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<document id="262482A7C841617080AEFAAC5B871BA1" ID-DOI="10.11646/zootaxa.3872.2.1" ID-GBIF-Dataset="e9b8d555-d661-4f1a-86e8-f2a0acc38798" ID-ISSN="1175-5326" ID-Zenodo-Dep="253224" ID-ZooBank="F9F81CC8-E033-46B7-B73B-9FB777DF4116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460926510372" checkinUser="plazi" docAuthor="Ponder, Winston F., Fukuda, Hiroshi &amp; Hallan, Anders" docDate="2014" docId="03CDAD6557723C2EFF05FBDCBE57AFAA" docLanguage="en" docName="zt03872p153.pdf" docOrigin="Zootaxa 3872 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clenchiella Abbott 1948" docType="treatment" docVersion="9" lastPageNumber="111" masterDocId="FFF4D51D57783C24FF92FFB2BB3EA911" masterDocTitle="A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)" masterLastPageNumber="153" masterPageNumber="101" pageNumber="111" updateTime="1698362634141" updateUser="plazi">
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<mods:title id="5F613CE6EEDD46BC3C040CBA8BDD3C1A">A review of the family Clenchiellidae (Mollusca: Caenogastropoda: Truncatelloidea)</mods:title>
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<mods:namePart id="6F8F4282BFBB38CDA638E5452110800A">Ponder, Winston F.</mods:namePart>
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<mods:namePart id="D6EB2EFB6608AB86FD4E5977A4D99CE9">Fukuda, Hiroshi</mods:namePart>
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<mods:namePart id="3C07DAC5422F29551B3FA802EA182DA4">Hallan, Anders</mods:namePart>
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<mods:title id="ED38C6B185C55FF3CCF4FB6D588C4F42">Zootaxa</mods:title>
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<mods:part id="FC957A5E4FD37466D21B287F8DD89E6D">
<mods:date id="649352673BC66B8B4063D9F9F920FA57">2014</mods:date>
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<mods:number id="8B4DE91B7DD43DE42EA3986451BDD48D">3872</mods:number>
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<mods:start id="9948FC22CA24F9003D279D0D52E1F2AE">101</mods:start>
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<treatment id="03CDAD6557723C2EFF05FBDCBE57AFAA" ID-DOI="http://doi.org/10.5281/zenodo.5631007" ID-GBIF-Taxon="119589590" ID-Zenodo-Dep="5631007" LSID="urn:lsid:plazi:treatment:03CDAD6557723C2EFF05FBDCBE57AFAA" httpUri="http://treatment.plazi.org/id/03CDAD6557723C2EFF05FBDCBE57AFAA" lastPageNumber="111" pageId="10" pageNumber="111">
<subSubSection id="C37E4FF857723C2EFF05FBDCBAF4AD99" box="[151,458,1134,1160]" pageId="10" pageNumber="111" type="nomenclature">
<paragraph id="8BDB1C7357723C2EFF05FBDCBAF4AD99" blockId="10.[151,458,1134,1160]" box="[151,458,1134,1160]" pageId="10" pageNumber="111">
<heading id="D093AB1F57723C2EFF05FBDCBAF4AD99" bold="true" box="[151,458,1134,1160]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
<taxonomicName id="4C6467F057723C2EFF05FBDCBAF4AD99" authority="Abbott, 1948" authorityName="Abbott" authorityYear="1948" box="[151,458,1134,1160]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157723C2EFF05FBDCBAF4AD99" bold="true" box="[151,458,1134,1160]" pageId="10" pageNumber="111">
<emphasis id="B910C06157723C2EFF05FBDCBA1CAD99" bold="true" box="[151,290,1134,1160]" italics="true" pageId="10" pageNumber="111">Clenchiella</emphasis>
<bibRefCitation id="EFF5618257723C2EFEBAFBDCBAF4AD99" author="Abbott" box="[296,458,1134,1160]" pageId="10" pageNumber="111" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott, 1948</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C37E4FF857723C2EFF05FB07BAFBADDA" box="[151,453,1205,1227]" pageId="10" pageNumber="111" type="reference_group">
<paragraph id="8BDB1C7357723C2EFF05FB07BAFBADDA" blockId="10.[151,868,1205,1258]" box="[151,453,1205,1227]" pageId="10" pageNumber="111">
<treatmentCitationGroup id="AB743B5D57723C2EFF05FB07BAFBADDA" box="[151,453,1205,1227]" pageId="10" pageNumber="111">
<taxonomicName id="4C6467F057723C2EFF05FB07BAFFADDA" authority="Abbott, 1948: 76" authorityName="Abbott" authorityPageNumber="76" authorityYear="1948" box="[151,449,1205,1227]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157723C2EFF05FB07BA32ADDA" box="[151,268,1205,1227]" italics="true" pageId="10" pageNumber="111">Clenchiella</emphasis>
<treatmentCitation id="0AC53A6257723C2EFE80FB07BAFFADDA" author="Abbott" box="[274,449,1205,1227]" page="76" pageId="10" pageNumber="111" year="1948">
<bibRefCitation id="EFF5618257723C2EFE80FB07BAA5ADDA" author="Abbott" box="[274,411,1205,1227]" pageId="10" pageNumber="111" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott, 1948</bibRefCitation>
: 76
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857723C2EFF05FB67B85AADFB" box="[151,868,1236,1258]" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="8BDB1C7357723C2EFF05FB67B85AADFB" blockId="10.[151,868,1205,1258]" box="[151,868,1236,1258]" pageId="10" pageNumber="111">
<typeStatus id="54DFA2D157723C2EFF05FB67BBF7ADFB" box="[151,201,1237,1258]" pageId="10" pageNumber="111">Type</typeStatus>
species:
<taxonomicName id="4C6467F057723C2EFEB4FB66B9B7ADFB" authority="Abbott, 1948" authorityName="Abbott" authorityYear="1948" box="[294,649,1236,1258]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="species" species="victoriae">
<emphasis id="B910C06157723C2EFEB4FB66BAC5ADFB" box="[294,507,1236,1258]" italics="true" pageId="10" pageNumber="111">Clenchiella victoriae</emphasis>
<bibRefCitation id="EFF5618257723C2EFD93FB66B9B7ADFB" author="Abbott" box="[513,649,1236,1258]" pageId="10" pageNumber="111" refString="Abbott, R. T. (1948) A new genus and species of Philippine Amnicolidae. The Nautilus, 61, 75 - 80, pl. 5." type="journal article" year="1948">Abbott, 1948</bibRefCitation>
</taxonomicName>
, original designation.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857723C2EFF05FAA4BFCDAF5E" pageId="10" pageNumber="111" type="diagnosis">
<paragraph id="8BDB1C7357723C2EFF05FAA4BFCDAF5E" blockId="10.[151,1437,1302,1723]" pageId="10" pageNumber="111">
<emphasis id="B910C06157723C2EFF05FAA4BA2FAC3E" bold="true" box="[151,273,1302,1327]" pageId="10" pageNumber="111">Diagnosis.</emphasis>
Shell large for family (
<quantity id="4C9CB19657723C2EFD8DFAA5B997AC3F" box="[543,681,1303,1327]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="10" pageNumber="111" unit="mm" value="1.75" valueMax="2.3" valueMin="1.2">1.22.3 mm</quantity>
in maximum diameter, although Abbott incorrectly gives up to
<quantity id="4C9CB19657723C2EFA1CFAA5BBFDAC43" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="111" unit="mm" value="4.0">4 mm</quantity>
for
<typeStatus id="54DFA2D157723C2EFF67FA89BA1BAC42" box="[245,293,1339,1363]" pageId="10" pageNumber="111">type</typeStatus>
species). Spire flat, umbilicus wide. Whorls typically with strong spiral mid-dorsal and mid-ventral keel and many distinct spiral cords (
<figureCitation id="135F00F657723C2EFDD3FAECB9BBAC66" box="[577,645,1374,1399]" captionStart="FIGURE 1" captionStartId="11.[151,250,1702,1724]" captionTargetBox="[163,1428,289,1686]" captionTargetId="figure@11.[163,1428,289,1687]" captionTargetPageId="11" captionText="FIGURE 1. Holotypes of species of Clenchiella with distinct keels (from left towards right: dorsal, lateral and ventral view respectively). A C, Clenchiella victoriae (USNM 488534; shell diameter 2.1 mm); D F, Clenchiella bicingulata n. sp. (AMS C. 462959; shell diameter 1.97 mm); G I, Clenchiella varicosa n. sp. (AMS C. 460738; shell diameter 1.96 mm); J L, Clenchiella iriomotensis n. sp. (AMS C. 445413; shell diameter 1.94 mm)." httpUri="https://zenodo.org/record/253225/files/figure.png" pageId="10" pageNumber="111">Fig. 1</figureCitation>
), keels subobsolete to absent in one species (
<figureCitation id="135F00F657723C2EFB0FFAECBFD8AC66" box="[1181,1254,1374,1399]" captionStart="FIGURE 2" captionStartId="12.[151,250,1700,1722]" captionTargetBox="[150,1436,985,1678]" captionTargetId="figure@12.[150,1436,985,1679]" captionTargetPageId="12" captionText="FIGURE 2. Type specimens of Clenchiella minutissima (from left towards right: dorsal, lateral and ventral view respectively). A C, holotype of Clenchiella papuensis, synonymised with Clenchiella minutissima herein (NHML, registration number not known; shell diameter 1.69 mm); D F, syntype of Clenchiella minutissima (MNHNP; shell diameter 1.70 mm)." httpUri="https://zenodo.org/record/253226/files/figure.png" pageId="10" pageNumber="111">Fig. 2</figureCitation>
). Aperture near circular, peristome simple, prosocline, external varix heavy, wide. Operculum near circular, thin, horny, of several whorls, nucleus near central; inner side lacks calcareous smear and has central projection (distinct peg to weak nipple-like protrusion). Head-foot usually with black pigment on cephalic tentacles and snout; posterior end of foot pointed.
<taxonomicName id="4C6467F057723C2EFF69FA5CBA72AF16" box="[251,332,1518,1543]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="10" pageNumber="111" phylum="Bryophyta" rank="genus">
<emphasis id="B910C06157723C2EFF69FA5CBA72AF16" box="[251,332,1518,1543]" italics="true" pageId="10" pageNumber="111">Radula</emphasis>
</taxonomicName>
as for family, cusps on inner outer marginal teeth slightly larger than those on outer marginals. Penis with two to four large, simple (
<emphasis id="B910C06157723C2EFDD8F9A6B953AF3A" box="[586,621,1556,1579]" italics="true" pageId="10" pageNumber="111">i.e.</emphasis>
with no apocrine glands) swellings on distal portion. Bursa copulatrix small to extremely large; bursal duct straight or strongly curved. Two posterior seminal receptacles.
</paragraph>
</subSubSection>
<subSubSection id="C37E4FF857723C2EFF55F9E8BE57AFAA" pageId="10" pageNumber="111" type="discussion">
<paragraph id="8BDB1C7357723C2EFF55F9E8BE57AFAA" blockId="10.[151,1437,1302,1723]" pageId="10" pageNumber="111">
<emphasis id="B910C06157723C2EFF55F9E8BA05AF62" bold="true" box="[199,315,1626,1651]" pageId="10" pageNumber="111">Remarks.</emphasis>
<taxonomicName id="4C6467F057723C2EFED1F9E8BAFAAF62" box="[323,452,1626,1651]" class="Gastropoda" family="Hydrobiidae" genus="Clenchiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="111" phylum="Mollusca" rank="genus">
<emphasis id="B910C06157723C2EFED1F9E8BAFAAF62" box="[323,452,1626,1651]" italics="true" pageId="10" pageNumber="111">Clenchiella</emphasis>
</taxonomicName>
is distinguished from the other three genera introduced below in having the following combination of characters: dorsal and ventral prominent to weak keels on the shell (absent in one species), a strong external apertural varix, a pigment band on each cephalic tentacle and the penis has a few distinct distal lobes.
</paragraph>
</subSubSection>
</treatment>
</document>