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<document id="3275D3B67C5A81101051AD8BC068C2EA" ID-CLB-Dataset="36817" ID-DOI="10.11646/zootaxa.3835.1.1" ID-GBIF-Dataset="7d11dc81-2b61-40fb-8eb1-fcdfb1547668" ID-ISSN="1175-5326" ID-Zenodo-Dep="286701" ID-ZooBank="5BBF2D80-F682-49E7-9B32-64E75ACAA6CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460455174602" checkinUser="plazi" docAuthor="Sartori, Michel" docDate="2014" docId="03A087A0433B4A14FF7AEABA95D35ACB" docLanguage="en" docName="zt03835p032.pdf" docOrigin="Zootaxa 3835 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Compsoneuria lieftincki Ulmer 1939" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="FF99FFD843314A1FFFEDE915942F5F7D" masterDocTitle="The concept of Compsoneuria Eaton, 1881 revisited in light of historical and new material from the Sunda Islands (Ephemeroptera: Heptageniidae: Ecdyonurinae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="11" updateTime="1698338546610" updateUser="ExternalLinkService">
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<mods:title id="474460CF45C0AC33BF7C882F9C8B6044">The concept of Compsoneuria Eaton, 1881 revisited in light of historical and new material from the Sunda Islands (Ephemeroptera: Heptageniidae: Ecdyonurinae)</mods:title>
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<treatment id="03A087A0433B4A14FF7AEABA95D35ACB" ID-DOI="http://doi.org/10.5281/zenodo.6124745" ID-GBIF-Taxon="127668968" ID-Zenodo-Dep="6124745" LSID="urn:lsid:plazi:treatment:03A087A0433B4A14FF7AEABA95D35ACB" httpUri="http://treatment.plazi.org/id/03A087A0433B4A14FF7AEABA95D35ACB" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="C313653D433B4A15FF7AEABA96445CB7" box="[151,619,943,970]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BB636B6433B4A15FF7AEABA96445CB7" blockId="10.[151,619,943,970]" box="[151,619,943,970]" pageId="10" pageNumber="11">
<heading id="D0FE81DA433B4A15FF7AEABA96445CB7" bold="true" box="[151,619,943,970]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C094D35433B4A15FF7AEABA96445CB7" ID-CoL="XHZV" authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" box="[151,619,943,970]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FF7AEABA96445CB7" bold="true" box="[151,619,943,970]" pageId="10" pageNumber="11">
<emphasis id="B97DEAA4433B4A15FF7AEABA95945CB4" bold="true" box="[151,443,943,969]" italics="true" pageId="10" pageNumber="11">Compsoneuria lieftincki</emphasis>
(
<bibRefCitation id="EF984B47433B4A15FE27EAA5964C5CB7" author="Ulmer, G." box="[458,611,944,970]" firstAuthor="Ulmer" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer, 1939</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313653D433B4A15FF7AEAED97355B56" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BB636B6433B4A15FF7AEAED904D5B71" blockId="10.[151,1122,1014,1067]" box="[151,1122,1014,1037]" pageId="10" pageNumber="11">
<taxonomicName id="4C094D35433B4A15FF7AEAED95DC5B71" authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" box="[151,499,1014,1037]" class="Insecta" family="Heptageniidae" genus="Heptagenia" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FF7AEAED95435B71" box="[151,364,1014,1037]" italics="true" pageId="10" pageNumber="11">Heptagenia lieftincki</emphasis>
<bibRefCitation id="EF984B47433B4A15FE9FEAE395DC5B71" author="Ulmer, G." box="[370,499,1014,1036]" firstAuthor="Ulmer" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer, 1939</bibRefCitation>
</taxonomicName>
, Arch. Hydrobiol. Suppl. 16, p. 571, male and female imagos
</paragraph>
<paragraph id="8BB636B6433B4A15FF7AED0397355B56" blockId="10.[151,1122,1014,1067]" box="[151,794,1045,1067]" pageId="10" pageNumber="11">
<taxonomicName id="4C094D35433B4A15FF7AED03966B5B56" ID-CoL="XHZV" authority="Webb et al., 2006" authorityName="Webb et al." authorityYear="2006" box="[151,580,1045,1067]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FF7AED0395A45B56" box="[151,395,1045,1067]" italics="true" pageId="10" pageNumber="11">Compsoneuria lieftincki</emphasis>
<bibRefCitation id="EF984B47433B4A15FE7CED00966B5B56" author="Webb, J. M. &amp; Braasch, D. &amp; McCafferty, W. P." box="[401,580,1045,1067]" etAl="et al." firstAuthor="Webb" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="55 - 68" part="1335" refId="ref16952" refString="Webb, J. M., Braasch, D. &amp; McCafferty, W. P. (2006) Reevaluation of the genera Compsoneuria Eaton and Trichogenia Braasch &amp; Soldan (Ephemeroptera: Heptageniidae). Zootaxa, 1335, 55 - 68" title="Reevaluation of the genera Compsoneuria Eaton and Trichogenia Braasch &amp; Soldan (Ephemeroptera: Heptageniidae)" type="journal article" year="2006">Webb et al., 2006</bibRefCitation>
</taxonomicName>
, Zootaxa 1335, p. 59
</paragraph>
</subSubSection>
<subSubSection id="C313653D433B4A15FF7AED4294C05BCA" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BB636B6433B4A15FF7AED4294C05BCA" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">
<emphasis id="B97DEAA4433B4A15FF7AED4295A85B0D" bold="true" box="[151,391,1111,1136]" pageId="10" pageNumber="11">Material examined.</emphasis>
<materialsCitation id="3B613CEB433B4A15FE78ED4D94C05BCA" ID-GBIF-Occurrence="4037950312" collectingDate="1935-07-06" collectionCode="ZMH" collectorName="Lieftinck" country="Indonesia" elevation="150" location="Tjibarangbang River" municipality="Djasinga" pageId="10" pageNumber="11" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="West Java" typeStatus="syntype">
<specimenCount id="9D0FFD3F433B4A15FE78ED4D963A5B0D" box="[405,533,1111,1136]" pageId="10" pageNumber="11">Two males</specimenCount>
and
<specimenCount id="9D0FFD3F433B4A15FDB7ED4D96F35B0D" box="[602,732,1111,1136]" pageId="10" pageNumber="11" type="female">one female</specimenCount>
<typeStatus id="54B28814433B4A15FD07ED4D97635B0D" box="[746,844,1112,1136]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
with subimaginal exuviae,
<collectingCountry id="F31E7626433B4A15FB71ED4291235B0D" box="[1180,1292,1111,1136]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
,
<collectingRegion id="49CDF854433B4A15FAF3ED4D91B85B12" box="[1310,1431,1112,1135]" country="Indonesia" name="Jawa Barat" pageId="10" pageNumber="11">West Java</collectingRegion>
,
<collectingMunicipality id="6BD2ACCC433B4A15FF7AED6994D15BE8" box="[151,254,1148,1173]" pageId="10" pageNumber="11">Djasinga</collectingMunicipality>
,
<location id="8ED6606D433B4A15FEE1ED6995D65BE8" LSID="urn:lsid:plazi:treatment:03A087A0433B4A14FF7AEABA95D35ACB:8ED6606D433B4A15FEE1ED6995D65BE8" box="[268,505,1148,1173]" country="Indonesia" municipality="Djasinga" name="Tjibarangbang River" pageId="10" pageNumber="11" stateProvince="West Java">Tjibarangbang River</location>
,
<quantity id="4CF19B53433B4A15FDEAED6996625BE8" box="[519,589,1148,1173]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="m" value="150.0">
<elevation id="0024D185433B4A15FDEAED6996625BE8" box="[519,589,1148,1173]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="m" value="150.0">150m</elevation>
</quantity>
., caught at light in the evening above the river,
<date id="FFB71076433B4A15FB90ED6990D55BE8" box="[1149,1274,1148,1173]" pageId="10" pageNumber="11" value="1935-07-06">
<collectingDate id="EFF3E99E433B4A15FB90ED6990D55BE8" box="[1149,1274,1148,1173]" pageId="10" pageNumber="11" value="1935-07-06">6.VII.1935</collectingDate>
</date>
,
<collectorName id="26FC5360433B4A15FAE4ED69915F5BE8" box="[1289,1392,1148,1173]" pageId="10" pageNumber="11">Lieftinck</collectorName>
leg [
<collectionCode id="ED18AE73433B4A15FF73EDB594CA5BCA" box="[158,229,1184,1207]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="10" pageNumber="11">ZMH</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C313653D433B4A15FF2AEDD195C859F0" pageId="10" pageNumber="11" type="description">
<paragraph id="8BB636B6433B4A15FF2AEDD195765A7D" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">
<emphasis id="B97DEAA4433B4A15FF2AEDD195765BA0" bold="true" box="[199,345,1220,1245]" pageId="10" pageNumber="11">Description.</emphasis>
<emphasis id="B97DEAA4433B4A15FE8CEDD195C55BA1" box="[353,490,1220,1245]" italics="true" pageId="10" pageNumber="11">Male imago</emphasis>
. (Note: Only significant characters are mentioned here; for a complete description see
<bibRefCitation id="EF984B47433B4A15FF2FEDF295635A7D" author="Ulmer, G." box="[194,332,1255,1280]" firstAuthor="Ulmer" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer 1939</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636B6433B4A15FF2AEC1997D55A35" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">All specimens faded. Forewings translucent, except costal and subcostal fields milky. Hindwings entirely colored in light brown, color presumably more pronounced on fresh material.</paragraph>
<paragraph id="8BB636B6433B4A15FF2AEC4190D95A10" blockId="10.[151,1437,1111,2037]" box="[199,1270,1364,1389]" pageId="10" pageNumber="11">
Foreleg similar to
<taxonomicName id="4C094D35433B4A15FE78EC4096025A10" box="[405,557,1364,1389]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B97DEAA4433B4A15FE78EC4096025A10" box="[405,557,1364,1389]" italics="true" pageId="10" pageNumber="11">C. spectabilis</emphasis>
</taxonomicName>
, with first segment of tarsus very short (
<bibRefCitation id="EF984B47433B4A15FC1FEC4190C55A10" author="Ulmer, G." box="[1010,1258,1364,1389]" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer, 1939, fig. 160</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636B6433B4A15FF2AEC6296AE5AA5" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">
Forewing (
<figureCitation id="13322A33433B4A15FEA8EC6295B65AED" box="[325,409,1399,1424]" captionStart="FIGURES 22 24" captionStartId="11.[151,264,1236,1259]" captionTargetBox="[178,1427,205,1201]" captionTargetId="figure@11.[158,1428,193,1216]" captionTargetPageId="11" captionText="FIGURES 22 24. Compsoneuria lieftincki (Ulmer), male imago. Fig. 22: Forewing; Fig. 23: Genitalia; Fig. 24: Penis structure in ventral (left) and dorsal (right) view." httpUri="https://zenodo.org/record/286706/files/figure.png" pageId="10" pageNumber="11">Fig. 22</figureCitation>
) with 12 crossveins in costal and subcostal fields; row of crossveins readily apparent in middle of radial-median field, crossveins on distal part of radial field less distinct. Overall number of crossveins greater than number in other species (&gt; 90).
</paragraph>
<paragraph id="8BB636B6433B4A15FF2AECF197FD5915" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">
Gonopods, styliger plate and genitalia characteristic of genus (
<figureCitation id="13322A33433B4A15FC69ECF197F75A81" box="[900,984,1508,1533]" captionStart="FIGURES 22 24" captionStartId="11.[151,264,1236,1259]" captionTargetBox="[178,1427,205,1201]" captionTargetId="figure@11.[158,1428,193,1216]" captionTargetPageId="11" captionText="FIGURES 22 24. Compsoneuria lieftincki (Ulmer), male imago. Fig. 22: Forewing; Fig. 23: Genitalia; Fig. 24: Penis structure in ventral (left) and dorsal (right) view." httpUri="https://zenodo.org/record/286706/files/figure.png" pageId="10" pageNumber="11">Fig. 23</figureCitation>
). Penis lobes (
<figureCitation id="13322A33433B4A15FB96ECF190E25A81" box="[1147,1229,1508,1533]" captionStart="FIGURES 22 24" captionStartId="11.[151,264,1236,1259]" captionTargetBox="[178,1427,205,1201]" captionTargetId="figure@11.[158,1428,193,1216]" captionTargetPageId="11" captionText="FIGURES 22 24. Compsoneuria lieftincki (Ulmer), male imago. Fig. 22: Forewing; Fig. 23: Genitalia; Fig. 24: Penis structure in ventral (left) and dorsal (right) view." httpUri="https://zenodo.org/record/286706/files/figure.png" pageId="10" pageNumber="11">Fig. 24</figureCitation>
) with small apical spine at base of well-developed apical sclerite, protruding outwards slightly, giving apex of lobe more concave appearance than found in other species; subapical spine and titillators slender; basal sclerite not easily observed, with only acute and long spine visible; lateral sclerite broad and quadrate.
</paragraph>
<paragraph id="8BB636B6433B4A15FF2AEF6195C859F0" blockId="10.[151,1437,1111,2037]" box="[199,487,1652,1677]" pageId="10" pageNumber="11">Nymphal stage unknown.</paragraph>
</subSubSection>
<subSubSection id="C313653D433B4A14FF2AEF8295D35ACB" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BB636B6433B4A15FF2AEF8297C95888" blockId="10.[151,1437,1111,2037]" pageId="10" pageNumber="11">
<emphasis id="B97DEAA4433B4A15FF2AEF82956359CD" bold="true" box="[199,332,1687,1712]" pageId="10" pageNumber="11">Discussion.</emphasis>
<bibRefCitation id="EF984B47433B4A15FEB5EF8295D959CD" author="Ulmer, G." box="[344,502,1687,1712]" firstAuthor="Ulmer" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer (1939)</bibRefCitation>
described this species in the genus
<taxonomicName id="4C094D35433B4A15FC44EF8C90E459CD" authority="Walsh, 1863" authorityName="Walsh" authorityYear="1863" box="[937,1227,1687,1712]" class="Insecta" family="Heptageniidae" genus="Heptagenia" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAA4433B4A15FC44EF8C900359CD" box="[937,1068,1689,1712]" italics="true" pageId="10" pageNumber="11">Heptagenia</emphasis>
Walsh, 1863
</taxonomicName>
, because the first segment of the hind tarsus is not longer than the second. As illustrated by
<bibRefCitation id="EF984B47433B4A15FC3DEFA990FF59A8" author="Ulmer, G." box="[976,1232,1724,1749]" journalOrPublisher="Archiv fur Hydrobiologie" pageId="10" pageNumber="11" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939">Ulmer (1939, fig. 161)</bibRefCitation>
, this first segment is subequal in length to the second one in
<taxonomicName id="4C094D35433B4A15FD6BEFF497225985" box="[646,781,1759,1784]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FD6BEFF497225985" box="[646,781,1759,1784]" italics="true" pageId="10" pageNumber="11">H. lieftincki</emphasis>
</taxonomicName>
. The presence of a transverse suture on the mesonotum, together with the median depression of the furcasterum of mesothorax subparallel clearly indicates that
<taxonomicName id="4C094D35433B4A15FAF7EE1091B35860" box="[1306,1436,1796,1821]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FAF7EE1091B35860" box="[1306,1436,1796,1821]" italics="true" pageId="10" pageNumber="11">C. lieftincki</emphasis>
</taxonomicName>
belongs to the subfamily Ecdyonurinae and therefore cannot be a member of the Heptageniinae where belongs the genus
<taxonomicName id="4C094D35433B4A15FF32EE58954D5819" box="[223,354,1869,1892]" class="Insecta" family="Heptageniidae" genus="Heptagenia" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAA4433B4A15FF32EE58954D5819" box="[223,354,1869,1892]" italics="true" pageId="10" pageNumber="11">Heptagenia</emphasis>
</taxonomicName>
. Consequently, the combination proposed by
<bibRefCitation id="EF984B47433B4A15FC8DEE5990005818" author="Webb, J. M. &amp; Braasch, D. &amp; McCafferty, W. P." box="[864,1071,1868,1893]" etAl="et al." firstAuthor="Webb" journalOrPublisher="Zootaxa" pageId="10" pageNumber="11" pagination="55 - 68" part="1335" refId="ref16952" refString="Webb, J. M., Braasch, D. &amp; McCafferty, W. P. (2006) Reevaluation of the genera Compsoneuria Eaton and Trichogenia Braasch &amp; Soldan (Ephemeroptera: Heptageniidae). Zootaxa, 1335, 55 - 68" title="Reevaluation of the genera Compsoneuria Eaton and Trichogenia Braasch &amp; Soldan (Ephemeroptera: Heptageniidae)" type="journal article" year="2006">Webb et al. (2006)</bibRefCitation>
is followed here.
<taxonomicName id="4C094D35433B4A15FB1AEE5894D658F5" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FB1AEE5894D658F5" italics="true" pageId="10" pageNumber="11">Compsoneuria lieftincki</emphasis>
</taxonomicName>
possesses genitalia in accordance with the actual concept of
<taxonomicName id="4C094D35433B4A15FC46EE65907F58FA" box="[939,1104,1904,1927]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B97DEAA4433B4A15FC46EE65907F58FA" box="[939,1104,1904,1927]" italics="true" pageId="10" pageNumber="11">Compsoneuria</emphasis>
</taxonomicName>
, but the forewing venation is intriguing since it possesses many more crossveins than in
<taxonomicName id="4C094D35433B4A15FCAAEE8097CC58D0" box="[839,995,1940,1965]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B97DEAA4433B4A15FCAAEE8097CC58D0" box="[839,995,1940,1965]" italics="true" pageId="10" pageNumber="11">C. spectabilis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C094D35433B4A15FC19EE80912A58D0" box="[1012,1285,1940,1965]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433B4A15FC19EE80912A58D0" box="[1012,1285,1940,1965]" italics="true" pageId="10" pageNumber="11">Compsoneuria lieftincki</emphasis>
</taxonomicName>
exhibits few crossveins in the costal and subcostal fields, as in
<taxonomicName id="4C094D35433B4A15FD27EEAC974C58AC" box="[714,867,1976,2001]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B97DEAA4433B4A15FD27EEAC974C58AC" box="[714,867,1976,2001]" italics="true" pageId="10" pageNumber="11">C. spectabilis</emphasis>
</taxonomicName>
, together with rows of crossveins in the radial and median fields, together with the reduced first tarsal segment of the forelegs.
</paragraph>
<caption id="DF76663E433A4A14FF7AEDC1960C5A77" httpUri="https://zenodo.org/record/286706/files/figure.png" pageId="11" pageNumber="12" targetBox="[178,1427,205,1201]" targetPageId="11">
<paragraph id="8BB636B6433A4A14FF7AEDC1960C5A77" blockId="11.[151,1436,1236,1290]" pageId="11" pageNumber="12">
<emphasis id="B97DEAA4433A4A14FF7AEDC1957C5B97" bold="true" box="[151,339,1236,1259]" pageId="11" pageNumber="12">FIGURES 2224.</emphasis>
<taxonomicName id="4C094D35433A4A14FEB4EDC396895B96" authority="Ulmer" authorityName="Ulmer" box="[345,678,1237,1259]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433A4A14FEB4EDC396625B96" box="[345,589,1237,1259]" italics="true" pageId="11" pageNumber="12">Compsoneuria lieftincki</emphasis>
(Ulmer)
</taxonomicName>
, male imago. Fig. 22: Forewing; Fig. 23: Genitalia; Fig. 24: Penis structure in ventral (left) and dorsal (right) view.
</paragraph>
</caption>
<paragraph id="8BB636B6433A4A14FF2AEC2495D35ACB" blockId="11.[151,1436,1329,1462]" pageId="11" pageNumber="12">
The female of
<taxonomicName id="4C094D35433A4A14FE9EEC2795D75A37" box="[371,504,1329,1354]" class="Insecta" family="Heptageniidae" genus="Compsoneuria" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="B97DEAA4433A4A14FE9EEC2795D75A37" box="[371,504,1329,1354]" italics="true" pageId="11" pageNumber="12">C. lieftincki</emphasis>
</taxonomicName>
described by
<bibRefCitation id="EF984B47433A4A14FD70EC2497BD5A37" author="Ulmer, G." box="[669,914,1329,1354]" journalOrPublisher="Archiv fur Hydrobiologie" pageId="11" pageNumber="12" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939" yearSuffix="p">Ulmer (1939, p. 573)</bibRefCitation>
does not belong to the same species or even genus as the male mentioned here, because of the wing venation with numerous crossveins in the costal and subcostal fields and the hind leg proportions, the latter being already mentioned by
<bibRefCitation id="EF984B47433A4A14FB6AEC6C91575AEF" author="Ulmer, G." box="[1159,1400,1401,1426]" journalOrPublisher="Archiv fur Hydrobiologie" pageId="11" pageNumber="12" pagination="443 - 692" part="16" refId="ref16811" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, 16, 443 - 692." title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln" type="journal article" year="1939" yearSuffix="p">Ulmer (1939, p.574)</bibRefCitation>
as something strange (“auffällig”).
</paragraph>
</subSubSection>
</treatment>
</document>