treatments-xml/data/03/8F/97/038F9706FFC25454BC4F34A838E21304.xml
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<document ID-DOI="10.15560/15.4.579" ID-ISSN="1809-127X" approvalRequired="75" approvalRequired_for_textStreams="74" approvalRequired_for_treatments="1" checkinTime="1630355267505" checkinUser="marcus" docAuthor="Taboada-Verona, Carlos, Sermeño-Correa, Carlos, Sierra-Serrano, Oscar &amp; Noriega, Jorge Ari" docDate="2019" docId="038F9706FFC25454BC4F34A838E21304" docLanguage="en" docName="CheckList.2019.15.4.579-594.pdf" docOrigin="Check List 15 (4)" docSource="http://dx.doi.org/10.15560/15.4.579" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Tomarus fossor" docType="treatment" docVersion="1" lastPageId="6" lastPageNumber="585" masterDocId="FFB6EF7EFFC75452BF62316F396F126F" masterDocTitle="Checklist of the superfamily Scarabaeoidea (Insecta, Coleoptera) in an urban area of the Caribbean Colombia" masterLastPageNumber="594" masterPageNumber="579" pageId="5" pageNumber="584" updateTime="1630355270070" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Checklist of the superfamily Scarabaeoidea (Insecta, Coleoptera) in an urban area of the Caribbean Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Taboada-Verona, Carlos</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sermeño-Correa, Carlos</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierra-Serrano, Oscar</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Noriega, Jorge Ari</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-07-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>15</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>579</mods:start>
<mods:end>594</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/15.4.579</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/15.4.579</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:038F9706FFC25454BC4F34A838E21304" httpUri="http://treatment.plazi.org/id/038F9706FFC25454BC4F34A838E21304" lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584">
<subSubSection box="[813,1175,1478,1502]" pageId="5" pageNumber="584" type="nomenclature">
<paragraph blockId="5.[813,1175,1478,1502]" box="[813,1175,1478,1502]" pageId="5" pageNumber="584">
<heading box="[813,1175,1478,1502]" fontSize="10" level="2" pageId="5" pageNumber="584" reason="2">
<taxonomicName authority="(Latreille, 1813)" baseAuthorityName="Latreille" baseAuthorityYear="1813" box="[813,1175,1478,1502]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="fossor">
<emphasis bold="true" box="[813,1175,1478,1502]" inLineHeading="true" pageId="5" pageNumber="584" reason="1">
<emphasis bold="true" box="[813,986,1478,1502]" italics="true" pageId="5" pageNumber="584">Tomarus fossor</emphasis>
(Latreille, 1813)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="584" type="description">
<paragraph blockId="5.[813,918,1512,1536]" box="[813,918,1512,1536]" pageId="5" pageNumber="584">
<figureCitation box="[813,918,1512,1536]" captionStart="Figure 4" captionStartId="4.[168,231,1984,2004]" captionTargetBox="[251,1321,182,1939]" captionTargetId="figure-86@4.[234,1353,160,1960]" captionTargetPageId="4" captionText="Figure 4. Beetles of superfamily Scarabaeoidea present at the University of Sucre campus. A. Passalus interruptus. B. Passalus interstitialis. C. Passalus punctiger.D. Gymnetis stellata. E. Hoplopyga liturata. F. Dyscinetus dubius.G. Euetheola humilis.H. Megasoma elephas.I. Phileurus didymus. J. Phileurus valgus. K. Strategus aloeus. L. Tomarus fossor.M. Liogenys quadridens. N. Phyllophaga menetriesi. O. Anomala valida. P. Leucothyreus cf. femoratus. Q. Pelidnota polita. R. Xenopelidnota anomala. Scale bars = 1 cm." pageId="5" pageNumber="584">Figure 4L</figureCitation>
</paragraph>
<paragraph blockId="5.[813,1421,1555,1813]" pageId="5" pageNumber="584">
<emphasis bold="true" box="[813,1102,1555,1579]" pageId="5" pageNumber="584">Geographic distribution.</emphasis>
<collectingCountry box="[1112,1203,1556,1580]" name="Antigua and Barbuda" pageId="5" pageNumber="584">Antigua</collectingCountry>
,
<collectingCountry box="[1217,1319,1556,1580]" name="Bahamas" pageId="5" pageNumber="584">Bahamas</collectingCountry>
,
<collectingCountry box="[1333,1415,1556,1580]" name="Bolivia" pageId="5" pageNumber="584">Bolivia</collectingCountry>
,
<collectingCountry box="[813,882,1589,1613]" name="Brazil" pageId="5" pageNumber="584">Brazil</collectingCountry>
,
<collectingCountry box="[893,1001,1589,1613]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
,
<collectingCountry box="[1011,1071,1589,1613]" name="Cuba" pageId="5" pageNumber="584">Cuba</collectingCountry>
,
<collectingCountry box="[1081,1175,1589,1613]" name="Curaçao" pageId="5" pageNumber="584">Curaçao</collectingCountry>
,
<collectingCountry box="[1186,1278,1589,1613]" name="Ecuador" pageId="5" pageNumber="584">Ecuador</collectingCountry>
,
<collectingCountry box="[1288,1415,1589,1613]" name="El Salvador" pageId="5" pageNumber="584">El Salvador</collectingCountry>
,
<collectingCountry box="[813,903,1622,1646]" name="Jamaica" pageId="5" pageNumber="584">Jamaica</collectingCountry>
,
<collectingCountry box="[913,1002,1623,1646]" name="Panama" pageId="5" pageNumber="584">Panama</collectingCountry>
,
<collectingCountry box="[1013,1146,1622,1646]" name="Puerto Rico" pageId="5" pageNumber="584">Puerto Rico</collectingCountry>
, and
<collectingCountry box="[1204,1316,1622,1646]" name="Venezuela" pageId="5" pageNumber="584">Venezuela</collectingCountry>
(
<bibRefCitation author="Endrodi S" pageId="5" pageNumber="584" pagination="1 - 145" refId="ref10219" refString="Endrodi S (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden 37: 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
,
<bibRefCitation author="Ratcliffe BC" box="[879,1036,1656,1680]" pageId="5" pageNumber="584" pagination="1 - 506" refId="ref14615" refString="Ratcliffe BC (2003) The Dynastinae scarab beetles of Costa Rica and Panama. Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003</bibRefCitation>
,
<bibRefCitation author="Ratcliffe BC &amp; Cave RD" box="[1048,1316,1656,1680]" pageId="5" pageNumber="584" pagination="1 - 424" refId="ref14699" refString="Ratcliffe BC, Cave RD (2006) The ynastinae scarab beetles of Hon- duras, Nicaragua and El Salvador. Bulletin of the University of Nebraska State Museum 21: 1 - 424." type="journal article" year="2006">Ratcliffe and Cave 2006</bibRefCitation>
,
<bibRefCitation author="Carvajal V &amp; Villamarin C &amp; Ortega AM" pageId="5" pageNumber="584" refId="ref9765" refString="Carvajal V, Villamarin C, Ortega AM (2011) Escarabajos del Ecuador. Principales generos. Serie Entomologia, No. 1. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito, 350 pp." type="book" year="2011">Carvajal et al. 2011</bibRefCitation>
). In
<collectingCountry box="[976,1085,1689,1713]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
:
<collectingRegion box="[1096,1208,1689,1713]" country="Colombia" name="Antioquia" pageId="5" pageNumber="584">Antioquia</collectingRegion>
,
<collectingRegion box="[1219,1322,1689,1713]" country="Colombia" name="Atlantico" pageId="5" pageNumber="584">Atlántico</collectingRegion>
, Bolívar,
<collectingRegion box="[814,888,1723,1747]" country="Colombia" name="Caldas" pageId="5" pageNumber="584">Caldas</collectingRegion>
,
<collectingRegion box="[900,968,1723,1747]" country="Colombia" name="Cauca" pageId="5" pageNumber="584">Cauca</collectingRegion>
,
<collectingRegion box="[980,1042,1723,1747]" country="Colombia" name="Cesar" pageId="5" pageNumber="584">Cesar</collectingRegion>
,
<collectingRegion box="[1054,1147,1723,1747]" country="Colombia" name="Cordoba" pageId="5" pageNumber="584">Córdoba</collectingRegion>
,
<collectingRegion box="[1158,1240,1723,1747]" country="Colombia" name="La Guajira" pageId="5" pageNumber="584">Guajira</collectingRegion>
,
<collectingRegion box="[1251,1370,1723,1747]" country="Colombia" name="Magdalena" pageId="5" pageNumber="584">Magdalena</collectingRegion>
, and Sucre (
<bibRefCitation author="Endrodi S" box="[886,1031,1756,1780]" pageId="5" pageNumber="584" pagination="1 - 145" refId="ref10219" refString="Endrodi S (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden 37: 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
,
<bibRefCitation author="Restrepo-Giraldo H &amp; Moron MA &amp; Vallejo F &amp; Pardo-Locarno LC &amp; Lopez-Avila A" box="[1039,1337,1756,1780]" pageId="5" pageNumber="584" pagination="239 - 263" refId="ref14897" refString="Restrepo-Giraldo H, Moron MA, Vallejo F, Pardo-Locarno LC, Lopez-Avila A (2003) Catalogo de Coleoptera Melolonthidae (Scarabaeidae Pleurosticti) de Colombia. Folia Entomologica Mexicana 42 (2): 239 - 263." type="journal article" year="2003">Restrepo-Giraldo et al. 2003</bibRefCitation>
, Lopez- Garcia et al. 2015).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584" type="materials_examined">
<paragraph blockId="5.[813,1420,1833,2091]" lastBlockId="6.[168,775,159,218]" lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584">
<emphasis bold="true" box="[813,1042,1833,1857]" pageId="5" pageNumber="584">Material examined.</emphasis>
<collectingCountry box="[1053,1161,1833,1857]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
, Sucre, Sincelejo, University of Sucre campus (
<geoCoordinate box="[1117,1243,1865,1890]" degrees="09" direction="north" minutes="18" orientation="latitude" pageId="5" pageNumber="584" precision="15" seconds="52" value="9.314445">09°1852″N</geoCoordinate>
,
<geoCoordinate box="[1262,1407,1865,1891]" degrees="075" direction="west" minutes="23" orientation="longitude" pageId="5" pageNumber="584" precision="15" seconds="18" value="-75.38833">075°2318″W</geoCoordinate>
),
<date box="[813,929,1900,1924]" pageId="5" pageNumber="584" value="2016-05-19">2016/05/19</date>
,
<collectionCode box="[942,1036,1900,1924]" pageId="5" pageNumber="584">MZUSU</collectionCode>
<accessionNumber box="[1044,1127,1900,1924]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E02501" pageId="5" pageNumber="584">E02501</accessionNumber>
-02 (
<specimenCount box="[1179,1209,1900,1924]" pageId="5" pageNumber="584" type="female">2♀</specimenCount>
).
<collectingCountry box="[1231,1339,1900,1924]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
, Sucre, Sincelejo, University of Sucre campus (
<geoCoordinate box="[1289,1415,1932,1957]" degrees="09" direction="north" minutes="18" orientation="latitude" pageId="5" pageNumber="584" precision="15" seconds="52" value="9.314445">09°1852″N</geoCoordinate>
,
<geoCoordinate box="[813,954,1965,1991]" degrees="075" direction="west" minutes="23" orientation="longitude" pageId="5" pageNumber="584" precision="15" seconds="18" value="-75.38833">075°2318″W</geoCoordinate>
).
<date box="[994,1112,1967,1991]" pageId="5" pageNumber="584" value="2016-05-27">2016/05/27</date>
,
<collectionCode box="[1140,1234,1967,1991]" pageId="5" pageNumber="584">MZUSU</collectionCode>
<accessionNumber box="[1259,1341,1967,1991]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E02503" pageId="5" pageNumber="584">E02503</accessionNumber>
(
<specimenCount box="[1374,1406,1967,1991]" pageId="5" pageNumber="584" type="male">1♂</specimenCount>
).
<collectingCountry box="[813,921,2000,2024]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
, Sucre, Sincelejo, University of Sucre campus (
<geoCoordinate box="[870,998,2032,2057]" degrees="09" direction="north" minutes="18" orientation="latitude" pageId="5" pageNumber="584" precision="15" seconds="52" value="9.314445">09°1852″N</geoCoordinate>
,
<geoCoordinate box="[1014,1159,2032,2058]" degrees="075" direction="west" minutes="23" orientation="longitude" pageId="5" pageNumber="584" precision="15" seconds="18" value="-75.38833">075°2318″W</geoCoordinate>
),
<date box="[1184,1301,2033,2057]" pageId="5" pageNumber="584" value="2016-05-30">2016/05/30</date>
/,
<collectionCode box="[1325,1419,2034,2058]" pageId="5" pageNumber="584">MZUSU</collectionCode>
<accessionNumber box="[813,899,2067,2091]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E02504" pageId="5" pageNumber="584">E02504</accessionNumber>
-06 (
<specimenCount box="[962,993,2067,2091]" pageId="5" pageNumber="584" type="female">3♀</specimenCount>
).
<collectingCountry box="[1026,1134,2067,2091]" name="Colombia" pageId="5" pageNumber="584">Colombia</collectingCountry>
, Sucre, Sincelejo, Uni- versity of Sucre campus (
<geoCoordinate box="[473,599,159,184]" degrees="09" direction="north" minutes="18" orientation="latitude" pageId="6" pageNumber="585" precision="15" seconds="52" value="9.314445">09°1852″N</geoCoordinate>
,
<geoCoordinate box="[618,763,159,184]" degrees="075" direction="west" minutes="23" orientation="longitude" pageId="6" pageNumber="585" precision="15" seconds="18" value="-75.38833">075°2318″W</geoCoordinate>
),
<date box="[168,286,194,218]" pageId="6" pageNumber="585" value="2016-08-06">2016/06/08</date>
/,
<collectionCode box="[304,398,194,218]" pageId="6" pageNumber="585">MZUSU</collectionCode>
<accessionNumber box="[405,491,194,218]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/E02507" pageId="6" pageNumber="585">E02507</accessionNumber>
-08 (
<specimenCount box="[539,570,194,218]" pageId="6" pageNumber="585" type="female">2♀</specimenCount>
).
</paragraph>
<paragraph blockId="6.[168,775,238,364]" pageId="6" pageNumber="585">
<emphasis bold="true" box="[168,330,238,262]" pageId="6" pageNumber="585">Identification.</emphasis>
This species can be distinguished from other
<taxonomicName box="[233,416,272,296]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[233,328,273,296]" italics="true" pageId="6" pageNumber="585">Tomarus</emphasis>
species
</taxonomicName>
by its small size, reddish color, pronotum without fovea nor tubercles, and deeper punctuation on pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>