, M. (1950) Beitrage zur Kenntnis der Agrotiden-Fauna Chinas aus den Ausbeuten Dr. H. Hone's (Beitrag zur Fauna Sinica). Mitteilungen der Munchner Entomologischen Gesellschaft, 40 (1), 1 - 174." type="journal article" year="1950">Draudt, 1950</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97DEAEAFFBCFFD9FD19FEBEFCE5FE80" box="[765,792,325,348]" italics="true" pageId="1" pageNumber="276">—</emphasis>
<taxonomicName id="4C094D7BFFBCFFD9FCFDFEBEFAC1FE80" authority="Saldaitis, Benedek &amp; Behounek, 2016" authorityName="Saldaitis, Benedek &amp; Behounek" authorityYear="2016" box="[793,1340,325,348]" class="Insecta" family="Noctuidae" genus="Polia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="hofer">
<emphasis id="B97DEAEAFFBCFFD9FCFDFEBEFC6DFE80" box="[793,912,325,348]" italics="true" pageId="1" pageNumber="276">Polia hofer</emphasis>
<bibRefCitation id="EF984B09FFBCFFD9FC7EFEBEFAC1FE80" author="Saldaitis" box="[922,1340,325,348]" pageId="1" pageNumber="276" refString="Saldaitis, A., Benedek, B. &amp; Behounek, G. (2016) A new Polia Ochsenheimer, 1816 species from China (Lepidoptera, Noctuidae, Hadenini), Zootaxa, 4093 (4), 577 - 582." type="journal article" year="2016">Saldaitis, Benedek &amp; Behounek, 2016</bibRefCitation>
</taxonomicName>
speciespair!) and the characteristically constructed and sclerotised, broad, laminal coverage (“cloak”) of the appendix bursae with a rolled proximal edge are all unique characters of the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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