treatments-xml/data/03/8F/19/038F191A560EFFFF60BEB61C886CFE3E.xml

<document id="CD292C7051CE9A55993891B14780D933" ID-CLB-Dataset="29956" ID-DOI="10.11646/zootaxa.4429.2.13" ID-GBIF-Dataset="bfbd1a77-236d-4192-b05b-23a247ffdadc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1283957" ID-ZooBank="58F3C349-9521-4DAF-8A05-D3AC050779D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528273989242" checkinUser="plazi" docAuthor="Grave, Sammy De &amp; Anker, Arthur" docDate="2018" docId="038F191A560EFFFF60BEB61C886CFE3E" docLanguage="en" docName="zootaxa.4429.2.13.pdf" docOrigin="Zootaxa 4429 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lysmata napoleoni Grave &amp; Anker 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="FFB66162560FFFF66029B5628C44FFCA" masterDocTitle="A new, distinctly coloured species of Lysmata Risso, 1816 (Malacostraca: Decapoda: Lysmatidae) from the south-central Atlantic" masterLastPageNumber="400" masterPageNumber="390" pageNumber="391" updateTime="1698547729141" updateUser="ExternalLinkService">
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<mods:title id="C919BE9CF56F6A66DD79864EFB8931B9">A new, distinctly coloured species of Lysmata Risso, 1816 (Malacostraca: Decapoda: Lysmatidae) from the south-central Atlantic</mods:title>
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<mods:namePart id="6EB13D78D8E6C38D9C0CFA2A04BC9684">Grave, Sammy De</mods:namePart>
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<mods:date id="203CCEF6EE042758C7B644F27F800346">2018</mods:date>
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<heading id="D0D11F60560EFFF760BEB61C8DA5FC52" bold="true" box="[151,481,894,920]" fontSize="11" level="1" pageId="1" pageNumber="391" reason="1">
<emphasis id="B952741E560EFFF760BEB61C8DA5FC52" bold="true" box="[151,481,894,920]" pageId="1" pageNumber="391">
<taxonomicName id="4C26D38F560EFFF760BEB61C8D3FFC52" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[151,379,894,920]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="391" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E560EFFF760BEB61C8D3FFC52" bold="true" box="[151,379,894,920]" italics="true" pageId="1" pageNumber="391">Lysmata napoleoni</emphasis>
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</heading>
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<paragraph id="8B99A80C560EFFF760BEB6C38D56FC70" blockId="1.[151,481,894,954]" box="[151,274,929,954]" pageId="1" pageNumber="391">
(
<figureCitation id="131DB489560EFFF760B6B6C38D4CFC70" box="[159,264,929,954]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1878,1900]" captionStartId-1="3.[151,250,1757,1779]" captionStartId-2="5.[151,250,1941,1963]" captionStartId-3="6.[151,250,1500,1522]" captionStartId-4="8.[151,250,1980,2002]" captionTargetBox-0="[157,1404,196,1850]" captionTargetBox-1="[155,1380,200,1724]" captionTargetBox-2="[167,1395,217,1889]" captionTargetBox-3="[151,1436,193,1478]" captionTargetBox-4="[151,1436,193,1959]" captionTargetId-0="figure@2.[151,1411,193,1858]" captionTargetId-1="figure@3.[151,1436,193,1735]" captionTargetId-2="figure@5.[161,1421,193,1913]" captionTargetId-3="figure@6.[151,1436,193,1478]" captionTargetId-4="figure@8.[151,1436,193,1959]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="8" captionText-0="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." captionText-1="FIGURE 2. Lysmata napoleoni sp. nov., (a) right first pereiopod, lateral view; b) same, ischium, mesial view; (c) same, chela and distal portion of carpus, ventromesial view; (d) right second pereiopod, lateral view; (e) same, chela, mesial view. All from paratype (pocl 5.2 mm), MZUSP 37298." captionText-2="FIGURE 3. Lysmata napoleoni sp. nov., (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298." captionText-3="FIGURE 4. Lysmata napoleoni sp. nov. on St. Helena Island, (a) paratype, OUMNH.ZC. 2018-04-004; (b) ovigerous hermaphrodite, not deposited; (c) – (d) paratype, OUMNH.ZC. 2018-04-004, showing association with the sea anemone Telmactis cricoides (Duchassaing). Photographs by Peter Wirtz (a, b, c) and Judith Brown (d)." captionText-4="FIGURE 5. Lysmata napoleoni sp. nov. from Ascension Island, specimens not collected, (a) colour pattern by day; (b) nocturnal cleaning behaviour. Photographs by Judith Brown." httpUri-0="https://zenodo.org/record/1283959/files/figure.png" httpUri-1="https://zenodo.org/record/1283961/files/figure.png" httpUri-2="https://zenodo.org/record/1283963/files/figure.png" httpUri-3="https://zenodo.org/record/1283965/files/figure.png" httpUri-4="https://zenodo.org/record/1283967/files/figure.png" pageId="1" pageNumber="391">Figs. 1–5</figureCitation>
)
</paragraph>
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<taxonomicName id="4C26D38F560EFFF760BEB6888CAFFC35" authorityName="Grave &amp; Anker" authorityYear="2018" box="[151,235,1002,1023]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E560EFFF760BEB6888CAFFC35" box="[151,235,1002,1023]" italics="true" pageId="1" pageNumber="391">Lysmata</emphasis>
</taxonomicName>
sp.— 
<treatmentCitation id="0A878E1D560EFFF7610FB6888D88FC35" author="" box="[294,460,1001,1023]" page="57" pageId="1" pageNumber="391">
<bibRefCitation id="EFB7D5FD560EFFF7610FB6888DEEFC35" box="[294,426,1001,1023]" pageId="1" pageNumber="391" refId="ref4507" refString="Brown, J. (2014) Marine life of St. Helena. Pisces Publications, Newbury, 220 pp." type="book">Brown 2014</bibRefCitation>
: 57
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(left photo).
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</paragraph>
</subSubSection>
<subSubSection id="C33CFB87560EFFF760BEB14E8C96FA3E" pageId="1" pageNumber="391" type="materials_examined">
<paragraph id="8B99A80C560EFFF760BEB14E8E22FAAE" blockId="1.[151,1437,1068,1993]" pageId="1" pageNumber="391">
<materialsCitation id="3B4EA251560EFFF760BEB14E8EC2FBA2" ID-GBIF-Occurrence="1848505589" collectingDate="2014-01-14" collectionCode="OUMNH" collectorName="Long Ledge &amp; J. Brown &amp; P. Wirtz" country="Saint Helena, Ascension and Tristan da Cunha" elevation="15" location="St. Helena" pageId="1" pageNumber="391" specimenCount="2" stateProvince="St. Helena" typeStatus="holotype">
<emphasis id="B952741E560EFFF760BEB14E8D82FB8F" bold="true" box="[151,454,1068,1093]" pageId="1" pageNumber="391">
<typeStatus id="549D16AE560EFFF760BEB14E8C96FB8F" box="[151,210,1068,1093]" pageId="1" pageNumber="391">Type</typeStatus>
material. 
<collectingRegion id="49E266EE560EFFF76165B14E8D82FB8F" box="[332,454,1068,1093]" country="Saint Helena, Ascension and Tristan da Cunha" name="Saint Helena" pageId="1" pageNumber="391">
<collectingCountry id="F331E89C560EFFF76165B14E8D82FB8F" box="[332,454,1068,1093]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="391">St. Helena</collectingCountry>
</collectingRegion>
</emphasis>
. 
<typeStatus id="549D16AE560EFFF761FDB14E8E05FB8F" box="[468,577,1068,1093]" pageId="1" pageNumber="391" type="holotype">Holotype</typeStatus>
: hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF7631FB14E8FD4FB8E" box="[822,912,1068,1093]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="391" unit="mm" value="5.8">5.8 mm</quantity>
), off 
<collectorName id="26D3CDDA560EFFF763E2B14F8812FB8F" box="[971,1110,1068,1093]" pageId="1" pageNumber="391">Long Ledge</collectorName>
, -15.945° -5.753°, 
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<quantity id="4CDE05E9560EFFF76505B14E8923FB8E" box="[1324,1383,1068,1093]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="391" unit="m" value="15.0">15 m</quantity>
</elevation>
, leg. 
<collectorName id="26D3CDDA560EFFF760BEB1328CBBFBAD" box="[151,255,1104,1127]" pageId="1" pageNumber="391">J. Brown</collectorName>
&amp; 
<collectorName id="26D3CDDA560EFFF76103B1328DCDFBA2" box="[298,393,1103,1128]" pageId="1" pageNumber="391">P. Wirtz</collectorName>
, 
<collectingDate id="EFDC7724560EFFF761B1B1328E42FBA2" box="[408,518,1104,1128]" pageId="1" pageNumber="391" value="2014-01-14">
<date id="FF988ECC560EFFF761B1B1328E42FBA2" box="[408,518,1104,1128]" pageId="1" pageNumber="391" value="2014-01-14">14.I.2014</date>
</collectingDate>
, 
<collectionCode id="ED3730C9560EFFF7623CB1328EC2FBA2" box="[533,646,1104,1128]" pageId="1" pageNumber="391">OUMNH</collectionCode>
</materialsCitation>
.ZC.2018-04-001 (fcn 
<date id="FF988ECC560EFFF763AAB12D8F98FBA2" box="[899,988,1103,1128]" pageId="1" pageNumber="391" value="14-01-14">140114</date>
/60/1&amp;2). Paratypes: ov. hermaphrodite (pocl 4.0 mm), James Bay Wharf steps, -15.921° -5.718°, 
<quantity id="4CDE05E9560EFFF76337B1168F24FB46" box="[798,864,1140,1164]" metricMagnitude="0" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="391" unit="m" value="9.8">9.8 m</quantity>
, leg. J. Brown, 
<date id="FF988ECC560EFFF76424B11688D4FB47" box="[1037,1168,1140,1165]" pageId="1" pageNumber="391" value="2013-09-06">06.IX.2013</date>
, OUMNH.ZC. 
<materialsCitation id="3B4EA251560EFFF7651FB1168D43FB1E" ID-GBIF-Occurrence="1848505948" collectingDate="06-09-13" collectingDateMax="2013-03-28" collectingDateMin="06-09-13" collectionCode="OUMNH" collectorName="J. Brown" country="Saint Helena, Ascension and Tristan da Cunha" elevation="10" location="St. Helena" pageId="1" pageNumber="391" specimenCount="1" stateProvince="St. Helena" typeStatus="holotype">
2018-04- 0 0 2 (fcn 
<collectingDate id="EFDC7724560EFFF7612FB1F58D1AFB7A" box="[262,350,1175,1200]" pageId="1" pageNumber="391" value="06-09-13">
<date id="FF988ECC560EFFF7612FB1F58D1AFB7A" box="[262,350,1175,1200]" pageId="1" pageNumber="391" value="06-09-13">130906</date>
</collectingDate>
/44/11); ov. hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF762C8B1FA8F04FB7A" box="[737,832,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="391" unit="mm" value="3.8">3.8 mm</quantity>
), same location, 
<elevation id="000B4F3F560EFFF76425B1FA8817FB65" box="[1036,1107,1176,1200]" metricMagnitude="0" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="391" unit="m" value="9.7">
<quantity id="4CDE05E9560EFFF76425B1FA8817FB65" box="[1036,1107,1176,1200]" metricMagnitude="0" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="391" unit="m" value="9.7">9.7 m</quantity>
</elevation>
, leg. 
<collectorName id="26D3CDDA560EFFF764B3B1FA8943FB65" box="[1178,1287,1176,1199]" pageId="1" pageNumber="391">J. Brown</collectorName>
, 
<collectingDate id="EFDC7724560EFFF7653EB1FA89D3FB7A" box="[1303,1431,1176,1200]" pageId="1" pageNumber="391" value="2013-03-28">
<date id="FF988ECC560EFFF7653EB1FA89D3FB7A" box="[1303,1431,1176,1200]" pageId="1" pageNumber="391" value="2013-03-28">28.III.2013</date>
</collectingDate>
, 
<collectionCode id="ED3730C9560EFFF760BEB1DE8D43FB1E" box="[151,263,1212,1236]" pageId="1" pageNumber="391">OUMNH</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B4EA251560EFFF76123B1DE8FCEFB32" ID-GBIF-Occurrence="1848505951" collectingDate="2014-01-19" collectingDateMax="28-03-13" collectingDateMin="2014-01-19" collectionCode="OUMNH" collectorName="J. Brown &amp; P. Wirtz" country="Saint Helena, Ascension and Tristan da Cunha" elevation="18" location="Buoys Hole" pageId="1" pageNumber="391" specimenCount="1" stateProvince="St. Helena" typeStatus="holotype">
ZC.2018-04-003 (fcn 
<collectingDate id="EFDC7724560EFFF76221B1DE8E1BFB1F" box="[520,607,1212,1237]" pageId="1" pageNumber="391" value="28-03-13">
<date id="FF988ECC560EFFF76221B1DE8E1BFB1F" box="[520,607,1212,1237]" pageId="1" pageNumber="391" value="28-03-13">130328</date>
</collectingDate>
/Q4/05); 4 hermaphrodites (pocl 3.5–5.0 mm), 
<location id="8EF9FED7560EFFF764A0B1DF8952FB1F" LSID="urn:lsid:plazi:treatment:038F191A560EFFFF60BEB61C886CFE3E:8EF9FED7560EFFF764A0B1DF8952FB1F" box="[1161,1302,1212,1237]" country="Saint Helena, Ascension and Tristan da Cunha" name="Buoys Hole" pageId="1" pageNumber="391" stateProvince="St. Helena">Buoys Hole</location>
, -15.911° - 5.688°, 
<elevation id="000B4F3F560EFFF760DDB1828D0EFB3D" box="[244,330,1248,1272]" metricMagnitude="1" metricUnit="m" metricValue="1.77" pageId="1" pageNumber="391" unit="m" value="17.7">
<quantity id="4CDE05E9560EFFF760DDB1828D0EFB3D" box="[244,330,1248,1272]" metricMagnitude="1" metricUnit="m" metricValue="1.77" pageId="1" pageNumber="391" unit="m" value="17.7">17.7 m</quantity>
</elevation>
, leg. 
<collectorName id="26D3CDDA560EFFF761B9B1828DBEFB3D" box="[400,506,1248,1271]" pageId="1" pageNumber="391">J. Brown</collectorName>
&amp; 
<collectorName id="26D3CDDA560EFFF76200B1828ECEFB32" box="[553,650,1247,1272]" pageId="1" pageNumber="391">P. Wirtz</collectorName>
, 
<collectingDate id="EFDC7724560EFFF762B2B1828F4CFB32" box="[667,776,1248,1272]" pageId="1" pageNumber="391" value="2014-01-19">
<date id="FF988ECC560EFFF762B2B1828F4CFB32" box="[667,776,1248,1272]" pageId="1" pageNumber="391" value="2014-01-19">19.I.2014</date>
</collectingDate>
, 
<collectionCode id="ED3730C9560EFFF76333B1828FCEFB32" box="[794,906,1248,1272]" pageId="1" pageNumber="391">OUMNH</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B4EA251560EFFF763A4B1828D43FA8A" ID-GBIF-Occurrence="1848505950" collectingDate="19-01-14" collectingDateMax="2014-01-23" collectingDateMin="19-01-14" collectionCode="OUMNH" collectorName="P. Wirtz &amp; J. Brown" country="Saint Helena, Ascension and Tristan da Cunha" location="Papa Nui" pageId="1" pageNumber="391" specimenCount="1" stateProvince="St. Helena" typeStatus="holotype">
ZC.2018-04-004 (fcn 
<collectingDate id="EFDC7724560EFFF764A2B1BD88A7FB32" box="[1163,1251,1247,1272]" pageId="1" pageNumber="391" value="19-01-14">
<date id="FF988ECC560EFFF764A2B1BD88A7FB32" box="[1163,1251,1247,1272]" pageId="1" pageNumber="391" value="19-01-14">140119</date>
</collectingDate>
/67/ 
<collectingDate id="EFDC7724560EFFF7653BB1BD8931FB32" box="[1298,1397,1247,1272]" pageId="1" pageNumber="391" value="2001-05-02">
<date id="FF988ECC560EFFF7653BB1BD8931FB32" box="[1298,1397,1247,1272]" pageId="1" pageNumber="391" value="2001-05-02">01-02-05</date>
</collectingDate>
); 1 hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF761BEB0668DBFFAD6" box="[407,507,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="391" unit="mm" value="4.1">4.1 mm</quantity>
), 
<location id="8EF9FED7560EFFF76230B0678ECFFAD7" LSID="urn:lsid:plazi:treatment:038F191A560EFFFF60BEB61C886CFE3E:8EF9FED7560EFFF76230B0678ECFFAD7" box="[537,651,1284,1309]" country="Saint Helena, Ascension and Tristan da Cunha" name="Papa Nui" pageId="1" pageNumber="391" stateProvince="St. Helena">Papa Nui</location>
wreck, -15.922° -5.72°, leg. 
<collectorName id="26D3CDDA560EFFF7642CB067882DFAD7" box="[1029,1129,1284,1309]" pageId="1" pageNumber="391">P. Wirtz</collectorName>
&amp; 
<collectorName id="26D3CDDA560EFFF76488B0678950FAD6" box="[1185,1300,1285,1308]" pageId="1" pageNumber="391">J. Brown</collectorName>
, 
<collectingDate id="EFDC7724560EFFF76503B06689D3FAD6" box="[1322,1431,1284,1308]" pageId="1" pageNumber="391" value="2014-01-23">
<date id="FF988ECC560EFFF76503B06689D3FAD6" box="[1322,1431,1284,1308]" pageId="1" pageNumber="391" value="2014-01-23">23.I.2014</date>
</collectingDate>
, 
<collectionCode id="ED3730C9560EFFF760BEB0458D43FA8A" box="[151,263,1319,1344]" pageId="1" pageNumber="391">OUMNH</collectionCode>
</materialsCitation>
.ZC.2018-04-005 (fcn 
<date id="FF988ECC560EFFF7622EB04A8E19FA8A" box="[519,605,1320,1344]" pageId="1" pageNumber="391" value="23-01-14">140123</date>
/74/03); 1 ov. hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF763D0B0458812FAF5" box="[1017,1110,1319,1344]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="391" unit="mm" value="5.2">5.2 mm</quantity>
, dissected), same collection data, MZUSP 37298 (fcn 
<date id="FF988ECC560EFFF76190B02E8E4AFAAE" box="[441,526,1356,1380]" pageId="1" pageNumber="391" value="23-01-14">140123</date>
/74/04).
</paragraph>
<paragraph id="8B99A80C560EFFF760EEB00D8C96FA3E" blockId="1.[151,1437,1068,1993]" pageId="1" pageNumber="391">
<materialsCitation id="3B4EA251560EFFF760EEB00D8F3EFA66" ID-GBIF-Occurrence="1848505949" collectingDate="2015-07-16" collectionCode="OUMNH" collectorName="P. Wirtz" country="Saint Helena, Ascension and Tristan da Cunha" elevation="15" location="Non-type" municipality="Two Hook" pageId="1" pageNumber="391" specimenCount="1">
<emphasis id="B952741E560EFFF760EEB00D8EC2FA42" bold="true" box="[199,646,1391,1416]" pageId="1" pageNumber="391">
<location id="8EF9FED7560EFFF760EEB00D8D71FA42" LSID="urn:lsid:plazi:treatment:038F191A560EFFFF60BEB61C886CFE3E:8EF9FED7560EFFF760EEB00D8D71FA42" box="[199,309,1391,1416]" country="Saint Helena, Ascension and Tristan da Cunha" municipality="Two Hook" name="Non-type" pageId="1" pageNumber="391">Non-type</location>
material. 
<collectingCountry id="F331E89C560EFFF7619AB00D8E6FFA42" box="[435,555,1391,1416]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="391">Ascension</collectingCountry>
Island.
</emphasis>
1 ov. hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF76397B00D885EFA42" box="[958,1050,1391,1416]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="391" unit="mm" value="5.2">5.2 mm</quantity>
), 1 hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF7651DB00D89D4FA42" box="[1332,1424,1391,1416]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="391" unit="mm" value="3.6">3.6 mm</quantity>
), 
<collectingMunicipality id="6BFD3276560EFFF760BEB0F78D4BFA67" box="[151,271,1428,1453]" pageId="1" pageNumber="391">Two Hook</collectingMunicipality>
dive site, 
<elevation id="000B4F3F560EFFF761A1B0F68D81FA66" box="[392,453,1428,1453]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="391" unit="m" value="15.0">
<quantity id="4CDE05E9560EFFF761A1B0F68D81FA66" box="[392,453,1428,1453]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="391" unit="m" value="15.0">15 m</quantity>
</elevation>
, leg. 
<collectorName id="26D3CDDA560EFFF7622CB0F78E27FA67" box="[517,611,1428,1453]" pageId="1" pageNumber="391">P. Wirtz</collectorName>
, 
<collectingDate id="EFDC7724560EFFF76246B0F68EBFFA67" box="[623,763,1428,1453]" pageId="1" pageNumber="391" value="2015-07-16">
<date id="FF988ECC560EFFF76246B0F68EBFFA67" box="[623,763,1428,1453]" pageId="1" pageNumber="391" value="2015-07-16">16.VII.2015</date>
</collectingDate>
, 
<collectionCode id="ED3730C9560EFFF76321B0F68F3EFA66" box="[776,890,1428,1452]" pageId="1" pageNumber="391">OUMNH</collectionCode>
</materialsCitation>
.ZC.2018-01-024; 1 hermaphrodite (pocl 
<quantity id="4CDE05E9560EFFF76563B0F689CBFA66" box="[1354,1423,1428,1453]" metricMagnitude="0" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="391" unit="m" value="3.6">3.6 m</quantity>
), Red Rock, -7.89435° -14.39475°, 
<quantity id="4CDE05E9560EFFF7620CB0D58E2EFA05" box="[549,618,1463,1488]" metricMagnitude="0" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="391" unit="m" value="8.5">8.5 m</quantity>
, leg. A. Richardson, 
<date id="FF988ECC560EFFF7634DB0D58FB4FA1A" box="[868,1008,1463,1488]" pageId="1" pageNumber="391" value="2015-07-20">20.VII.2015</date>
, OUMNH.ZC.2018-01-025 (fcn C20 009).
</paragraph>
</subSubSection>
<subSubSection id="C33CFB87560EFFF260EEB09D8EA1F8BA" lastPageId="4" lastPageNumber="394" pageId="1" pageNumber="391" type="description">
<paragraph id="8B99A80C560EFFF760EEB09D8E90F906" blockId="1.[151,1437,1068,1993]" pageId="1" pageNumber="391">
<emphasis id="B952741E560EFFF760EEB09D8D1DF9D2" bold="true" box="[199,345,1535,1560]" pageId="1" pageNumber="391">Description.</emphasis>
Rostrum (
<figureCitation id="131DB489560EFFF761FCB09D8E33F9D2" box="[469,631,1535,1560]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1a, b, j–l</figureCitation>
) elongate, stout, about four times as long as carapace, reaching midlength of second article of antennular peduncle; dorsal margin with five teeth (
<typeStatus id="549D16AE560EFFF76434B34688C1F9F7" box="[1053,1157,1572,1597]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
), most posterior tooth at about 0.25 of carapace length, third tooth close to post-orbital margin or slightly more anterior; ventral margin with two or three teeth (three in 
<typeStatus id="549D16AE560EFFF761C6B30E8E11F94F" box="[495,597,1644,1669]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
), relatively distal, tip not subdivided. Carapace smooth, with rounded posteroventral margin (
<figureCitation id="131DB489560EFFF7618CB3ED8DBDF962" box="[421,505,1679,1704]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1c</figureCitation>
); pterygostomial angle with small, acute tooth (
<figureCitation id="131DB489560EFFF76406B3ED88C1F962" box="[1071,1157,1679,1704]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1b</figureCitation>
); antennal tooth strong, acute, falling short of middle of cornea (
<figureCitation id="131DB489560EFFF76273B3D68E83F906" box="[602,711,1716,1741]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1a–c</figureCitation>
).
</paragraph>
<paragraph id="8B99A80C560EFFF760EEB3B58FC2F803" blockId="1.[151,1437,1068,1993]" pageId="1" pageNumber="391">
First three pleonites ventrally rounded; fourth somite with square (about 90°) posterolateral angle; fifth somite (
<figureCitation id="131DB489560EFFF760B6B39E8CABF8DF" box="[159,239,1788,1813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1d</figureCitation>
) with well-developed, distally projecting posterolateral angle; sixth somite about 1.5 as long as fifth, with square posteroventral angle (
<figureCitation id="131DB489560EFFF761C9B27D8E77F8F2" box="[480,563,1823,1848]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1d</figureCitation>
); posterolateral margin of sixth pleuron with well-developed polygonal lobe above articulation with uropod, posteromedial part developed into broad tooth. Telson (
<figureCitation id="131DB489560EFFF76459B22688FBF896" box="[1136,1215,1860,1885]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="1" pageNumber="391">Fig. 1e</figureCitation>
) about 2.5 times as long as wide, tapering posteriorly; dorsal surface with two pairs of spines, approximately inserted at 0.4 and 0.6 of telson length, respectively (
<typeStatus id="549D16AE560EFFF761FFB2EE8E7AF86F" box="[470,574,1932,1957]" pageId="1" pageNumber="391" type="holotype">holotype</typeStatus>
); posterior margin medially bluntly produced, furnished with two pairs of spiniform setae, mesial pair about four times as long as lateral pair.
</paragraph>
<caption id="DF59F884560DFFF460BEB2348E80F82D" httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="2" pageNumber="392" startId="2.[151,250,1878,1900]" targetBox="[157,1404,196,1850]" targetPageId="2">
<paragraph id="8B99A80C560DFFF460BEB2348E80F82D" blockId="2.[151,1437,1878,2023]" pageId="2" pageNumber="392">
<emphasis id="B952741E560DFFF460BEB2348D49F8A6" bold="true" box="[151,269,1878,1900]" pageId="2" pageNumber="392">FIGURE 1</emphasis>
. 
<taxonomicName id="4C26D38F560DFFF46132B23A8D98F8A6" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[283,476,1878,1901]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="392" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E560DFFF46132B23A8D98F8A6" box="[283,476,1878,1901]" italics="true" pageId="2" pageNumber="392">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C965560DFFF461CAB2358E70F8A6" box="[483,564,1879,1900]" pageId="2" pageNumber="392" rank="species">
<emphasis id="B952741E560DFFF461CAB2358E70F8A6" bold="true" box="[483,564,1879,1900]" pageId="2" pageNumber="392">sp. nov.</emphasis>
</taxonomicNameLabel>
, (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004.
</paragraph>
</caption>
<caption id="DF59F884560CFFF560BEB3BF8E76F8FA" httpUri="https://zenodo.org/record/1283961/files/figure.png" pageId="3" pageNumber="393" startId="3.[151,250,1757,1779]" targetBox="[155,1380,200,1724]" targetPageId="3">
<paragraph id="8B99A80C560CFFF560BEB3BF8E76F8FA" blockId="3.[151,1436,1756,1840]" pageId="3" pageNumber="393">
<emphasis id="B952741E560CFFF560BEB3BF8D57F938" bold="true" box="[151,275,1757,1779]" pageId="3" pageNumber="393">FIGURE 2.</emphasis>
<taxonomicName id="4C26D38F560CFFF56133B3BC8D9EF938" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[282,474,1756,1779]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="393" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E560CFFF56133B3BC8D9EF938" box="[282,474,1756,1779]" italics="true" pageId="3" pageNumber="393">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C965560CFFF561CBB3BF8E76F938" box="[482,562,1757,1778]" pageId="3" pageNumber="393" rank="species">
<emphasis id="B952741E560CFFF561CBB3BF8E76F938" bold="true" box="[482,562,1757,1778]" pageId="3" pageNumber="393">sp. nov.</emphasis>
</taxonomicNameLabel>
, (a) right first pereiopod, lateral view; b) same, ischium, mesial view; (c) same, chela and distal portion of carpus, ventromesial view; (d) right second pereiopod, lateral view; (e) same, chela, mesial view. All from paratype (pocl 5.2 mm), MZUSP 37298.
</paragraph>
</caption>
<paragraph id="8B99A80C560CFFF260EEB2358FEDFF1F" blockId="3.[151,1436,1879,2012]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="394" pageId="3" pageNumber="393">
Antennular peduncle (
<figureCitation id="131DB489560CFFF561EDB2358E70F8BA" box="[452,564,1879,1904]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="3" pageNumber="393">Fig. 1a, b</figureCitation>
) about 0.7 times as long as scaphocerite; first article about 2.2–2.5 times as long as second; third article about half as long as second; distodorsal margins of second and third articles furnished with few small spinules; stylocerite reaching distal margin of first article; lateral antennular flagellum long; accessory branch (
<figureCitation id="131DB489560CFFF56147B2A18DF9F816" box="[366,445,1987,2012]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="3" pageNumber="393">Fig. 1f</figureCitation>
) with 11 free divisions (
<typeStatus id="549D16AE560CFFF562C8B2A18F0DF816" box="[737,841,1987,2012]" pageId="3" pageNumber="393" type="holotype">holotype</typeStatus>
), fused portion with nine divisions (
<typeStatus id="549D16AE560CFFF564DBB2A1891EF816" box="[1266,1370,1987,2012]" pageId="3" pageNumber="393" type="holotype">holotype</typeStatus>
), free portion about 1.7 times as long as fused portion (
<typeStatus id="549D16AE560BFFF262E1B5F58F74FF7A" box="[712,816,151,176]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
). Scaphocerite about 3.5 times as long as wide; lateral margin straight; distolateral tooth overreaching distal margin of blade.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB5BD8E8BFEF5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
Third maxilliped (
<figureCitation id="131DB489560BFFF261BDB5BD8DA1FF32" box="[404,485,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="4" pageNumber="394">
Fig. 
<quantity id="4CDE05E9560BFFF261E0B5828DA1FF32" box="[457,485,224,248]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="4" pageNumber="394" unit="g" value="1.0">1g</quantity>
</figureCitation>
) overreaching scaphocerite by distal half of ultimate article; exopod about 0.5 times as long as antepenultimate article of endopod; penultimate article about 0.4 times as long as ultimate article; tip with three subdistal and one distal spiniform setae.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB42E8D56FE1A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
First pereiopod (
<figureCitation id="131DB489560BFFF26156B42E8DADFEAE" box="[383,489,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1757,1779]" captionTargetBox="[155,1380,200,1724]" captionTargetId="figure@3.[151,1436,193,1735]" captionTargetPageId="3" captionText="FIGURE 2. Lysmata napoleoni sp. nov., (a) right first pereiopod, lateral view; b) same, ischium, mesial view; (c) same, chela and distal portion of carpus, ventromesial view; (d) right second pereiopod, lateral view; (e) same, chela, mesial view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283961/files/figure.png" pageId="4" pageNumber="394">Fig. 2a–c</figureCitation>
) reaching past scaphocerite by length of fingers when extended; ischium with row of spinules along ventral margin; merus about 1.2 times as long as carpus; chela about 1.2 times as long as carpus, with palm about 1.6 times as long as fingers; tips of both pollex and dactylus simple; carpo-propodal brush well developed.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB4BE8DFAFD4F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
Second pereiopod (
<figureCitation id="131DB489560BFFF26188B4BE8E4AFE3E" box="[417,526,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1757,1779]" captionTargetBox="[155,1380,200,1724]" captionTargetId="figure@3.[151,1436,193,1735]" captionTargetPageId="3" captionText="FIGURE 2. Lysmata napoleoni sp. nov., (a) right first pereiopod, lateral view; b) same, ischium, mesial view; (c) same, chela and distal portion of carpus, ventromesial view; (d) right second pereiopod, lateral view; (e) same, chela, mesial view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283961/files/figure.png" pageId="4" pageNumber="394">Fig. 2d, e</figureCitation>
) slender, when fully extended overreaching scaphocerite by distal 2/3 of carpus, ending in small, simple chelae; right and left pereiopod subequal, left second pereiopod proportionally longer than right by only about 0.2 of length (
<typeStatus id="549D16AE560BFFF26236B7468EC3FDF7" box="[543,647,548,573]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
); ischium with two–three weak, distal annulations; merus subdivided into 14 (right) and 15 (left) divisions (
<typeStatus id="549D16AE560BFFF2626EB7258EEBFDAA" box="[583,687,583,608]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
); carpus twice as long as merus, subdivided into 24 (right) and 25 (left) divisions (
<typeStatus id="549D16AE560BFFF26163B70E8DF6FD4F" box="[330,434,620,645]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB7ED895DFC02" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
Third to fifth pereiopods generally similar, decreasing in length (
<figureCitation id="131DB489560BFFF263F7B7ED886EFD62" box="[990,1066,655,680]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1963]" captionTargetBox="[167,1395,217,1889]" captionTargetId="figure@5.[161,1421,193,1913]" captionTargetPageId="5" captionText="FIGURE 3. Lysmata napoleoni sp. nov., (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283963/files/figure.png" pageId="4" pageNumber="394">Fig. 3</figureCitation>
). Third pereiopod (
<figureCitation id="131DB489560BFFF26530B7ED89D0FD62" box="[1305,1428,655,680]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1963]" captionTargetBox="[167,1395,217,1889]" captionTargetId="figure@5.[161,1421,193,1913]" captionTargetPageId="5" captionText="FIGURE 3. Lysmata napoleoni sp. nov., (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283963/files/figure.png" pageId="4" pageNumber="394">Fig. 3a, b</figureCitation>
) overreaching scaphocerite by entire length of propodus; merus about 3.4 times as long as ischium, mesially with five spines (
<typeStatus id="549D16AE560BFFF26105B7B58DD5FD3A" box="[300,401,727,752]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
) and additional two spines present along ventral margin on right side, six and three, respectively, spines on left side; carpus about half as long as merus, with four spiniform setae on ventral margin (
<typeStatus id="549D16AE560BFFF260B6B67D8D43FCF2" box="[159,263,799,824]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
); propodus about 1.4 times as long as carpus, ventrally armed (
<typeStatus id="549D16AE560BFFF263E8B67D8863FCF2" box="[961,1063,799,824]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
) with six spiniform setae on right side, nine on left side, distal of those paired; dactylus about 0.2 as long as propodus, biunguiculate, with dorsal unguis larger than ventral one; flexor margin with additional three spinules. Fourth pereiopod (
<figureCitation id="131DB489560BFFF264F8B6058966FC4A" box="[1233,1314,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1963]" captionTargetBox="[167,1395,217,1889]" captionTargetId="figure@5.[161,1421,193,1913]" captionTargetPageId="5" captionText="FIGURE 3. Lysmata napoleoni sp. nov., (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283963/files/figure.png" pageId="4" pageNumber="394">Fig. 3c</figureCitation>
) generally similar to third, with four spines in mesial series on merus (
<typeStatus id="549D16AE560BFFF2637FB6EE8FF8FC6F" box="[854,956,908,933]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
, both sides) and single ventral one. Fifth pereiopod (
<figureCitation id="131DB489560BFFF2613FB6CD8D3BFC02" box="[278,383,943,968]" captionStart="FIGURE 3" captionStartId="5.[151,250,1941,1963]" captionTargetBox="[167,1395,217,1889]" captionTargetId="figure@5.[161,1421,193,1913]" captionTargetPageId="5" captionText="FIGURE 3. Lysmata napoleoni sp. nov., (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283963/files/figure.png" pageId="4" pageNumber="394">Fig. 3d–f</figureCitation>
) similar, with one (right) or two (left) spines in mesial series on merus (
<typeStatus id="549D16AE560BFFF2648CB6CD8949FC02" box="[1189,1293,943,968]" pageId="4" pageNumber="394" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB6B68CA0FBFF" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
Uropod (
<figureCitation id="131DB489560BFFF26119B6B68D38FC27" box="[304,380,980,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="4" pageNumber="394">
Fig. 
<date id="FF988ECC560BFFF26141B6B68D38FC27" box="[360,380,980,1005]" pageId="4" pageNumber="394">1i</date>
</figureCitation>
) with relatively slender exopod and endopod, latter with well-developed diaeresis bearing three spaced acute or subacute teeth, lateral-most strongest, distolateral tooth and adjacent spiniform seta both strong.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB15D8DABFAB2" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
Colour pattern (
<figureCitation id="131DB489560BFFF26157B15D8D94FB92" box="[382,464,1087,1112]" captionStart="FIGURE 4" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Lysmata napoleoni sp. nov. on St. Helena Island, (a) paratype, OUMNH.ZC. 2018-04-004; (b) ovigerous hermaphrodite, not deposited; (c) – (d) paratype, OUMNH.ZC. 2018-04-004, showing association with the sea anemone Telmactis cricoides (Duchassaing). Photographs by Peter Wirtz (a, b, c) and Judith Brown (d)." httpUri="https://zenodo.org/record/1283965/files/figure.png" pageId="4" pageNumber="394">Figs. 4</figureCitation>
, 
<figureCitation id="131DB489560BFFF261F6B15D8DABFB92" box="[479,495,1087,1112]" captionStart="FIGURE 5" captionStartId="8.[151,250,1980,2002]" captionTargetBox="[151,1436,193,1959]" captionTargetId="figure@8.[151,1436,193,1959]" captionTargetPageId="8" captionText="FIGURE 5. Lysmata napoleoni sp. nov. from Ascension Island, specimens not collected, (a) colour pattern by day; (b) nocturnal cleaning behaviour. Photographs by Judith Brown." httpUri="https://zenodo.org/record/1283967/files/figure.png" pageId="4" pageNumber="394">5</figureCitation>
): body mostly deep red, except some semi-transparent areas on anterior carapace flanks, two conspicuous white transverse bands, both quite irregular in shape, one running across posterior margin of carapace and another across middle of third pleonite, as well as series of white spots or patches of various sizes, on dorsal surface of carapace and on pleon, in latter case typically arranged in transverse row of spaced spots (for example, as dorsal, subdorsal, lateral and ventrolateral), or forming two continuing bands running ventrolaterally on each side of pleon, from third pleomere to sixth pleonite; ventral base of each eye peduncle and area between them also white; antennular and antennal peduncles, their flagella, first to fifth pereiopods and tail fan mostly deep red or reddish, with occasional larger white spots or bands, latter especially well marked on telson and uropodal margin; fresh eggs orange-red.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB0E688BCF99A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
<emphasis id="B952741E560BFFF260EEB0E68DAFFA57" bold="true" box="[199,491,1412,1437]" pageId="4" pageNumber="394">Morphological variation</emphasis>
. In terms of rostral variation, 
<taxonomicName id="4C26D38F560BFFF26369B0E78F8AFA57" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[832,974,1412,1437]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E560BFFF26369B0E78F8AFA57" box="[832,974,1412,1437]" italics="true" pageId="4" pageNumber="394">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C965560BFFF263FFB0E68875FA57" box="[982,1073,1412,1437]" pageId="4" pageNumber="394" rank="species">
<emphasis id="B952741E560BFFF263FFB0E68875FA57" bold="true" box="[982,1073,1412,1437]" pageId="4" pageNumber="394">sp. nov.</emphasis>
</taxonomicNameLabel>
exhibits little variation, with all examined specimens possessing five dorsal teeth, although the position of the second and third proximal teeth differs slightly between specimens; approximately half of the specimens possess two ventral teeth, the other half have three (
<figureCitation id="131DB489560BFFF2610AB08D8DEFF9C2" box="[291,427,1519,1544]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="4" pageNumber="394">Fig. 1c, j–l</figureCitation>
). The length of the free versus fused portion of the accessory branch of the antennal flagellum varies between 1.1 and 2.2, with a median value of about 1.7; the number of divisions in the free part varies between 10 and 13, whereas the number of divisions in the fused part varies between 6 and 9.
</paragraph>
<paragraph id="8B99A80C560BFFF260EEB33E8EA1F8BA" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">The comparative lengths of the left and right second pereiopods differs slightly, with sometimes one being longer (left or right), and on occasion both being of approximately the same length. In terms of subdivision of the second pereiopod, the number of divisions of the merus ranges from 13 to 18, with less variation seen in the carpus (25–27). Typically, there is a slight difference (1–2 divisions) between the left and the right sides. The number of spines on the merus (combined for mesial and ventral series) varies significantly on the third pereiopod (3–9) with one individual having no spines at all. Less variation in this feature exists on the fourth (2–5) and fifth (1–2) pereiopods. The number of ventral spiniform setae on the third pereiopod propodus is also not constant, ranging from 6 to 10, with the distal one always being paired</paragraph>
</subSubSection>
<subSubSection id="C33CFB87560BFFF260EEB21E8FEBF872" pageId="4" pageNumber="394" type="etymology">
<paragraph id="8B99A80C560BFFF260EEB21E8FEBF872" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="394">
<emphasis id="B952741E560BFFF260EEB21E8D0AF85F" bold="true" box="[199,334,1916,1941]" pageId="4" pageNumber="394">Etymology.</emphasis>
The species is named after Napoléon Bonaparte (
<date id="FF988ECC560BFFF26354B21E8845F85F" box="[893,1025,1916,1941]" pageId="4" pageNumber="394" value="1769" valueMax="1821">1769–1821</date>
), arguably St. Helena’s most famous resident, from his exile to the island in 1815 to his death there in 1821.
</paragraph>
</subSubSection>
<subSubSection id="C33CFB87560BFFF260EEB2A68F71F817" box="[199,821,1988,2013]" pageId="4" pageNumber="394" type="materials_examined">
<paragraph id="8B99A80C560BFFF260EEB2A68F71F817" blockId="4.[151,1437,151,2013]" box="[199,821,1988,2013]" pageId="4" pageNumber="394">
<materialsCitation id="3B4EA251560BFFF260EEB2A68F71F817" ID-GBIF-Occurrence="1848505590" box="[199,821,1988,2013]" country="Saint Helena, Ascension and Tristan da Cunha" location="St. Helena" pageId="4" pageNumber="394" specimenCount="1" typeStatus="holotype">
<emphasis id="B952741E560BFFF260EEB2A68D21F817" bold="true" box="[199,357,1988,2013]" pageId="4" pageNumber="394">
<typeStatus id="549D16AE560BFFF260EEB2A68D46F817" box="[199,258,1988,2013]" pageId="4" pageNumber="394">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F331E89C560BFFF26145B2A68DA4F817" box="[364,480,1988,2013]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="4" pageNumber="394">St. Helena</collectingCountry>
Island, south-central Atlantic.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF59F884560AFFF360BEB2F78F56F822" httpUri="https://zenodo.org/record/1283963/files/figure.png" pageId="5" pageNumber="395" startId="5.[151,250,1941,1963]" targetBox="[167,1395,217,1889]" targetPageId="5">
<paragraph id="8B99A80C560AFFF360BEB2F78F56F822" blockId="5.[151,1436,1941,2024]" pageId="5" pageNumber="395">
<emphasis id="B952741E560AFFF360BEB2F78D56F860" bold="true" box="[151,274,1941,1963]" pageId="5" pageNumber="395">FIGURE 3.</emphasis>
<taxonomicName id="4C26D38F560AFFF36131B2F48D9CF861" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[280,472,1941,1963]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="395" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E560AFFF36131B2F48D9CF861" box="[280,472,1941,1963]" italics="true" pageId="5" pageNumber="395">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C965560AFFF361F7B2F78E6BF860" box="[478,559,1941,1962]" pageId="5" pageNumber="395" rank="species">
<emphasis id="B952741E560AFFF361F7B2F78E6BF860" bold="true" box="[478,559,1941,1962]" pageId="5" pageNumber="395">sp. nov.</emphasis>
</taxonomicNameLabel>
, (a) right third pereiopod, lateral view; (b) same, dactylus, lateral view; (c) right fourth pereiopod, lateral view; (d) right fifth pereiopod, lateral view; (e) same, dactylus, lateral view; (f) left fifth pereiopod, merus, lateral view. All from paratype (pocl 5.2 mm), MZUSP 37298.
</paragraph>
</caption>
<caption id="DF59F8845609FFF060BEB0BE880BF9E5" httpUri="https://zenodo.org/record/1283965/files/figure.png" pageId="6" pageNumber="396" startId="6.[151,250,1500,1522]" targetBox="[151,1436,193,1478]" targetPageId="6">
<paragraph id="8B99A80C5609FFF060BEB0BE880BF9E5" blockId="6.[151,1436,1500,1583]" pageId="6" pageNumber="396">
<emphasis id="B952741E5609FFF060BEB0BE8D5CFA3B" bold="true" box="[151,280,1500,1522]" pageId="6" pageNumber="396">FIGURE 4.</emphasis>
<taxonomicName id="4C26D38F5609FFF0610CB0BF8DAFFA38" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[293,491,1500,1522]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5609FFF0610CB0BF8DAFFA38" box="[293,491,1500,1522]" italics="true" pageId="6" pageNumber="396">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655609FFF061D1B0BE8E0AFA3B" box="[504,590,1500,1521]" pageId="6" pageNumber="396" rank="species">
<emphasis id="B952741E5609FFF061D1B0BE8E0AFA3B" bold="true" box="[504,590,1500,1521]" pageId="6" pageNumber="396">sp. nov.</emphasis>
</taxonomicNameLabel>
on St. Helena Island, (a) paratype, OUMNH.ZC. 2018-04-004; (b) ovigerous hermaphrodite, not deposited; (c) – (d) paratype, OUMNH.ZC. 2018-04-004, showing association with the sea anemone 
<emphasis id="B952741E5609FFF060BEB37B8CB3F9E5" box="[151,247,1561,1583]" italics="true" pageId="6" pageNumber="396">Telmactis</emphasis>
<taxonomicName id="4C26D38F5609FFF060D4B37B8DB2F9E5" authority="(Duchassaing)" authorityName="Duchassaing" baseAuthorityName="Duchassaing" baseAuthorityYear="1850" box="[253,502,1561,1583]" class="Anthozoa" family="Isophelliidae" genus="Telmatactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="396" phylum="Cnidaria" rank="species" species="cricoides">
<emphasis id="B952741E5609FFF060D4B37B8D1EF9E5" box="[253,346,1561,1583]" italics="true" pageId="6" pageNumber="396">cricoides</emphasis>
(Duchassaing)
</taxonomicName>
. Photographs by Peter Wirtz (a, b, c) and Judith Brown (d).
</paragraph>
</caption>
<subSubSection id="C33CFB875609FFF060EEB3348D03F959" pageId="6" pageNumber="396" type="distribution">
<paragraph id="8B99A80C5609FFF060EEB3348D03F959" blockId="6.[151,1436,1622,2007]" pageId="6" pageNumber="396">
<emphasis id="B952741E5609FFF060EEB3348D24F9A5" bold="true" box="[199,352,1622,1647]" pageId="6" pageNumber="396">Distribution.</emphasis>
Currently, the species is known only from 
<collectingRegion id="49E266EE5609FFF0636EB3358FF8F9A5" box="[839,956,1622,1647]" country="Saint Helena, Ascension and Tristan da Cunha" name="Saint Helena" pageId="6" pageNumber="396">
<collectingCountry id="F331E89C5609FFF0636EB3358FF8F9A5" box="[839,956,1622,1647]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="396">St. Helena</collectingCountry>
</collectingRegion>
and 
<collectingRegion id="49E266EE5609FFF063DDB334882DF9A5" box="[1012,1129,1622,1647]" country="Saint Helena, Ascension and Tristan da Cunha" name="Ascension" pageId="6" pageNumber="396">
<collectingCountry id="F331E89C5609FFF063DDB334882DF9A5" box="[1012,1129,1622,1647]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="6" pageNumber="396">Ascension</collectingCountry>
</collectingRegion>
Island, in the south-central Atlantic Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C33CFB875609FFF160EEB3FC8D20FDF7" lastPageId="7" lastPageNumber="397" pageId="6" pageNumber="396" type="biology_ecology">
<paragraph id="8B99A80C5609FFF060EEB3FC8F93F88D" blockId="6.[151,1436,1622,2007]" pageId="6" pageNumber="396">
<emphasis id="B952741E5609FFF060EEB3FC8D6FF97D" bold="true" box="[199,299,1694,1719]" pageId="6" pageNumber="396">Ecology.</emphasis>
All specimens of 
<taxonomicName id="4C26D38F5609FFF061D6B3FD8E92F97D" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[511,726,1694,1719]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5609FFF061D6B3FD8E92F97D" box="[511,726,1694,1719]" italics="true" pageId="6" pageNumber="396">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655609FFF062F6B3FC8F7FF97D" box="[735,827,1694,1719]" pageId="6" pageNumber="396" rank="species">
<emphasis id="B952741E5609FFF062F6B3FC8F7FF97D" bold="true" box="[735,827,1694,1719]" pageId="6" pageNumber="396">sp. nov.</emphasis>
</taxonomicNameLabel>
were collected while scuba diving at 
<quantity id="4CDE05E95609FFF064C6B3FC8936F97C" box="[1263,1394,1694,1719]" metricMagnitude="1" metricUnit="m" metricValue="1.31" metricValueMax="1.77" metricValueMin="0.85" pageId="6" pageNumber="396" unit="m" value="13.1" valueMax="17.7" valueMin="8.5">8.5–17.7 m</quantity>
, on hard or mixed (rocky-sandy) bottoms. The new species appears to be always associated with – or at least is found in the vicinity of – the sea anemone 
<taxonomicName id="4C26D38F5609FFF0626EB3848FA1F935" authority="(Duchassaing)" authorityName="Duchassaing" baseAuthorityName="Duchassaing" baseAuthorityYear="1850" box="[583,997,1766,1791]" class="Anthozoa" family="Isophelliidae" genus="Telmatactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="396" phylum="Cnidaria" rank="species" species="cricoides">
<emphasis id="B952741E5609FFF0626EB3848F7DF935" box="[583,825,1766,1791]" italics="true" pageId="6" pageNumber="396">Telmatactis cricoides</emphasis>
(Duchassaing)
</taxonomicName>
(P. Wirtz, pers. comm.), although the exact nature of this association remains unknown. Photographic evidence (
<figureCitation id="131DB4895609FFF063D3B268880AF8E9" box="[1018,1102,1802,1827]" captionStart="FIGURE 5" captionStartId="8.[151,250,1980,2002]" captionTargetBox="[151,1436,193,1959]" captionTargetId="figure@8.[151,1436,193,1959]" captionTargetPageId="8" captionText="FIGURE 5. Lysmata napoleoni sp. nov. from Ascension Island, specimens not collected, (a) colour pattern by day; (b) nocturnal cleaning behaviour. Photographs by Judith Brown." httpUri="https://zenodo.org/record/1283967/files/figure.png" pageId="6" pageNumber="396">Fig. 5b</figureCitation>
) shows that the new species engages in nocturnal fish cleaning behaviour (see also 
<bibRefCitation id="EFB7D5FD5609FFF062D5B24D8F8FF88D" box="[764,971,1838,1863]" pageId="6" pageNumber="396" refId="ref4529" refString="Brown, J., Downes, R. J., Mrowicki, R. J., Nolan, E. L., Richardson, A. J., Swinnen, F. &amp; Wirtz, P. (2017) New records of marine invertebrates from Ascension Island (central Atlantic). Arquipelago, 33, 71 - 79." type="journal article">
Brown 
<emphasis id="B952741E5609FFF06366B24D8FC3F88D" box="[847,903,1838,1863]" italics="true" pageId="6" pageNumber="396">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8B99A80C5609FFF160EEB2308FCCFED6" blockId="6.[151,1436,1622,2007]" lastBlockId="7.[151,1437,151,1977]" lastPageId="7" lastPageNumber="397" pageId="6" pageNumber="396">
<emphasis id="B952741E5609FFF060EEB2308D85F8A1" bold="true" box="[199,449,1874,1899]" pageId="6" pageNumber="396">Taxonomic remarks.</emphasis>
On account of the length of the free portion of the accessory branch of the lateral antennular flagellum, 
<taxonomicName id="4C26D38F5609FFF061B1B2158E36F845" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[408,626,1910,1935]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5609FFF061B1B2158E36F845" box="[408,626,1910,1935]" italics="true" pageId="6" pageNumber="396">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655609FFF06257B2148E99F845" box="[638,733,1910,1935]" pageId="6" pageNumber="396" rank="species">
<emphasis id="B952741E5609FFF06257B2148E99F845" bold="true" box="[638,733,1910,1935]" pageId="6" pageNumber="396">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the so-called “long-branch group” of the genus 
<taxonomicName id="4C26D38F5609FFF060BEB2F98DA2F879" authority="(Fiedler et al. 2010)" authorityName="Fiedler et al." authorityYear="2010" box="[151,486,1946,1971]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5609FFF060BEB2F98CB0F878" box="[151,244,1947,1970]" italics="true" pageId="6" pageNumber="396">Lysmata</emphasis>
(
<bibRefCitation id="EFB7D5FD5609FFF0612CB2F88D9AF879" box="[261,478,1946,1971]" pageId="6" pageNumber="396" refId="ref5090" refString="Fiedler, G. C., Rhyne, A. L., Segawa, R., Aotsuka, T. &amp; Schizas, N. V. (2010) The evolution of euhermaphroditism in caridean shrimps: a molecular perspective of sexual systems and systematics. BMC Evolutionary Biology, 10 (e 297), 1 - 14. https: // doi. org / 10.1186 / 1471 - 2148 - 10 - 297" type="journal article">
Fiedler 
<emphasis id="B952741E5609FFF06176B2F98DDEF879" box="[351,410,1946,1971]" italics="true" pageId="6" pageNumber="396">et al.</emphasis>
2010
</bibRefCitation>
)
</taxonomicName>
, which at least partly corresponds to the “cosmopolitan clade” in the phylogeny of 
<bibRefCitation id="EFB7D5FD5609FFF060BEB2DD8D74F81D" box="[151,304,1983,2007]" pageId="6" pageNumber="396" refId="ref4444" refString="Baeza, J. A. (2010) Molecular systematics of peppermint and cleaner shrimps: phylogeny and taxonomy of the genera Lysmata and Exhippolysmata (Crustacea: Caridea: Hippolytidae). Zoological Journal of the Linnean Society, 160, 254 - 265. https: // doi. org / 10.1111 / j. 1096 - 3642.2009.00605. x" type="journal article">Baeza (2010)</bibRefCitation>
. In the eastern Atlantic, only three further species belong to this clade, viz. 
<taxonomicName id="4C26D38F5609FFF06452B2DD88A3F81C" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[1147,1255,1983,2006]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5609FFF06452B2DD88A3F81C" box="[1147,1255,1983,2006]" italics="true" pageId="6" pageNumber="396">L. moorei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C26D38F5609FFF164DCB2DD8D1EFF7A" authority="Dohrn &amp; Holthuis, 1950" authorityName="Dohrn &amp; Holthuis" authorityYear="1950" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="7" lastPageNumber="397" order="Decapoda" pageId="6" pageNumber="396" phylum="Arthropoda" rank="species" species="nilita">
<emphasis id="B952741E5609FFF064DCB2DD8909F81D" box="[1269,1357,1982,2007]" italics="true" pageId="6" pageNumber="396">L. nilita</emphasis>
<bibRefCitation id="EFB7D5FD5609FFF1657DB2DC8D1EFF7A" lastPageId="7" lastPageNumber="397" pageId="6" pageNumber="396" refId="ref4959" refString="Dohrn, P. F. R. &amp; Holthuis, L. B. (1950) Lysmata nilita, a new species of prawn (Crustacea Decapoda) from the western Mediterranean. Pubblicazioni della Stazione Zoologica di Napoli, 22, 339 - 347." type="journal article">Dohrn &amp; Holthuis, 1950</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C26D38F5608FFF161B8B5FB8E74FF7A" baseAuthorityName="Risso" baseAuthorityYear="1816" box="[401,560,151,176]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF161B8B5FB8E74FF7A" box="[401,560,151,176]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
. The western Atlantic 
<taxonomicName id="4C26D38F5608FFF16318B5FB88CFFF7A" authority="(Kingsley, 1879)" authorityName="Kingsley" authorityYear="1879" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[817,1163,151,176]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF16318B5FB8F83FF7A" box="[817,967,151,176]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
(
<bibRefCitation id="EFB7D5FD5608FFF163FFB5F588C7FF7A" box="[982,1155,151,176]" pageId="7" pageNumber="397" refId="ref5474" refString="Kingsley, J. S. (1879) List of the North American Crustacea belonging to the suborder Caridea. Bulletin of the Essex Institute, 10 [for 1878], 53 - 71." type="journal article">Kingsley, 1879</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C26D38F5608FFF164EBB5FB8D89FF1E" authority="Rhyne, Calado &amp; dos Santos, 2012" authorityName="Rhyne, Calado &amp; dos Santos" authorityYear="2012" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="jundalini">
<emphasis id="B952741E5608FFF164EBB5FB8900FF7A" box="[1218,1348,151,176]" italics="true" pageId="7" pageNumber="397">L. jundalini</emphasis>
Rhyne, Calado &amp; dos Santos, 2012
</taxonomicName>
also belong to this clade (
<bibRefCitation id="EFB7D5FD5608FFF162D0B5DE884AFF1E" box="[761,1038,188,213]" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz 2000</bibRefCitation>
; 
<bibRefCitation id="EFB7D5FD5608FFF16432B5DE88AFFF1E" box="[1051,1259,188,213]" pageId="7" pageNumber="397" refId="ref5775" refString="Rhyne, A. L., Calado, R. &amp; dos Santos, A. (2012) Lysmata jundalini, a new peppermint shrimp (Decapoda, Caridea, Hippolytidae) from the western Atlantic. Zootaxa, 3579, 71 - 79." type="journal article">
Rhyne 
<emphasis id="B952741E5608FFF16445B5DF88E2FF1F" box="[1132,1190,188,213]" italics="true" pageId="7" pageNumber="397">et al.</emphasis>
2012
</bibRefCitation>
). All remaining Atlantic species of 
<taxonomicName id="4C26D38F5608FFF16151B5838D91FF32" box="[376,469,225,248]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5608FFF16151B5838D91FF32" box="[376,469,225,248]" italics="true" pageId="7" pageNumber="397">Lysmata</emphasis>
</taxonomicName>
can be easily differentiated from the new species by their very short or unguiform accessory branch (see 
<bibRefCitation id="EFB7D5FD5608FFF161BBB4668E2FFED6" box="[402,619,260,285]" pageId="7" pageNumber="397" refId="ref4405" refString="Anker, A. &amp; Cox, D. (2011) A new species of the shrimp genus Lysmata Risso, 1816 (Crustacea, Decapoda) from Guam. Micronesica, 41, 197 - 214." type="journal article">Anker &amp; Cox 2011</bibRefCitation>
; 
<bibRefCitation id="EFB7D5FD5608FFF1625EB4668F38FED6" box="[631,892,260,285]" pageId="7" pageNumber="397" refId="ref5552" refString="Prakash, S. &amp; Baeza, J. A. (2017) A new species of Lysmata Risso, 1816 (Crustacea, Decapoda, Lysmatidae) from the Gulf of Mexico. Zootaxa, 4363 (4), 576 - 582. https: // doi. org / 10.11646 / zootaxa. 4363.4.10" type="journal article">Prakash &amp; Baeza 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8B99A80C5608FFF160EEB4458D20FDF7" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="397">
Morphologically, 
<taxonomicName id="4C26D38F5608FFF161B8B44B8E2CFE8A" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[401,616,295,320]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF161B8B44B8E2CFE8A" box="[401,616,295,320]" italics="true" pageId="7" pageNumber="397">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF16258B4458E89FE8A" box="[625,717,295,320]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF16258B4458E89FE8A" bold="true" box="[625,717,295,320]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
may be separated from both 
<taxonomicName id="4C26D38F5608FFF16436B44B88FCFE8A" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[1055,1208,295,320]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF16436B44B88FCFE8A" box="[1055,1208,295,320]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C26D38F5608FFF164DBB44B8933FE8A" authorityName="Rhyne, Calado &amp; dos Santos" authorityYear="2012" box="[1266,1399,295,320]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="jundalini">
<emphasis id="B952741E5608FFF164DBB44B8933FE8A" box="[1266,1399,295,320]" italics="true" pageId="7" pageNumber="397">L. jundalini</emphasis>
</taxonomicName>
by the number of fused and free divisions in the accessory branch of the lateral antennular flagellum, which is 17–18 (fused) and 3–4 (free) in 
<taxonomicName id="4C26D38F5608FFF16199B4138E03FE42" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[432,583,367,392]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF16199B4138E03FE42" box="[432,583,367,392]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
</taxonomicName>
, and 18–24 (fused) and 3 (free) in 
<taxonomicName id="4C26D38F5608FFF163F9B4138817FE42" authorityName="Rhyne, Calado &amp; dos Santos" authorityYear="2012" box="[976,1107,367,392]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="jundalini">
<emphasis id="B952741E5608FFF163F9B4138817FE42" box="[976,1107,367,392]" italics="true" pageId="7" pageNumber="397">L. jundalini</emphasis>
</taxonomicName>
, versus 10–13 (free) and 6–9 (fused) in 
<taxonomicName id="4C26D38F5608FFF16125B4F78DDFFE67" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[268,411,404,429]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF16125B4F78DDFFE67" box="[268,411,404,429]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF1618AB4F68DBAFE67" box="[419,510,404,429]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF1618AB4F68DBAFE67" bold="true" box="[419,510,404,429]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, as well as by the relative proportions of these two parts, with the free part being less than 0.15 of the fused part in both 
<taxonomicName id="4C26D38F5608FFF16272B4DB8EB0FE1A" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[603,756,439,464]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF16272B4DB8EB0FE1A" box="[603,756,439,464]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C26D38F5608FFF16307B4DB8FF7FE1A" authorityName="Rhyne, Calado &amp; dos Santos" authorityYear="2012" box="[814,947,439,464]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="jundalini">
<emphasis id="B952741E5608FFF16307B4DB8FF7FE1A" box="[814,947,439,464]" italics="true" pageId="7" pageNumber="397">L. jundalini</emphasis>
</taxonomicName>
vs. being much longer in 
<taxonomicName id="4C26D38F5608FFF164CBB4DB8936FE1A" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[1250,1394,439,464]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF164CBB4DB8936FE1A" box="[1250,1394,439,464]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF16553B4D58C8CFE3F" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF16553B4D58C8CFE3F" bold="true" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
(1.15–2.17) (see 
<bibRefCitation id="EFB7D5FD5608FFF161A5B4BE8EDAFE3E" box="[396,670,476,501]" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz 2000</bibRefCitation>
; 
<bibRefCitation id="EFB7D5FD5608FFF16280B4BE8F30FE3E" box="[681,884,476,501]" pageId="7" pageNumber="397" refId="ref5775" refString="Rhyne, A. L., Calado, R. &amp; dos Santos, A. (2012) Lysmata jundalini, a new peppermint shrimp (Decapoda, Caridea, Hippolytidae) from the western Atlantic. Zootaxa, 3579, 71 - 79." type="journal article">
Rhyne 
<emphasis id="B952741E5608FFF162D0B4BF8F75FE3F" box="[761,817,476,501]" italics="true" pageId="7" pageNumber="397">et al.</emphasis>
2012
</bibRefCitation>
). On the same basis, 
<taxonomicName id="4C26D38F5608FFF16476B4BF88A9FE3F" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[1119,1261,476,501]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF16476B4BF88A9FE3F" box="[1119,1261,476,501]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF164DDB4BE8909FE3F" box="[1268,1357,476,501]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF164DDB4BE8909FE3F" bold="true" box="[1268,1357,476,501]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="4C26D38F5608FFF16159B7638D8CFDD2" authorityName="Dohrn &amp; Holthuis" authorityYear="1950" box="[368,456,511,536]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="nilita">
<emphasis id="B952741E5608FFF16159B7638D8CFDD2" box="[368,456,511,536]" italics="true" pageId="7" pageNumber="397">L. nilita</emphasis>
</taxonomicName>
, in which the free part of the accessory branch is less than 0.33 of the fused part (
<bibRefCitation id="EFB7D5FD5608FFF1657AB49D8D1CFDF7" pageId="7" pageNumber="397" refId="ref4959" refString="Dohrn, P. F. R. &amp; Holthuis, L. B. (1950) Lysmata nilita, a new species of prawn (Crustacea Decapoda) from the western Mediterranean. Pubblicazioni della Stazione Zoologica di Napoli, 22, 339 - 347." type="journal article">Dohrn &amp; Holthuis 1950</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33CFB875608FFF160EEB72B88E2F9BF" pageId="7" pageNumber="397" type="reference_group">
<paragraph id="8B99A80C5608FFF160EEB72B8ED8FC27" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="397">
<taxonomicName id="4C26D38F5608FFF160EEB72B8D3BFDAA" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[199,383,585,608]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF160EEB72B8D3BFDAA" box="[199,383,585,608]" italics="true" pageId="7" pageNumber="397">Lysmata moorei</emphasis>
</taxonomicName>
is a relatively poorly described species, which shows considerable variation in size and morphology between the western, central and eastern Atlantic populations (De Grave &amp; Anker, unpublished data). Although 
<taxonomicName id="4C26D38F5608FFF16125B7F38D3FFD62" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[268,379,657,680]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF16125B7F38D3FFD62" box="[268,379,657,680]" italics="true" pageId="7" pageNumber="397">L. moorei</emphasis>
</taxonomicName>
exhibits similar proportions of the fused vs. free part of the accessory branch to the new species, a number of other morphological characteristics clearly differentiate the two species. In 
<taxonomicName id="4C26D38F5608FFF164D0B7D7892CFD06" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[1273,1384,693,716]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF164D0B7D7892CFD06" box="[1273,1384,693,716]" italics="true" pageId="7" pageNumber="397">L. moorei</emphasis>
</taxonomicName>
, the scaphocerite is relatively short and wide, with a short distolateral tooth, not overreaching the distal margin of the blade (
<bibRefCitation id="EFB7D5FD5608FFF160CBB79E8DC1FCDE" box="[226,389,764,789]" pageId="7" pageNumber="397" refId="ref5710" refString="Rathbun, M. J. (1901) Investigations of the Aquatic Resources and Fisheries of Porto Rico by the United States Fish Commission Steamer Fish Hawk in 1899. The Brachyura and Macrura of Porto Rico. Bulletin of the United States Fish Commission, 20 [for 1900, preprint dated 1901, published in 1902], 1 - 127." type="journal article">Rathbun 1901</bibRefCitation>
), whereas in 
<taxonomicName id="4C26D38F5608FFF16231B79F8EE1FCDF" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[536,677,764,789]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF16231B79F8EE1FCDF" box="[536,677,764,789]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF16285B79E8F42FCDF" box="[684,774,764,789]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF16285B79E8F42FCDF" bold="true" box="[684,774,764,789]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, it is rather elongate and narrow, with the distolateral tooth clearly overreaching the blade (
<figureCitation id="131DB4895608FFF16223B67D8E1AFCF2" box="[522,606,799,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="7" pageNumber="397">Fig. 1a</figureCitation>
). The number of carpal divisions in the second pereiopod is about 17 (
<bibRefCitation id="EFB7D5FD5608FFF160B7B6268D7BFC96" box="[158,319,836,861]" pageId="7" pageNumber="397" refId="ref5710" refString="Rathbun, M. J. (1901) Investigations of the Aquatic Resources and Fisheries of Porto Rico by the United States Fish Commission Steamer Fish Hawk in 1899. The Brachyura and Macrura of Porto Rico. Bulletin of the United States Fish Commission, 20 [for 1900, preprint dated 1901, published in 1902], 1 - 127." type="journal article">Rathbun 1901</bibRefCitation>
) vs. considerably higher (25–27) in the new species (
<figureCitation id="131DB4895608FFF163BCB6268FADFC97" box="[917,1001,836,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1757,1779]" captionTargetBox="[155,1380,200,1724]" captionTargetId="figure@3.[151,1436,193,1735]" captionTargetPageId="3" captionText="FIGURE 2. Lysmata napoleoni sp. nov., (a) right first pereiopod, lateral view; b) same, ischium, mesial view; (c) same, chela and distal portion of carpus, ventromesial view; (d) right second pereiopod, lateral view; (e) same, chela, mesial view. All from paratype (pocl 5.2 mm), MZUSP 37298." httpUri="https://zenodo.org/record/1283961/files/figure.png" pageId="7" pageNumber="397">Fig. 2d</figureCitation>
). In addition, only a single post-orbital tooth is present in 
<taxonomicName id="4C26D38F5608FFF16142B60B8D9CFC4A" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[363,472,873,896]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF16142B60B8D9CFC4A" box="[363,472,873,896]" italics="true" pageId="7" pageNumber="397">L. moorei</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB7D5FD5608FFF161C1B6058EC8FC4A" box="[488,652,871,896]" pageId="7" pageNumber="397" refId="ref5710" refString="Rathbun, M. J. (1901) Investigations of the Aquatic Resources and Fisheries of Porto Rico by the United States Fish Commission Steamer Fish Hawk in 1899. The Brachyura and Macrura of Porto Rico. Bulletin of the United States Fish Commission, 20 [for 1900, preprint dated 1901, published in 1902], 1 - 127." type="journal article">Rathbun 1901</bibRefCitation>
; Fransen 1991), whilst two are present in 
<taxonomicName id="4C26D38F5608FFF16445B60B88BFFC4A" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[1132,1275,871,896]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF16445B60B88BFFC4A" box="[1132,1275,871,896]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF1652AB605891AFC4A" box="[1283,1374,871,896]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF1652AB605891AFC4A" bold="true" box="[1283,1374,871,896]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="131DB4895608FFF16547B6058CF1FC6F" captionStart="FIGURE 1" captionStartId="2.[151,250,1878,1900]" captionTargetBox="[157,1404,196,1850]" captionTargetId="figure@2.[151,1411,193,1858]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata napoleoni sp. nov., (a) frontal region, dorsal view; (b) same, lateral view; (c) carapace, lateral view; (d) fifth and sixth pleonites, lateral view; (e) telson, dorsal view; (f) lateral antennular flagellum, accessory branch, lateral view; (g) third maxilliped, lateral view; (h) same, tip, dorsomesial view; (i) uropod, dorsal view; (j), (k) and (l) rostrum, lateral view. From (a) – (i) paratype (pocl 5.2 mm), MZUSP 37298; (j) holotype (pocl 5.8 mm), OUMNH.ZC.2018-04-001; (k) and (l) paratypes (cl 3.5–5.0 mm), OUMNH.ZC.2018-04-004." httpUri="https://zenodo.org/record/1283959/files/figure.png" pageId="7" pageNumber="397">Fig. 1b</figureCitation>
). In the material of 
<taxonomicName id="4C26D38F5608FFF1618AB6EF8E56FC6E" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[419,530,909,932]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF1618AB6EF8E56FC6E" box="[419,530,909,932]" italics="true" pageId="7" pageNumber="397">L. moorei</emphasis>
</taxonomicName>
from 
<collectingCountry id="F331E89C5608FFF16277B6EE8EA2FC6F" box="[606,742,908,933]" name="Cape Verde" pageId="7" pageNumber="397">Cape Verde</collectingCountry>
, these features are confirmed: all specimens harbour only a single post-orbital tooth, a relatively short scaphocerite, with the distal tooth not overreaching the blade, and the second pereiopod with 16-17 carpal divisions.
</paragraph>
<paragraph id="8B99A80C5608FFF160EEB69A883BFAE5" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="397">
In overall morphology, 
<taxonomicName id="4C26D38F5608FFF161CFB69B8E87FBDA" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[486,707,1015,1040]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF161CFB69B8E87FBDA" box="[486,707,1015,1040]" italics="true" pageId="7" pageNumber="397">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF162FBB6958F77FBDA" box="[722,819,1015,1040]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF162FBB6958F77FBDA" bold="true" box="[722,819,1015,1040]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be closest to the widely distributed 
<emphasis id="B952741E5608FFF165ACB69B8D5BFBFF" italics="true" pageId="7" pageNumber="397">
<taxonomicName id="4C26D38F5608FFF165ACB69B8D5FFBFF" baseAuthorityName="Risso" baseAuthorityYear="1816" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">L. seticaudata</taxonomicName>
,
</emphasis>
for which there are surprisingly few good illustrations in recent taxonomic literature (e.g. 
<bibRefCitation id="EFB7D5FD5608FFF1651CB17E8D7AFB92" pageId="7" pageNumber="397" refId="ref4959" refString="Dohrn, P. F. R. &amp; Holthuis, L. B. (1950) Lysmata nilita, a new species of prawn (Crustacea Decapoda) from the western Mediterranean. Pubblicazioni della Stazione Zoologica di Napoli, 22, 339 - 347." type="journal article">Dohrn &amp; Holthuis 1950</bibRefCitation>
; Zariquiey-Álvarez 1968; 
<bibRefCitation id="EFB7D5FD5608FFF1625AB15D8FCBFB92" box="[627,911,1087,1112]" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz 2000</bibRefCitation>
). Nevertheless, based on rostral morphology, proportions of the scaphocerite, as well as the armature of the ambulatory pereiopods, 
<taxonomicName id="4C26D38F5608FFF16449B10788AAFBB7" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[1120,1262,1124,1149]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF16449B10788AAFBB7" box="[1120,1262,1124,1149]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF164DCB106890BFBB7" box="[1269,1359,1124,1149]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF164DCB106890BFBB7" bold="true" box="[1269,1359,1124,1149]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C26D38F5608FFF165ACB1078D5CFB6A" baseAuthorityName="Risso" baseAuthorityYear="1816" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF165ACB1078D5CFB6A" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
appear to be very similar. Although the drawings of the antennule of 
<taxonomicName id="4C26D38F5608FFF1644BB1EB8D0CFB0F" authority="in Dohrn &amp; Holthuis (1950)" authorityName="in Dohrn &amp; Holthuis" authorityYear="1950" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF1644BB1EB8941FB6A" box="[1122,1285,1159,1184]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
in 
<bibRefCitation id="EFB7D5FD5608FFF1651AB1E58D0CFB0F" pageId="7" pageNumber="397" refId="ref4959" refString="Dohrn, P. F. R. &amp; Holthuis, L. B. (1950) Lysmata nilita, a new species of prawn (Crustacea Decapoda) from the western Mediterranean. Pubblicazioni della Stazione Zoologica di Napoli, 22, 339 - 347." type="journal article">Dohrn &amp; Holthuis (1950)</bibRefCitation>
</taxonomicName>
and 
<bibRefCitation id="EFB7D5FD5608FFF16157B1CE8EDBFB0E" box="[382,671,1196,1221]" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz (2000)</bibRefCitation>
are somewhat conflicting, they agree in that the number of divisions in the fixed part of the accessory branch of the antennule is 15 or more, contrasting to 
<quantity id="4CDE05E95608FFF16477B1AD88E2FB22" box="[1118,1190,1231,1256]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="7" pageNumber="397" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
the new species. This distinguishing feature is herein confirmed using a large sample from 
<collectingRegion id="49E266EE5608FFF163E2B1968860FAC7" box="[971,1060,1268,1293]" country="France" name="Corse" pageId="7" pageNumber="397">
<collectingCountry id="F331E89C5608FFF163E2B1968860FAC7" box="[971,1060,1268,1293]" name="France" pageId="7" pageNumber="397">Corsica</collectingCountry>
</collectingRegion>
, in which the number of carpal divisions ranges from 19 to 27, thus allowing a morphological separation of the two taxa.
</paragraph>
<paragraph id="8B99A80C5608FFF160EEB05F88E2F9BF" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="397">
<taxonomicName id="4C26D38F5608FFF160EEB05F8DDAFA9F" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[199,414,1340,1365]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF160EEB05F8DDAFA9F" box="[199,414,1340,1365]" italics="true" pageId="7" pageNumber="397">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF16181B05E8E40FA9F" box="[424,516,1340,1365]" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF16181B05E8E40FA9F" bold="true" box="[424,516,1340,1365]" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
displays a striking colour pattern (
<figureCitation id="131DB4895608FFF163B0B05E8FAFFA9E" box="[921,1003,1340,1365]" captionStart="FIGURE 4" captionStartId="6.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@6.[151,1436,193,1478]" captionTargetPageId="6" captionText="FIGURE 4. Lysmata napoleoni sp. nov. on St. Helena Island, (a) paratype, OUMNH.ZC. 2018-04-004; (b) ovigerous hermaphrodite, not deposited; (c) – (d) paratype, OUMNH.ZC. 2018-04-004, showing association with the sea anemone Telmactis cricoides (Duchassaing). Photographs by Peter Wirtz (a, b, c) and Judith Brown (d)." httpUri="https://zenodo.org/record/1283965/files/figure.png" pageId="7" pageNumber="397">Figs. 4</figureCitation>
, 
<figureCitation id="131DB4895608FFF163D3B05E884CFA9F" box="[1018,1032,1340,1365]" captionStart="FIGURE 5" captionStartId="8.[151,250,1980,2002]" captionTargetBox="[151,1436,193,1959]" captionTargetId="figure@8.[151,1436,193,1959]" captionTargetPageId="8" captionText="FIGURE 5. Lysmata napoleoni sp. nov. from Ascension Island, specimens not collected, (a) colour pattern by day; (b) nocturnal cleaning behaviour. Photographs by Judith Brown." httpUri="https://zenodo.org/record/1283967/files/figure.png" pageId="7" pageNumber="397">5</figureCitation>
) and may be distinguished at once from all Atlantic congeners for which the colour pattern is known. For instance, the colour pattern of the morphologically closest Atlantic species, 
<taxonomicName id="4C26D38F5608FFF1625BB0E78F56FA57" authorityName="in Dohrn &amp; Holthuis" authorityYear="1950" box="[626,786,1412,1437]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF1625BB0E78F56FA57" box="[626,786,1412,1437]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
(see photograph in 
<bibRefCitation id="EFB7D5FD5608FFF163D0B0E688A8FA57" box="[1017,1260,1412,1437]" pageId="7" pageNumber="397" refId="ref5273" refString="Gonzalez Perez J. A., (1995) Catalogo de los crustaceos decapodos de las Islas Canarias. Gambas. Langostas. Cangrejos. Publicaciones Turquesa S. L., Santa Cruz de Tenerife, 282 pp." type="book">González Pérez 1995</bibRefCitation>
) and 
<taxonomicName id="4C26D38F5608FFF16507B0E789D8FA56" baseAuthorityName="Rathbun" baseAuthorityYear="1901" box="[1326,1436,1413,1436]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B952741E5608FFF16507B0E789D8FA56" box="[1326,1436,1413,1436]" italics="true" pageId="7" pageNumber="397">L. moorei</emphasis>
</taxonomicName>
(see photograph in Brown 2015) are quite distinct from that of the new species. It is important to note that the colour patterns of several Atlantic species of 
<taxonomicName id="4C26D38F5608FFF16299B0AF8F49FA2E" box="[688,781,1485,1508]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5608FFF16299B0AF8F49FA2E" box="[688,781,1485,1508]" italics="true" pageId="7" pageNumber="397">Lysmata</emphasis>
</taxonomicName>
(
<taxonomicName id="4C26D38F5608FFF1630AB0AF88B0FA2F" authority="Holthuis &amp; Maurin, 1952" authorityName="Holthuis &amp; Maurin" authorityYear="1952" box="[803,1268,1484,1509]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="uncicornis">
<emphasis id="B952741E5608FFF1630AB0AF8FFFFA2E" box="[803,955,1485,1508]" italics="true" pageId="7" pageNumber="397">L. uncicornis</emphasis>
Holthuis &amp; Maurin, 1952
</taxonomicName>
; 
<taxonomicName id="4C26D38F5608FFF1652DB0AF8D6DF9C2" authority="Chace, 1972" authorityName="Chace" authorityYear="1972" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="anchisteus">
<emphasis id="B952741E5608FFF1652DB0AF89D8FA2F" box="[1284,1436,1484,1509]" italics="true" pageId="7" pageNumber="397">L. anchisteus</emphasis>
<bibRefCitation id="EFB7D5FD5608FFF160BEB08D8D6DF9C2" box="[151,297,1519,1544]" pageId="7" pageNumber="397" refId="ref4645" refString="Chace, F. A. Jr. (1972) Thc shrimps of the Smithsonian-Bredin Caribbean Expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179." type="journal article">Chace, 1972</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C26D38F5608FFF16111B0938EABF9C2" authority="Crosnier &amp; Forest, 1973" authorityName="Crosnier &amp; Forest" authorityYear="1973" box="[312,751,1519,1544]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="stenolepis">
<emphasis id="B952741E5608FFF16111B0938D8DF9C2" box="[312,457,1519,1544]" italics="true" pageId="7" pageNumber="397">L. stenolepis</emphasis>
<bibRefCitation id="EFB7D5FD5608FFF161FAB08D8EABF9C2" box="[467,751,1519,1544]" pageId="7" pageNumber="397" refId="ref4691" refString="Crosnier, A. &amp; Forest, J. (1973) Les crevettes profondes de l'Atlantique oriental tropical. Faune Tropicale, 19, 1 - 409." type="journal article">Crosnier &amp; Forest, 1973</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C26D38F5608FFF162D7B093883AF9C2" authority="Prakash &amp; Baeza 2017" authorityName="Prakash &amp; Baeza" authorityYear="2017" box="[766,1150,1519,1544]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="baueri">
<emphasis id="B952741E5608FFF162D7B0938F2CF9C2" box="[766,872,1519,1544]" italics="true" pageId="7" pageNumber="397">L. baueri</emphasis>
<bibRefCitation id="EFB7D5FD5608FFF1635AB08D883AF9C2" box="[883,1150,1519,1544]" pageId="7" pageNumber="397" refId="ref5552" refString="Prakash, S. &amp; Baeza, J. A. (2017) A new species of Lysmata Risso, 1816 (Crustacea, Decapoda, Lysmatidae) from the Gulf of Mexico. Zootaxa, 4363 (4), 576 - 582. https: // doi. org / 10.11646 / zootaxa. 4363.4.10" type="journal article">Prakash &amp; Baeza 2017</bibRefCitation>
</taxonomicName>
) are presently unknown. These, however do not belong to the “long-branch group” and therefore, can be separated from 
<taxonomicName id="4C26D38F5608FFF164CBB3778936F9E7" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[1250,1394,1556,1581]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF164CBB3778936F9E7" box="[1250,1394,1556,1581]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655608FFF16553B3768C8CF99A" pageId="7" pageNumber="397" rank="species">
<emphasis id="B952741E5608FFF16553B3768C8CF99A" bold="true" pageId="7" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
on the basis of the antennular morphology alone. Nevertheless, an effort should be made in the future to recollect and document the colour pattern of these species to facilitate easy field recognition.
</paragraph>
</subSubSection>
<subSubSection id="C33CFB875608FFFF60EEB31D886CFE3E" lastPageId="9" lastPageNumber="399" pageId="7" pageNumber="397" type="discussion">
<paragraph id="8B99A80C5608FFF160EEB31D895BF873" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="397">
<emphasis id="B952741E5608FFF160EEB31D8D01F952" bold="true" box="[199,325,1663,1688]" pageId="7" pageNumber="397">Discussion</emphasis>
. The present description raises to four the number of 
<taxonomicName id="4C26D38F5608FFF16397B3E3885FF952" box="[958,1051,1665,1688]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5608FFF16397B3E3885FF952" box="[958,1051,1665,1688]" italics="true" pageId="7" pageNumber="397">Lysmata</emphasis>
</taxonomicName>
species known from the isolated islands in the south-central Atlantic. This includes the record of 
<taxonomicName id="4C26D38F5608FFF163AAB3C78859F977" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[899,1053,1700,1725]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF163AAB3C78859F977" box="[899,1053,1700,1725]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
</taxonomicName>
from two locations at 
<collectingRegion id="49E266EE5608FFF1650EB3C689D8F977" box="[1319,1436,1700,1725]" country="Saint Helena, Ascension and Tristan da Cunha" name="Ascension" pageId="7" pageNumber="397">
<collectingCountry id="F331E89C5608FFF1650EB3C689D8F977" box="[1319,1436,1700,1725]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="397">Ascension</collectingCountry>
</collectingRegion>
Island by Manning &amp; Chace (1990), which was later corrected to “probably 
<taxonomicName id="4C26D38F5608FFF163D0B3AB8CA1F8CE" authority="&quot; in d'Udekem d'Acoz (2000)" authorityName="&quot; in d'Udekem d'Acoz" authorityYear="2000" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF163D0B3AB88DDF92A" box="[1017,1177,1735,1760]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
” in 
<bibRefCitation id="EFB7D5FD5608FFF164E2B3A58CA1F8CE" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz (2000)</bibRefCitation>
</taxonomicName>
. Despite extensive fieldwork by several teams on 
<collectingRegion id="49E266EE5608FFF1636DB38E8FFDF8CF" box="[836,953,1772,1797]" country="Saint Helena, Ascension and Tristan da Cunha" name="Ascension" pageId="7" pageNumber="397">
<collectingCountry id="F331E89C5608FFF1636DB38E8FFDF8CF" box="[836,953,1772,1797]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="397">Ascension</collectingCountry>
</collectingRegion>
Island, forming the basis of the revised checklist published by 
<bibRefCitation id="EFB7D5FD5608FFF161B6B2738EE3F8E2" box="[415,679,1807,1832]" pageId="7" pageNumber="397" refId="ref4884" refString="De Grave, S., Anker, A., Dworschak, P. C., Clark, P. F. &amp; Wirtz, P. (2017) An updated checklist of the marine Decapoda of Ascension Island, central Atlantic Ocean. Journal of the Marine Biological Association of the United Kingdom, 97, 759 - 770. https: // doi. org / 10.1017 / S 0025315414001295" type="journal article">
De Grave 
<emphasis id="B952741E5608FFF1623FB2738E15F8E2" box="[534,593,1807,1832]" italics="true" pageId="7" pageNumber="397">et al.</emphasis>
(2017)
</bibRefCitation>
, no specimens identifiable as 
<taxonomicName id="4C26D38F5608FFF1642BB27388E6F8E2" authorityName="&quot; in d'Udekem d'Acoz" authorityYear="2000" box="[1026,1186,1807,1832]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF1642BB27388E6F8E2" box="[1026,1186,1807,1832]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
were encountered. In addition, no records exist for 
<taxonomicName id="4C26D38F5608FFF161C9B2578E3AF887" authorityName="&quot; in d'Udekem d'Acoz" authorityYear="2000" box="[480,638,1844,1869]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF161C9B2578E3AF887" box="[480,638,1844,1869]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
south of the Canary Islands (
<bibRefCitation id="EFB7D5FD5608FFF163EFB2568898F886" box="[966,1244,1844,1869]" pageId="7" pageNumber="397" refId="ref5007" refString="d'Udekem d'Acoz, C. (2000) Redescription of Lysmata intermedia (Kingsley, 1879) based on topotypical specimens, with remarks on Lysmata seticaudata (Risso, 1816) (Decapoda, Caridea, Hippolytidae). Crustaceana, 73, 719 - 735. https: // doi. org / 10.1163 / 156854000504750" type="journal article">d’Udekem d’Acoz 2000</bibRefCitation>
). Given the close morphological similarity between 
<taxonomicName id="4C26D38F5608FFF16230B23B8EFCF8BA" authorityName="&quot; in d'Udekem d'Acoz" authorityYear="2000" box="[537,696,1879,1904]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="seticaudata">
<emphasis id="B952741E5608FFF16230B23B8EFCF8BA" box="[537,696,1879,1904]" italics="true" pageId="7" pageNumber="397">L. seticaudata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C26D38F5608FFF162C6B23B8F39F8BA" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[751,893,1879,1904]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5608FFF162C6B23B8F39F8BA" box="[751,893,1879,1904]" italics="true" pageId="7" pageNumber="397">L. napoleoni</emphasis>
</taxonomicName>
<emphasis id="B952741E5608FFF163ADB2358FA2F8BA" bold="true" box="[900,998,1879,1904]" pageId="7" pageNumber="397">
<taxonomicNameLabel id="A261C9655608FFF163ADB2358FA4F8BA" box="[900,992,1879,1904]" pageId="7" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
and in the absence of any colour notes for the two specimens forming the basis of the record from 
<collectingRegion id="49E266EE5608FFF1634EB21E8F98F85F" box="[871,988,1916,1941]" country="Saint Helena, Ascension and Tristan da Cunha" name="Ascension" pageId="7" pageNumber="397">
<collectingCountry id="F331E89C5608FFF1634EB21E8F98F85F" box="[871,988,1916,1941]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="397">Ascension</collectingCountry>
</collectingRegion>
Island, we postulate that the material reported by Manning &amp; Chace (1990) as 
<taxonomicName id="4C26D38F5608FFF1624DB2C38EBFF873" baseAuthorityName="Kingsley" baseAuthorityYear="1879" box="[612,763,1952,1977]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="397" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B952741E5608FFF1624DB2C38EBFF873" box="[612,763,1952,1977]" italics="true" pageId="7" pageNumber="397">L. intermedia</emphasis>
</taxonomicName>
may potentially refer to the present new species.
</paragraph>
<caption id="DF59F8845607FFFE60BEB2DE8EBAF83B" httpUri="https://zenodo.org/record/1283967/files/figure.png" pageId="8" pageNumber="398" startId="8.[151,250,1980,2002]" targetBox="[151,1436,193,1959]" targetPageId="8">
<paragraph id="8B99A80C5607FFFE60BEB2DE8EBAF83B" blockId="8.[151,1436,1980,2033]" pageId="8" pageNumber="398">
<emphasis id="B952741E5607FFFE60BEB2DE8D53F818" bold="true" box="[151,279,1980,2002]" pageId="8" pageNumber="398">FIGURE 5.</emphasis>
<taxonomicName id="4C26D38F5607FFFE610BB2DF8DA3F818" authority="Grave &amp; Anker, 2018" authorityName="Grave &amp; Anker" authorityYear="2018" box="[290,487,1980,2002]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="398" phylum="Arthropoda" rank="species" species="napoleoni" status="sp. nov.">
<emphasis id="B952741E5607FFFE610BB2DF8DA3F818" box="[290,487,1980,2002]" italics="true" pageId="8" pageNumber="398">Lysmata napoleoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261C9655607FFFE61DAB2DF8E0CF818" box="[499,584,1981,2002]" pageId="8" pageNumber="398" rank="species">
<emphasis id="B952741E5607FFFE61DAB2DF8E0CF818" bold="true" box="[499,584,1981,2002]" pageId="8" pageNumber="398">sp. nov.</emphasis>
</taxonomicNameLabel>
from Ascension Island, specimens not collected, (a) colour pattern by day; (b) nocturnal cleaning behaviour. Photographs by Judith Brown.
</paragraph>
</caption>
<paragraph id="8B99A80C5606FFFF60EEB5F5886CFE3E" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="399">
Colour patterns are of high taxonomic value in the genus 
<taxonomicName id="4C26D38F5606FFFF63B7B5FB8FBFFF7A" box="[926,1019,153,176]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5606FFFF63B7B5FB8FBFFF7A" box="[926,1019,153,176]" italics="true" pageId="9" pageNumber="399">Lysmata</emphasis>
</taxonomicName>
, and are largely species specific, representing the most practical and easiest field diagnostic character in most groups (Fielder 
<emphasis id="B952741E5606FFFF6525B5DF8908FF1F" box="[1292,1356,188,213]" italics="true" pageId="9" pageNumber="399">et al.</emphasis>
2010). Therefore, we advise against further descriptions of new taxa in 
<taxonomicName id="4C26D38F5606FFFF6348B5838FFAFF32" box="[865,958,225,248]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5606FFFF6348B5838FFAFF32" box="[865,958,225,248]" italics="true" pageId="9" pageNumber="399">Lysmata</emphasis>
</taxonomicName>
without colour pattern information, unless they exhibit truly unique species-level morphological differences. We also council against erecting new species based on single specimens (e.g. 
<bibRefCitation id="EFB7D5FD5606FFFF622CB44A8EF2FE8A" box="[517,694,295,320]" pageId="9" pageNumber="399" refId="ref5219" refString="Gan, Z. &amp; Li, X. (2016) Lysmata leptodactylus, a new species of lysmatid shrimp (Crustacea: Decapoda: Caridea) from China. Zootaxa, 4138 (1), 181 - 188. https: // doi. org / 10.11646 / zootaxa. 4138.1.9" type="journal article">Gan &amp; Li 2016</bibRefCitation>
; 
<bibRefCitation id="EFB7D5FD5606FFFF62EAB4458F8DFE8A" box="[707,969,295,320]" pageId="9" pageNumber="399" refId="ref5552" refString="Prakash, S. &amp; Baeza, J. A. (2017) A new species of Lysmata Risso, 1816 (Crustacea, Decapoda, Lysmatidae) from the Gulf of Mexico. Zootaxa, 4363 (4), 576 - 582. https: // doi. org / 10.11646 / zootaxa. 4363.4.10" type="journal article">Prakash &amp; Baeza 2017</bibRefCitation>
; 
<bibRefCitation id="EFB7D5FD5606FFFF63FFB44A88EFFE8A" box="[982,1195,295,320]" pageId="9" pageNumber="399" refId="ref5963" refString="Wang, Y. - R. &amp; Sha, Z. - L. (2018) Description of two new species of Lysmata Risso, 1816 (Decapoda, Lysmatidae) from the China seas, with remarks on Lysmata vittata (Stimpson 1860). Zootaxa, 4392 (1), 28 - 40. https: // doi. org / 10.11646 / zootaxa. 4392.1.2" type="journal article">Wang &amp; Sha 2018</bibRefCitation>
), as has equally been done in recent years. All 
<taxonomicName id="4C26D38F5606FFFF6193B42F8E5CFEAE" box="[442,536,333,356]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5606FFFF6193B42F8E5CFEAE" box="[442,536,333,356]" italics="true" pageId="9" pageNumber="399">Lysmata</emphasis>
</taxonomicName>
species are rather variable in the traditional morphological characters used to differentiate them, such as the meral spination on the ambulatory pereiopods and the number of carpal divisions the second pereiopod, as amply demonstrated by the 
<typeStatus id="549D16AE5606FFFF6295B4F78EA8FE67" box="[700,748,405,429]" pageId="9" pageNumber="399">type</typeStatus>
series of 
<taxonomicName id="4C26D38F5606FFFF6371B4F78FA1FE67" authorityName="Grave &amp; Anker" authorityYear="2018" box="[856,997,404,429]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="399" phylum="Arthropoda" rank="species" species="napoleoni">
<emphasis id="B952741E5606FFFF6371B4F78FA1FE67" box="[856,997,404,429]" italics="true" pageId="9" pageNumber="399">L. napoleoni</emphasis>
</taxonomicName>
. A large part of future taxonomic work in 
<taxonomicName id="4C26D38F5606FFFF609DB4DB8D55FE1A" box="[180,273,441,464]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="B952741E5606FFFF609DB4DB8D55FE1A" box="[180,273,441,464]" italics="true" pageId="9" pageNumber="399">Lysmata</emphasis>
</taxonomicName>
will be trying to match these “ 
<emphasis id="B952741E5606FFFF62A7B4D58EB3FE1A" box="[654,759,439,464]" italics="true" pageId="9" pageNumber="399">mihi itch</emphasis>
” descriptions (see 
<bibRefCitation id="EFB7D5FD5606FFFF63F4B4D588C8FE1A" box="[989,1164,439,464]" pageId="9" pageNumber="399" refId="ref5065" refString="Evenhuis, N. (2008) The &quot; mihi itch &quot; - a brief history. Zootaxa, 1890, 5 - 68." type="journal article">Evenhuis 2008</bibRefCitation>
) with newly collected, sequenceable and photo-vouchered material, to confirm the validity of these taxa.
</paragraph>
</subSubSection>
</treatment>
</document>