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<document id="03985D5C04B91C549E116D8C2067579A" ID-CLB-Dataset="29009" ID-DOI="10.11646/zootaxa.4459.3.8" ID-GBIF-Dataset="98e47881-cd39-4b69-a385-2d5fe5bf56cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458903" ID-ZooBank="0F8B9138-8D58-4D03-BE91-45294DD963CD" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539290289176" checkinUser="plazi" docAuthor="Mariño-Pérez, Ricardo, Fontana, Paolo &amp; Woller, Derek A." docDate="2018" docId="03F387C3FF964B1934E3A8C4E7526BFF" docLanguage="en" docName="zootaxa.4459.3.8.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pedies andreae Mariño-Pérez &amp; Fontana &amp; Woller 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="559" masterDocId="FFCAFFBBFF924B113474AB39E3626D28" masterDocTitle="Studies in Mexican Grasshoppers: A new species of Pedies Saussure (Acrididae: Melanoplinae) with comments on the unusual blue coloration on the abdomens of females and live specimen images for three other congenerics" masterLastPageNumber="564" masterPageNumber="551" pageNumber="555" updateTime="1698578931779" updateUser="ExternalLinkService">
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<mods:title id="035ABD3B9F90717BC54437BAC7DD6D76">Studies in Mexican Grasshoppers: A new species of Pedies Saussure (Acrididae: Melanoplinae) with comments on the unusual blue coloration on the abdomens of females and live specimen images for three other congenerics</mods:title>
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<mods:namePart id="68906F4F07DF7A743316C2D31E5EAF8D">Mariño-Pérez, Ricardo</mods:namePart>
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<mods:namePart id="9A0AA9EFDC1C42AB070AD4B04B6E94A6">Fontana, Paolo</mods:namePart>
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<mods:namePart id="5B650C4245B6C3B8796218F1A05FFBD7">Woller, Derek A.</mods:namePart>
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<treatment id="03F387C3FF964B1934E3A8C4E7526BFF" ID-DOI="http://doi.org/10.5281/zenodo.5993397" ID-GBIF-Taxon="148692878" ID-Zenodo-Dep="5993397" LSID="urn:lsid:plazi:treatment:03F387C3FF964B1934E3A8C4E7526BFF" httpUri="http://treatment.plazi.org/id/03F387C3FF964B1934E3A8C4E7526BFF" lastPageId="8" lastPageNumber="559" pageId="4" pageNumber="555">
<subSubSection id="C340655EFF964B1534E3A8C4E77D6910" pageId="4" pageNumber="555" type="nomenclature">
<paragraph id="8BE536D5FF964B1534E3A8C4E003693F" blockId="4.[151,1055,1021,1080]" box="[151,865,1021,1047]" pageId="4" pageNumber="555">
<heading id="D0AD81B9FF964B1534E3A8C4E003693F" bold="true" box="[151,865,1021,1047]" fontSize="11" level="1" pageId="4" pageNumber="555" reason="1">
<emphasis id="B92EEAC7FF964B1534E3A8C4E003693F" bold="true" box="[151,865,1021,1047]" pageId="4" pageNumber="555">
<taxonomicName id="4C5A4D56FF964B1534E3A8C4E22C693F" ID-CoL="8Q2Y8" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[151,334,1021,1047]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF964B1534E3A8C4E22C693F" bold="true" box="[151,334,1021,1047]" italics="true" pageId="4" pageNumber="555">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF964B153521A8C4E2D1693F" box="[341,435,1021,1047]" pageId="4" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
Fontana, Mariño-Pérez, &amp; Woller
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE536D5FF964B1534E3AF26E77D6910" blockId="4.[151,1055,1021,1080]" box="[151,1055,1055,1080]" pageId="4" pageNumber="555">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="4C5A4D56FF964B15367DAF26E1EB6910" box="[521,649,1055,1080]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:502398
</paragraph>
</subSubSection>
<subSubSection id="C340655EFF964B1534E3AF5EE3946B50" pageId="4" pageNumber="555" type="diagnosis">
<paragraph id="8BE536D5FF964B1534E3AF5EE7A66855" blockId="4.[151,1437,1127,2019]" pageId="4" pageNumber="555">
<emphasis id="B92EEAC7FF964B1534E3AF5EE27169A8" bold="true" box="[151,275,1127,1152]" pageId="4" pageNumber="555">Diagnosis:</emphasis>
Different from congeneric species for the shape of male cerci that are elongate, apically thin and subcylindrical, and relatively short (
<figureCitation id="13612A50FF964B15365FAFB5E1D6698C" box="[555,692,1164,1189]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Figs. 3A&amp;B</figureCitation>
and
<figureCitation id="13612A50FF964B15369FAFB5E05E698C" box="[747,828,1164,1188]" captionStart="FIGURE 10" captionStartId="11.[151,250,1496,1518]" captionTargetBox="[151,1435,249,1440]" captionTargetId="figure@11.[151,1435,233,1440]" captionTargetPageId="11" captionText="FIGURE 10. Genitalia comparison of three Pedies species that appear to be the most closely related based on morphology and geography (see Table 1). External genitalia: A. P. andreae sp. nov., left lateral view; B. P. andreae sp. nov., dorsal view; C. P. capotamius, left lateral view; D. P. capotamius, dorsal view; E. P. malinchensis, left lateral view; F. P. malinchensis, dorsal view; Internal genitalia -ectophallus/endophallus: G. P. andreae sp. nov., left lateral view; H. P. andreae sp. nov., dorsal view; I. P. capotamius, left lateral view; J. P. capotamius, dorsal view; K. P. malinchensis, left lateral view; L. P. malinchensis, dorsal view. *black arrow = sheath of aedeagus, green arrow = dorsal valves of aedeagus, blue arrow = ventral valves of aedeagus [all drawings reprinted from Cigliano and Otte, 2003]" httpUri="https://zenodo.org/record/1458923/files/figure.png" pageId="4" pageNumber="555">10AF</figureCitation>
), and for the phallic complex resembling a lizard head with a deep, open “mouth” formed by the ventral valves of aedeagus when viewed in profile (
<figureCitation id="13612A50FF964B1530A5AF96E65769E0" box="[1233,1333,1199,1224]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Figs. 3C</figureCitation>
,
<figureCitation id="13612A50FF964B153137AF89E60469E0" box="[1347,1382,1200,1224]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="4" pageNumber="555">4C</figureCitation>
, and
<figureCitation id="13612A50FF964B1534E3AFEDE39669C4" box="[151,244,1236,1260]" captionStart="FIGURE 10" captionStartId="11.[151,250,1496,1518]" captionTargetBox="[151,1435,249,1440]" captionTargetId="figure@11.[151,1435,233,1440]" captionTargetPageId="11" captionText="FIGURE 10. Genitalia comparison of three Pedies species that appear to be the most closely related based on morphology and geography (see Table 1). External genitalia: A. P. andreae sp. nov., left lateral view; B. P. andreae sp. nov., dorsal view; C. P. capotamius, left lateral view; D. P. capotamius, dorsal view; E. P. malinchensis, left lateral view; F. P. malinchensis, dorsal view; Internal genitalia -ectophallus/endophallus: G. P. andreae sp. nov., left lateral view; H. P. andreae sp. nov., dorsal view; I. P. capotamius, left lateral view; J. P. capotamius, dorsal view; K. P. malinchensis, left lateral view; L. P. malinchensis, dorsal view. *black arrow = sheath of aedeagus, green arrow = dorsal valves of aedeagus, blue arrow = ventral valves of aedeagus [all drawings reprinted from Cigliano and Otte, 2003]" httpUri="https://zenodo.org/record/1458923/files/figure.png" pageId="4" pageNumber="555">10G,I,K</figureCitation>
). Although this latter characteristic is shared with
<taxonomicName id="4C5A4D56FF964B153752AFECE0A169C4" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[806,963,1237,1260]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="capotamius">
<emphasis id="B92EEAC7FF964B153752AFECE0A169C4" box="[806,963,1237,1260]" italics="true" pageId="4" pageNumber="555">P. capotamius</emphasis>
</taxonomicName>
, the “upper jaw” (ventral portion) is more prominent here coupled with the presence of longer cerci in
<taxonomicName id="4C5A4D56FF964B153725AFC0E0926827" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[849,1008,1272,1296]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="capotamius">
<emphasis id="B92EEAC7FF964B153725AFC0E0926827" box="[849,1008,1272,1296]" italics="true" pageId="4" pageNumber="555">P. capotamius</emphasis>
</taxonomicName>
. Compared with
<taxonomicName id="4C5A4D56FF964B1530CDAFC0E60B6838" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[1209,1385,1272,1296]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF964B1530CDAFC0E60B6838" box="[1209,1385,1272,1296]" italics="true" pageId="4" pageNumber="555">P. malinchensis</emphasis>
</taxonomicName>
, the most similar species by geographic distribution (
<quantity id="4CA29B30FF964B1536B7AE25E072681D" box="[707,784,1308,1333]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="555" unit="km" value="40.0">40 km</quantity>
, different mountain system see
<figureCitation id="13612A50FF964B1530E7AE25E7BA681C" box="[1171,1240,1308,1333]" captionStart="FIGURE 7" captionStartId="8.[151,250,961,983]" captionTargetBox="[151,1436,185,956]" captionTargetId="figure@8.[151,1436,185,956]" captionTargetPageId="8" captionText="FIGURE 7. Map displaying known geographic distributions of Pedies andreae sp. nov. (red triangle) and all 12 previouslydescribed species of Pedies (in alpha order by specific name): P. capotamius (black square), P. cerropotosi (yellow triangle), P. chicoensis (fucsia sand clock), P. comiamus (gray pentagon), P. huacochaus (green cross), P. huarus (yellow star), P. malinchensis (purple plus sign), P. monarca (blue triangle), P. siyonamius (orange rhombus), P. tabeius (white circle), P. tericercus (brown inverted triangle), and P. virescens (blue star)." httpUri="https://zenodo.org/record/1458917/files/figure.png" pageId="4" pageNumber="555">Fig. 7</figureCitation>
), the cerci are of similar length but different form (wider in the second half in
<taxonomicName id="4C5A4D56FF964B15373DAE78E0956870" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[841,1015,1344,1368]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF964B15373DAE78E0956870" box="[841,1015,1344,1368]" italics="true" pageId="4" pageNumber="555">P. malinchensis</emphasis>
</taxonomicName>
) and the “jaws” of the ventral valves of aedeagus are less developed overall. In
<tableCitation id="C6D8036EFF964B153605AE5DE1A66854" box="[625,708,1380,1405]" captionStart="TABLE 1" captionStartId="6.[151,239,152,175]" captionText="TABLE 1. Comparison of three Pedies species that appear to be the most closely related based on genitalia morphology and geography." pageId="4" pageNumber="555">Table 1</tableCitation>
and
<figureCitation id="13612A50FF964B15368EAE5DE00A6854" box="[762,872,1380,1405]" captionStart="FIGURE 10" captionStartId="11.[151,250,1496,1518]" captionTargetBox="[151,1435,249,1440]" captionTargetId="figure@11.[151,1435,233,1440]" captionTargetPageId="11" captionText="FIGURE 10. Genitalia comparison of three Pedies species that appear to be the most closely related based on morphology and geography (see Table 1). External genitalia: A. P. andreae sp. nov., left lateral view; B. P. andreae sp. nov., dorsal view; C. P. capotamius, left lateral view; D. P. capotamius, dorsal view; E. P. malinchensis, left lateral view; F. P. malinchensis, dorsal view; Internal genitalia -ectophallus/endophallus: G. P. andreae sp. nov., left lateral view; H. P. andreae sp. nov., dorsal view; I. P. capotamius, left lateral view; J. P. capotamius, dorsal view; K. P. malinchensis, left lateral view; L. P. malinchensis, dorsal view. *black arrow = sheath of aedeagus, green arrow = dorsal valves of aedeagus, blue arrow = ventral valves of aedeagus [all drawings reprinted from Cigliano and Otte, 2003]" httpUri="https://zenodo.org/record/1458923/files/figure.png" pageId="4" pageNumber="555">Figure 10</figureCitation>
, we compare the three species.
</paragraph>
<paragraph id="8BE536D5FF964B1534B3AEBEE3946B50" blockId="4.[151,1437,1127,2019]" pageId="4" pageNumber="555">
<emphasis id="B92EEAC7FF964B1534B3AEBEE2336888" bold="true" box="[199,337,1415,1440]" pageId="4" pageNumber="555">Coloration:</emphasis>
(
<figureCitation id="13612A50FF964B153515AEBEE2A96888" box="[353,459,1415,1440]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1315,1337]" captionStartId-1="2.[151,250,1607,1629]" captionStartId-2="3.[151,250,1223,1245]" captionTargetBox-0="[151,1436,193,1294]" captionTargetBox-1="[161,1436,409,1585]" captionTargetBox-2="[151,1423,225,1165]" captionTargetId-0="figure@1.[151,1436,193,1294]" captionTargetId-1="figure@2.[151,1436,409,1585]" captionTargetId-2="figure@3.[151,1436,215,1171]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Live specimens of Pedies andreae sp. nov. from type habitat on 3-XII-2011: A. male; B. female; C. and D. Type habitat: grassland with forest patches in Mexico, Puebla, La Cañada, near Libres, [19.510436, -97.773011] (WGS84). 2820 masl. 3-XII-2011." captionText-1="FIGURE 2. Pedies andreae sp. nov. holotype male: A. Habitus, left lateral view; B. Thorax, dorsal view." captionText-2="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri-0="https://zenodo.org/record/1458905/files/figure.png" httpUri-1="https://zenodo.org/record/1458907/files/figure.png" httpUri-2="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Figs. 13</figureCitation>
) Antennae brown, living males with bluish coloration in region just behind and below eyes (
<figureCitation id="13612A50FF964B1534ACAE95E24C68EC" box="[216,302,1452,1477]" captionStart="FIGURE 1" captionStartId="1.[151,250,1315,1337]" captionTargetBox="[151,1436,193,1294]" captionTargetId="figure@1.[151,1436,193,1294]" captionTargetPageId="1" captionText="FIGURE 1. Live specimens of Pedies andreae sp. nov. from type habitat on 3-XII-2011: A. male; B. female; C. and D. Type habitat: grassland with forest patches in Mexico, Puebla, La Cañada, near Libres, [19.510436, -97.773011] (WGS84). 2820 masl. 3-XII-2011." httpUri="https://zenodo.org/record/1458905/files/figure.png" pageId="4" pageNumber="555">Fig. 1A</figureCitation>
) (not obvious in preserved specimens), side of pronotum and metanotum sides bluish in male, brown and greenish in females. Discum of pronotum, tegmina, fore and middle legs brownish in both sexes. Abdomen mostly yellow with blackish spots on sides of each dorsal tergite in male, greenish with a continuous wide dark brown band on each side in females. Blue coloration present on the ventral side of some female abomens, most obvious in live specimens. Lower exterior region of hind femora and tibiae yellow in males, reddish-orange in females.
</paragraph>
</subSubSection>
<subSubSection id="C340655EFF964B1934B3ADBDE7EC69F7" lastPageId="8" lastPageNumber="559" pageId="4" pageNumber="555" type="description">
<paragraph id="8BE536D5FF964B1534B3ADBDE3B36A07" blockId="4.[151,1437,1127,2019]" pageId="4" pageNumber="555">
<emphasis id="B92EEAC7FF964B1534B3ADBDE28E6BB5" bold="true" box="[199,492,1668,1693]" pageId="4" pageNumber="555">Pronotum and Tegmina:</emphasis>
(
<figureCitation id="13612A50FF964B15358FADBDE12B6BB4" box="[507,585,1668,1693]" captionStart="FIGURE 2" captionStartId="2.[151,250,1607,1629]" captionTargetBox="[161,1436,409,1585]" captionTargetId="figure@2.[151,1436,409,1585]" captionTargetPageId="2" captionText="FIGURE 2. Pedies andreae sp. nov. holotype male: A. Habitus, left lateral view; B. Thorax, dorsal view." httpUri="https://zenodo.org/record/1458907/files/figure.png" pageId="4" pageNumber="555">Figs. 2</figureCitation>
and
<figureCitation id="13612A50FF964B15360BADBDE1EF6BB5" box="[639,653,1668,1693]" captionStart="FIGURE 5" captionStartId="6.[151,250,1987,2009]" captionTargetBox="[162,1436,928,1965]" captionTargetId="figure@6.[151,1436,928,1965]" captionTargetPageId="6" captionText="FIGURE 5. Pedies andreae sp. nov. paratype female specimen: A. Habitus, left lateral view; B. Thorax, dorsal view." httpUri="https://zenodo.org/record/1458913/files/figure.png" pageId="4" pageNumber="555">5</figureCitation>
) Pronotum in both sexes straight in lateral view, scarcely tectiform in transversal section. Anterior and posterior margin of pronotum almost straight in both sexes. Tegmina in both sexes lateral, not attingent on dorsum, regularly elliptical with widely rounded apex, reaching the middle of the 1
<taxonomicName id="4C5A4D56FF964B1531E6ADF2E26F6A23" pageId="4" pageNumber="555" rank="subSpecies" subSpecies="abdominal">st abdominal</taxonomicName>
tergites; venation dense with raised longitudinal (almost parallel) veins and less prominent transversal ones.
</paragraph>
<paragraph id="8BE536D5FF964B1434B3AC03E17E6EA8" blockId="4.[151,1437,1127,2019]" lastBlockId="5.[151,1437,151,896]" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
<emphasis id="B92EEAC7FF964B1534B3AC03E1446A7B" bold="true" box="[199,550,1850,1875]" pageId="4" pageNumber="555">Terminalia: Male, external.</emphasis>
Furcula subtriangular and relatively short (
<figureCitation id="13612A50FF964B153021AC03E7DA6A7B" box="[1109,1208,1850,1875]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Fig. 3B</figureCitation>
). Supra-anal plate subtriangular, with acute apex and convex lateral sides; surface almost plain with shallow, median groove that extends apically for almost entire length and is deepest basally (
<figureCitation id="13612A50FF964B153711ACBBE0DE6AB3" box="[869,956,1922,1947]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Fig. 3B</figureCitation>
). Cerci compressed laterally and lightly Sshaped (stretched anteroposteriorly), with apical halves subcylindrical and tapering to rounded points that curve slightly inwards; extend about 3/4 the length of subgenital plate (
<figureCitation id="13612A50FF964B1537E6ACF3E77B6ACB" box="[914,1049,1994,2019]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="4" pageNumber="555">Fig. 3A&amp;B</figureCitation>
). Subgenital plate hemiconical in lateral view and strongly pointed at apex (
<figureCitation id="13612A50FF974B1436D3ABAEE0506D98" box="[679,818,151,176]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="5" pageNumber="556">Fig. 3A&amp;B</figureCitation>
).
<emphasis id="B92EEAC7FF974B143739ABAEE7E96D98" bold="true" box="[845,1163,151,176]" pageId="5" pageNumber="556">Internal phallic complex:</emphasis>
Overall, typical for a melanopline, with the following unique characters: Epiphallus: ancorae relatively elongate, subtriangular, and gently curving ventrally; lophi prominent, subrectangular, and with internal corners the highest (
<figureCitation id="13612A50FF974B143094ABE6E6096DD0" box="[1248,1387,223,248]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="5" pageNumber="556">Figs. 3C&amp;D</figureCitation>
and
<figureCitation id="13612A50FF974B1434E3AA3DE3D96C34" box="[151,187,260,284]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556">4B</figureCitation>
); post-epiphallic lobe (
<bibRefCitation id="EFCB4B24FF974B1435A5AA3DE06C6C35" box="[465,782,260,285]" pageId="5" pageNumber="556" refId="ref4720" refString="Gurney, A. B. &amp; Buxton, G. M. (1968) Grasshoppers of the genus Karokia and of the saltator, immunis, and harperi species groups of the genus Melanoplus in California and Oregon (Orthoptera: Acrididae). Occasional Papers of the Bureau of Entomology, California Department of Agriculture, 13, 1 - 56." type="journal article">Gurney and Buxton, 1968</bibRefCitation>
;
<bibRefCitation id="EFCB4B24FF974B143755AA3DE0826C34" box="[801,992,260,285]" pageId="5" pageNumber="556" refId="ref4319" refString="Carbonell, C. S. (2000) Taxonomy and a study of the phallic complex, including its muscles, of Paulinia acuminata (Acrididae, Pauliniinae) and Marellia remipes (Acrididae, incertae sedis). Journal of Orthoptera Research, 9, 161 - 180. https: // doi. org / 10.2307 / 3503647" type="journal article">Carbonell, 2000</bibRefCitation>
;
<bibRefCitation id="EFCB4B24FF974B143787AA3DE79E6C34" box="[1011,1276,260,285]" pageId="5" pageNumber="556" refId="ref4380" refString="Carbonell, C. S., Mesa, A. &amp; Condutta, V. L. (1980) Systematics and Karyotype of the Grasshopper Genera Omalotettix and Abracris. Proceedings of the Academy of Natural Sciences of Philadelphia, 132, 279 - 284." type="journal article">
Carbonell
<emphasis id="B92EEAC7FF974B14301BAA3CE7C56C34" box="[1135,1191,260,284]" italics="true" pageId="5" pageNumber="556">et al</emphasis>
., 1980
</bibRefCitation>
) only mildly wrinkled, with two prominent and elongate sclerotized portions that angle anteriorly towards the center (
<figureCitation id="13612A50FF974B143140AA1EE6F26C68" box="[1332,1424,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556">Fig. 4A</figureCitation>
). Ectophallus: rami prominent, and relatively wide, narrowing posteriorly and extending well below valves of aedeagus (
<figureCitation id="13612A50FF974B143579AA56E2F36CA0" box="[269,401,367,392]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556">Fig. 4A&amp;C</figureCitation>
). Sheath of aedeagus taking the form of two halves that do not meet, each containing a lobe that broadly attaches to apices of rami and are covered in raised microstructures resembling those on the postepiphallic lobe; posterior portion of each with a deep, but broad, envagination resembling the &quot;lizard jaws&quot; of the ventral valves of aedeagus that are emerging from the center of each lobe; lower posterior apex of each extending further posteriorly (
<figureCitation id="13612A50FF974B14350BAAC6E1776F30" box="[383,533,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556">Fig. 4A,C,D</figureCitation>
). Endophallus: arch of aedeagus well-developed. Dorsal valves of aedeagus hidden within sheath of aedeagus and not visible without dissection; do not meet flexures and about ½ the length of ventral valves, relatively wide and slightly separated basodorsally just beyond the arch. Ventral valves of aedeagus meet flexures and are twice as long as the dorsal valves, robust throughout with unusual apices that are sclerotized, and covered in raised microstructures; strongly resemble the head of a lizard when viewed laterally, with an open, deep &quot;mouth&quot; that has both &quot;jaws developed&quot;, upper/ventral more-so (where the eye would be); in posterodorsal view, the apices of the upper jaws are lightly excavated (
<figureCitation id="13612A50FF974B143745A9EEE0DC6FD8" box="[817,958,727,752]" captionStart="FIGURE 3" captionStartId="3.[151,250,1223,1245]" captionTargetBox="[151,1423,225,1165]" captionTargetId="figure@3.[151,1436,215,1171]" captionTargetPageId="3" captionText="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." httpUri="https://zenodo.org/record/1458909/files/figure.png" pageId="5" pageNumber="556">Figs. 3C&amp;D</figureCitation>
and
<figureCitation id="13612A50FF974B14378FA9E1E7346FD8" box="[1019,1110,728,752]" captionStart="FIGURE 4" captionStartId="5.[151,250,1976,1998]" captionTargetBox="[162,1420,975,1920]" captionTargetId="figure@5.[151,1436,959,1932]" captionTargetPageId="5" captionText="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556">4A,C,D</figureCitation>
).
<emphasis id="B92EEAC7FF974B143018A9EEE65D6FD8" bold="true" box="[1132,1343,727,752]" pageId="5" pageNumber="556">Female, external:</emphasis>
Female supra-anal plate subtriangular, elongated, with acutely rounded apex and surface almost plain (
<figureCitation id="13612A50FF974B14309EA9C5E62A6E3D" box="[1258,1352,764,789]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[362,1221,194,1836]" captionTargetId="figure@7.[362,1225,193,1851]" captionTargetPageId="7" captionText="FIGURE 6. Pedies andreae sp. nov. paratype female: External genitalia: A. Dorsal view; B. Left lateral view; C. Ventral view." httpUri="https://zenodo.org/record/1458915/files/figure.png" pageId="5" pageNumber="556">Fig. 6A</figureCitation>
). Cerci subconical, flattened longer 1.5 times as wide at base (
<figureCitation id="13612A50FF974B14376EA826E0C36E10" box="[794,929,799,824]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[362,1221,194,1836]" captionTargetId="figure@7.[362,1225,193,1851]" captionTargetPageId="7" captionText="FIGURE 6. Pedies andreae sp. nov. paratype female: External genitalia: A. Dorsal view; B. Left lateral view; C. Ventral view." httpUri="https://zenodo.org/record/1458915/files/figure.png" pageId="5" pageNumber="556">Fig. 6A&amp;B</figureCitation>
). Ovipositor quite long; ventral valves with strong tooth on the lower margin (
<figureCitation id="13612A50FF974B143651A87DE1E26E75" box="[549,640,836,861]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[362,1221,194,1836]" captionTargetId="figure@7.[362,1225,193,1851]" captionTargetPageId="7" captionText="FIGURE 6. Pedies andreae sp. nov. paratype female: External genitalia: A. Dorsal view; B. Left lateral view; C. Ventral view." httpUri="https://zenodo.org/record/1458915/files/figure.png" pageId="5" pageNumber="556">Fig. 6B</figureCitation>
). Subgenital plate with posterior margin subtriangular, acute apex on concave lateral sides (
<figureCitation id="13612A50FF974B1435FAA85EE16D6EA8" box="[398,527,871,896]" captionStart="FIGURE 6" captionStartId="7.[151,250,1872,1894]" captionTargetBox="[362,1221,194,1836]" captionTargetId="figure@7.[362,1225,193,1851]" captionTargetPageId="7" captionText="FIGURE 6. Pedies andreae sp. nov. paratype female: External genitalia: A. Dorsal view; B. Left lateral view; C. Ventral view." httpUri="https://zenodo.org/record/1458915/files/figure.png" pageId="5" pageNumber="556">Fig. 6B&amp;C</figureCitation>
).
</paragraph>
<caption id="DF25665DFF974B1434E3AC81E63C6AC5" httpUri="https://zenodo.org/record/1458911/files/figure.png" pageId="5" pageNumber="556" startId="5.[151,250,1976,1998]" targetBox="[162,1420,975,1920]" targetPageId="5">
<paragraph id="8BE536D5FF974B1434E3AC81E63C6AC5" blockId="5.[151,1436,1976,2029]" pageId="5" pageNumber="556">
<emphasis id="B92EEAC7FF974B1434E3AC81E2766AE6" bold="true" box="[151,276,1976,1998]" pageId="5" pageNumber="556">FIGURE 4.</emphasis>
<taxonomicName id="4C5A4D56FF974B143569AC81E2D96AE6" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[285,443,1976,1998]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF974B143569AC81E2D96AE6" box="[285,443,1976,1998]" italics="true" pageId="5" pageNumber="556">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF974B1435B7AC81E1746AE6" box="[451,534,1976,1998]" pageId="5" pageNumber="556" rank="species">
<emphasis id="B92EEAC7FF974B1435B7AC81E1746AE6" bold="true" box="[451,534,1976,1998]" pageId="5" pageNumber="556">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype male: Internal genitalia (KOH-cleared):
<emphasis id="B92EEAC7FF974B14306BAC81E75A6AE6" bold="true" box="[1055,1080,1976,1998]" pageId="5" pageNumber="556">A.</emphasis>
Complete phallic complex, dorsal view;
<emphasis id="B92EEAC7FF974B1434A1ACEEE38E6AC5" bold="true" box="[213,236,2007,2029]" pageId="5" pageNumber="556">B.</emphasis>
Epiphallus, dorsal view;
<emphasis id="B92EEAC7FF974B14359AACEEE1656AC5" bold="true" box="[494,519,2007,2029]" pageId="5" pageNumber="556">C.</emphasis>
Ectophallus/endophallus, left lateral view;
<emphasis id="B92EEAC7FF974B1437CBACEEE0BA6AC5" bold="true" box="[959,984,2007,2029]" pageId="5" pageNumber="556">D.</emphasis>
Ectophallus/endophallus, dorsal view.
</paragraph>
</caption>
<caption id="DF25665DFF944B1734E3ABA1E2586DE7" pageId="6" pageNumber="557">
<paragraph id="8BE536D5FF944B1734E3ABA1E2586DE7" blockId="6.[151,1435,151,207]" pageId="6" pageNumber="557">
<emphasis id="B92EEAC7FF944B1734E3ABA1E2686D87" bold="true" box="[151,266,152,175]" pageId="6" pageNumber="557">TABLE 1.</emphasis>
Comparison of three
<taxonomicName id="4C5A4D56FF944B173580ABAEE1596D87" box="[500,571,151,175]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC7FF944B173580ABAEE1596D87" box="[500,571,151,175]" italics="true" pageId="6" pageNumber="557">Pedies</emphasis>
</taxonomicName>
species that appear to be the most closely related based on genitalia morphology and geography.
</paragraph>
</caption>
<paragraph id="8BE536D5FF944B1735FEABD3E65C6E7D" pageId="6" pageNumber="557">
<table id="F95AC475FF94B4EE34EBABD3E6076E7D" box="[159,1381,234,853]" gridcols="4" gridrows="8" pageId="6" pageNumber="557">
<tr id="356A3497FF94B4EE34EBABD3E6076C28" box="[159,1381,234,256]" gridrow="0" pageId="6" pageNumber="557" rowspan-0="1">
<th id="76BB5DEBFF94B4EE35FEABD3E1B16C28" box="[394,723,234,256]" gridcol="1" gridrow="0" pageId="6" pageNumber="557">
<taxonomicName id="4C5A4D56FF944B1735FEABD3E1446C28" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[394,550,234,256]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF944B1735FEABD3E1446C28" box="[394,550,234,256]" italics="true" pageId="6" pageNumber="557">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF944B173658ABD3E11F6C28" box="[556,637,234,256]" pageId="6" pageNumber="557" rank="species">
<emphasis id="B92EEAC7FF944B173658ABD3E11F6C28" bold="true" box="[556,637,234,256]" pageId="6" pageNumber="557">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="76BB5DEBFF94B4EE3692ABD3E74A6C28" box="[742,1064,234,256]" gridcol="2" gridrow="0" pageId="6" pageNumber="557">
<taxonomicName id="4C5A4D56FF944B173692ABD3E0C76C28" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[742,933,234,256]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="capotamius">
<emphasis id="B92EEAC7FF944B173692ABD3E0C76C28" box="[742,933,234,256]" italics="true" pageId="6" pageNumber="557">Pedies capotamius</emphasis>
</taxonomicName>
</th>
<th id="76BB5DEBFF94B4EE304FABD3E6076C28" box="[1083,1381,234,256]" gridcol="3" gridrow="0" pageId="6" pageNumber="557">
<taxonomicName id="4C5A4D56FF944B173048ABD3E66B6C28" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[1084,1289,234,256]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF944B173048ABD3E66B6C28" box="[1084,1289,234,256]" italics="true" pageId="6" pageNumber="557">Pedies malinchensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356A3497FF94B4EE34EBAA2DE6076C02" box="[159,1381,276,298]" gridrow="1" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBAA2DE2126C02" box="[159,368,276,298]" gridcol="0" gridrow="1" pageId="6" pageNumber="557">Cerci length</th>
<td id="76BB5DEBFF94B4EE35FEAA2DE1B16C02" box="[394,723,276,298]" gridcol="1" gridrow="1" pageId="6" pageNumber="557">shorter (Fig. 10A&amp;B)</td>
<td id="76BB5DEBFF94B4EE3692AA2DE74A6C02" box="[742,1064,276,298]" gridcol="2" gridrow="1" pageId="6" pageNumber="557">longer (Fig. 10C&amp;D)</td>
<td id="76BB5DEBFF94B4EE304FAA2DE6076C02" box="[1083,1381,276,298]" gridcol="3" gridrow="1" pageId="6" pageNumber="557">shorter (Fig. 10E&amp;F)</td>
</tr>
<tr id="356A3497FF94B4EE34EBAA04E6076C5A" box="[159,1381,317,370]" gridrow="2" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBAA04E2126C5A" box="[159,368,317,370]" gridcol="0" gridrow="2" pageId="6" pageNumber="557">Cerci form</th>
<td id="76BB5DEBFF94B4EE35FEAA04E1B16C5A" box="[394,723,317,370]" gridcol="1" gridrow="2" pageId="6" pageNumber="557">basal half tapering and subcylindrical (Fig. 10A&amp;B)</td>
<td id="76BB5DEBFF94B4EE3692AA04E74A6C5A" box="[742,1064,317,370]" gridcol="2" gridrow="2" pageId="6" pageNumber="557">basal half wide and flattened (Fig. 10C&amp;D)</td>
<td id="76BB5DEBFF94B4EE304FAA04E6076C5A" box="[1083,1381,317,370]" gridcol="3" gridrow="2" pageId="6" pageNumber="557">basal half wide and flattened (Fig. 10E&amp;F)</td>
</tr>
<tr id="356A3497FF94B4EE34EBAABFE6076C92" box="[159,1381,390,442]" gridrow="3" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBAABFE2126C92" box="[159,368,390,442]" gridcol="0" gridrow="3" pageId="6" pageNumber="557">Subgenital plate</th>
<td id="76BB5DEBFF94B4EE35FEAABFE1B16C92" box="[394,723,390,442]" gridcol="1" gridrow="3" pageId="6" pageNumber="557">apex strongly pointed (Fig. 10A&amp;B)</td>
<td id="76BB5DEBFF94B4EE3692AABFE74A6C92" box="[742,1064,390,442]" gridcol="2" gridrow="3" pageId="6" pageNumber="557">apex rounded (Fig. 10C&amp;D)</td>
<td id="76BB5DEBFF94B4EE304FAABFE6076C92" box="[1083,1381,390,442]" gridcol="3" gridrow="3" pageId="6" pageNumber="557">apex moderately pointed (Fig. 10E&amp;F)</td>
</tr>
<tr id="356A3497FF94B4EE34EBAAF7E6076F09" box="[159,1381,462,545]" gridrow="4" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBAAF7E2126F09" box="[159,368,462,545]" gridcol="0" gridrow="4" pageId="6" pageNumber="557">Sheath of aedeagus (in profile)</th>
<td id="76BB5DEBFF94B4EE35FEAAF7E1B16F09" box="[394,723,462,545]" gridcol="1" gridrow="4" pageId="6" pageNumber="557">
upper portion in-between
<taxonomicName id="4C5A4D56FF944B1736E4AAF6E29D6F2A" authorityName="Cigliano &amp; Otte" authorityYear="2003" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="capotamius">
<emphasis id="B92EEAC7FF944B1736E4AAF6E29D6F2A" italics="true" pageId="6" pageNumber="557">P. capotamius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D56FF944B173644AAD4E1AF6F2A" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[560,717,492,514]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF944B173644AAD4E1AF6F2A" box="[560,717,492,514]" italics="true" pageId="6" pageNumber="557">P. malinchensis</emphasis>
</taxonomicName>
(Fig. 10G - black arrow)
</td>
<td id="76BB5DEBFF94B4EE3692AAF7E74A6F09" box="[742,1064,462,545]" gridcol="2" gridrow="4" pageId="6" pageNumber="557">upper portion highly constricted (Fig. 10I - black arrow)</td>
<td id="76BB5DEBFF94B4EE304FAAF7E6076F09" box="[1083,1381,462,545]" gridcol="3" gridrow="4" pageId="6" pageNumber="557">upper portion globose (Fig. 10K - black arrow)</td>
</tr>
<tr id="356A3497FF94B4EE34EBA90DE6076F41" box="[159,1381,564,617]" gridrow="5" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBA90DE2126F41" box="[159,368,564,617]" gridcol="0" gridrow="5" pageId="6" pageNumber="557">Dorsal valves of aedeagus</th>
<td id="76BB5DEBFF94B4EE35FEA90DE1B16F41" box="[394,723,564,617]" gridcol="1" gridrow="5" pageId="6" pageNumber="557">not visible (Fig. 10G - green arrow)</td>
<td id="76BB5DEBFF94B4EE3692A90DE74A6F41" box="[742,1064,564,617]" gridcol="2" gridrow="5" pageId="6" pageNumber="557">visible (Fig. 10I - green arrow)</td>
<td id="76BB5DEBFF94B4EE304FA90DE6076F41" box="[1083,1381,564,617]" gridcol="3" gridrow="5" pageId="6" pageNumber="557">visible and slightly arched (Fig. 10K - green arrow)</td>
</tr>
<tr id="356A3497FF94B4EE34EBA945E6076FC6" box="[159,1381,636,750]" gridrow="6" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBA945E2126FC6" box="[159,368,636,750]" gridcol="0" gridrow="6" pageId="6" pageNumber="557">Ventral valves of aedeagus (in profile)</th>
<td id="76BB5DEBFF94B4EE35FEA945E1B16FC6" box="[394,723,636,750]" gridcol="1" gridrow="6" pageId="6" pageNumber="557">resembles lizard head; open, deep mouth with both jaws developed, upper/ventral more- so (Fig. 10G - blue arrow)</td>
<td id="76BB5DEBFF94B4EE3692A945E74A6FC6" box="[742,1064,636,750]" gridcol="2" gridrow="6" pageId="6" pageNumber="557">resembles lizard head; open, deep mouth with both jaws developed, upper/ventral less-so (Fig. 10I - blue arrow)</td>
<td id="76BB5DEBFF94B4EE304FA945E6076FC6" box="[1083,1381,636,750]" gridcol="3" gridrow="6" pageId="6" pageNumber="557">resembles lizard head; open, shallow mouth (Fig. 10K - blue arrow)</td>
</tr>
<tr id="356A3497FF94B4EE34EBA83BE6076E7D" box="[159,1381,770,853]" gridrow="7" pageId="6" pageNumber="557">
<th id="76BB5DEBFF94B4EE34EBA83BE2126E7D" box="[159,368,770,853]" gridcol="0" gridrow="7" pageId="6" pageNumber="557">Geography</th>
<td id="76BB5DEBFF94B4EE35FEA83BE1B16E7D" box="[394,723,770,853]" gridcol="1" gridrow="7" pageId="6" pageNumber="557">
close proximity to
<taxonomicName id="4C5A4D56FF944B17363DA83AE16C6E1E" authorityName="Cigliano &amp; Otte" authorityYear="2003" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF944B17363DA83AE16C6E1E" italics="true" pageId="6" pageNumber="557">P. malinchensis</emphasis>
</taxonomicName>
(slightly southwest) (Fig. 7 - red triangle)
</td>
<td id="76BB5DEBFF94B4EE3692A83BE74A6E7D" box="[742,1064,770,853]" gridcol="2" gridrow="7" pageId="6" pageNumber="557">
some distance west from both
<taxonomicName id="4C5A4D56FF944B173060A83AE05A6E1E" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="B92EEAC7FF944B173060A83AE05A6E1E" italics="true" pageId="6" pageNumber="557">P. andreae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D56FF944B17371DA81BE7646E1E" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[873,1030,800,823]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF944B17371DA81BE7646E1E" box="[873,1030,800,823]" italics="true" pageId="6" pageNumber="557">P. malinchensis</emphasis>
</taxonomicName>
(Fig. 7 - black square)
</td>
<td id="76BB5DEBFF94B4EE304FA83BE6076E7D" box="[1083,1381,770,853]" gridcol="3" gridrow="7" pageId="6" pageNumber="557">
close proximity to
<taxonomicName id="4C5A4D56FF944B17308EA83AE6076E30" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[1274,1381,770,792]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="B92EEAC7FF944B17308EA83AE6076E30" box="[1274,1381,770,792]" italics="true" pageId="6" pageNumber="557">P. andreae</emphasis>
</taxonomicName>
(slightly northeast) (Fig. 7 - purple plus sign)
</td>
</tr>
</table>
</paragraph>
<caption id="DF25665DFF944B1734E3ACFAE65F6AF1" box="[151,1341,1987,2009]" httpUri="https://zenodo.org/record/1458913/files/figure.png" pageId="6" pageNumber="557" startId="6.[151,250,1987,2009]" targetBox="[162,1436,928,1965]" targetPageId="6">
<paragraph id="8BE536D5FF944B1734E3ACFAE65F6AF1" blockId="6.[151,1341,1987,2009]" box="[151,1341,1987,2009]" pageId="6" pageNumber="557">
<emphasis id="B92EEAC7FF944B1734E3ACFAE2706AF1" bold="true" box="[151,274,1987,2009]" pageId="6" pageNumber="557">FIGURE 5.</emphasis>
<taxonomicName id="4C5A4D56FF944B17356DACFAE2D76AF1" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[281,437,1987,2009]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="557" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF944B17356DACFAE2D76AF1" box="[281,437,1987,2009]" italics="true" pageId="6" pageNumber="557">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF944B1735CFACFAE1696AF1" box="[443,523,1987,2009]" pageId="6" pageNumber="557" rank="species">
<emphasis id="B92EEAC7FF944B1735CFACFAE1696AF1" bold="true" box="[443,523,1987,2009]" pageId="6" pageNumber="557">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype female specimen:
<emphasis id="B92EEAC7FF944B17375CACFAE0226AF1" bold="true" box="[808,832,1987,2009]" pageId="6" pageNumber="557">A.</emphasis>
Habitus, left lateral view;
<emphasis id="B92EEAC7FF944B17303AACFAE7046AF1" bold="true" box="[1102,1126,1987,2009]" pageId="6" pageNumber="557">B.</emphasis>
Thorax, dorsal view.
</paragraph>
</caption>
<caption id="DF25665DFF954B1634E3AC69E6FF6A4E" box="[151,1437,1872,1894]" httpUri="https://zenodo.org/record/1458915/files/figure.png" pageId="7" pageNumber="558" startId="7.[151,250,1872,1894]" targetBox="[362,1221,194,1836]" targetPageId="7">
<paragraph id="8BE536D5FF954B1634E3AC69E6FF6A4E" blockId="7.[151,1437,1872,1894]" box="[151,1437,1872,1894]" pageId="7" pageNumber="558">
<emphasis id="B92EEAC7FF954B1634E3AC69E2706A4E" bold="true" box="[151,274,1872,1894]" pageId="7" pageNumber="558">FIGURE 6.</emphasis>
<taxonomicName id="4C5A4D56FF954B16356CAC69E2D66A4E" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[280,436,1872,1894]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="558" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF954B16356CAC69E2D66A4E" box="[280,436,1872,1894]" italics="true" pageId="7" pageNumber="558">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF954B1635CEAC69E1686A4E" box="[442,522,1872,1894]" pageId="7" pageNumber="558" rank="species">
<emphasis id="B92EEAC7FF954B1635CEAC69E1686A4E" bold="true" box="[442,522,1872,1894]" pageId="7" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype female: External genitalia:
<emphasis id="B92EEAC7FF954B1637F4AC69E0FB6A4E" bold="true" box="[896,921,1872,1894]" pageId="7" pageNumber="558">A.</emphasis>
Dorsal view;
<emphasis id="B92EEAC7FF954B163053AC69E75C6A4E" bold="true" box="[1063,1086,1872,1894]" pageId="7" pageNumber="558">B.</emphasis>
Left lateral view;
<emphasis id="B92EEAC7FF954B16308CAC69E6726A4E" bold="true" box="[1272,1296,1872,1894]" pageId="7" pageNumber="558">C.</emphasis>
Ventral view.
</paragraph>
</caption>
<paragraph id="8BE536D5FF954B1634B3ACB7E72A6AC7" blockId="7.[151,1436,1934,2031]" pageId="7" pageNumber="558">
<emphasis id="B92EEAC7FF954B1634B3ACB7E17F6A8F" bold="true" box="[199,541,1934,1959]" pageId="7" pageNumber="558">Male measurements (in mm)</emphasis>
(n=3): Body length “Entire body” 18.8019.55 (19.14 ± 0.37); pronotum length 3.944.04 (4.01 ± 0.05); prozona length 2.442.53 (2.47 ± 0.05); metazona length 1.501.59 (1.53 ± 0.05); hind femur length 9.8710.34 (10.15 ± 0.24) and tegmina length 3.103.29 (3.22 ± 0.10).
</paragraph>
<caption id="DF25665DFF9A4B1934E3A8F8E0406979" httpUri="https://zenodo.org/record/1458917/files/figure.png" pageId="8" pageNumber="559" startId="8.[151,250,961,983]" targetBox="[151,1436,185,956]" targetPageId="8">
<paragraph id="8BE536D5FF9A4B1934E3A8F8E0406979" blockId="8.[151,1437,961,1105]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934E3A8F8E2766EFF" bold="true" box="[151,276,961,983]" pageId="8" pageNumber="559">FIGURE 7.</emphasis>
Map displaying known geographic distributions of
<taxonomicName id="4C5A4D56FF9A4B193758A8F8E0AB6EFF" authority="Mariño-Pérez &amp; Fontana &amp; Woller, 2018" authorityName="Mariño-Pérez &amp; Fontana &amp; Woller" authorityYear="2018" box="[812,969,961,983]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="andreae" status="sp. nov.">
<emphasis id="B92EEAC7FF9A4B193758A8F8E0AB6EFF" box="[812,969,961,983]" italics="true" pageId="8" pageNumber="559">Pedies andreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BCFF9A4B1937A5A8F8E7416EFF" box="[977,1059,961,983]" pageId="8" pageNumber="559" rank="species">
<emphasis id="B92EEAC7FF9A4B1937A5A8F8E7416EFF" bold="true" box="[977,1059,961,983]" pageId="8" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
(red triangle) and all 12 previouslydescribed species of
<taxonomicName id="4C5A4D56FF9A4B19351DA8E6E2CF6EDD" box="[361,429,991,1013]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC7FF9A4B19351DA8E6E2CF6EDD" box="[361,429,991,1013]" italics="true" pageId="8" pageNumber="559">Pedies</emphasis>
</taxonomicName>
(in alpha order by specific name):
<taxonomicName id="4C5A4D56FF9A4B19377BA8D9E0FF6EDD" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[783,925,992,1013]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="capotamius">
<emphasis id="B92EEAC7FF9A4B19377BA8D9E0FF6EDD" box="[783,925,992,1013]" italics="true" pageId="8" pageNumber="559">P. capotamius</emphasis>
</taxonomicName>
(black square),
<taxonomicName id="4C5A4D56FF9A4B193034A8D9E7AF6EDD" authorityName="Fontana &amp; Buzzetti" authorityYear="2007" box="[1088,1229,992,1013]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="cerropotosi">
<emphasis id="B92EEAC7FF9A4B193034A8D9E7AF6EDD" box="[1088,1229,992,1013]" italics="true" pageId="8" pageNumber="559">P. cerropotosi</emphasis>
</taxonomicName>
(yellow triangle),
<taxonomicName id="4C5A4D56FF9A4B1931FDA8D9E263693C" authorityName="Cigliano &amp; Otte" authorityYear="2003" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="chicoensis">
<emphasis id="B92EEAC7FF9A4B1931FDA8D9E263693C" italics="true" pageId="8" pageNumber="559">P. chicoensis</emphasis>
</taxonomicName>
(fucsia sand clock),
<taxonomicName id="4C5A4D56FF9A4B193598A8C6E111693C" box="[492,627,1023,1044]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="comiamus">
<emphasis id="B92EEAC7FF9A4B193598A8C6E111693C" box="[492,627,1023,1044]" italics="true" pageId="8" pageNumber="559">P. comiamus</emphasis>
</taxonomicName>
(gray pentagon),
<taxonomicName id="4C5A4D56FF9A4B19374EA8C6E0B0693C" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[826,978,1022,1044]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="huacochaus">
<emphasis id="B92EEAC7FF9A4B19374EA8C6E0B0693C" box="[826,978,1022,1044]" italics="true" pageId="8" pageNumber="559">P. huacochaus</emphasis>
</taxonomicName>
(green cross),
<taxonomicName id="4C5A4D56FF9A4B193008A8C6E780693C" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[1148,1250,1022,1044]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="huarus">
<emphasis id="B92EEAC7FF9A4B193008A8C6E780693C" box="[1148,1250,1022,1044]" italics="true" pageId="8" pageNumber="559">P. huarus</emphasis>
</taxonomicName>
(yellow star),
<taxonomicName id="4C5A4D56FF9A4B1931FDA8C6E279691A" authorityName="Cigliano &amp; Otte" authorityYear="2003" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="malinchensis">
<emphasis id="B92EEAC7FF9A4B1931FDA8C6E279691A" italics="true" pageId="8" pageNumber="559">P. malinchensis</emphasis>
</taxonomicName>
(purple plus sign),
<taxonomicName id="4C5A4D56FF9A4B193586AF27E10A691B" authorityName="Buzzetti, Barrientos Lozano &amp; Fontana" authorityYear="2010" box="[498,616,1054,1075]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="monarca">
<emphasis id="B92EEAC7FF9A4B193586AF27E10A691B" box="[498,616,1054,1075]" italics="true" pageId="8" pageNumber="559">P. monarca</emphasis>
</taxonomicName>
(blue triangle),
<taxonomicName id="4C5A4D56FF9A4B19376CAF27E0C6691B" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[792,932,1054,1075]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="siyonamius">
<emphasis id="B92EEAC7FF9A4B19376CAF27E0C6691B" box="[792,932,1054,1075]" italics="true" pageId="8" pageNumber="559">P. siyonamius</emphasis>
</taxonomicName>
(orange rhombus),
<taxonomicName id="4C5A4D56FF9A4B193008AF27E780691A" authorityName="Cigliano &amp; Otte" authorityYear="2003" box="[1148,1250,1052,1075]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="tabeius">
<emphasis id="B92EEAC7FF9A4B193008AF27E780691A" box="[1148,1250,1052,1075]" italics="true" pageId="8" pageNumber="559">P. tabeius</emphasis>
</taxonomicName>
(white circle),
<taxonomicName id="4C5A4D56FF9A4B1931FDAF27E3996979" authorityName="Cigliano &amp; Otte" authorityYear="2003" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="tericercus">
<emphasis id="B92EEAC7FF9A4B1931FDAF27E3996979" italics="true" pageId="8" pageNumber="559">P. tericercus</emphasis>
</taxonomicName>
(brown inverted triangle), and
<taxonomicName id="4C5A4D56FF9A4B193643AF05E1CF6979" authorityName="Saussure" authorityYear="1861" box="[567,685,1084,1105]" class="Insecta" family="Acrididae" genus="Pedies" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="559" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B92EEAC7FF9A4B193643AF05E1CF6979" box="[567,685,1084,1105]" italics="true" pageId="8" pageNumber="559">P. virescens</emphasis>
</taxonomicName>
(blue star).
</paragraph>
</caption>
<paragraph id="8BE536D5FF9A4B1934B3AF47E7EC69F7" blockId="8.[151,1437,1150,1751]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934B3AF47E12069BF" bold="true" box="[199,578,1150,1175]" pageId="8" pageNumber="559">Female measurements (in mm)</emphasis>
(n=8): Body length “Entire body” 22.0925.00 (24.09 ± 0.87); pronotum length 4.514.88 (4.74 ± 0.12); prozona length 2.723.00 (2.92 ± 0.10); metazona length 1.691.88 (1.82 ± 0.06); hind femur length 11.7514.00 (12.90 ± 0.63) and tegmina length 3.764.51 (4.24 ± 0.29).
</paragraph>
</subSubSection>
<subSubSection id="C340655EFF9A4B1934B3AFD3E0B6680E" pageId="8" pageNumber="559" type="etymology">
<paragraph id="8BE536D5FF9A4B1934B3AFD3E0B6680E" blockId="8.[151,1437,1150,1751]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934B3AFD3E233682B" bold="true" box="[199,337,1258,1283]" pageId="8" pageNumber="559">Etymology:</emphasis>
This species is dedicated to our colleague Laura Andrea Abela-Posada who helped us to collect the
<typeStatus id="54E18877FF9A4B1934B4AE36E392680F" box="[192,240,1295,1319]" pageId="8" pageNumber="559">type</typeStatus>
material. The specific name is a female noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C340655EFF9A4B1934B3AE0BE63D68D7" pageId="8" pageNumber="559" type="materials_examined">
<paragraph id="8BE536D5FF9A4B1934B3AE0BE0AA68BA" blockId="8.[151,1437,1150,1751]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934B3AE0BE25F6863" bold="true" box="[199,317,1330,1355]" pageId="8" pageNumber="559">
<typeStatus id="54E18877FF9A4B1934B3AE0BE25A6863" box="[199,312,1330,1355]" pageId="8" pageNumber="559" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<figureCitation id="13612A50FF9A4B1935E0AE0BE1676862" box="[404,517,1330,1355]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1607,1629]" captionStartId-1="3.[151,250,1223,1245]" captionStartId-2="5.[151,250,1976,1998]" captionTargetBox-0="[161,1436,409,1585]" captionTargetBox-1="[151,1423,225,1165]" captionTargetBox-2="[162,1420,975,1920]" captionTargetId-0="figure@2.[151,1436,409,1585]" captionTargetId-1="figure@3.[151,1436,215,1171]" captionTargetId-2="figure@5.[151,1436,959,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 2. Pedies andreae sp. nov. holotype male: A. Habitus, left lateral view; B. Thorax, dorsal view." captionText-1="FIGURE 3. Pedies andreae sp. nov. holotype male: External genitalia: A. Left lateral view; B. Dorsal view; Internal genitalia (dry): C. Left lateral view; D. Dorsal view." captionText-2="FIGURE 4. Pedies andreae sp. nov. holotype male: Internal genitalia (KOH-cleared): A. Complete phallic complex, dorsal view; B. Epiphallus, dorsal view; C. Ectophallus/endophallus, left lateral view; D. Ectophallus/endophallus, dorsal view." httpUri-0="https://zenodo.org/record/1458907/files/figure.png" httpUri-1="https://zenodo.org/record/1458909/files/figure.png" httpUri-2="https://zenodo.org/record/1458911/files/figure.png" pageId="8" pageNumber="559">Figs. 24</figureCitation>
).
<collectingRegion id="499EF837FF9A4B193668AE0BE1176863" box="[540,629,1330,1355]" country="Mexico" name="Mexico" pageId="8" pageNumber="559">
<collectingCountry id="F34D7645FF9A4B193668AE0BE1176863" box="[540,629,1330,1355]" name="Mexico" pageId="8" pageNumber="559">Mexico</collectingCountry>
</collectingRegion>
,
<collectingRegion id="499EF837FF9A4B1936F7AE0BE1B06863" box="[643,722,1330,1355]" country="Mexico" name="Puebla" pageId="8" pageNumber="559">Puebla</collectingRegion>
, La
<collectingCountry id="F34D7645FF9A4B19377DAE0BE0026863" box="[777,864,1330,1355]" name="Canada" pageId="8" pageNumber="559">Cañada</collectingCountry>
, near Libres, [19.510436, -97.773011] (WGS84).
<quantity id="4CA29B30FF9A4B1934E3AE6EE2706847" box="[151,274,1366,1391]" metricMagnitude="3" metricUnit="m" metricValue="2.82" pageId="8" pageNumber="559" unit="m" value="2820.0">2820 masl</quantity>
.
<date id="FFE41015FF9A4B193554AE6EE2C16847" box="[288,419,1367,1391]" pageId="8" pageNumber="559" value="2010-12-08">8-XII-2010</date>
. Grassland with
<taxonomicName id="4C5A4D56FF9A4B19361DAE6EE1CB6846" box="[617,681,1367,1390]" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B19361DAE6EE1CB6846" box="[617,681,1367,1390]" italics="true" pageId="8" pageNumber="559">Pinus</emphasis>
</taxonomicName>
,
<emphasis id="B92EEAC7FF9A4B1936CEAE6EE0566846" box="[698,820,1367,1390]" italics="true" pageId="8" pageNumber="559">
<taxonomicName id="4C5A4D56FF9A4B1936CEAE6EE0526846" authorityName="Linnaeus" authorityYear="1753" box="[698,816,1367,1390]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">Cupressus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C5A4D56FF9A4B193707AE6EE0B36846" box="[883,977,1367,1390]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B193707AE6EE0B36846" box="[883,977,1367,1390]" italics="true" pageId="8" pageNumber="559">Quercus</emphasis>
</taxonomicName>
patches of forest. Coll. Paolo Fontana, Ricardo Mariño-Pérez, and Laura Andrea Abela-Posada.
<emphasis id="B92EEAC7FF9A4B19376DAE43E0A368BB" bold="true" box="[793,961,1402,1427]" pageId="8" pageNumber="559">CNIN-UNAM</emphasis>
.
</paragraph>
<paragraph id="8BE536D5FF9A4B1934B3AEA7E63D68D7" blockId="8.[151,1437,1150,1751]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934B3AEA7E1D6689F" bold="true" box="[199,692,1438,1463]" pageId="8" pageNumber="559">Additional Type Material: CNIN-UNAM</emphasis>
. 6 paratypes (1 male, 5 females). Same data as holotype.
<emphasis id="B92EEAC7FF9A4B19313EAEA7E61C689F" bold="true" box="[1354,1406,1438,1463]" pageId="8" pageNumber="559">CPF</emphasis>
. 4 paratypes (1 male, 3 females). Same data as holotype.
<emphasis id="B92EEAC7FF9A4B19368BAEFBE01268F3" bold="true" box="[767,880,1474,1499]" pageId="8" pageNumber="559">TAMUIC</emphasis>
. 2 paratypes (both female) in ethanol. Same data as holotype, but
<date id="FFE41015FF9A4B193524AEDEE2B168D7" box="[336,467,1511,1535]" pageId="8" pageNumber="559" value="2011-12-03">3-XII-2011</date>
. Coll. Paolo Fontana, Ricardo Mariño-Pérez, Derek A. Woller, and Paola Tirello.
</paragraph>
</subSubSection>
<subSubSection id="C340655EFF9A4B1934B3AD33E7526BFF" pageId="8" pageNumber="559" type="description">
<paragraph id="8BE536D5FF9A4B1934B3AD33E7526BFF" blockId="8.[151,1437,1150,1751]" pageId="8" pageNumber="559">
<emphasis id="B92EEAC7FF9A4B1934B3AD33E1CA6B0B" bold="true" box="[199,680,1546,1571]" pageId="8" pageNumber="559">Geographic Distribution and Habitat:</emphasis>
This species was collected in the Aztec district of the east subprovince of the Trans-Mexican Volcanic Belt province. It is on its edge, very close to the Carso Huasteco subprovince of the Sierra Madre Oriental province. Both provinces belong to the Mexican Transition Zone, which harbors both Neartic and Neotropical biota, plus endemic biota (
<bibRefCitation id="EFCB4B24FF9A4B19371EAD4EE77B6BA7" box="[874,1049,1655,1679]" pageId="8" pageNumber="559" refId="ref5033" refString="Morrone, J. J. (2017) Neotropical Biogeography: Regionalization and Evolution. CRC Press, Boca Raton, 282 pp." type="book">Morrone, 2017</bibRefCitation>
). The habitat in which this species was found (
<figureCitation id="13612A50FF9A4B193550ADA3E2CB6B9A" box="[292,425,1690,1715]" captionStart="FIGURE 1" captionStartId="1.[151,250,1315,1337]" captionTargetBox="[151,1436,193,1294]" captionTargetId="figure@1.[151,1436,193,1294]" captionTargetPageId="1" captionText="FIGURE 1. Live specimens of Pedies andreae sp. nov. from type habitat on 3-XII-2011: A. male; B. female; C. and D. Type habitat: grassland with forest patches in Mexico, Puebla, La Cañada, near Libres, [19.510436, -97.773011] (WGS84). 2820 masl. 3-XII-2011." httpUri="https://zenodo.org/record/1458905/files/figure.png" pageId="8" pageNumber="559">Fig. 1C&amp;D</figureCitation>
) can be described as mixed vegetation with patches of forest (
<taxonomicName id="4C5A4D56FF9A4B1930EEADA2E7B86B9A" box="[1178,1242,1691,1714]" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B1930EEADA2E7B86B9A" box="[1178,1242,1691,1714]" italics="true" pageId="8" pageNumber="559">Pinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A4D56FF9A4B193099ADA3E6036B99" authorityName="Linnaeus" authorityYear="1753" box="[1261,1377,1690,1713]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B193099ADA3E6036B99" box="[1261,1377,1690,1713]" italics="true" pageId="8" pageNumber="559">Cupressus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5A4D56FF9A4B1934E3AD86E3976BFE" box="[151,245,1727,1750]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B1934E3AD86E3976BFE" box="[151,245,1727,1750]" italics="true" pageId="8" pageNumber="559">Quercus</emphasis>
</taxonomicName>
), with native grasses, such as
<taxonomicName id="4C5A4D56FF9A4B19363CAD87E1876BFE" authorityName="Schreber" authorityYear="1789" box="[584,741,1726,1750]" class="Magnoliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="8" pageNumber="559" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAC7FF9A4B19363CAD87E1876BFE" box="[584,741,1726,1750]" italics="true" pageId="8" pageNumber="559">Muhlenbergia</emphasis>
</taxonomicName>
sp. and crops, such as maize.
</paragraph>
</subSubSection>
</treatment>
</document>