treatments-xml/data/03/A0/87/03A087E00E52FFA9FF7FFC16FE54F87D.xml

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<document id="120D49587EF553FCA1CE399DCA12DC08" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.bibliography_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent &amp; Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E52FFA9FF7FFC16FE54F87D" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sindhiluta" docType="treatment" docVersion="11" lastPageNumber="128" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="128" updateTime="1729174488204" updateUser="julia">
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<mods:title id="70C77A992C5E42121A6E11A2750C78C5">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="E71B3AB5E08AF552BDE028922E8C4D28">Merle, Didier</mods:namePart>
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<mods:namePart id="59737DFDFA61627486BE5B603891C0E6">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="F4B0A025B0307E18FDA5673FECA70DBA">Métais, Grégoire</mods:namePart>
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<mods:namePart id="689C607E854CB346110A19EB539C3F36">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="E74DAAB0DD457BB668BBAC0C79D0C5B3">Lashari, Rafiq A.</mods:namePart>
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<mods:namePart id="7D8544721014E51AA25E50895487C636">Brohi, Imdad A.</mods:namePart>
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<mods:namePart id="4415E0477F548368F6FC78E5B83AA75D">Solangi, Sarfraz H.</mods:namePart>
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<mods:namePart id="EA4115352C6C5026C97A4E99DFEB5B46">Marivaux, Laurent</mods:namePart>
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<mods:namePart id="191EBB814FC6F6B19E1EE16E0575E67C">Welcomme, Jean-Loup</mods:namePart>
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<mods:date id="5DFD0E4E4F7900459E7CEA9174D9BF8C">2014</mods:date>
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<treatment id="03A087E00E52FFA9FF7FFC16FE54F87D" ID-DOI="http://doi.org/10.5281/zenodo.6137862" ID-GBIF-Taxon="119509272" ID-Zenodo-Dep="6137862" LSID="urn:lsid:zoobank.org:act:47584485-3B39-4E1B-BF7F-DF422771B498" httpUri="http://treatment.plazi.org/id/03A087E00E52FFA9FF7FFC16FE54F87D" lastPageNumber="128" pageId="27" pageNumber="128">
<subSubSection id="C313657D0E52FFA9FF7FFC16FE38FBB9" box="[151,482,1015,1042]" pageId="27" pageNumber="128" type="nomenclature">
<paragraph id="8BB636F60E52FFA9FF7FFC16FE38FBB9" blockId="27.[151,482,1015,1042]" box="[151,482,1015,1042]" pageId="27" pageNumber="128">
<heading id="D0FE819A0E52FFA9FF7FFC16FE38FBB9" bold="true" box="[151,482,1015,1042]" fontSize="11" level="1" pageId="27" pageNumber="128" reason="1">
<emphasis id="B97DEAE40E52FFA9FF7FFC16FE38FBB9" bold="true" box="[151,482,1015,1042]" pageId="27" pageNumber="128">
Genus
<taxonomicName id="4C094D750E52FFA9FF06FC19FEB7FBBA" authority="Merle et al., 2014" box="[238,365,1015,1041]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FF06FC19FEB7FBBA" bold="true" box="[238,365,1015,1041]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
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gen. nov.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C313657D0E52FFA9FF7FFBD0FBB4FBFF" box="[151,1134,1086,1108]" pageId="27" pageNumber="128" type="materials_examined">
<paragraph id="8BB636F60E52FFA9FF7FFBD0FBB4FBFF" blockId="27.[151,1134,1086,1108]" box="[151,1134,1086,1108]" pageId="27" pageNumber="128">
<emphasis id="B97DEAE40E52FFA9FF7FFBD0FEF8FBFF" bold="true" box="[151,290,1086,1108]" pageId="27" pageNumber="128">
<typeStatus id="54B288540E52FFA9FF7FFBD0FF16FBFF" box="[151,204,1086,1108]" pageId="27" pageNumber="128">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C094D750E52FFA9FEC1FBD0FDD7FBFF" box="[297,525,1086,1108]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="lakhraensis">
<emphasis id="B97DEAE40E52FFA9FEC1FBD0FDD7FBFF" box="[297,525,1086,1108]" italics="true" pageId="27" pageNumber="128">Sindhiluta lakhraensis</emphasis>
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nov. sp. Early Eocene, Lakhra Formation (Sindh,
<collectingCountry id="F31E76660E52FFA9FBE0FBD0FBB9FBFF" box="[1032,1123,1086,1108]" name="Pakistan" pageId="27" pageNumber="128">Pakistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E52FFA9FF7FFB6FFA99FB31" box="[151,1347,1153,1178]" pageId="27" pageNumber="128" type="etymology">
<paragraph id="8BB636F60E52FFA9FF7FFB6FFA99FB31" blockId="27.[151,1437,1153,2006]" box="[151,1347,1153,1178]" pageId="27" pageNumber="128">
<emphasis id="B97DEAE40E52FFA9FF7FFB6FFEC4FB31" bold="true" box="[151,286,1153,1178]" pageId="27" pageNumber="128">Etymology.</emphasis>
From Sindh (province of
<collectingCountry id="F31E76660E52FFA9FDA9FB6FFD78FB31" box="[577,674,1153,1178]" name="Pakistan" pageId="27" pageNumber="128">Pakistan</collectingCountry>
) and
<emphasis id="B97DEAE40E52FFA9FD37FB6FFCD1FB31" box="[735,779,1153,1178]" italics="true" pageId="27" pageNumber="128">luta</emphasis>
, the two last syllables of
<taxonomicName id="4C094D750E52FFA9FBC0FB6FFBB5FB31" box="[1064,1135,1153,1178]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FBC0FB6FFBB5FB31" box="[1064,1135,1153,1178]" italics="true" pageId="27" pageNumber="128">Voluta</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E52FFA9FF2FFB4BFAEBFAD9" pageId="27" pageNumber="128" type="diagnosis">
<paragraph id="8BB636F60E52FFA9FF2FFB4BFABDFAAD" blockId="27.[151,1437,1153,2006]" pageId="27" pageNumber="128">
<emphasis id="B97DEAE40E52FFA9FF2FFB4BFE9BFB15" bold="true" box="[199,321,1189,1214]" pageId="27" pageNumber="128">Diagnosis.</emphasis>
Shape elongate, high-spired. First teleoconch whorls subcarinate; last whorl smooth, poorly terraced at shoulder, not excavated anteriorly. Costae on spire rounded, and orthocline, fading out progressively over last whorl. Outer lip not thickened externally. Four strong, oblique columellar folds. Parietal callus thick.
</paragraph>
<paragraph id="8BB636F60E52FFA9FF2FFAFFFAEBFAD9" blockId="27.[151,1437,1153,2006]" pageId="27" pageNumber="128">
<emphasis id="B97DEAE40E52FFA9FF2FFAFFFE4BFA81" bold="true" box="[199,401,1297,1322]" pageId="27" pageNumber="128">Included species.</emphasis>
<taxonomicName id="4C094D750E52FFA9FE7FFAFFFD48FA81" box="[407,658,1297,1322]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="lakhraensis" status="sp. nov.">
<emphasis id="B97DEAE40E52FFA9FE7FFAFFFD48FA81" box="[407,658,1297,1322]" italics="true" pageId="27" pageNumber="128">Sindhiluta lakhraensis</emphasis>
</taxonomicName>
<emphasis id="B97DEAE40E52FFA9FD71FAFFFD29FA81" bold="true" box="[665,755,1297,1322]" pageId="27" pageNumber="128">sp. nov.</emphasis>
(Lakhra Formation, Ypresian, Sindh Province,
<collectingCountry id="F31E76660E52FFA9FAECFAFFFABFFA81" box="[1284,1381,1297,1322]" name="Pakistan" pageId="27" pageNumber="128">Pakistan</collectingCountry>
) and
<taxonomicName id="4C094D750E52FFA9FF7FFADBFE20FAE5" authority="Rouault, 1850" authorityName="Rouault" authorityYear="1850" box="[151,506,1333,1358]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E52FFA9FF7FFADBFE9FFAE6" box="[151,325,1333,1358]" italics="true" pageId="27" pageNumber="128">Voluta prevosti</emphasis>
Rouault, 1850
</taxonomicName>
[=
<taxonomicName id="4C094D750E52FFA9FDD8FADBFBA8FAE5" authority="Doncieux, 1908" authorityName="Doncieux" authorityYear="1908" box="[560,1138,1333,1358]" class="Gastropoda" family="Volutidae" genus="Psephaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="atacica" subGenus="Eopsephaea">
<emphasis id="B97DEAE40E52FFA9FDD8FADBFC72FAE6" box="[560,936,1333,1358]" italics="true" pageId="27" pageNumber="128">Psephaea (Eopsephaea) atacica</emphasis>
<bibRefCitation id="EF984B070E52FFA9FC5DFADBFBA8FAE5" author="Doncieux" box="[949,1138,1333,1358]" firstAuthor="Doncieux" pageId="27" pageNumber="128" refString="Doncieux, L. (1908) Catalogue descriptif des fossiles nummulitiques de l'Aude et de l'Herault. 2 eme partie: Corbieres septentrionales. Annales de l'Universite de Lyon, n. ser. I, 22, 1 - 250." type="journal article" year="1908">Doncieux, 1908</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C094D750E52FFA9FB6DFADBFE1DFAD9" authority="Doncieux, 1908" authorityName="Doncieux" authorityYear="1908" class="Gastropoda" family="Volutidae" genus="Psephaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="rabetensis" subGenus="Eopsephaea">
<emphasis id="B97DEAE40E52FFA9FB6DFADBFED0FAD9" italics="true" pageId="27" pageNumber="128">Psephaea (Eopsephaea) rabetensis</emphasis>
<bibRefCitation id="EF984B070E52FFA9FEF9FAB7FE1DFAD9" author="Doncieux" box="[273,455,1369,1394]" firstAuthor="Doncieux" pageId="27" pageNumber="128" refString="Doncieux, L. (1908) Catalogue descriptif des fossiles nummulitiques de l'Aude et de l'Herault. 2 eme partie: Corbieres septentrionales. Annales de l'Universite de Lyon, n. ser. I, 22, 1 - 250." type="journal article" year="1908">Doncieux, 1908</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C094D750E52FFA9FE3BFAB7FC52FAD9" authority="Cossmann, 1923" authorityName="Cossmann" authorityYear="1923" box="[467,904,1369,1394]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="ogormani">
<emphasis id="B97DEAE40E52FFA9FE3BFAB7FD1EFADA" box="[467,708,1369,1394]" italics="true" pageId="27" pageNumber="128">Volutilithes ogormani</emphasis>
Cossmann, 1923
</taxonomicName>
] (Ypresian, Aquitaine Basin,
<collectingCountry id="F31E76660E52FFA9FB3BFAB4FAFFFAD9" box="[1235,1317,1370,1394]" name="France" pageId="27" pageNumber="128">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E52FFA9FF2FFA93FE54F87D" pageId="27" pageNumber="128" type="discussion">
<paragraph id="8BB636F60E52FFA9FF2FFA93FE54F87D" blockId="27.[151,1437,1153,2006]" pageId="27" pageNumber="128">
<emphasis id="B97DEAE40E52FFA9FF2FFA93FE96FA3D" bold="true" box="[199,332,1405,1430]" pageId="27" pageNumber="128">Discussion.</emphasis>
<bibRefCitation id="EF984B070E52FFA9FEB1FA93FD4BFA3D" author="Pilsbry and Olsson" box="[345,657,1405,1430]" firstAuthor="Pilsbry" pageId="27" pageNumber="128" refString="Pilsbry, H. A. &amp; Olsson A. A. (1954) Systems of the Volutidae. Bulletins of American Paleontology, 35 (152), 1 - 307." type="journal article" year="1954">Pilsbry and Olsson (1954)</bibRefCitation>
when introducing the new subfamily
<taxonomicName id="4C094D750E52FFA9FBB7FA93FB2DFA3D" box="[1119,1271,1405,1430]" class="Gastropoda" family="Volutidae" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="subFamily" subFamily="Volutilithinae">Volutilithinae</taxonomicName>
included two genera:
<taxonomicName id="4C094D750E52FFA9FF18FA4FFEB4FA11" box="[240,366,1441,1466]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FF18FA4FFEB4FA11" box="[240,366,1441,1466]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D750E52FFA9FE4CFA4CFD71FA11" authority="Conrad, 1855" authorityName="Conrad" authorityYear="1855" box="[420,683,1441,1466]" class="Gastropoda" family="Volutidae" genus="Lapparia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FE4CFA4CFDD6FA12" box="[420,524,1442,1465]" italics="true" pageId="27" pageNumber="128">Lapparia</emphasis>
<bibRefCitation id="EF984B070E52FFA9FDFBFA4FFD71FA11" author="Conrad" box="[531,683,1441,1466]" firstAuthor="Conrad" pageId="27" pageNumber="128" refString="Conrad, T. A. (1855) Description of one Tertiary and eight new Cretaceous fossils from Texas. Proceedings of the Academy of Natural Sciences of Philadelphia, 7, 268 - 269." type="journal article" year="1855">Conrad, 1855</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54B288540E52FFA9FD52FA4CFD36FA11" box="[698,748,1442,1466]" pageId="27" pageNumber="128">type</typeStatus>
species:
<taxonomicName id="4C094D750E52FFA9FCB9FA4CFB4BFA11" authority="Conrad, 1854" authorityName="Conrad" authorityYear="1854" box="[849,1169,1441,1466]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="dumosa">
<emphasis id="B97DEAE40E52FFA9FCB9FA4CFC35FA11" box="[849,1007,1441,1466]" italics="true" pageId="27" pageNumber="128">Mitra dumosa</emphasis>
Conrad, 1854
</taxonomicName>
, Middle Eocene,
<collectingCountry id="F31E76660E52FFA9FABCFA4CFA4BFA11" box="[1364,1425,1442,1466]" name="United States of America" pageId="27" pageNumber="128">USA</collectingCountry>
). Subsequent authors such as
<bibRefCitation id="EF984B070E52FFA9FE27FA2BFCD2FA75" author="Pacaud &amp; Le Renard" box="[463,776,1477,1502]" firstAuthor="Pacaud" pageId="27" pageNumber="128" pagination="151 - 187" refId="ref30075" refString="Pacaud, J. - M. &amp; Le Renard, J. (1996) Revision des mollusques paleogenes du Bassin de Paris (IV, Liste systematique reactualisee). Cossmanniana, 3 (4), 151 - 187." type="journal article" year="1996">Pacaud &amp; Le Renard (1996)</bibRefCitation>
and
<bibRefCitation id="EF984B070E52FFA9FCD6FA2BFBCAFA75" author="Merle" box="[830,1040,1477,1502]" etAl="et al." firstAuthor="Merle" pageId="27" pageNumber="128" pagination="182 - 227" refId="ref29876" refString="Merle, D., Pacaud, J. - M., Kriloff, A. &amp; Loubry, P. (2008) Les motifs colores residuels des coquilles lutetiennes du bassin de Paris. In: Merle, D. (Ed.), Stratotype Lutetien. Collection Patrimoine geologique, Biotope, Meze, MNHN, Paris, BRGM Orleans, pp. 182 - 227." type="book chapter" year="2008">
Merle
<emphasis id="B97DEAE40E52FFA9FC6FFA28FC65FA75" box="[903,959,1477,1502]" italics="true" pageId="27" pageNumber="128">et al.</emphasis>
(2008)
</bibRefCitation>
also included
<emphasis id="B97DEAE40E52FFA9FB58FA2BFA8AFA75" box="[1200,1360,1477,1502]" italics="true" pageId="27" pageNumber="128">Pseudaulicina</emphasis>
in this subfamily.
<taxonomicName id="4C094D750E52FFA9FEF0FA07FE51F9A9" box="[280,395,1513,1538]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FEF0FA07FE51F9A9" box="[280,395,1513,1538]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
shares characters of the axial sculpture with
<taxonomicName id="4C094D750E52FFA9FC43FA07FBF3F9A9" box="[939,1065,1513,1538]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FC43FA07FBF3F9A9" box="[939,1065,1513,1538]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
and columellar characters with
<emphasis id="B97DEAE40E52FFA9FF7FF9E3FEEDF98D" box="[151,311,1549,1574]" italics="true" pageId="27" pageNumber="128">Pseudaulicina</emphasis>
and
<taxonomicName id="4C094D750E52FFA9FE87F9E0FE0DF98E" box="[367,471,1550,1573]" class="Gastropoda" family="Volutidae" genus="Lapparia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FE87F9E0FE0DF98E" box="[367,471,1550,1573]" italics="true" pageId="27" pageNumber="128">Lapparia</emphasis>
</taxonomicName>
and therefore is assigned to the
<taxonomicName id="4C094D750E52FFA9FCAEF9E3FC38F98D" box="[838,994,1549,1574]" class="Gastropoda" family="Volutidae" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="subFamily" subFamily="Volutilithinae">Volutilithinae</taxonomicName>
. Rounded and subcarinate costae and a lack of spiral sculpture are characters usually found in species of
<taxonomicName id="4C094D750E52FFA9FC73F9DFFBC3F9E1" box="[923,1049,1585,1610]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FC73F9DFFBC3F9E1" box="[923,1049,1585,1610]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
, such as:
<taxonomicName id="4C094D750E52FFA9FB7BF9DCFF02F9C5" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="angustus">
<emphasis id="B97DEAE40E52FFA9FB7BF9DCFACCF9E2" box="[1171,1302,1586,1609]" italics="true" pageId="27" pageNumber="128">V. angustus</emphasis>
(Deshayes, 1835)
</taxonomicName>
from the Ypresian of the Paris Basin,
<taxonomicName id="4C094D750E52FFA9FD66F9B8FBDDF9C5" authority="Rouault, 1850" authorityName="Rouault" authorityYear="1850" box="[654,1031,1621,1646]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="deshayesianus">
<emphasis id="B97DEAE40E52FFA9FD66F9B8FC97F9C5" box="[654,845,1621,1646]" italics="true" pageId="27" pageNumber="128">V. deshayesianus</emphasis>
(Rouault, 1850)
</taxonomicName>
from the Ypresian of the Aquitaine Basin, and
<taxonomicName id="4C094D750E52FFA9FEFFF994FDB6F939" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" box="[279,620,1657,1682]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="species" species="torulosus">
<emphasis id="B97DEAE40E52FFA9FEFFF994FE46F939" box="[279,412,1657,1682]" italics="true" pageId="27" pageNumber="128">V. torulosus</emphasis>
(Deshayes, 1835)
</taxonomicName>
and
<emphasis id="B97DEAE40E52FFA9FD4DF994FCFCF93A" box="[677,806,1658,1681]" italics="true" pageId="27" pageNumber="128">V. c os t ar i u s</emphasis>
(
<bibRefCitation id="EF984B070E52FFA9FCD1F997FC3DF939" author="Lamarck" box="[825,999,1657,1682]" firstAuthor="Lamarck" pageId="27" pageNumber="128" refString="Lamarck, J. - B. (1803) Memoires sur les fossiles des environs de Paris comprenant la determination des especes qui appartiennent aux animaux marins sans vertebres, et dont la plupart sont figures dans la collection des velins du Museum. Annales du Museum d'Histoire naturelle, 1, 474 - 479." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
) from the Lutetian of the Paris Basin. However,
<taxonomicName id="4C094D750E52FFA9FEE3F973FEA4F91D" box="[267,382,1693,1718]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FEE3F973FEA4F91D" box="[267,382,1693,1718]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
differs obviously from
<taxonomicName id="4C094D750E52FFA9FD64F973FCD0F91D" box="[652,778,1693,1718]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FD64F973FCD0F91D" box="[652,778,1693,1718]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
in two characters. First,
<taxonomicName id="4C094D750E52FFA9FBCCF973FB4DF91D" box="[1060,1175,1693,1718]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FBCCF973FB4DF91D" box="[1060,1175,1693,1718]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
lacks costae on its last whorls, whereas they are present on
<taxonomicName id="4C094D750E52FFA9FDAEF92FFD1EF971" box="[582,708,1729,1754]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FDAEF92FFD1EF971" box="[582,708,1729,1754]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
. Second,
<taxonomicName id="4C094D750E52FFA9FCD3F92FFC74F971" box="[827,942,1729,1754]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FCD3F92FFC74F971" box="[827,942,1729,1754]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
has four strong columellar folds, whereas members of
<taxonomicName id="4C094D750E52FFA9FEC3F90BFE73F955" box="[299,425,1765,1790]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FEC3F90BFE73F955" box="[299,425,1765,1790]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
have only two or three weaker columellar folds. The number and development of the columellar folds are more nearly similar to those of
<emphasis id="B97DEAE40E52FFA9FCC1F8E7FC13F889" box="[809,969,1801,1826]" italics="true" pageId="27" pageNumber="128">Pseudaulicina</emphasis>
than to
<taxonomicName id="4C094D750E52FFA9FBD6F8E7FB66F889" box="[1086,1212,1801,1826]" class="Gastropoda" family="Volutidae" genus="Volutilithes" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FBD6F8E7FB66F889" box="[1086,1212,1801,1826]" italics="true" pageId="27" pageNumber="128">Volutilithes</emphasis>
</taxonomicName>
. Nevertheless, the persistence of the costae on the last whorls, associated with variably developed shoulder spines, an excavated base and the presence of spiral sculpture in most species of
<emphasis id="B97DEAE40E52FFA9FCC8F8BFFC1AF8C1" box="[800,960,1873,1898]" italics="true" pageId="27" pageNumber="128">Pseudaulicina</emphasis>
contrast greatly with the morphology of
<taxonomicName id="4C094D750E52FFA9FF7FF89BFED0F825" box="[151,266,1909,1934]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FF7FF89BFED0F825" box="[151,266,1909,1934]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
.
<collectingCountry id="F31E76660E52FFA9FEF2F898FEE3F825" box="[282,313,1910,1934]" name="American Samoa" pageId="27" pageNumber="128">As</collectingCountry>
in
<taxonomicName id="4C094D750E52FFA9FE89F89BFE0EF825" box="[353,468,1909,1934]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E52FFA9FE89F89BFE0EF825" box="[353,468,1909,1934]" italics="true" pageId="27" pageNumber="128">Sindhiluta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C094D750E52FFA9FE0CF898FD96F826" box="[484,588,1910,1933]" class="Gastropoda" family="Volutidae" genus="Lapparia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="128" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E52FFA9FE0CF898FD96F826" box="[484,588,1910,1933]" italics="true" pageId="27" pageNumber="128">Lapparia</emphasis>
</taxonomicName>
has four strong columellar folds, the most anterior being weaker, but it is distinguished by its axial sculpture, which continues to persist down the shell and finishes with the appearance of acute shoulder spines.
</paragraph>
</subSubSection>
</treatment>
</document>