treatments-xml/data/03/CD/87/03CD87CC6500C137FF16FDBBFD6DB8D1.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3854814" ID-GBIF-Dataset="40effc56-2cc7-40ae-9592-c7baafeccef1" ID-GBIF-Taxon="163976991" ID-ISSN="1313–2970" ID-Zenodo-Dep="3854814" approvalRequired="53" approvalRequired_for_bibRefs="1" approvalRequired_for_textStreams="51" approvalRequired_for_treatments="1" checkinTime="1587997431956" checkinUser="plazi" docAuthor="Lourenco, Wilson &amp; Duhem, Bernard" docDate="2009" docId="03CD87CC6500C137FF16FDBBFD6DB8D1" docLanguage="en" docName="ZK_article_2004.pdf" docOrigin="ZooKeys 14" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Pantobuthus complicatus Lourenco &amp; Duhem 2009, sp. n." docType="treatment" docVersion="6" lastPageId="11" lastPageNumber="48" masterDocId="FFF4FFB46508C13CFF9AFFF7FFFEBB75" masterDocTitle="Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan" masterLastPageNumber="54" masterPageNumber="37" pageId="8" pageNumber="45" updateTime="1636548053746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lourenco, Wilson</mods:namePart>
<mods:affiliation>Museum national d Histoire naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Duhem, Bernard</mods:namePart>
<mods:affiliation>Muséum national d'Histoire naturelle, Paris ,, France</mods:affiliation>
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<mods:date>2009</mods:date>
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<mods:number>2009-07-10</mods:number>
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<mods:number>14</mods:number>
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<mods:start>37</mods:start>
<mods:end>54</mods:end>
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<mods:identifier type="DOI">10.3897/zookeys.14.212</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3854814" ID-GBIF-Taxon="163976991" ID-Zenodo-Dep="3854814" LSID="urn:lsid:plazi:treatment:03CD87CC6500C137FF16FDBBFD6DB8D1" httpUri="http://treatment.plazi.org/id/03CD87CC6500C137FF16FDBBFD6DB8D1" lastPageId="11" lastPageNumber="48" pageId="8" pageNumber="45">
<subSubSection box="[140,510,588,615]" pageId="8" pageNumber="45" type="nomenclature">
<paragraph blockId="8.[140,968,588,686]" box="[140,510,588,615]" pageId="8" pageNumber="45">
<heading bold="true" box="[140,510,588,615]" fontSize="11" level="2" pageId="8" pageNumber="45" reason="0">
<emphasis bold="true" box="[140,510,588,615]" pageId="8" pageNumber="45">
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[140,437,588,615]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis bold="true" box="[140,437,588,615]" italics="true" pageId="8" pageNumber="45">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[445,510,589,615]" pageId="8" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="45" type="description">
<paragraph blockId="8.[140,968,588,686]" lastBlockId="8.[140,1108,729,1672]" pageId="8" pageNumber="45">
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urn:lsid:zoobank.org:act: 
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Figs 
<figureCitation box="[192,207,659,686]" captionStart="Figures 1-8" captionStartId="4.[140,222,1558,1580]" captionTargetBox="[310,943,177,1536]" captionTargetId="figure@4.[336,919,851,1538]" captionTargetPageId="4" captionText="Figures 1-8. Carapace and tergites I-II. 1. Saharobuthus elegans sp. n., male holotype. 2. Compsobuthus acutecarinatus, female holotype. 3. Leiurus quinquestriatus, male. 4. Mesobuthus eupeus persicus, female. 5. Pantobuthus complicatus sp. n., female holotype. 6. Odontobuthus doriae, female. 7. Buthus draa, female. 8. Hottentotta schach, male (scales = 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770927" httpUri="https://zenodo.org/record/3770927/files/figure.png" pageId="8" pageNumber="45">5</figureCitation>
, 
<figureCitation box="[212,319,659,686]" captionStart-0="Figures 18-28" captionStart-1="Figures 29-32. 29" captionStartId-0="9.[140,222,1526,1548]" captionStartId-1="10.[140,222,1590,1612]" captionTargetBox-0="[156,1080,195,1494]" captionTargetBox-1="[164,1038,207,1546]" captionTargetId-0="figure@9.[140,1108,182,1497]" captionTargetId-1="figure@10.[140,1108,180,1565]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figures 18-28. Pantobuthus complicatus sp. n., female holotype. 18. Chelicerae, dorsal aspect. 19. Sternum, genital operculum and pectines. 20. Cutting edge of movable finger with rows of granules. 21. Leg IV, showing tibial and pedal spurs. 22. Idem, detail of tarsi. 23-25. Chela, dorso-external and ventral aspects and fixed finger in detail. 26-27. Patella, dorsal and external aspects. 28. Femur, dorsal aspect (scales = 3 mm)." captionText-1="Figures 29-32. 29. Pedipalp, dorsal aspect, Vachoniolus globimanus, male holotype (scale = 5 mm). 30- 32. Metasoma and telson, lateral aspect. 30. Saharobuthus elegans sp. n., male holotype. 31. Pantobuthus complicatus sp. n., female holotype. 32. Odontobuthus doriae, female (scales 30-32 = 10 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.3770931" figureDoi-1="http://doi.org/10.5281/zenodo.3770933" httpUri-0="https://zenodo.org/record/3770931/files/figure.png" httpUri-1="https://zenodo.org/record/3770933/files/figure.png" pageId="8" pageNumber="45">18-28,31</figureCitation>
<emphasis bold="true" box="[140,267,729,755]" pageId="8" pageNumber="45">Diagnosis:</emphasis>
as for the new genus.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="45" type="materials_examined">
<paragraph blockId="8.[140,1108,729,1672]" pageId="8" pageNumber="45">
<emphasis bold="true" box="[185,358,765,791]" pageId="8" pageNumber="45">Type material:</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610752301" collectingDate="1971-11" collectorName="C. Naumann" country="Afghanistan" county="Dasht-e-Leili" elevation="480" location="North" municipality="East of Vic Shiberghan" pageId="8" pageNumber="45" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry box="[366,505,765,791]" name="Afghanistan" pageId="8" pageNumber="45">Afghanistan</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:03CD87CC6500C137FF16FDBBFD6DB8D1:8EBB60016500C134FD9FFD0AFDB5B862" box="[517,587,765,791]" country="Afghanistan" county="Dasht-e-Leili" municipality="East of Vic Shiberghan" name="North" pageId="8" pageNumber="45">North</location>
range, 
<collectingMunicipality box="[674,939,765,792]" pageId="8" pageNumber="45">East of Vic Shiberghan</collectingMunicipality>
, 
<collectingCounty box="[951,1103,765,791]" pageId="8" pageNumber="45">Dasht-e-Leili</collectingCounty>
, sandy/loess steppe (rather than desert), 
<quantity box="[584,661,800,827]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="45" unit="m" value="480.0">
<elevation box="[584,661,800,827]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="45" unit="m" value="480.0">480 m</elevation>
</quantity>
, 
<date box="[672,769,800,826]" bridgedPair="/" pageId="8" pageNumber="45" value="1971-11">
<collectingDate box="[672,769,800,826]" bridgedPair="/" pageId="8" pageNumber="45" value="1971-11">XI/1971</collectingDate>
</date>
(
<collectorName box="[786,939,800,827]" pageId="8" pageNumber="45">C. Naumann</collectorName>
), 
<specimenCount box="[959,1055,800,826]" pageId="8" pageNumber="45" type="female">1 female</specimenCount>
<typeStatus pageId="8" pageNumber="45">holotype</typeStatus>
. Deposited in the collection of the Muséum national d’Histoire naturelle, Paris
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="45" type="etymology">
<paragraph blockId="8.[140,1108,729,1672]" pageId="8" pageNumber="45">
<emphasis bold="true" box="[185,323,871,897]" pageId="8" pageNumber="45">Etymology:</emphasis>
The specific name (from Latin 
<emphasis box="[684,798,871,897]" italics="true" pageId="8" pageNumber="45">complicare</emphasis>
) means ‘consisting of parts intricately combined’ in reference to the puzzling characteristics of the new species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="48" pageId="8" pageNumber="45" type="description">
<paragraph blockId="8.[140,1108,729,1672]" box="[185,930,941,968]" pageId="8" pageNumber="45">
Description based on female 
<typeStatus box="[515,614,941,967]" pageId="8" pageNumber="45">holotype</typeStatus>
(Measurements in 
<tableCitation box="[830,913,941,967]" captionStart="Table 1" captionStartId="5.[142,209,166,190]" captionTargetBox="[149,1077,244,1475]" captionTargetId="graphics@5.[141,1106,806,874]" captionTargetPageId="5" captionText="Table 1. Morphometric values (in mm) of the male holotype of Saharobuthus elegans sp. n., and the female holotype of Pantobuthus complicatus sp. n." httpUri="http://table.plazi.org/id/DF1B6652650DC139FF14FF51FD25BBAB" pageId="8" pageNumber="45" tableUuid="DF1B6652650DC139FF14FF51FD25BBAB">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="8.[140,1108,729,1672]" pageId="8" pageNumber="45">
Coloration. Basically yellowish with some light grey zones. Prosoma: carapace yellowish, paler anteriorly; carinae slightly reddish; eyes marked by dark to blackish pigment. Mesosoma: yellowish with tergites marked with greyish confluent zones; only the median carinae are dark. Metasoma: segments I to 
<collectionCode box="[691,711,1082,1108]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="45" type="Museum">V</collectionCode>
yellowish with reddish carinae; vesicle yellowish with vestigial reddish zones laterally; aculeus yellowish at the base and dark reddish at its extremity. Venter yellowish; pectines pale yellow. Chelicerae yellowish without any variegated spots; fingers yellowish with reddish teeth. Pedipalps: yellowish with carinae reddish; chela fingers with the oblique rows of granules reddish. Legs pale yellow.
</paragraph>
<paragraph blockId="8.[140,1108,729,1672]" pageId="8" pageNumber="45">
Morphology. Carapace moderately granular; anterior margin with a very weak median concavity, almost straight. Carinae moderately marked; anterior median strongly granular, central median and posterior median carinae moderately granular, with a moderately to weakly marked ‘lyre’ configuration. All furrows moderate. Median ocular tubercle at the centre of carapace. Eyes separated by more than two ocular diameters. Three pairs of lateral eyes situated in a posterior position in relation to the anterior edge; the first two of moderate size, the last vestigial. Sternum triangular and short; wider than long. Mesosoma: tergites moderately granular. Th ree longitudinal carinae weakly crenulate in tergites I to 
<collectionCode box="[506,539,1540,1566]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="45">VI</collectionCode>
; lateral carinae reduced in tergites I to III. Tergite VII pentacarinate. Venter: genital operculum divided longitudinally in two semi-oval plates. Pectines: pectinal tooth count 24-23; middle basal lamella of the pectines not dilated. Sternites without granules, smooth with elongated spiracles; four carinae on
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770931" ID-Zenodo-Dep="3770931" httpUri="https://zenodo.org/record/3770931/files/figure.png" pageId="9" pageNumber="46" startId="9.[140,222,1526,1548]" targetBox="[156,1080,195,1494]" targetPageId="9">
<paragraph blockId="9.[140,1108,1526,1676]" pageId="9" pageNumber="46">
<emphasis bold="true" box="[140,296,1526,1548]" pageId="9" pageNumber="46">Figures 1 8-28.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[302,520,1526,1547]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis box="[302,520,1526,1547]" italics="true" pageId="9" pageNumber="46">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[527,578,1527,1548]" pageId="9" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
, female holotype. 
<emphasis bold="true" box="[748,781,1526,1548]" pageId="9" pageNumber="46">1 8.</emphasis>
Chelicerae, dorsal aspect. 
<emphasis bold="true" box="[1025,1058,1526,1548]" pageId="9" pageNumber="46">1 9.</emphasis>
Sternum, genital operculum and pectines. 
<emphasis bold="true" box="[495,528,1558,1580]" pageId="9" pageNumber="46">20.</emphasis>
Cutting edge of movable finger with rows of granules. 
<emphasis bold="true" box="[1035,1068,1558,1580]" pageId="9" pageNumber="46">2 1.</emphasis>
Leg IV, showing tibial and pedal spurs. 
<emphasis bold="true" box="[477,510,1590,1612]" pageId="9" pageNumber="46">22.</emphasis>
Idem, detail of tarsi. 
<emphasis bold="true" box="[718,786,1590,1612]" pageId="9" pageNumber="46">23-25.</emphasis>
Chela, dorso-external and ventral aspects and fixed finger in detail. 
<emphasis bold="true" box="[460,527,1622,1644]" pageId="9" pageNumber="46">26-27.</emphasis>
Patella, dorsal and external aspects. 
<emphasis bold="true" box="[874,907,1622,1644]" pageId="9" pageNumber="46">28.</emphasis>
Femur, dorsal aspect (scales = 3 mm).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770933" ID-Zenodo-Dep="3770933" httpUri="https://zenodo.org/record/3770933/files/figure.png" pageId="10" pageNumber="47" startId="10.[140,222,1590,1612]" targetBox="[164,1038,207,1546]" targetPageId="10">
<paragraph blockId="10.[140,1108,1590,1676]" pageId="10" pageNumber="47">
<emphasis bold="true" box="[140,333,1590,1612]" pageId="10" pageNumber="47">Figures 29-32. 29.</emphasis>
Pedipalp, dorsal aspect, 
<taxonomicName authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" box="[561,773,1590,1611]" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="globimanus">
<emphasis box="[561,773,1590,1611]" italics="true" pageId="10" pageNumber="47">Vachoniolus globimanus</emphasis>
</taxonomicName>
, male holotype (scale = 5 mm). 
<emphasis bold="true" pageId="10" pageNumber="47">30- 32.</emphasis>
Metasoma and telson, lateral aspect. 
<emphasis bold="true" box="[519,552,1622,1644]" pageId="10" pageNumber="47">30.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[558,746,1622,1643]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis box="[558,746,1622,1643]" italics="true" pageId="10" pageNumber="47">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[752,804,1623,1644]" pageId="10" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
, male holotype. 
<emphasis bold="true" box="[958,990,1622,1644]" pageId="10" pageNumber="47">3 1.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis italics="true" pageId="10" pageNumber="47">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[248,300,1655,1676]" pageId="10" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
, female holotype. 
<emphasis bold="true" box="[470,503,1654,1676]" pageId="10" pageNumber="47">32.</emphasis>
<taxonomicName baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[509,697,1654,1675]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="doriae">
<emphasis box="[509,697,1654,1675]" italics="true" pageId="10" pageNumber="47">Odontobuthus doriae</emphasis>
</taxonomicName>
, female (scales 30-32 = 10 mm).
</paragraph>
</caption>
<subSection pageId="11" pageNumber="48" type="multiple">
<paragraph blockId="11.[140,1108,167,932]" pageId="11" pageNumber="48">
sternite VII; other sternites acarinated and with two vestigial furrows. Metasoma: segments I to III with 10 crenulated carinae; segment IV with 8 crenulated carinae; ventral carinae strongly marked on segments II to IV with lobate denticles; the first four segments with a smooth dorsal depression; segment 
<collectionCode box="[750,770,273,299]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="48" type="Museum">V</collectionCode>
with five carinae; the latero-
</paragraph>
<paragraph blockId="11.[140,1108,167,932]" pageId="11" pageNumber="48">
ventral carinae crenulate with 2-3 lobate denticles posteriorly; the most posterior very large; ventral median carina slightly divided posteriorly; anal arc composed of 7-8 ventral teeth, and two lateral lobes. Intercarinal spaces moderately granular. Telson with a strongly globular vesicle and some granulations on the lateral and ventral surfaces; aculeus curved and much shorter than the vesicle, without a subaculear tooth. Cheliceral dentition as defined by 
<bibRefCitation author="Vachon" box="[486,657,483,510]" firstAuthor="Vachon" pageId="11" pageNumber="48" pagination="161 - 166" refId="ref7658" refString="Vachon M (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris 2 e ser. 35 (2): 161 - 166." type="journal article" year="1963">Vachon (1963)</bibRefCitation>
for the family 
<taxonomicName authorityName="C. L. Koch" authorityYear="1837" box="[831,937,483,509]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="48" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
; external distal and internal distal teeth approximately the same length; basal teeth on movable finger small but not fused; ventral aspect of both fingers and manus covered with long dense setae. Pedipalps: femur pentacarinate; patella with 7-8 carinae; chela with only vestigial carinae; all faces weakly granular. Fixed and movable fingers with 9-10 oblique rows of granules. Internal and external accessory granules present, strong; three accessory granules on the distal end of the movable finger next to the terminal denticle. Legs: tarsi with two series of short spine-like setae; internal series with 5-7 setae, external reduced to 1-3 setae or totally absent; tibial spur strong on legs III and IV; pedal spurs moderate to strong on legs I to IV. Trichobothriotaxy: trichobothrial pattern of 
<typeStatus box="[1052,1107,801,827]" pageId="11" pageNumber="48">Type</typeStatus>
A, orthobothriotaxic as defined by 
<bibRefCitation author="Vachon" box="[540,715,835,862]" firstAuthor="Vachon" pageId="11" pageNumber="48" pagination="857 - 958" refId="ref7700" refString="Vachon M (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser. n ° 140, Zool. 104: 857 - 958." type="journal article" year="1974">Vachon (1974)</bibRefCitation>
. Dorsal trichobothria of femur arranged in β (beta) configuration (
<bibRefCitation author="Vachon" box="[523,689,870,897]" firstAuthor="Vachon" pageId="11" pageNumber="48" pagination="1597 - 1599" refId="ref7764" refString="Vachon M (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus de l'Academie des Sciences, Paris, ser. D 281: 1597 - 1599." type="journal article" year="1975">Vachon, 1975</bibRefCitation>
). Trichobothrium 
<emphasis bold="true" box="[899,930,871,897]" pageId="11" pageNumber="48">db</emphasis>
of chela totally displaced over the internal face of fixed finger.
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>