237 lines
29 KiB
XML
237 lines
29 KiB
XML
<document id="2215968827B9166C586249A23063FB86" ID-DOI="10.5281/zenodo.180740" ID-GBIF-Dataset="727b5424-a42a-493c-ae17-fbad8934319b" ID-ISSN="1175-5326" ID-Zenodo-Dep="180740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459813896422" checkinUser="plazi" docAuthor="Ernst, Raffael, Agyei, Alex C. & Rödel, Mark-Oliver" docDate="2008" docId="03A0878DFF89FF7AA4BFFA5A36696AD9" docLanguage="en" docName="zt01697p068.pdf" docOrigin="Zootaxa 1697" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Arthroleptis krokosua Ernst, Agyei & Rödel, 2008, sp. nov." docType="treatment" docVersion="7" lastPageNumber="62" masterDocId="FF99FFF5FF8BFF7EA428FFD7323E6D70" masterDocTitle="A new giant species of Arthroleptis (Amphibia: Anura: Arthroleptidae) from the Krokosua Hills Forest Reserve, south-western Ghana" masterLastPageNumber="68" masterPageNumber="58" pageNumber="60" updateTime="1698221336504" updateUser="plazi">
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<mods:titleInfo id="27E242CF25F88F90A248E1A06D089434">
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<mods:title id="C5FAA64AC437CBC42EC4F63D285FAC13">A new giant species of Arthroleptis (Amphibia: Anura: Arthroleptidae) from the Krokosua Hills Forest Reserve, south-western Ghana</mods:title>
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<mods:name id="4859EC27171C3AE87AF1FB9300A1710E" type="personal">
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<mods:roleTerm id="25EC1A78A3B4A1880AA4B6D0EE6D7493">Author</mods:roleTerm>
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<mods:namePart id="4F11543490E9537B3A64C23461548D12">Ernst, Raffael</mods:namePart>
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<mods:roleTerm id="D805938F87AE3C5D8C203AAE5F2BBB32">Author</mods:roleTerm>
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<mods:namePart id="1E4E0848A3E84D9D11B4C3DCD0A6C716">Agyei, Alex C.</mods:namePart>
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<mods:name id="4E7A369553FE1257C1D887BC6980E68D" type="personal">
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<mods:roleTerm id="D7C9E1E9CB668A27A0D499B410A0A25B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5CBAB7DAE5748EED848B413F3087131D">Rödel, Mark-Oliver</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="6316105675A52791DDD077F55E2F6237">text</mods:typeOfResource>
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<mods:title id="BDB1F7F1A90AF52B481498D8300A12D7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5BE52B2D4FB085514229A2517CF9F747">
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<mods:date id="8283963AFB19E83F8F8C8B9B87AA8DD7">2008</mods:date>
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<mods:detail id="886DC2B784698C4AF4A42F787D2335C7" type="volume">
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<mods:number id="0B389B9DC2508AB8AECD3303ACB2CC6E">1697</mods:number>
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<mods:classification id="E75CE34356FD2336E57C147278FD2F3F">journal article</mods:classification>
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<mods:identifier id="9ED20ADBD82CDE96ABAC8EFF5ED21FD3" type="DOI">10.5281/zenodo.180740</mods:identifier>
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<mods:identifier id="EB789805902FFDE1D352F48462BDCCF5" type="GBIF-Dataset">727b5424-a42a-493c-ae17-fbad8934319b</mods:identifier>
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<mods:identifier id="49D24B6D9E8CE053B1728D1A71EB86A7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4288BCF4EDC3F12A352F919A2FE921A5" type="Zenodo-Dep">180740</mods:identifier>
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<treatment id="03A0878DFF89FF7AA4BFFA5A36696AD9" ID-DOI="http://doi.org/10.5281/zenodo.6229240" ID-GBIF-Taxon="119354446" ID-Zenodo-Dep="6229240" LSID="urn:lsid:plazi:treatment:03A0878DFF89FF7AA4BFFA5A36696AD9" httpUri="http://treatment.plazi.org/id/03A0878DFF89FF7AA4BFFA5A36696AD9" lastPageId="4" lastPageNumber="62" pageId="2" pageNumber="60">
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<subSubSection id="C3136510FF89FF7CA4BFFA5A32C368BA" pageId="2" pageNumber="60" type="nomenclature">
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<paragraph id="8BB6369BFF89FF7CA4BFFA5A303A68D7" blockId="2.[151,516,1421,1482]" box="[151,516,1421,1447]" pageId="2" pageNumber="60">
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<heading id="D0FE81F7FF89FF7CA4BFFA5A303A68D7" bold="true" box="[151,516,1421,1447]" fontSize="11" level="1" pageId="2" pageNumber="60" reason="1">
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<emphasis id="B97DEA89FF89FF7CA4BFFA5A303A68D7" bold="true" box="[151,516,1421,1447]" pageId="2" pageNumber="60">
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<taxonomicName id="4C094D18FF89FF7CA4BFFA5A33A168D7" ID-CoL="GYR5" box="[151,415,1421,1447]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="60" phylum="Chordata" rank="species" species="krokosua" status="sp. nov.">
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<emphasis id="B97DEA89FF89FF7CA4BFFA5A33A168D7" bold="true" box="[151,415,1421,1447]" italics="true" pageId="2" pageNumber="60">Arthroleptis krokosua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57F2FF89FF7CA58FFA5A303A68D7" box="[423,516,1421,1447]" pageId="2" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB6369BFF89FF7CA4BFFA6732C368BA" blockId="2.[151,516,1421,1482]" box="[151,253,1456,1482]" pageId="2" pageNumber="60">
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<figureCitation id="13322A1EFF89FF7CA4BFFA6732C368BA" box="[151,253,1456,1482]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,254,1005,1029]" captionStartId-1="3.[151,254,1614,1638]" captionStartId-2="4.[151,254,1179,1203]" captionTargetBox-0="[471,1129,193,960]" captionTargetBox-1="[497,1046,1126,1583]" captionTargetBox-2="[550,1036,477,1154]" captionTargetId-0="figure@3.[450,1137,96,981]" captionTargetId-1="figure@3.[494,1093,1124,1591]" captionTargetId-2="figure@4.[550,1036,474,1156]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Dorsal (a), ventral (b), and latero-ventral (c) view of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst" captionText-1="FIGURE 2. Right hand and plantar surface of right foot of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst." captionText-2="FIGURE 3. Arthroleptis krokosua sp. nov. holotype (SMNS 12555) in life. Photographs R. Ernst" httpUri-0="https://zenodo.org/record/180741/files/figure.png" httpUri-1="https://zenodo.org/record/180742/files/figure.png" httpUri-2="https://zenodo.org/record/180743/files/figure.png" pageId="2" pageNumber="60">Figs 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136510FF89FF7CA4BFFA2D37246BFC" pageId="2" pageNumber="60" type="materials_examined">
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<paragraph id="8BB6369BFF89FF7CA4BFFA2D37246BFC" blockId="2.[151,1437,1530,2036]" pageId="2" pageNumber="60">
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<emphasis id="B97DEA89FF89FF7CA4BFFA2D332E6B64" bold="true" box="[151,272,1530,1556]" pageId="2" pageNumber="60">
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<typeStatus id="54B28839FF89FF7CA4BFFA2D33356B64" box="[151,267,1530,1556]" pageId="2" pageNumber="60" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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An adult male (
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<collectionCode id="ED18AE5EFF89FF7CA5FAFA2D301A6B64" LSID="urn:lsid:biocol.org:col:34840" box="[466,548,1530,1556]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="60">SMNS</collectionCode>
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12555; GB:
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<accessionNumber id="945AAB78FF89FF7CA6E8FA2D317C6B64" box="[704,834,1530,1556]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU350211" pageId="2" pageNumber="60">EU350211</accessionNumber>
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; field number KHA0103) collected by A.C. Agyei and R. Ernst on
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<date id="FFB7105BFF89FF7CA57CF9F530166B4C" box="[340,552,1570,1596]" pageId="2" pageNumber="60" value="2003-11-08">8 November 2003</date>
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on a steep hill, SE of the base camp (at village of Mmem), in the
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<taxonomicName id="4C094D18FF89FF7CA102F9F537A26B4C" box="[1322,1436,1570,1596]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="krokosua">Krokosua</taxonomicName>
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Hills Forest Reserve, south western
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<collectingCountry id="F31E760BFF89FF7CA660F99D30AA6B14" box="[584,660,1610,1636]" name="Ghana" pageId="2" pageNumber="60">Ghana</collectingCountry>
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(
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<geoCoordinate id="EE3D505CFF89FF7CA68FF99D31006B14" box="[679,830,1610,1636]" direction="north" orientation="latitude" pageId="2" pageNumber="60" precision="1" value="6.5995">6°35.970’ N</geoCoordinate>
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,
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<geoCoordinate id="EE3D505CFF89FF7CA764F99D31D66B14" box="[844,1000,1610,1636]" direction="west" orientation="longitude" pageId="2" pageNumber="60" precision="1" value="-2.8450832">2°50.705’ W</geoCoordinate>
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; GPS geodetic datum: WGS84; altitude ca.
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<quantity id="4CF19B7EFF89FF7CA4D1F9A5337D6BFC" box="[249,323,1650,1676]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="60" unit="m" value="300.0">300 m</quantity>
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asl). The habitat consisted of dry forest with a thick leaf litter on a granitic outcrop.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136510FF89FF7CA4EDF94D30256A84" pageId="2" pageNumber="60" type="etymology">
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<paragraph id="8BB6369BFF89FF7CA4EDF94D32F26A5C" blockId="2.[151,1437,1530,2036]" pageId="2" pageNumber="60">
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<emphasis id="B97DEA89FF89FF7CA4EDF94D336C6BC4" bold="true" box="[197,338,1690,1716]" pageId="2" pageNumber="60">Etymology.</emphasis>
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The species is named after the
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<typeStatus id="54B28839FF89FF7CA6E6F94D313E6BC4" box="[718,768,1690,1716]" pageId="2" pageNumber="60">type</typeStatus>
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locality
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<taxonomicName id="4C094D18FF89FF7CA742F94D31E26BC4" box="[874,988,1690,1716]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="66" phylum="Chordata" rank="species" species="krokosua">Krokosua</taxonomicName>
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Hills. The name was chosen to highlight the extraordinary importance of the location in the context of the conservation of Globally Significant Biodiversity Areas (GSBA) in the Western Region,
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<collectingCountry id="F31E760BFF89FF7CA6DEF93D31786A74" box="[758,838,1770,1796]" name="Ghana" pageId="2" pageNumber="60">Ghana</collectingCountry>
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. The specific epithet is treated as a noun in apposition.
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</paragraph>
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<paragraph id="8BB6369BFF89FF7CA4EDF8ED30256A84" blockId="2.[151,1437,1530,2036]" pageId="2" pageNumber="60">
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<emphasis id="B97DEA89FF89FF7CA4EDF8ED30036A24" bold="true" box="[197,573,1850,1876]" pageId="2" pageNumber="60">Adult definition and diagnosis</emphasis>
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. A large
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<taxonomicName id="4C094D18FF89FF7CA680F8EC310A6A24" box="[680,820,1851,1876]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="60" phylum="Chordata" rank="genus">
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<emphasis id="B97DEA89FF89FF7CA680F8EC310A6A24" box="[680,820,1851,1876]" italics="true" pageId="2" pageNumber="60">Arthroleptis</emphasis>
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</taxonomicName>
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, snout-vent length (SVL)
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<quantity id="4CF19B7EFF89FF7CA040F8ED36ED6A24" box="[1128,1235,1850,1876]" metricMagnitude="-2" metricUnit="m" metricValue="4.33" pageId="2" pageNumber="60" unit="mm" value="43.3">43.3 mm</quantity>
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; sturdy and compact, broad head;
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<taxonomicName id="4C094D18FF89FF7CA540F8B430386A0C" box="[360,518,1891,1916]" class="Amphibia" family="Arthroleptidae" genus="Astylosternus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="60" phylum="Chordata" rank="genus">
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<emphasis id="B97DEA89FF89FF7CA540F8B430386A0C" box="[360,518,1891,1916]" italics="true" pageId="2" pageNumber="60">Astylosternus</emphasis>
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</taxonomicName>
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-like habitus (
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<figureCitation id="13322A1EFF89FF7CA680F8B530D06A0C" box="[680,750,1890,1916]" captionStart="FIGURE 1" captionStartId="3.[151,254,1005,1029]" captionTargetBox="[471,1129,193,960]" captionTargetId="figure@3.[450,1137,96,981]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal (a), ventral (b), and latero-ventral (c) view of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst" httpUri="https://zenodo.org/record/180741/files/figure.png" pageId="2" pageNumber="60">Fig. 1</figureCitation>
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a–c,
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<figureCitation id="13322A1EFF89FF7CA705F8B5314D6A0C" box="[813,883,1890,1916]" captionStart="FIGURE 3" captionStartId="4.[151,254,1179,1203]" captionTargetBox="[550,1036,477,1154]" captionTargetId="figure@4.[550,1036,474,1156]" captionTargetPageId="4" captionText="FIGURE 3. Arthroleptis krokosua sp. nov. holotype (SMNS 12555) in life. Photographs R. Ernst" httpUri="https://zenodo.org/record/180743/files/figure.png" pageId="2" pageNumber="60">Fig. 3</figureCitation>
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); larger and more compact body shape than all other known West African
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<taxonomicName id="4C094D18FF89FF7CA5C9F85C30506AD4" box="[481,622,1931,1956]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="60" phylum="Chordata" rank="genus">
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<emphasis id="B97DEA89FF89FF7CA5C9F85C30506AD4" box="[481,622,1931,1956]" italics="true" pageId="2" pageNumber="60">Arthroleptis</emphasis>
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</taxonomicName>
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species; well developed maxillary teeth; inner metatarsal tubercles elongated, reaching 80.9% of the length of first toe; tips of toes are slightly expanded into discs and tips of fingers are slightly swollen (
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<figureCitation id="13322A1EFF89FF7CA5EDF80D30326A84" box="[453,524,2010,2036]" captionStart="FIGURE 2" captionStartId="3.[151,254,1614,1638]" captionTargetBox="[497,1046,1126,1583]" captionTargetId="figure@3.[494,1093,1124,1591]" captionTargetPageId="3" captionText="FIGURE 2. Right hand and plantar surface of right foot of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst." httpUri="https://zenodo.org/record/180742/files/figure.png" pageId="2" pageNumber="60">Fig. 2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF766613FF88FF7DA4BFFC3A33EA6957" httpUri="https://zenodo.org/record/180741/files/figure.png" pageId="3" pageNumber="61" targetBox="[471,1129,193,960]" targetPageId="3">
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<paragraph id="8BB6369BFF88FF7DA4BFFC3A33EA6957" blockId="3.[151,1436,1005,1063]" pageId="3" pageNumber="61">
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<emphasis id="B97DEA89FF88FF7DA4BFFC3A33246975" bold="true" box="[151,282,1005,1029]" pageId="3" pageNumber="61">FIGURE 1.</emphasis>
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Dorsal (a), ventral (b), and latero-ventral (c) view of the
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<taxonomicName id="4C094D18FF88FF7DA7BBFC3936426975" box="[915,1148,1006,1029]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="61" phylum="Chordata" rank="species" species="krokosua" status="sp. nov.">
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<emphasis id="B97DEA89FF88FF7DA7BBFC3936426975" box="[915,1148,1006,1029]" italics="true" pageId="3" pageNumber="61">Arthroleptis krokosua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57F2FF88FF7DA0ADFC3A36E26975" box="[1157,1244,1005,1029]" pageId="3" pageNumber="61" rank="species">
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<emphasis id="B97DEA89FF88FF7DA0ADFC3A36E26975" bold="true" box="[1157,1244,1005,1029]" pageId="3" pageNumber="61">sp. nov.</emphasis>
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</taxonomicNameLabel>
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holotype (SMNS 12555). Photographs R. Ernst
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</paragraph>
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</caption>
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<caption id="DF766613FF88FF7DA4BFF99933BC6BF8" httpUri="https://zenodo.org/record/180742/files/figure.png" pageId="3" pageNumber="61" targetBox="[497,1046,1126,1583]" targetPageId="3">
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<paragraph id="8BB6369BFF88FF7DA4BFF99933BC6BF8" blockId="3.[151,1437,1614,1672]" pageId="3" pageNumber="61">
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<emphasis id="B97DEA89FF88FF7DA4BFF99933276B16" bold="true" box="[151,281,1614,1638]" pageId="3" pageNumber="61">FIGURE 2.</emphasis>
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Right hand and plantar surface of right foot of the
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<taxonomicName id="4C094D18FF88FF7DA768F99836196B16" box="[832,1063,1615,1638]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="61" phylum="Chordata" rank="species" species="krokosua" status="sp. nov.">
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<emphasis id="B97DEA89FF88FF7DA768F99836196B16" box="[832,1063,1615,1638]" italics="true" pageId="3" pageNumber="61">Arthroleptis krokosua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57F2FF88FF7DA007F99936BB6B16" box="[1071,1157,1614,1638]" pageId="3" pageNumber="61" rank="species">
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<emphasis id="B97DEA89FF88FF7DA007F99936BB6B16" bold="true" box="[1071,1157,1614,1638]" pageId="3" pageNumber="61">sp. nov.</emphasis>
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</taxonomicNameLabel>
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holotype (SMNS 12555). Photographs R. Ernst.
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</paragraph>
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</caption>
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<subSubSection id="C3136510FF88FF7AA4EDF909300D6A79" lastPageId="4" lastPageNumber="62" pageId="3" pageNumber="61" type="description">
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<paragraph id="8BB6369BFF88FF7AA4EDF90937336CD2" blockId="3.[151,1436,1758,2025]" lastBlockId="4.[151,1437,152,418]" lastPageId="4" lastPageNumber="62" pageId="3" pageNumber="61">
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<emphasis id="B97DEA89FF88FF7DA4EDF90930276B88" bold="true" box="[197,537,1758,1784]" pageId="3" pageNumber="61">
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Description of the
|
||
<typeStatus id="54B28839FF88FF7DA586F90930276B88" box="[430,537,1758,1784]" pageId="3" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. A male (SMNS 12555) with SVL of
|
||
<quantity id="4CF19B7EFF88FF7DA7C8F90836736B89" box="[992,1101,1759,1785]" metricMagnitude="-2" metricUnit="m" metricValue="4.33" pageId="3" pageNumber="61" unit="mm" value="43.3">43.3 mm</quantity>
|
||
; sturdy compact body shape (
|
||
<figureCitation id="13322A1EFF88FF7DA4B7F8D032DD6A51" box="[159,227,1799,1825]" captionStart="FIGURE 1" captionStartId="3.[151,254,1005,1029]" captionTargetBox="[471,1129,193,960]" captionTargetId="figure@3.[450,1137,96,981]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal (a), ventral (b), and latero-ventral (c) view of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst" httpUri="https://zenodo.org/record/180741/files/figure.png" pageId="3" pageNumber="61">Fig. 1</figureCitation>
|
||
a, c); head width
|
||
<quantity id="4CF19B7EFF88FF7DA59CF8D0301F6A51" box="[436,545,1799,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="3" pageNumber="61" unit="mm" value="19.7">19.7 mm</quantity>
|
||
; interorbital distance
|
||
<quantity id="4CF19B7EFF88FF7DA708F8D031406A51" box="[800,894,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="61" unit="mm" value="5.9">5.9 mm</quantity>
|
||
; distance from eye to snout tip
|
||
<quantity id="4CF19B7EFF88FF7DA0DBF8D0376F6A51" box="[1267,1361,1799,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="61" unit="mm" value="5.8">5.8 mm</quantity>
|
||
; nares located lateral on snout, hardly visible from above and closer to tip of snout than to anterior corner of eyes; snout rounded, only slightly oval (
|
||
<figureCitation id="13322A1EFF88FF7DA605F880304A6A01" box="[557,628,1879,1905]" captionStart="FIGURE 1" captionStartId="3.[151,254,1005,1029]" captionTargetBox="[471,1129,193,960]" captionTargetId="figure@3.[450,1137,96,981]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal (a), ventral (b), and latero-ventral (c) view of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst" httpUri="https://zenodo.org/record/180741/files/figure.png" pageId="3" pageNumber="61">Fig. 1</figureCitation>
|
||
, b); well developed maxillary teeth, first two teeth on either maxilla particularly prominent; mandibular teeth absent; canthus rostralis rounded but distinct; loreal region slightly concave; round tympanum comparatively large (3.0 mm), 61.2% of eye diameter (
|
||
<quantity id="4CF19B7EFF88FF7DA059F87036EF6AB1" box="[1137,1233,1959,1985]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="61" unit="mm" value="4.8">4.8 mm</quantity>
|
||
) but not distinct, same colour as surrounding area; upper arm length
|
||
<quantity id="4CF19B7EFF88FF7DA6C4F818316A6A99" box="[748,852,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="61" unit="mm" value="12.3">12.3 mm</quantity>
|
||
(28.4% of SVL), forearm length
|
||
<quantity id="4CF19B7EFF88FF7DA0F1F818377E6A99" box="[1241,1344,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="61" unit="mm" value="11.5">11.5 mm</quantity>
|
||
(26.5% of SVL); fingers without webbing; longest finger
|
||
<quantity id="4CF19B7EFF8FFF7AA6F5FF4F31076DC2" box="[733,825,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="62" unit="mm" value="7.5">7.5 mm</quantity>
|
||
; relative finger length: 4<1=2<3; subarticular tubercles well defined, large and present on all fingers (
|
||
<figureCitation id="13322A1EFF8FFF7AA6CAFF1731176DAA" box="[738,809,192,218]" captionStart="FIGURE 2" captionStartId="3.[151,254,1614,1638]" captionTargetBox="[497,1046,1126,1583]" captionTargetId="figure@3.[494,1093,1124,1591]" captionTargetPageId="3" captionText="FIGURE 2. Right hand and plantar surface of right foot of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst." httpUri="https://zenodo.org/record/180742/files/figure.png" pageId="4" pageNumber="62">Fig. 2</figureCitation>
|
||
); tibula-fibula (
|
||
<quantity id="4CF19B7EFF8FFF7AA7F7FF1736756DAA" box="[991,1099,192,218]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="4" pageNumber="62" unit="mm" value="22.8">22.8 mm</quantity>
|
||
) 52.6% of SVL; femur (
|
||
<quantity id="4CF19B7EFF8FFF7AA14FFF1732F96C72" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="4" pageNumber="62" unit="mm" value="21.7">21.7 mm</quantity>
|
||
) 50.1% of SVL; tibiale-fibulare including longest toe
|
||
<quantity id="4CF19B7EFF8FFF7AA764FF3F31886C72" box="[844,950,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="4" pageNumber="62" unit="mm" value="31.5">31.5 mm</quantity>
|
||
(72.7% of SVL); terminal toe phalanges slightly expanded into discs; subarticular tubercles on toes distinct but smaller than those on fingers; single inner metatarsal tubercle (
|
||
<quantity id="4CF19B7EFF8FFF7AA5F9FEEF300E6C22" box="[465,560,312,338]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="62" unit="mm" value="2.7">2.7 mm</quantity>
|
||
) slightly more than three quarters of first toe length; relative toe length: 1<2<5<3<4; foot webbing rudimental, almost absent (
|
||
<figureCitation id="13322A1EFF8FFF7AA703FEB731496C0A" box="[811,887,352,378]" captionStart="FIGURE 2" captionStartId="3.[151,254,1614,1638]" captionTargetBox="[497,1046,1126,1583]" captionTargetId="figure@3.[494,1093,1124,1591]" captionTargetPageId="3" captionText="FIGURE 2. Right hand and plantar surface of right foot of the Arthroleptis krokosua sp. nov. holotype (SMNS 12555). Photographs R. Ernst." httpUri="https://zenodo.org/record/180742/files/figure.png" pageId="4" pageNumber="62">Fig. 2</figureCitation>
|
||
); skin on dorsum, ventrum and legs smooth, except for urostyle-cloacal region, which is porous to sulcate; gular region smooth, without folds.
|
||
</paragraph>
|
||
<caption id="DF766613FF8FFF7AA4BFFB4C369869C3" box="[151,1190,1179,1203]" httpUri="https://zenodo.org/record/180743/files/figure.png" pageId="4" pageNumber="62" targetBox="[550,1036,477,1154]" targetPageId="4">
|
||
<paragraph id="8BB6369BFF8FFF7AA4BFFB4C369869C3" blockId="4.[151,1190,1179,1203]" box="[151,1190,1179,1203]" pageId="4" pageNumber="62">
|
||
<emphasis id="B97DEA89FF8FFF7AA4BFFB4C332669C3" bold="true" box="[151,280,1179,1203]" pageId="4" pageNumber="62">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C094D18FF8FFF7AA537FB4B303969C3" box="[287,519,1180,1203]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="62" phylum="Chordata" rank="species" species="krokosua" status="sp. nov.">
|
||
<emphasis id="B97DEA89FF8FFF7AA537FB4B303969C3" box="[287,519,1180,1203]" italics="true" pageId="4" pageNumber="62">Arthroleptis krokosua</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E57F2FF8FFF7AA625FB4C305D69C3" box="[525,611,1179,1203]" pageId="4" pageNumber="62" rank="species">
|
||
<emphasis id="B97DEA89FF8FFF7AA625FB4C305D69C3" bold="true" box="[525,611,1179,1203]" pageId="4" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype (SMNS 12555) in life. Photographs R. Ernst
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB6369BFF8FFF7AA4EDFAD933376BC9" blockId="4.[151,1437,1294,1961]" pageId="4" pageNumber="62">
|
||
<emphasis id="B97DEA89FF8FFF7AA4EDFAD9303B6858" bold="true" box="[197,517,1294,1320]" pageId="4" pageNumber="62">
|
||
Colour of
|
||
<typeStatus id="54B28839FF8FFF7AA56CFAD9338E6858" box="[324,432,1294,1320]" pageId="4" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
Dorsal colouration generally light brown to beige without clearly marked patterns (
|
||
<figureCitation id="13322A1EFF8FFF7AA4CCFAE033116821" box="[228,303,1335,1361]" captionStart="FIGURE 3" captionStartId="4.[151,254,1179,1203]" captionTargetBox="[550,1036,477,1154]" captionTargetId="figure@4.[550,1036,474,1156]" captionTargetPageId="4" captionText="FIGURE 3. Arthroleptis krokosua sp. nov. holotype (SMNS 12555) in life. Photographs R. Ernst" httpUri="https://zenodo.org/record/180743/files/figure.png" pageId="4" pageNumber="62">Fig. 3</figureCitation>
|
||
). Head from snout tip to interorbital cream white, forming a bright triangle; interorbital and occipital region slightly darker than rest of dorsum; slightly darker triangular shaped pattern in sacral region; legs with dark banding; one main band on femur, an additional band at level of femur-tibula-fibula joint; three bands on shanks; inside of entire hind extremity dark; only inside of lower arm dark; peculiar row of dark lateral blotches (
|
||
<figureCitation id="13322A1EFF8FFF7AA515FA0033BA6881" box="[317,388,1495,1521]" captionStart="FIGURE 3" captionStartId="4.[151,254,1179,1203]" captionTargetBox="[550,1036,477,1154]" captionTargetId="figure@4.[550,1036,474,1156]" captionTargetPageId="4" captionText="FIGURE 3. Arthroleptis krokosua sp. nov. holotype (SMNS 12555) in life. Photographs R. Ernst" httpUri="https://zenodo.org/record/180743/files/figure.png" pageId="4" pageNumber="62">Fig. 3</figureCitation>
|
||
); the anterior set each bordered by thin white line; first blotch on left side partially fused with the dark tympanal patch; the latter starts behind the eye and broadens subsequently, covering the entire tympanum and extending dorsally and posteriorly; entire patch bordered by thin white line; throat completely dark; dark pigmentation continues across chest and fades caudally; irregular white spots distributed across venter, starting at level of chest (
|
||
<figureCitation id="13322A1EFF8FFF7AA608F9A030576BE1" box="[544,617,1655,1681]" captionStart="FIGURE 3" captionStartId="4.[151,254,1179,1203]" captionTargetBox="[550,1036,477,1154]" captionTargetId="figure@4.[550,1036,474,1156]" captionTargetPageId="4" captionText="FIGURE 3. Arthroleptis krokosua sp. nov. holotype (SMNS 12555) in life. Photographs R. Ernst" httpUri="https://zenodo.org/record/180743/files/figure.png" pageId="4" pageNumber="62">Fig. 3</figureCitation>
|
||
); femur irregularly marbled; distinct dark labial bars underneath eye and naris.
|
||
</paragraph>
|
||
<paragraph id="8BB6369BFF8FFF7AA4EDF911300D6A79" blockId="4.[151,1437,1294,1961]" pageId="4" pageNumber="62">
|
||
<emphasis id="B97DEA89FF8FFF7AA4EDF91130BE6B90" bold="true" box="[197,640,1734,1760]" pageId="4" pageNumber="62">
|
||
Colour of
|
||
<typeStatus id="54B28839FF8FFF7AA560F911338A6B90" box="[328,436,1734,1760]" pageId="4" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
Nearly no colour difference between life and preserved specimen. Colour in ethanol only more faded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136510FF8FFF7AA4EDF8C136696AD9" pageId="4" pageNumber="62" type="distribution">
|
||
<paragraph id="8BB6369BFF8FFF7AA4EDF8C136696AD9" blockId="4.[151,1437,1294,1961]" pageId="4" pageNumber="62">
|
||
<emphasis id="B97DEA89FF8FFF7AA4EDF8C133C36A40" bold="true" box="[197,509,1814,1840]" pageId="4" pageNumber="62">Distribution and habitat.</emphasis>
|
||
The species is so far only known from the
|
||
<typeStatus id="54B28839FF8FFF7AA7D2F8C036136A41" box="[1018,1069,1815,1841]" pageId="4" pageNumber="62">type</typeStatus>
|
||
locality at
|
||
<taxonomicName id="4C094D18FF8FFF7AA087F8C0371F6A41" box="[1199,1313,1815,1841]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="66" phylum="Chordata" rank="species" species="krokosua">Krokosua</taxonomicName>
|
||
Hills Forest Reserve. There it was found to be associated with uphill moist semi-deciduous forest. The microhabitat in which it was encountered was characterized by a comparatively thick and dense leaf litter. We recorded no open bodies of water, neither lentic nor lotic, in direct vicinity of the
|
||
<typeStatus id="54B28839FF8FFF7AA797F85831CF6AD9" box="[959,1009,1935,1961]" pageId="4" pageNumber="62">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |